taxonID	type	description	language	source
7E7FDE5BFF98FFC2FF5DFBB7FAEF6179.taxon	materials_examined	Holotype: NMBE 1061645, field number HJ 12 – 125 (Figs. 2 – 5), GenBank accession number KJ 488546, from the summit trail between Church Camp and the summit of Gunung Murud, near the intersection with Sungei Murud, Gunung Murud, Pulong Tau National Park, Sarawak, East Malaysia (Borneo), altitude 2,152 m asl (N 03 ° 54.721 ', E 115 ° 29.954, GPS WGS 84), coll. Y. M. Pui, A. Jankowski and S. T. Hertwig, 28 February 2012. Adult female. Paratype: NMBE 1066153, field number SH 13 – 291 (Fig. 3 C, D), GenBank accession number KJ 488547, 25 m away from the locality of holotype at the summit trail, coll. Y. M. Pui, T. Einecke, T. Keilholz and S. T. Hertwig, 10 March 2013. Adult female.	en	Hertwig, Stefan T., Min, Pui Yong, Haas, Alexander, Das, Indraneil (2014): Dressed in black. A New Ansonia Stoliczka, 1870 (Lissamphibia: Anura: Bufonidae) from Gunung Murud, Sarawak, East Malaysia (Borneo). Zootaxa 3814 (3): 419-431, DOI: 10.11646/zootaxa.3814.3.9
7E7FDE5BFF98FFC2FF5DFBB7FAEF6179.taxon	diagnosis	Diagnosis. The new species is assigned to the genus Ansonia based on its relatively slender body, long legs, indistinct subarticular tubercles, membraneous digitial webbing, the absence of parotoid glands, tympanum visible (Inger 1960, 1966) and a robustly supported phylogenetic hypothesis (Fig. 1). Ansonia vidua sp. nov. can be differentiated from all other congeners by the following combination of characters: SVL 33.5 and 34.3 mm in the two females known; body not flattened; canthus rostralis slightly swollen; two low longitudinal interorbital ridges present; dorsum of body, flanks and upper surfaces of the limbs covered with numerous small, regular, low, rounded warts, resulting in a velvet-like appearance, only warts on scapular and temporal regions slightly enlarged; warts on dorsum, head and limbs terminating in a single fine, keratinous spine; first finger short, not reaching tip of second finger when adpressed; no sharp tarsal ridge; no skin flaps on posterior thigh near vent; third and fifth toe with 2 ½ phalanges free of web medially; colouration of the dorsal surface of head and body uniformly black-brown; limbs dark brown with lighter areas on joints of the limbs and phalanges; ventral surface uniformly dark grey to yellowish brown; dorsal and ventral parts of body and limbs completely patternless.	en	Hertwig, Stefan T., Min, Pui Yong, Haas, Alexander, Das, Indraneil (2014): Dressed in black. A New Ansonia Stoliczka, 1870 (Lissamphibia: Anura: Bufonidae) from Gunung Murud, Sarawak, East Malaysia (Borneo). Zootaxa 3814 (3): 419-431, DOI: 10.11646/zootaxa.3814.3.9
7E7FDE5BFF98FFC2FF5DFBB7FAEF6179.taxon	description	Description of holotype. Habitus slender; body as wide as head (Figs. 2 – 3 A, B), body not dorsoventrally flattened; head slightly longer than wide (Table 2); HWAJ / TBL 0.62; rostrum truncate in dorsal view, tip of snout rounded, longer than canthus, oblique in lateral view, projecting beyond tip of mandible, longitudinal, medial groove on the dorsal face of snout, rostral length greater than eye diameter; nostril lateral, closer to snout tip than eye (Fig. 4); distinct vertical ridge between tip of snout and centre of maxilla (Figs. 3 A, B, 4); canthus swollen with rounded edge, slightly arcuate in lateral view; lores vertical and concave; eye diameter greater than eye-nostril distance; interorbital distance subequal to upper eyelid (Table 2); tympanum distinct, eye diameter 1.9 times greater than tympanic diameter (Fig. 4, Table 2); two low, slightly curvestridges on head forming a parenthetical shape “ () ” extends from the anterior aspect of the interorbital region to a point approximately in line with the centre of the tympani (Fig. 3 A), one supratympanic ridge in parotoid region (Fig. 2 and 3 A). Tips of fingers IV and V weakly expanded, spatulate, without distinct discs; first finger (II) much shorter than second (III), only reaching middle of second phalange of second finger; subarticular tubercles low; palm smooth with large, distinct rounded palmar tubercle. Tips of all toes rounded; fifth toe projecting farther than third; toes not fully webbed, first toe with one phalange free of web, second with 1 ½ phalanges free of web medially, third and fifth with 2 ½ phalanges free of web medially, fourth toe with 4 phalanges free of web medially; subarticular tubercles indistinct; two metatarsal tubercles, both oval and slightly raised (Fig. 5); no tarsal fold (Fig. 3). Left foot slightly deformed, toes shortened (Fig. 3 A, B). Skin of sides, back and dorsal surfaces of limbs uniformly covered with small, low, rounded tubercles; shagreened skin has a characteristic velvet-like appearance; tubercles not arranged in distinct rows, tubercles on top of head and eyelids smaller (Figs. 2 – 4), slightly enlarged tubercles on scapular and temporal region and posterior of tympanum (Figs. 2 – 4); one row of enlarged, spineless tubercles with darker pigmentation beneath mandibular symphysis (Fig. 3 B), followed posteriorly by few smaller, spineless, irregularly arranged tubercles (Fig. 3 B); tubercles on dorsal surfaces of head, body, and limbs tipped with one small, regularly shaped dark brown spine, tubercles on sides, venter and throat without spines (Fig. 3 A); no skin flaps on posterior thigh near vent (Fig. 3 A, B). Colour in preservative uniformly dark brown above, slightly lighter areas on sides and on joints of limbs and phalanges; venter uniformly brownish grey, throat and chest uniformly yellowish brown (Fig. 3 A, B). Colour in life uniformly black-brown above, limbs dark brown with lighter areas on joints of the limbs and phalanges (Fig. 2). Iris black with small irregular intense red spots (Fig. 2). Variation. In the paratype (Table 2, Fig. 3 C, D), skin on sides is slightly smoother, abdomen and throat finely granular, tubercles smaller and indistinct, only a single row of enlarged tubercles with darker pigmentation present posterior of the mandibular symphysis. Skin fold on rear thigh near vent is a preservation artifact (Fig. 3 C, D). Lighter areas on sides larger in preservative, colour of throat changes from light yellowish grey to yellowish brown anteriorly (Fig. 3 C, D).	en	Hertwig, Stefan T., Min, Pui Yong, Haas, Alexander, Das, Indraneil (2014): Dressed in black. A New Ansonia Stoliczka, 1870 (Lissamphibia: Anura: Bufonidae) from Gunung Murud, Sarawak, East Malaysia (Borneo). Zootaxa 3814 (3): 419-431, DOI: 10.11646/zootaxa.3814.3.9
7E7FDE5BFF98FFC2FF5DFBB7FAEF6179.taxon	etymology	Etymology. The species name vidua means´widow´and refers to the uniform brownish-black colouration in life of this species, which is traditionally the colour of choice by widows. We suggest the English vernacular name ‘ Murud Black Slender Toad’ for the species. Ecological notes. The type series of Ansonia vidua sp. nov., consisting of two female individuals, was collected on two expeditions to the Pulong Tau National Park in 2012 and 2013. Males and larval stages of the new species are, as yet, unknown. Both specimens were found at night, resting on the leaves of lower vegetation at the edge of the summit trail to Gunung Murud, ca. two hours walk from Church Camp, a few meters from the edge of a small stream. The type locality is situated between this unnamed small side stream and the Sugei Murud that runs eastward from near the summit of Gunung Murud down to the plateau of the Kelabit Highlands. The type locality is situated just below the summit ridge of the Gunung Murud massif. In this undisturbed montane mossy forest (Beaman 1999; Beaman & Anderson 1997; Mjöberg 1925), the following species of frogs have been recorded in sympatry: Limnonectes cf. kuhlii, Philautus mjobergi, P. cf. petersi, Pelophryne murudensis and Leptobrachium montanum. The amphibian fauna of Gunung Murud has been described by Das (2005, 2008). The discovery of a new and possibly endemic species from the Gunung Murud massif underlines the importance of the Pulong Tau National Park in the protection of the highly diverse montane amphibian fauna of Sarawak and that of Borneo.	en	Hertwig, Stefan T., Min, Pui Yong, Haas, Alexander, Das, Indraneil (2014): Dressed in black. A New Ansonia Stoliczka, 1870 (Lissamphibia: Anura: Bufonidae) from Gunung Murud, Sarawak, East Malaysia (Borneo). Zootaxa 3814 (3): 419-431, DOI: 10.11646/zootaxa.3814.3.9
