identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
7D431D7FFFD9985F3C22FC65565FF0E4.text	7D431D7FFFD9985F3C22FC65565FF0E4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stygiopontius lomonosovi Ivanenko and Martinez Arbizu	<div><p>Stygiopontius lomonosovi Ivanenko and Martínez Arbizu, new species</p><p>Figs 3–9</p><p>Type material. Dissected holotype Ψ mounted on 3 slides (SMF 31035), dissected allotype ♂ mounted on 5 slides (SMF 31036), 2 paratype ΨΨ mounted on 2 slides (SMF 31037and SMF 31038), and 1 paratype ♂ mounted on 1 slide (SMF 31039).</p><p>Type locality. Mid­Atlantic Ridge, Logachev­1 hydrothermal field, IRINA II site, station 35­ GTV, 14°45.19’N, 44°58.75’W, depth 3019 m. Periphery of the smoker complex covered by Bathymodiolus puteoserpentis (Fig. 1 A). Collected 25 January 2004.</p><p>Etymology. The specific epithet is derived from the family name of the outstanding Russian scientist Michail V. Lomonosov (1711–1765), one of the founders of the Moscow State University established in 1755. It is the Latinized, masculine, genitive form of Lomonosov.</p><p>Female holotype. Body (Fig. 3 A) with broad flattened prosome. Total length, excluding caudal setae, 0.93 mm, greatest width 0.50 mm. Prosome with ventrolateral folds as shown for male in Fig. 7 B. Urosome (Fig. 3 B) 5­segmented, consisting of fifth pedigerous somite, genital double­somite and 3 postgenital somites. First somite of urosome trapezoidal in dorsal view. Lateral expansions in anterior part of genital doublesomite with dorsolateral genital fields bearing small seta (Fig. 3 B–C). Posteroventral margin of the genital double­somite with lateral spiniform processes (Fig. 3 B &amp; D). Anal opening near posterior margin of the last abdominal somite (Fig. 3 E).</p><p>Rostrum weakly developed, oral cone short and robust as in male (Fig. 7 B).</p><p>Caudal ramus (Fig. 3 B, E &amp; F) short, with 6 setae, outermost terminal seta smooth, innermost terminal seta with setules on inner side (2 dorsal setae lost in holotype).</p><p>Antennule (Fig. 4 A) 12­segmented: first segment with 1 seta; second segment the longest, with 14 unequal setae; segment 3 subdivided, with 10 unequal setae, 2 setae very small; segment 4 short with 2 setae; segment 5 double, with 4 (2 pairs) of setae; segments 6–9 each with 2 setae, 1 seta near the middle, and another seta near the distal edge; segment 10 with 2 setae and 1 aesthetasc near the distal edge; segment 11 bearing six setae; segment 12 with 8 setae. All setae, except distal seta of segment 5, smooth.</p><p>Antenna (Fig. 4 B): small coxa without ornamentation, elongate basis with setules on outer and inner margins. Exopod small, 1­segmented, with 3 smooth setae. Endopod 2­ segmented; segment 1 elongate and unarmed, with setules on outer and inner margins; segment 2 short, ornamented with setules, armed with 5 setae: 4 terminal setae (3 elongate of different lengths and 1 short) and 1 short subterminal seta.</p><p>Mandible (Fig. 4 C): gnathobase stylet­like and flattened; terminal teeth of unequal size, proximoventral margin with a row of small teeth.</p><p>Maxillule (Fig. 4 D–E): inner lobe with a row of slender setules on inner margin and 5 terminal setae, 1 short and 4 long; outer lobe articulate, with 3 long, terminal and 1 short, subterminal setae. All long setae ornamented with setules of different lengths.</p><p>Maxilla (Fig. 5 A) 2­segmented: segment 1 wide; segment 2 elongate, with flattened spoon­like tip ornamented with setules. Long inner seta near juncture of segments; its distal part ornamented with setules.</p><p>Maxilliped (Fig. 5 B): syncoxa with 1 long, inner, bipinnate seta; basis with shorter, inner seta along inner margin of the segment, ornamented with spinules. Endopod 3­ segmented: segment 1 with 2 short, posterior setae; segment 2 with 1 long, posterior seta ornamented with spinules; segment 3 bearing 2 terminal setae with setules on inner margin, 1 seta very long and thick.</p><p>Swimming legs 1–4 (Figs 5 C–D, 6A–C) with 3­segmented rami, except for leg 4, with 2­segmented endopod. Formula for the armature of legs 1–4 in Table 1. Distal endopodal segment of legs 1–3 and distal exopodal segment of leg 3 ornamented with groups of slender posterior setules as shown in the figures. Leg 1: inner seta of basis slender and smooth; outer spine of exopodal segment 2 small and curved proximally; 1 outer spine of first exopodal segment and 3 outer spines of third ornamented with long subterminal setule; this setule absent on spines of other swimming legs. Leg 4: inner seta of exopodal segment 1 smooth in proximal part and ornamented with small setules in distal part (Fig. 6 C). Intercoxal sclerites of legs 3–4 expanded. Distal exopodal segment of leg 4 with 2 outer spines.</p><p>Leg 5 (Fig. 6 D) indistinctly 2­segmented; segment 1 with 1 long outer seta; segment 2 armed with 3 setae of different lengths (lost seta arrowed).</p><p>Leg 6 (Fig. 3 C) represented by a flap covering gonopores and bearing 1 small seta. Color of living specimens unknown.</p><p>Egg sacs not observed.</p><p>Male allotype. Differs from female as follows:</p><p>Body (Fig. 7 A–B): total length of allotype, excluding caudal setae, 1.03 mm, greatest width 0.56 mm. Shield of cephalothorax and tergites of 2 anterior metasomites with small pointed posterolateral processes, also present in female, but not visible in dorsal view.</p><p>Urosome (Fig. 7 C–E) 6­segmented, consisting of fifth pedigerous somite, genital somite, and 4 postgenital somites. Genital somite with pair of posterior genital flaps on ventral side (Fig. 7 D) and posterodorsal prominence visible from lateral view (Fig. 7 E). First abdominal somite corresponding to abdominal somite included in the female genital double­somite and possessing posterolateral spiniform processes.</p><p>Antennule (Fig. 8 A–D) 12­segmented, geniculate between segments 10 and 11; formula of setation as follows: 1, 2, 12, 8, 2, 2, 4, 2, 2, 4, 4+aesthetasc, 12. Segments 2 and 3 correspond to segment 2 of female antennule. Distal part of segment 3 with long curved process bearing subterminal seta (Fig. 8 B). Segment 4 with 3 incomplete arthrodial membranes indicating a segment complex. Segment 5 with ventral process and curved stout spine on it (Fig. 8 C). Two setae on segment 10 and 3 setae of segment 11 modified to short flattened element (Fig. 8 D, arrowed).</p><p>Maxilliped (Fig. 8 E–F). Syncoxa prolonged medioventrally into terminal clawlike process with subterminal barbed seta. Inner seta of basis shorter than that of female. Segment 2 of endopod with 2 posterior setae, 1 distal and 1 middle (the last is absent in female).</p><p>Distal endopodal segment of leg 1 (Fig. 9 A) and distal exopodal segment of leg 3 (Fig. 9 C) with group of slender posterior setules. Leg 2 (Fig. 9 B): distal endopodal segment of leg 2 with 1 outer, 2 terminal, and 1 distomedial spine (formula I,II,I,2), outermost terminal spine curved inwardly; coxal seta short.</p><p>Leg 5 (Figs 7 D, 9D) located ventrally. Basis fused with somite represented by 1 outer seta, exopodal segment articulate, armed with 3 setae.</p><p>Leg 6 (Fig. 7 D–E) represented by genital flap bearing 2 long posterior setae.</p><p>Colour unknown.</p><p>Remarks. Stygiopontius lomonosovi n. sp. can be easily assigned to the genus Stygiopontius by the combination of the following characters: the endopod of leg 1 is 3­ segmented; the endopod of leg 4 is 2­segmented, its first segment unarmed and its second segment with two setae (one terminal and one inner) (Ivanenko &amp; Ferrari 2003). The new species is similar to S. mirus Humes, 1996 and S. latulus Humes, 1996 (described from the Snake Pit (23ºN) of the MAR) sharing with the new species the unusual medioventrally prolonged syncoxa of the maxilliped. The new species can be distinguished from both S. mirus and S. latulus by the absence of one spine on the distal exopodal segment of leg 4: armature formula II,I,4, instead of III,I,4 as found in S. mirus and S. latulus . Five other congeners possess only two outer spines on the third exopodal segment of leg 4, viz. S. cinctiger Humes, 1987; S. mucroniferus Humes, 1987; S. teres Humes, 1996; S. rimivagus Humes, 1997; S. verruculatus Humes 1987 . In this group, only females are known for the first three species and only males are described for the last two species. The following attributes distinguish these species from S. lomonosovi n. sp.: the maxilliped of S. rimivagus and S. verruculatus lacks an elongation of the syncoxa; leg 5 of S. teres is short and 1­segmented, its prosome moderately slender; the coxa and basis of maxilliped of S.</p><p>mucroniferus are armed with a stout seta; the second abdominal somite of S. cinctiger is shorter.</p><p>Males of 12 valid species included in the genus Stygiopontius have been described so far. Only males of S. lomonosovi sp. n. and five other species ( S. mirus, S. latulus, S. brevispina Humes, 1991, S. lauensis Humes, 1991, and S. rimivagus) were recorded in the Lau Basin of the West Pacific and the MAR. They are characterized by reduction of two inner setae on the exopod of leg 5 (the segment is fused with the sixth thoracic somite in S. latulus). The exopod is armed with three setae instead of a maximum of five setae present in males of congeners. Further investigations of unknown males of dirivultid species are necessary to clarify if the reduction of the setae on the exopod of leg 5 can be used as a character to define a monophyletic lineage of Stygiopontius distributed in both the West Pacific and the MAR.</p></div>	https://treatment.plazi.org/id/7D431D7FFFD9985F3C22FC65565FF0E4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ivanenko, Viatcheslav N.;Arbizu, Pedro Martínez;Stecher, Jens	Ivanenko, Viatcheslav N., Arbizu, Pedro Martínez, Stecher, Jens (2006): Copepods of the family Dirivultidae (Siphonostomatoida) from deep­sea hydrothermal vent fields on the Mid­Atlantic Ridge at 14 ºN and 5 ºS. Zootaxa 1277: 1-21, DOI: 10.5281/zenodo.173351
7D431D7FFFD3985F3C22FD1557C2F763.text	7D431D7FFFD3985F3C22FD1557C2F763.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stygiopontius cladarus Humes 1996	<div><p>Stygiopontius cladarus Humes, 1996</p><p>Material examined. 6 ΨΨ, Mid­Atlantic Ridge, Turtle Pits site, chimney “Tower”, samples 123 ROV­5 and 123 ROV­6, 4°48.6’S, 12°22.4’W, depth 2992 m, washing of alvinocaridid shrimp Rimicaris sp., 11 April 2005 (Fig. 1 B).</p><p>Differential diagnosis. Anterior part of genital double­somite swollen ventrally in anterior part. Caudal ramus subquadrate. Coxae of legs 1 and 4 without inner seta. Basis of leg 1 with mammilliform inner margin. Segment 3 of exopod of leg 4 with 3 outer spines (III,I,4). Leg 5 of female 2­segmented armed with 1 and 3 setae. Endopodal segment 3 of male leg 2 armed with 4 spines and 2 setae (formula I,II,I,2). Leg 5 of male 1­segmented, exopod bearing 5 setae.</p><p>Remarks. S. cladarus was previously recorded in washing of Rimicaris specimens from the Snake Pit site at 23°N (Humes 1996) and from plankton over a hydrothermal field of the Broken Spur field at 29°N (Ivanenko 1998).</p></div>	https://treatment.plazi.org/id/7D431D7FFFD3985F3C22FD1557C2F763	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ivanenko, Viatcheslav N.;Arbizu, Pedro Martínez;Stecher, Jens	Ivanenko, Viatcheslav N., Arbizu, Pedro Martínez, Stecher, Jens (2006): Copepods of the family Dirivultidae (Siphonostomatoida) from deep­sea hydrothermal vent fields on the Mid­Atlantic Ridge at 14 ºN and 5 ºS. Zootaxa 1277: 1-21, DOI: 10.5281/zenodo.173351
7D431D7FFFD398403C22FB9F570CF16C.text	7D431D7FFFD398403C22FB9F570CF16C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stygiopontius pectinatus Humes 1987	<div><p>Stygiopontius pectinatus Humes, 1987</p><p>Material examined. 39 ΨΨ, Mid­Atlantic Ridge, Turtle Pits site, chimney “Tower”, samples 123 ROV­5 and 123 ROV­6, 4°48.6’S, 12°22.4’W, depth 2992 m, washing of alvinocaridid shrimps ( Rimicaris sp.), 11 April 2005 (Fig. 1 B). 316 ΨΨ, 2 copepodids (stage 1), 3 nauplii (described by Ivanenko, Martínez Arbizu &amp; Stecher, submitted), Mid­ Atlantic Ridge, Red Lion site, chimney "Shrimps Farm", sample 146 ROV­6, 4°47.82’S, 12°22.60’W, depth 3048 m, washing of alvinocaridid shrimps ( Rimicaris sp.) (Fig. 2 A–B), 16 April 2005.</p><p>Differential diagnosis. Inner margin of claw of both antenna and maxilliped pectinate. Segment 3 of exopod of leg 4 with 3 outer spines (formula III,I,4). Segment 2 of endopod of leg 4 with short terminal spine (armature formula of the segment I,1).</p><p>Remarks. S. pectinatus is a widespread species with females reported from three hydrothermal vent fields of the MAR and one site of the Mariana Back­Arc Basin (West Pacific); males are unknown. The females were found in the branchial chambers and in washings of alvinocaridid shrimps ( Rimicaris exoculata and Chorocaris chacei) from TAG at 26°N and Snake Pit at 23°N (Humes 1987; Humes 1996). They were also found in plankton over Broken Spur at 29°N (Ivanenko 1998) and in washings of tubes of the polychaete Alvinella pompejana from the Marianna Back­Arc Basin (Humes 1990a).</p><p>Our observation of the alvinocaridid shrimps collected at 5ºS and the oral cone and the swimming legs of S. pectinatus collected on these shrimps supports the suggestion of Humes (1996) that females of this dirivultid live in the shrimps’ branchial chamber and feed on chemoautotrophic bacteria growing on the shrimp’s feeding appendages and the inner surface of branchial chamber (Van Dover et al. 1988; Segonzac et al.1993; Gebruk et al. 2000b). It is not clear if the copepods clean the branchial chamber off fouling bacteria or utilize bacterial food of the specialized shrimps.</p></div>	https://treatment.plazi.org/id/7D431D7FFFD398403C22FB9F570CF16C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ivanenko, Viatcheslav N.;Arbizu, Pedro Martínez;Stecher, Jens	Ivanenko, Viatcheslav N., Arbizu, Pedro Martínez, Stecher, Jens (2006): Copepods of the family Dirivultidae (Siphonostomatoida) from deep­sea hydrothermal vent fields on the Mid­Atlantic Ridge at 14 ºN and 5 ºS. Zootaxa 1277: 1-21, DOI: 10.5281/zenodo.173351
7D431D7FFFCC98403C22FDE05734F561.text	7D431D7FFFCC98403C22FDE05734F561.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aphotopontius atlanteus Humes 1996	<div><p>Aphotopontius atlanteus Humes, 1996</p><p>Syn. A. temperatus Humes, 1997</p><p>Material examined: 2 ΨΨ, Mid­Atlantic Ridge, Logachev­1 hydrothermal field, IRINA II site, sample 38 ROV­4, 14°45.19’N, 44°48.75’W, depth 3036 m, musselbed of Bathymodiolus puteoserpentis, southwest of smoker­complex, 26 January 2004. 27 ΨΨ, sample 35 GTV, 14°45.19’N, 44°58.75’W, depth 3019 m, musselbed of Bathymodiolus puteoserpentis near smoker­complex, 26 January 2004 (Fig. 1 A).</p><p>Differential diagnosis. Anterior part of genital double­somite laterally expanded and rounded. Caudal rami with ratio 1.9: 1 in female, 1.4: 1 in male. Leg 1 with inner coxal seta and with rounded inner margin of basis. Endopodal segment 3 of leg 2 with 2 terminal setae (formula 1,2,3) in female and 2 terminal spines (1,II,3) in male.</p><p>Remarks. The copepods of the genus Aphotopontius are not known in the western Pacific, but are common in the eastern Pacific: Nine of 10 species were recorded from eight sites situated from the equator (Galapagos Rift) to the 49°N (Explorer Ridge). A second species of Aphotopontius from the MAR, A. forcipatus Humes, 1987, was described from the Explorer Ridge and later recorded from the Snake Pit on the MAR (Humes 1996), the Gorda Ridge (Humes 1990b) and the Juan de Fuca Ridge (Humes &amp; Huys 1992). The species can be distinguished from Aphotopontius atlanteus by the elongate caudal ramus (ratio 5.6:1) among other attributes.</p><p>Aphotopontius atlanteus was previously recorded in many samples from Lucky Strike and Menez Gwen at 37°N (Humes 1996; Humes 1997; Humes &amp; Segonzac 1998). Aphotopontius temperatus Humes, 1997 described from Lucky Strike was synonymized with A. atlanteus by Ivanenko and Defaye (2006b).</p></div>	https://treatment.plazi.org/id/7D431D7FFFCC98403C22FDE05734F561	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ivanenko, Viatcheslav N.;Arbizu, Pedro Martínez;Stecher, Jens	Ivanenko, Viatcheslav N., Arbizu, Pedro Martínez, Stecher, Jens (2006): Copepods of the family Dirivultidae (Siphonostomatoida) from deep­sea hydrothermal vent fields on the Mid­Atlantic Ridge at 14 ºN and 5 ºS. Zootaxa 1277: 1-21, DOI: 10.5281/zenodo.173351
7D431D7FFFCD98413C22FF455646F11B.text	7D431D7FFFCD98413C22FF455646F11B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rimipontius mediospinifer Humes 1996	<div><p>Rimipontius mediospinifer Humes, 1996</p><p>Material examined. 2 ΨΨ, Mid­Atlantic Ridge, Logachev­1 hydrothermal field, IRINA II site, sample 35­ GTV, 14°45.19’N, 44°58.75’W, depth 3019 m, washing from Bathymodiolus puteoserpentis, 25 January 2004 (Fig. 1 A).</p><p>Differential diagnosis. Dorsal surface of genital female double­somite with stout median crest ending with spiniform posterior process. Caudal rami with 5 setae, innermost terminal seta absent. Maxilla with inner sinuous seta. Endopod of leg 4 is 2­segmented and armed with 1 terminal spine (formula 0­0; I,0).</p><p>Remarks. The species is the type and only species of the genus Rimipontius Humes, 1996 . It was previously recorded from the Logachev field (Humes &amp; Segonzac 1998), in washings of Rimicaris from Snake Pit site (Humes 1996) and in plankton over the Broken Spur (Ivanenko 1998).</p></div>	https://treatment.plazi.org/id/7D431D7FFFCD98413C22FF455646F11B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ivanenko, Viatcheslav N.;Arbizu, Pedro Martínez;Stecher, Jens	Ivanenko, Viatcheslav N., Arbizu, Pedro Martínez, Stecher, Jens (2006): Copepods of the family Dirivultidae (Siphonostomatoida) from deep­sea hydrothermal vent fields on the Mid­Atlantic Ridge at 14 ºN and 5 ºS. Zootaxa 1277: 1-21, DOI: 10.5281/zenodo.173351
