taxonID	type	description	language	source
7C5887A3FFE1B25FFF6AF82C32D0FB25.taxon	materials_examined	Type species: Diochus nanus Erichson, 1839: 300, fixed by monotypy. Syn.: Rhegmatocerus Motschulsky, 1858: 657 (species included: conicollis, punctipennis, antennatus); Gemminger and Harold, 1868: 608 (synonym of Diochus); Bernhauer and Schubert, 1914: 319 (synonym of Diochus); Cameron, 1932: 44 (synonym of Diochus); Blackwelder, 1943: 455 (synonym of Diochus; type species: punctipennis); Coiffait, 1972: 368 (synonym of Diochus); Smetana, 1982: 27 (synonym of Diochus); Smetana, 2004: 624 (synonym of Diochus); Löbl & Löbl, 2015: 1007 (synonym of Diochus). Type species: Rhegmatocerus punctipennis Motschulsky, 1858 fixed by subsequent designation by Blackwelder, 1943: 455.	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFE1B25FFF6AF82C32D0FB25.taxon	diagnosis	Diagnosis. the genus Diochus can be distinguished from all other genera within the tribe Diochini by the following characteristics: a) body slender, more or less fusiform (Fig. 3), usually small to medium in body-size (3 – 6 mm); b) head small, bearing two paired punctures quadrately located on disc (Fig. 1); a medial elevation between eyes only present in male (Fig. 4 A, 5 A, 6 A), sometimes underdeveloped or missing at all (Fig. 7 A); c) maxillary palpus markedly long, penultimate and segment II extremely long and of same length; penultimate obconical; last segment small and acicular (Fig. 5 P); d) labial palpus short, last segment slender and stalked (Fig. 6 M); e) neck nearly 1 / 3 width of head, with furrow or groove on both dorsal and ventral surface, also with three pairs of cervical sclerites (Fig. 4 K – L); f) pronotum oval shaped, anterior angles abruptly bended ventrally, with five paired punctures on surface (Fig. 1); g) elytra not overlapping at suture; normally with paired adsuture and admedian rows, each composed of five punctures (Fig. 1); h) protarsi somewhat dilated; i) antesternal plate transparently sclerotized, bearing numerous bubble-like protrusions (Fig. 2); j) superior line of hypomeron bending toward prosternum before anterior angle of pronotum, but not joining inferior line (Fig. 2); k) distinct intercoxal apophysis and transverse ridge on prosternum (Fig. 2); l) obvious longitudinal ridge, transverse ridge, oblique furrow on mesoventrites (Fig. 2); m) discrimen of metaventrites distinctly elevated (Fig. 2); n) aedeagus sclerotized to various extent: totally sclerotized (Fig. 4 - 1 E), partly membranous (Fig. 5 - 1 E, 6 - 1 E), or occasionally totally membranous (Fig. 7 - 1 E); parameres symmetrical and extraordinarily thin (Fig. 5 - 1 E, 6 - 1 E); possessing different kinds of sperm pump (Fig. 4 - 1 G, 5 - 1 G, 6 - 1 G, 7 - 1 G); o) female spermetheca tube-like, apically dilated (Fig. 4 - 1 J, 5 - 1 J, 6 - 1 J); segment IX symmetrical and narrowing into stylus-like apical portion with sharp spine-like apex (Fig. 4 - 1 O, H); tergite X distinctly broad (Fig. 4 - 1 I, 5 - 1 I, 6 - 1 I), sternite X atrophied (Fig. 4 - 1 L, 6 - 1 L), or absent (Fig. 5 - 1; Fig. 7 - 1).	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFE5B254FF6AFB173020FCC0.taxon	description	(Fig. 3 B; Fig. 4 A – P; Fig. 4 - 1 A – O)	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFE5B254FF6AFB173020FCC0.taxon	materials_examined	Type material. Holotype: male, CHINA: Fujian: Shanghang co.: Meihua Shan: Shuangche (E 116.8568 °, N 25.3411 °), 475 m, 06. XI. 2008, Wang Fengyan collected (IZ-CAS). Paratypes: Hainan: Ledong co.: Jianfeng Ling (E 108.8792 °, N 18.7211 °): Tianchi, one male, one female, 1000 m, 13. XI. 2008, Zhao Zongyi collected (IZ-CAS); Yulingu, one male, 900 m, 23. XI. 2008, Zhao Zongyi collected (IZ-CAS); Qiongzhong co.: Limu Shan (E 109.8016 °, N 19.2349 °), Sanquling, one male, 630 m, 04. XII. 2007, Yang Zhuo collected; Lingshui Lizu autonomous co., Diaoluoshan (E 109.9160 °, N 18.6598 °), one male, 930 m, 30. VII. 2004, Wu Jie & Chen Yongjie (IZ-CAS); Zhejiang: Lanao co.: Fengyang Shan (E 119.1989 °, N 27.9381 °), Fengyangjian, one male, 1400 m, 01. VIII. 2008, Xu Jinhua collected (IZ-CAS); Guangxi: Fangchenggang City: Fulong co. (E 107.9518 °, N 21.8146 °), four males, one female, 660 m, 21. IV. 1998, Zhou Haisheng collected (IZ-CAS); Yunnan: Mengla co.: Menglun (E 101.2490 °, N 21.9375 °), one female, 550 m, 08. II. 2004, Wu Jie & Bai Daiyuan collected (IZ-CAS); Longmen village, Xiaoniupeng (E 101.3252 °, N 21.3095 °), one female, 1010 m, 06. X. 2010, Zhou Yu-Lingzi collected (IZ-CAS); Jinghong City: Jinghong Nature Reserve (E 100.82263 °, N 20.0665 °), one female, 800 m, 26. II. 2004, Wu Jie collected (IZ-CAS).	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFE5B254FF6AFB173020FCC0.taxon	description	Description. Measurement (n = 10). BL = (5.1 – 5.2) mm, FL = (2.8 – 2.9) mm, HL = (0.6 – 0.7) mm, HW = (0.5 – 0.6) mm, PL = (0.9 – 1.0) mm, PW = (0.8 – 0.9) mm, EL = (0.9 – 1.0) mm, EW = (1.1 – 1.2) mm. Body nearly fusiform shaped, large sized (Fig. 3 B). Body dark brown, except elytra darker and abdominal intersegmental membrane yellowish brown (Fig. 4 O). Legs brown, tarsi distinctly lighter. Antennae brown, apex of each antennomere lighter. Maxillary and labial palpi brown. Head (Fig. 4 A, D). Oblong in shape (HL to HW ratio 1.41), tempora (behind eyes) straight and paralleled, posterior angles rounded. Dorsal integument shiny, extensively bearing micropunctures and microstriae, also with 5 – 7 irregular, scattered punctures around each eye, and numerous smaller punctures on deflexed portion of tempus and near the posterior margin. Each side of cranium with characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three ocular punctures near dorsal margin of eye, temporal puncture at the basal 1 / 6, also with occipital puncture at the basal 1 / 5; in addition, disc surface bearing two additional paired punctures (Fig. 1). Frontal furrows and anteocular furrows absent, a longitudinal linear elevation present between eyes (Fig. 1). Eye medium sized, distinctly protruding laterad, longitudinal diameter slightly shorter than half the length of tempora (eye: tempora = 0.22: 0.45 mm). Epistoma not flattened, abruptly oblique, anterior margin slightly emarginate medially. Distance between antennal insertions 0.17 mm being distinctly longer than distance from antenna to eye (0.09 mm). Ventral surface (Fig. 4 D) with scattered punctures, interspaces between them ca. 4 punctures’ diameter. Gular sutures deep, confluent at basal 1 / 4, then extended to base. Antennae (Fig. 4 G). Total length 1.6 mm. Scape rod-shaped, slightly thickened apically, 0.29 mm, shorter than two subsequent antennomeres combined; antennomere II elongate, 0.16 mm; III 0.19 mm, distinctly longer than II; IV 0.13 mm, IV to X gradually shortened, IV observably longer than X; XI 0.16 mm, distinctly longer than X. Mouthparts (Fig. 4 H – J, P). Labrum subrectangular (Fig. 4 H), lateral margin slightly wave-shaped, anterior margin sinuous. Mandibles falciform, left one with three small teeth on inner edge, right one with a small tooth (Fig. 4 I – J). Neck (Fig. 4 K – L). Surface shiny and nearly trapezoidal shaped, only bearing shallow microsculpture, with a bisinuate basal furrow near posterior margin (Fig. 4 K), width of 0.22 mm, wider than 1 / 3 width of head. Ventral surface with deep gular sutures to form obvious triangular region, which missing transverse median furrow (Fig. 4 L). Prothorax (Fig. 4 B, E). Pronotum elliptical (PL to PW ratio 1.19), markedly wider than head, also distinctly longer. Anterior region near anterior angles deflexed and narrowed, lateral margins widened posteriad to basal 1 / 3 (with the widest at apical 1 / 3), then convergent backwards, anterior and posterior angles broadly rounded. Dorsal surface glossy extensively with micropunctures, but without any microsculpture. Each side of the midline with symmetrical five large lateral punctures, with additional 13 – 15 punctures near each margin (including anterior, lateral and posterior margins). Prosternum bearing a distinct transverse ridge between basisternum and furcasternum, which angularly backwards (Fig. 4 E). Pterothorax (Fig. 4 F, N). Mesoscutellum triangular and small (Fig. 4 N), surface shiny, bearing obvious microstriae, but without any punctures. The transverse ridge on mesoventrites substraight (Fig. 4 F); the longitudinal ridge distinct; the paired oblique furrows short but deep; the carina (limiting mesocoxal cavity) slightly arcuate. The discrimen region on metaventrite normally elevated. Elytra (Fig. 4 C). Transverse and short (EL to EW ratio 0.84), distinctly wider than pronotum, and shorter. Humeri well developed, lateral margins divergent posteriorly, posterior margin not rounded, but obliquely truncate. Dorsal integument shiny, flattened, without microsculpture; each elytron with a row of 5 punctures along suture, a row of 5 punctures in median, also with 4 – 5 rows of punctures on deflexed portion. Legs (Fig. 4 M). First four segments of protarsi stout, wider than those of meso- and metatarsi. Last segment of protarsi longer than II – IV together; that of meso- slightly longer than II and III together; that of meta- longer than II and III together. Abdomen (Fig. 4 O) Broadest at segment V. Tergites III – VII covered with dense and obvious transverse microstriae; each tergite with dense yellow pubescence, also with a row of darker and coarser setae at posterior margin of III – VI. Tergites III – VII with a basal impression near anterior margin, respectively, each one only possessing one basal transverse carina; surface shiny, bearing polygonal reticulum, but without any punctures. Tergite VII possessing a transparent palisade fringe on posterior margin. All abdominal sternites shiny, with microstriae and punctures as those on tergites. Male (Fig. 4 - 1 A – G, K – M). Posterior margin of tergite VIII sinuate (Fig. 4 - 1 A), that of sternite VIII slightly triangularly incised (Fig. 4 - 1 B), sometimes more distinctly (Fig. 4 - 1 K), also bearing coarser setae. Tergite IX symmetrical, connected mediobasally (Fig. 4 - 1 C), sharpened apically. Sternite IX symmetrical, widest near the middle, with paired longitudinal furrow near each lateral margin, and basal margin wide and subtruncate (Fig. 4 - 1 D); sometimes basal 1 / 2 distinctly wider (Fig. 4 - 1 M). Tergite X symmetrical, triangular, with curved basal margin (Fig. 4 - 1 C). Aedeagus normally sclerotized, elongate, large in size (Fig. 4 E, F), ca. 0.6 mm long, ventral surface with scattered dots. Parameres symmetrical and slightly curved, thin and slightly shorter than 1 / 3 times the length of basal bulb; paired sclerites attached to base of parameres in ventral surface. Internal sac possessing inner sclerites composed of three parts: two paired sclerites at top (one soft and light color; the other horn-like shaped and black color), paired C-shaped sclerites in centre, and also rectangular sclerite in centre (Fig. 4 - 1 E, L). Sperm pump spirally coiled, thinned toward apex (Fig. 4 - 1 G). Female (Fig. 4 - 1 H – J, N – O). Tergite IX bearing numerous setae, elongate and sharply pointed apically (Fig. 4 - 1 O). Sternite IX symmetrical, each integrated, and with an observable concaved margin on apical 1 / 3 (Fig. 4 - 1 H). Tergite X nearly triangular, basal margin curved (Fig. 4 - 1 I). Sternite X slender and rhomboidal (Fig. 4 - 1 N). Outer capsule of spermatheca with parallel sides and sharp apical margin, and tube with two apical dilated lobes (Fig. 4 - 1 J).	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFE5B254FF6AFB173020FCC0.taxon	distribution	Distribution. Fujian, Hainan, Zhejiang, Guangxi, Yunnan.	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFE5B254FF6AFB173020FCC0.taxon	etymology	Etymology. The specific epithet is derived from the combination of Latin adverb “ bis ” (twice) and the adjective “ cornutus ” (having horns), which refer to the paired horn-like sclerites of internal sac.	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFE5B254FF6AFB173020FCC0.taxon	discussion	Remarks. The short, paired parameres and paired sclerites present on the apex of aedeagus are the similar to those of D. conicollis (Motschulsky), but the shape of the aedeagus is elongate, and the apical sclerites are hornlike. The body size is bigger, with a longer head; the outer capsule of the female spermatheca possesses parallel sides and a sharp apex. The shape of the spermatheca in D. punctipennis (Motschulsky) (Female, 1 syntype examined, MMZ) is similar to this species, but the ratio of spermatheca length / width in D. bicornutus is smaller and the outer capsule is parallel-sided.	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFE9B251FF6AFCED33D2F969.taxon	description	(Fig. 3 D; Fig. 5 A – P; Fig. 5 - 1 A – J)	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFE9B251FF6AFCED33D2F969.taxon	materials_examined	Type material. Holotype: male, CHINA: Guangxi: Napo co., Defushuiyuanlin Nature Reserve (E 105.7965 °, N 23.3148 °), 1400 m, 0 4. IV. 1998, Zhou Haisheng collected (IZ-CAS). Paratypes: Guangxi: Napo co., Defushuiyuanlin Nature Reserve (E 105.7965 °, N 23.3148 °), 3 males, 4 females, 1400 m, 0 4. IV. 1998, Zhou Haisheng collected (IZ-CAS).	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFE9B251FF6AFCED33D2F969.taxon	description	Description. Measurement (n = 5). BL = (4.2 – 4.3) mm, FL = (2.1 – 2.2) mm, HL = (0.5 – 0.6) mm, HW = (0.4 – 0.5) mm, PL = (0.7 – 0.8) mm, PW = (0.6 – 0.7) mm, EL = (0.6 – 0.7) mm, EW = (0.8 – 0.9) mm. Body nearly fusiform shaped, medium sized (Fig. 3 D). Body dark brown, abdominal intersegmental membrane yellowish brown. Legs entirely brown, except tarsi lighter. Antennae light brown. Maxillary and labial palpi yellowish brown. Head (Fig. 5 A, D). Rounded triangular in shape (HL to HW ratio 1.23), tempora (behind eyes) distinctly widened backwards, posterior angles rounded. Dorsal integument shiny, extensively bearing microstriae, without punctures around each eye, but bearing scattered smaller punctures on deflexed portion of tempus and near the posterior margin. Each side of cranium with characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three lateral punctures near dorsal margin of eye, also with occipital puncture at the basal 1 / 4, but temporal puncture unidentified; besides, disc surface bearing two additional paired punctures. Frontal furrows and anteocular furrows absent, a small round elevation present between eyes. Eye medium sized, distinctly protruding laterad, longitudinal diameter slightly shorter than half the length of tempora (eye: tempora = 0.16: 0.34 mm). Epistoma not flattened, abruptly oblique, anterior margin emarginate medially. Distance between antennal insertions 0.17 mm being distinctly longer than distance from antenna to eye (0.09 mm). Ventral surface (Fig. 5 D) with scattered punctures, interspaces between them over 4 punctures’ diameter. Gular sutures deep, gradually convergent, but not confluent, then divergent to neck region. Antennae (Fig. 5 G). Total length 1.2 mm. Scape rod-shaped, slightly thickened apically, 0.17 mm, shorter than two subsequent antennomeres combined; antennomere II elongate, 0.12 mm; III 0.13 mm, slightly longer than II; IV 0.09 mm, IV – X becoming shortened in length; XI 0.14 mm, distinctly longer than X. Mouthparts (Fig. 5 H – J). Labrum not transverse, nearly pentagon; lateral margin straight, not in waved shape; anterior margin distinctly shorter than lateral margins (Fig. 5 H). Mandibles falciform, left one with one sharp tooth on inner edge, right one without tooth (Fig. 5 I – J). Neck (Fig. 5 K – L). Cylindrical, shiny, first half of dorsal surface depressed, with a distinct groove (Fig. 5 K); only bearing shallow microsculpture, width of 0.17 mm, slightly wider than 1 / 3 width of head. Ventral surface with gular sutures to form obvious triangular region, with deep transverse median furrow (Fig. 4 L). Prothorax (Fig. 5 B, E). Pronotum elliptical (PL to PW ratio 1.09), wider and longer than head. Anterior region near anterior angles deflexed, lateral margins paralleled, not widened from anterior angles to the middle, also not convergent backwards, anterior and posterior angles broadly rounded. Dorsal surface glossy extensively with micropunctures, but without any microsculpture. Each side of the midline with symmetrical five large lateral punctures, with additional 13 – 15 punctures near each margins (including anterior, lateral and posterior margins). Prosternum bearing an observable transverse ridge between basisternum and furcasternum, angularly backwards (Fig. 5 E). Pterothorax (Fig. 5 F, N). Mesoscutellum slightly sharp triangular and small (Fig. 5 N), surface shiny, bearing some light microstriae, but without any punctures. The transverse ridge on mesoventrites substraight (Fig. 5 F); the longitudinal ridge distinct; the paired oblique furrows not distinct; the carina (limiting mesocoxal cavity) possessing wave-shaped median part. The discrimen region on metaventrite distinctly elevated. Elytra (Fig. 5 C). Transverse and short (EL to EW ratio 0.81), shorter and wider than pronotum. Humeri well developed, lateral margins divergent posteriorly, hind margin not rounded, but obliquely truncate. Dorsal integument shiny, flattened, without microsculpture; each elytron with a row of 5 punctures along suture, a row of 5 punctures in median, also with 4 – 5 rows of punctures on deflexed portion. Legs (Fig. 5 M). First four segments of protarsi stout, wider than those of meso- and metatarsi slender. Last segment of protarsi longer than II – IV together; that of meso- as long as II and III together; that of meta- longer than II and III together. Abdomen (Fig. 5 O). Broadest at segment V. Tergites III – VII covered with dense and obvious transverse microstriae; each tergite with dense and brown pubescence, posterior margin of III – VI without darker and coarser setae. Tergites III – VII with a basal impression near anterior margin, respectively, each one only possessing one basal transverse carina; surface shiny, without any punctures or microsculpture. Tergite VII possessing a transparent palisade fringe on posterior margin. All abdominal sternites shiny, with microstriae and punctures as those on tergites. Male (Fig. 5 - 1 A – G). Posterior margin of tergite VIII arcuate backwards (Fig. 5 - 1 A), that of sternite VIII distinctly bisinuate (Fig. 5 - 1 B). Tergite IX symmetrical, connected mediobasally, sharpened apically. Sternite IX symmetrical, widest near the apical 1 / 3, basal margin deeply emarginate, apical margin nearly subtruncate (Fig. 5 - 1 D). Tergite X symmetrical, in reverse trapezoidal shape (Fig. 5 - 1 C). Aedeagus large sized and bottle-shaped (Fig. 5 E, F), ca. 0.8 mm long, ventral surface normally sclerotized, with part membrane; dorsal surface fully membranous. Parameres symmetrical and soft, extraordinarily thin and long, rolled from ventral to dorsal side (Fig. 5 - 1 E, F). Internal sac membranous, without sclerotized structures. Sperm pump folded basally, spiral toward apex (Fig. 5 - 1 G). Female (Fig. 5 - 1 H – J). Tergite IX bearing numerous setae, elongate and sharply pointed apically. Sternite IX symmetrical, each integrated, and with a deep concaved margin in the middle (Fig. 5 - 1 H). Tergite X broad, basal margin curved (Fig. 5 - 1 I). Sternite X missing. Tube of spermatheca short, with an apical circle (Fig. 5 - 1 J).	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFE9B251FF6AFCED33D2F969.taxon	distribution	Distribution. Guangxi.	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFE9B251FF6AFCED33D2F969.taxon	etymology	Etymology. The specific epithet is derived from the Latin word “ ampullaceus ” (a flask swelled in middle) and refers to the bottle-shaped aedeagus.	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFE9B251FF6AFCED33D2F969.taxon	discussion	Remarks. Although this new species is similar in the round elevation between the eyes to D. pulchellus Cameron, it can be easily recognized by the dilated protarsi. This new species is distinguished from its congeners by the combination of the following characters: 1) sparsely distributed punctures on head; 2) different number of mandible teeth; 3) neck with both a dorsal and ventral groove; 4) transverse elytra; 5) aedeagus large sized and bottle-shaped (Fig. 5 E, F); parameres symmetrical and soft, extraordinarily thin and long, curving from ventral to dorsal side (Fig. 5 - 1 E, F).	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFECB24DFF6AF8B5377CFCED.taxon	description	(Fig. 3 E; Fig. 6 A – O; Fig. 6 - 1 A – L)	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFECB24DFF6AF8B5377CFCED.taxon	materials_examined	Type material. Holotype: male, CHINA: Hainan: Ledong co., Wuzhishan City (E 109.5169 °, N 18.7751 °), 785 m, 08. VII. 2004, Wu Jie & Chen Yongjie collected (IZ-CAS). Paratypes: Yunnan: Mengla co.: Longmen village, Xiaoniupeng (E 101.3252 °, N 21.3095 °), one male, 4 females, 1010 m, 06. X. 2010, Zhou Yu-Lingzi collected (IZ- CAS); Menglun Xipian (E 101.2489 °, N 21.9353 °), one female, 720 m, 16. II. 2004, Wu Jie collected (IZ-CAS); Menglun (E 101.2490 °, N 21.9375 °), one female, 860 m, 11. II. 2004, Wu Jie & Bai Daiyuan collected (IZ-CAS); Hainan: Ledong co., Wuzhishan City (E 109.5169 °, N 18.7751 °), one female, 770 m, 07. VII. 2004, Wu Jie & Chen Yongjie collected (IZ-CAS); Lingshuilizu autonomous co., Diaoluoshan (E 109.9160 °, N 18.6598 °), one female, 920 m, 28. VII. 2004, Wu Jie & Chen Yongjie (IZ-CAS).	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFECB24DFF6AF8B5377CFCED.taxon	description	Description. Measurement (N = 5). BL = (3.9 – 4.0) mm, FL = (2.2 – 2.3) mm, HL = (0.5 – 0.6) mm, HW = (0.4 – 0.5) mm, PL = (0.7 – 0.8) mm, PW = (0.6 – 0.7) mm, EL = (0.7 – 0.8) mm, EW = (0.8 – 0.9) mm. Body nearly fusiform shaped, small sized (Fig. 3 E). Body entirely dark brown; abdominal intersegmental membrane yellowish brown. Antennae dark brown. Leg, maxillary and labial palpi yellowish brown. Head (Fig. 6 A, D). Rectangular in shape (HL to HW ratio 1.21), tempora (behind eyes) distinctly widened backwards, posterior angles rounded. Dorsal integument shiny, extensively bearing microsculpture mixed by polygonal reticulum (mainly on frontal region) and microstriae, without punctures around each eye, but bearing scattered smaller punctures on deflexed portion of tempus and near the posterior margin. Each side of cranium with characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three lateral punctures near dorsal margin of eye, also with occipital puncture and temporal puncture near posterior margin; besides, disc surface bearing two additional paired punctures. Frontal furrows and anteocular furrows absent, a little round elevation present between eyes. Eye medium sized, distinctly protruding laterad, longitudinal diameter equal to half the length of tempora (eye: tempora = 0.17: 0.34 mm). Epistoma not flattened, abruptly oblique, anterior margin subtruncate. Distance between antennal insertions 0.15 mm being distinctly longer than distance from antenna to eye (0.07 mm). Ventral surface (Fig. 6 D) with scattered punctures, interspaces between them over 4 punctures’ diameter. Gular sutures deep, confluent at basal 1 / 4, then extended to neck region. Antennae (Fig. 6 G). Total length 1.1 mm. Scape rod-shaped, slightly thickened apically, 0.19 mm, slightly shorter than two subsequent antennomeres combined; antennomeres II and III equal in length of 0.10 mm; IV – VII of rectangular shape, subequal in length of 0.09 mm; VIII of square shape, IX and X transverse and becoming shortened, sometimes VIII – X square in shape and with same length; XI 0.14 mm, distinctly longer than X. Mouthparts (Fig. 6 H – J). Labrum transverse, anterior margin slightly emarginate medially; lateral margin arcuate (Fig. 6 H). Mandibles falciform, left one with three small teeth on inner edge, right one with two sharper teeth (Fig. 6 I – J). Neck (Fig. 6 K – L). Cylindrical, shiny, apical 1 / 3 of dorsal surface depressed, with a distinct groove; only bearing shallow microsculpture, width of 0.17 mm, slightly wider than 1 / 3 width of head (Fig. 6 K). Ventral surface with gular sutures to form obvious triangular region, with deep transverse median furrow (Fig. 6 L). Prothorax (Fig. 6 B). Pronotum rectangular (PL to PW ratio 1.19), distinctly wider and longer than head. Anterior region near anterior angles deflexed, lateral margins arcuate, with widest at basal 1 / 3. Dorsal surface glossy extensively with micropunctures, but without any microsculpture. Each side of the midline with symmetrical five large lateral punctures, with additional 13 – 15 punctures near each margins (including anterior, lateral and posterior margins). Prosternum bearing an observable transverse ridge between basisternum and furcasternum, angularly backwards (Fig. 6 E). Pterothorax (Fig. 6 F, N). Mesoscutellum slightly sharp triangular and medium sized (Fig. 6 N), surface shiny, bearing some light microstriae, but without any punctures. The transverse ridge on mesoventrites wave-shaped (Fig. 6 F); the longitudinal ridge distinct; the paired oblique furrows long but not deep; the carina (limiting mesocoxal cavity) possessing median horizontal part. The discrimen region on metaventrite distinctly elevated. Elytra (Fig. 6 C, F). Rectangular and long (EL to EW ratio 0.84), slightly longer than pronotum, but distinctly wider. Humeri well developed, lateral margins divergent posteriorly, hind margin not rounded, but obliquely truncate. Dorsal integument shiny, flattened, without microsculpture; each elytron with a row of 5 punctures along suture, a row of 5 punctures in median, also with 4 – 5 rows of punctures on deflexed portion. Legs. First four segments of protarsi stout, wider than those of meso- and metatarsi slender. Last segment of protarsi longer than II – IV together; that of meso- as long as II and III together; that of meta- longer than II and III together. Abdomen (Fig. 6 O). Broadest at segment V. Tergites III – VII covered with dense and transverse microstriae; each tergite with dense and brown pubescence, posterior margin of III – VI without darker and coarser setae. Tergites III – VII with a basal impression near anterior margin, respectively, each one only possessing one basal transverse carina; surface shiny, with microsculpture composed of polygonal reticulum. Tergite VII possessing a transparent palisade fringe on posterior margin. All abdominal sternites shiny, with microstriae and punctures as those on tergites. Male (Fig. 6 - 1 A – G). Posterior margin of tergite VIII arcuate backwards (Fig. 6 - 1 A), that of sternite VIII distinctly bisinuate (Fig. 6 - 1 B). Tergite IX symmetrical, connected mediobasally, sharpened apically. Sternite IX symmetrical, widest at middle, basal margin deeply emarginate, apical margin nearly subtruncate (Fig. 6 - 1 D). Tergite X symmetrical and triangular (Fig. 6 - 1 C). Aedeagus medium sized (Fig. 6 E, F), ca. 0.4 mm long, dorsal and ventral surface bearing obvious membranous part, apical margin bisinuate, basal margin waved. Parameres symmetrical, extraordinarily thin and long, cross at dorsal side (Fig. 6 - 1 E, F). Internal sac with symmetrical and sclerotized structures. Sperm pump short and folded basally, thinned toward apex (Fig. 6 - 1 G). Female (Fig. 6 - 1 H – L). Tergite IX bearing numerous setae, stout, with a sharp apical point (Fig. 6 - 1 K). Sternite IX symmetrical, each two-segmented (Fig. 6 - 1 H). Tergite X nearly triangular, basal margin distinctly curved (Fig. 6 - 1 I). Sternite X slender and pentagonal (Fig. 6 - 1 L). Tube of spermatheca long, with an apical circle (Fig. 6 - 1 J).	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFECB24DFF6AF8B5377CFCED.taxon	distribution	Distribution. Hainan, Yunnan.	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFECB24DFF6AF8B5377CFCED.taxon	etymology	Etymology. The specific epithet is derived from the combinations of Latin adverb “ bis ” (twice) and the adjective “ segmentatus ” (composed of strips, bands), and refers to female sternite IX seemed two segmented.	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFECB24DFF6AF8B5377CFCED.taxon	discussion	Remarks. This new species is distinguished from its congeners by the following characteristics: 1) polygonal reticulum on frontal region; 2) a little round elevation on head; 3) longer elytra; 4) number of the mandible teeth; 5) transverse ridge on mesoventrite wave-shaped (Fig. 6 F); 6) dorsal and ventral surface of aedeagus bearing obvious membranous part, apical margin bisinuate, basal margin waved. Although the shape of antennae is similar to that of D. antennatus (Motschulsky) (per. comm. with Jiri Janak), the paired parameres are extremely longer, and the outer margin of aedeagus is quite different: basal margin as waved-like, and apical margin sharply concaved.	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFF0B24EFF6AFC3C313EFA15.taxon	description	(Fig. 3 F; Fig. 7 A – O; Fig. 7 - 1 A – F)	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFF0B24EFF6AFC3C313EFA15.taxon	materials_examined	Type material. Holotype: male, CHINA: Fujian: Jiangle co., Longxishan, Yujiaping (E 117.2980 °, N 26.4911 °), 21. VI. 1991, Wang Huifu collected (IZ-CAS). Paratype: Fujian: Jiangle co., Longxishan, Yujiaping (E 117.2980 °, N 26.4911 °), one male, 21. VI. 1991, Wang Huifu collected (IZ-CAS).	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFF0B24EFF6AFC3C313EFA15.taxon	description	Description. Measurement. BL = (3.4 – 3.5) mm, FL = (1.6 – 3.5) mm, HL = (0.4 – 0.5) mm, HW = (0.4 – 0.5) mm, PL = (0.5 – 0.6) mm, PW = (0.4 – 0.5) mm, EL = (0.5 – 0.6) mm, EW = (0.5 – 0.6) mm. Body nearly fusiform shaped, small sized (Fig. 3 F). Body entirely brown, except antennae, legs, maxillary and labial palpi light brown. Head (Fig. 7 A, D). Nearly triangular (HL to HW ratio 0.96), tempora (behind eyes) extraordinarily widened backwards, posterior angles rounded. Dorsal integument shiny, extensively bearing microsculpture mixed by polygonal reticulum (mainly on frontal region) and microstriae, without punctures around each eye, but bearing scattered smaller punctures on deflexed portion of tempus and larger punctures near the posterior margin. Each side of cranium with characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three lateral punctures near dorsal margin of eye, but occipital puncture and temporal puncture unidentified; besides, disc surface bearing two additional paired punctures. Frontal furrows and anteocular furrows absent, without elevation between eyes. Eye small, smaller than 1 / 3 length of temple (eye: tempora = 0.08: 0.29 mm), flat, not protruding laterad. Epistoma broadly emarginate. Distance between antennal insertions 0.15 mm being distinctly longer than distance from antenna to eye (0.07 mm). Ventral side shiny and smooth, each side with two large-sized punctures along lateral margin, 3 – 4 small punctures near base. Gular plate distinctly broad, gular sutures widely separated (Fig. 7 D). Antennae (Fig. 7 G). Total length 1.0 mm. Scape rod-shaped, slightly thickened apically, 0.19 mm, longer than two subsequent antennomeres combined; antennomere II elongate, subequal length to III, 0.08 mm; IV and V rectangular, subequal in length (0.07 mm); VI to X square in shape, slightly shorter than length of V; XI 0.10 mm, distinctly longer than X. Mouthparts (Fig. 7 H – J). Labrum not transverse, but nearly pentagon (Fig. 7 H). Mandibles falciform and slender; left one with two sharp teeth on inner edge, right one also bearing two closer teeth (Fig. 7 I – J). Neck (Fig. 7 K – L). Cylindrical, width of 0.14 mm, narrower than 1 / 3 width of head; shiny, only bearing observable microsculpture, with a bisinuate basal furrow near posterior margin (Fig. 7 K). Ventral surface missing triangular region, with observable transverse median furrow (Fig. 7 L). Prothorax (Fig. 7 H – J). Pronotum quite elongate (PL to PW ratio 1.25), distinctly longer than head, but of same width. Anterior region near anterior angles not very deflexed, lateral margins slightly divergent backwards, with widest at basal 1 / 3, posterior angles broadly rounded. Dorsal surface glossy extensively with micropunctures, but without any microsculpture. Each side of the midline with symmetrical five large lateral punctures, with additional 13 – 15 punctures near each margins (including anterior, lateral and posterior margins). Prosternum bearing observable transverse ridge between basisternum and furcasternum, slightly arcuate backwards (Fig. 7 E). Pterothorax (Fig. 7 F, N). Mesoscutellum slightly sharp triangular and small (Fig. 7 N), surface shiny, bearing some light microstriae, but without any punctures. The transverse ridge on mesoventrites in wave shape (Fig. 7 F); the longitudinal ridge not distinct; the paired oblique furrows short but deep; the carina limiting mesocoxal cavity curved, not wave-shaped. The discrimen region on metaventrite normally elevated. Elytra (Fig. 7 C). Transverse and short (EL to EW ratio 0.86), shorter and wider than pronotum. Humeri well developed, lateral margins divergent posteriorly, hind margin not rounded, but obliquely truncate. Dorsal integument shiny, flattened, without microsculpture; each elytron with a row of 5 punctures along suture, a row of 5 punctures in median, also with 4 – 5 rows of punctures on deflexed portion. Legs (Fig. 7 M). First four segments of protarsi heart-shaped, dilated, and those of meso- and metatarsi slender. Last segment of protarsi equal to length of II – IV, that of meso-, meta- equal to length of each II – III. Abdomen (Fig. 7 O). Broadest at segment V. Tergites III – VII covered with dense and obvious transverse microstriae; each tergite with dense and brown pubescence, posterior margin of III – VI with coarser brown setae. Tergites III – VII with a basal impression near anterior margin, respectively, each one only possessing one basal transverse carina; surface shiny, without any punctures or microsculpture. Tergite VII possessing a transparent palisade fringe on posterior margin. All abdominal sternites shiny, with microstriae and punctures as those on tergites. Male (Fig. 7 - 1 A – F). Posterior margin of tergite VIII slightly arcuate backwards (Fig. 7 - 1 A), that of sternite VIII broadly emarginate (Fig. 7 - 1 B). Tergite IX symmetrical, connected mediobasally, sharpened apically. Sternite IX symmetrical, widest at basal 1 / 3, basal margin emarginate, a small tooth at each hind angle (Fig. 7 - 1 D). Tergite X in reuleaux triangular shape (Fig. 7 - 1 C). Aedeagus small sized (Fig. 7 E, F), 0.35 mm long, dorsal and ventral surface mostly membranous. Parameres symmetrical, small and horizontal (Fig. 7 - 1 E). Internal sac with asymmetrical and sclerotized structures (Fig. 7 - 1 E). Sperm pump relatively long, folded basally, and coiled apically (Fig. 7 - 1 F). Female. Unknown.	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFF0B24EFF6AFC3C313EFA15.taxon	etymology	Etymology. The specific epithet is derived from the Latin adjective word “ membranaceus ” (of membrane) and refers to the membranous aedeagus.	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFF0B24EFF6AFC3C313EFA15.taxon	distribution	Distribution. Fujian.	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFF0B24EFF6AFC3C313EFA15.taxon	discussion	Remarks. Although this new species is similar in head shape and coloration to the Indian species D. ochraceus Cameron (Female, 1 syntype examined, BMNH), it can easily be distinguished by the following combination of characters: 1) distinctly dilated tempora; 2) lack of elevation on head; 3) smaller eyes; 4) widely separated gular sutures; 5) elongate mandibles; 6) fully membranous aedeagus.	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFF3B249FF6AF9A43678FF00.taxon	description	(Fig. 8) Motschulsky, 1858: 659 (Rhegmatocerus; Type locality: Indes orientales); Bernhauer and Schubert, 1914: 319 (catalog); Bernhauer, 1922: 231 (Formosa); Cameron, 1932: 46 (Penang; Malay Peninsula); Shibata, 1973: 130 (Taiwan); Coiffait, 1982: 27 (Nepal); Herman, 2001: 2444 (catalog); Smetana, 2004: 624 (Taiwan; Nepal); Löbl & Löbl, 2015: 1007 (Palaearctic catalog; Australian Region). Syn.: Diochus indicus Kraatz, 1859: 113 (Type locality: India orientalis); Bernhauer and Schubert, 1914: 319 (synonym of D. antennatus); Cameron, 1931: 360 (New Guinea); Cameron, 1932: 46 (synonym of D. antennatus); Smetana, 2004: 624 (synonym of D. antennatus); Löbl & Löbl, 2015: 1007 (Palaearctic catalog; synonym of D. antennatus).	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFF3B249FF6AF9A43678FF00.taxon	materials_examined	Material examined: None.	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFF3B249FF6AF9A43678FF00.taxon	distribution	Distribution. China (Taiwan); “ East Indies ”, Nepal, Malaysia, Indonesia, New Guinea. Remarks. The syntypes are missing (per. comm. with Dr. Arnaldo Bordoni, Dr. Jiri Janak, and Dr. Aleksey Gusakov). However, the characters of the aedeagus and male genital segments (Fig. 8, cf. Jiri’s drafted drawing on a male syntype) could be easily recognized as different from congeneric species.	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFF5B245FF6AFF2833C2FB10.taxon	description	(Fig. 3 C; Fig. 9 A – O; Fig. 9 - 1 A – L) Motschulsky, 1858: 658 (Rhegmatocerus; Type locality: Indes orientales); Bernhauer, 1902: 32 (Diochus; Ceylon); Bernhauer and Schubert, 1914: 319 (catalog); Bernhauer, 1922: 231 (Formosa); Cameron, 1932: 46 (Ceylon; India); Shibata, 1973: 130 (Taiwan); Coiffait, 1977: 206 (Bhutan); Coiffait, 1981: 329 (Andaman Islands); Coiffait, 1982: 27 (Nepal); Herman, 2001: 2445 (catalog); Smetana, 2004: 624 (Hongkong, Taiwan; Nepal, India, Bhutan); Löbl & Löbl, 2015: 1007 (Palaearctic catalog; Hongkong, Taiwan; Nepal, India, Bhutan). Syn.: Diochus major Kraatz, 1859: 113 (Type locality: India orientalis; Ceylan); Bernhauer, 1902: 32 (synonym of D. conicollis); Bernhauer and Schubert, 1914: 319 (synonym of D. conicollis); Cameron, 1932: 46 (synonym of D. conicollis); Smetana, 2004: 624 (synonym of D. conicollis); Löbl & Löbl, 2015: 1007 (Palaearctic catalog; synonym of D. antennatus).	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFF5B245FF6AFF2833C2FB10.taxon	materials_examined	Type material examined: Lectotype [designated here]: male, “ ♂ ” / “ Ind. or ” / “ Diochus conicollis ”, “ ‘ (Motschulsky, 1858) ’ ”, “ LECTOTYPE ”, “ Det. Zhou Yulingzi 2016 ”; Paralectotypes: female, “ Rhegmatocerus ”, “ conicollis ”, “ Mots ”, “ Ind. or. Birma ” / “ Diochus conicollis ”, “ ‘ (Motschulsky, 1858) ’ ”, “ Det. Zhou Yulingzi 2016 ”; female, “ ♀ ” / “ Ind. or ” / “ Diochus conicollis ”, “ ‘ (Motschulsky, 1858) ’ ”, “ Det. Zhou Yulingzi 2016 ”. (ZMM).	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFF5B245FF6AFF2833C2FB10.taxon	description	Description. Measurement (n = 3). BL = (4.9 – 5.1) mm, FL = (2.0 – 2.2) mm, HL = (0.5 – 0.6) mm, HW = (0.4 – 0.5) mm, PL = (0.7 – 0.8) mm, PW = (0.6 – 0.7) mm, EL = (0.6 – 0.7) mm, EW = (0.7 – 0.8) mm. Body nearly fusiform shaped, large sized (Fig. 3 C). Body dark brown, except elytra darker and abdominal intersegmental membrane yellowish brown (Fig. 9 O). Legs brown, tarsi distinctly lighter. Antennae brown, apex of each antennomere lighter. Maxillary and labial palpi brown. Head (Fig. 9 A, D). Oblong in shape (HL to HW ratio 1.3), tempora (behind eyes) straight and paralleled, posterior angles rounded. Dorsal integument shiny, extensively bearing micropunctures and microstriae, also with 5 – 7 irregular, scattered punctures around each eye, and numerous smaller punctures on deflexed portion of tempus and near the posterior margin. Each side of cranium with characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three ocular punctures near dorsal margin of eye, temporal puncture at the basal 1 / 5, also with occipital puncture at the basal 1 / 4; in addition, disc surface bearing two additional paired punctures with front pair closer. Frontal furrows and anteocular furrows absent, a longitudinal linear elevation present between eyes. Eye medium sized, equal to half the length of tempora (eye: tempora = 0.15: 0.30 mm), distinctly protruding laterad. Epistoma not flattened, abruptly oblique, anterior margin emarginate medially. Distance between antennal insertions 0.15 mm being distinctly longer than distance from antenna to eye (0.09 mm). Ventral surface (Fig. 9 D) with scattered punctures, interspaces between them ca. 4 punctures’ diameter. Gular sutures deep, confluent at near base. Antennae (Fig. 9 G). Total length 1.5 mm. Scape rod-shaped, slightly thickened apically, 0.20 mm, shorter than two subsequent antennomeres combined; antennomere II elongate, 0.10 mm; III 0.15 mm, distinctly longer than II; IV 0.11 mm, IV to X gradually shortened, IV slightly longer than X; XI 0.12 mm, distinctly longer than X. Mouthparts (Fig. 9 H – J, P). Labrum transverse (Fig. 9 H), lateral margin rounded. Mandibles falciform, left one with three small teeth on inner edge, right one with two small tooth (Fig. 9 I – J). Neck (Fig. 9 K – L). Surface shiny and nearly trapezoidal shaped, only bearing shallow microsculpture, with a bisinuate basal furrow near posterior margin (Fig. 9 K), width of 0.19 mm, obviously wider than 1 / 3 width of head. Ventral surface with deep gular sutures to form obvious triangular region, which missing transverse median furrow (Fig. 9 L). Prothorax (Fig. 9 B, E). Pronotum elliptical (PL to PW ratio 1.2), distinctly longer and wider than head. Anterior region near anterior angles deflexed and narrowed, lateral margins widened posteriad to basal 1 / 3 (with the widest at middle), then convergent backwards, anterior and posterior angles broadly rounded. Dorsal surface glossy extensively with micropunctures, but without any microsculpture. Each side of the midline with symmetrical five large lateral punctures, with additional 13 – 15 punctures near each margins (including anterior, lateral and posterior margins). Prosternum bearing observable transverse ridge between basisternum and furcasternum, which angularly backwards (Fig. 9 E). Pterothorax (Fig. 9 E – F, N). Mesoscutellum triangular and small (Fig. 9 N). Surface shiny, bearing obvious microstriae, but without any punctures. The transverse ridge on mesoventrites substraight (Fig. 9 F); the longitudinal ridge distinct; the paired oblique furrows short but deep; the carina (limiting mesocoxal cavity) slightly arcuate. The discrimen region on metaventrite normally elevated. Elytra (Fig. 9 C). Transverse and short (EL to EW ratio 0.85), distinctly shorter than pronotum, but wider. Humeri well developed, lateral margins divergent posteriorly, hind margin not rounded, but obliquely truncate. Dorsal integument shiny, flattened, without microsculpture; each elytron with a row of 5 punctures along suture, a row of 5 punctures in median, also with 4 – 5 rows of punctures on deflexed portion. Legs (Fig. 9 M). First four segments of protarsi stout, wider than those of meso- and metatarsi slender. Last segment of protarsi longer than II – IV together; that of mesotarsi meta- longer than II and III together. Abdomen (Fig. 9 O). Broadest at segment V. Tergites III – VII covered with dense and obvious transverse microstriae; each tergite with dense yellow pubescence, with a row of darker and coarser setae at posterior margin of III – VI. Tergites III – VII with a basal impression near anterior margin, respectively, each one only possessing one basal transverse carina; surface shiny, bearing polygonal reticulum, but without any punctures. The intersegmental membrane bearing transparent yellow fenestra. All abdominal sternites shiny, with microstriae and punctures as those on tergites. Male (Fig. 9 - 1 A – H). Posterior margin of tergite VIII arcuate (Fig. 9 - 1 A), that of sternite VIII slightly emarginate medially (Fig. 9 - 1 B), also bearing coarser setae. Tergite IX symmetrical, connected mediobasally (Fig. 9 - 1 C), sharpened apically. Sternite IX symmetrical, widest near the middle, with paired longitudinal furrow near each lateral margin, and basal margin wide and subtruncate (Fig. 9 - 1 D). Tergite X symmetrical, small and triangular (Fig. 9 - 1 C). Aedeagus asymmetrical, fully membranous ventrally and normally sclerotized dorsally, large in size (Fig. 9 E – G), ca. 0.77 mm long. Parameres symmetrical and slightly curved, thin and slightly 1 / 3 times the length of basal bulb. Internal sac possessing inner sclerites composed of two parts: two paired sclerites at top (one soft and light color; the other X-shaped and brown color), some black sclerites in centre. Sperm pump coiled, thinned toward apex (Fig. 9 - 1 H). Female (Fig. 9 - 1 I – L). Tergite IX bearing numerous setae, elongate and bluntly pointed apically (Fig. 9 - 1 L). Sternite IX symmetrical, each integrated, and with an observable concaved margin in middle (Fig. 9 - 1 I – J). Tergite X nearly triangular, basal margin curved (Fig. 9 - 1 L). Sternite X missing. Outer capsule of spermatheca apically swollen, and with more rounded apex margin; tube long, with an apical dilated lobe (Fig. 9 - 1 K).	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFF5B245FF6AFF2833C2FB10.taxon	distribution	Distribution. China (Hongkong, Taiwan); Sri Lanka, India, Bhutan, Andaman Islands.	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFF5B245FF6AFF2833C2FB10.taxon	discussion	Remarks. This species is distinguished from its congeners by the combination of the following characters: 1) left mandible with three small teeth on inner edge, right with two small tooth (Fig. 9 I – J); 2) dark coarser setae on the posterior margin of abdominal sternite VIII; 3) pear-like aedeagus with apical paired sclerites; 4) outer capsule of spermatheca swollen.	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFF8B244FF6AFAA83396FAAD.taxon	description	(Fig. 3 A) Cameron, 1930: 206 (Type locality: Kobe); Scheerpeltz, 1933: 1324 (catalog); Nakane, 1963: 90 (habitus photograph; Japan); Herman, 2001: 2446 (Diochus; catalog); Smetana, 2004: 624 (Palaearctic catalog; Jilin, Liaoning; Japan, South Korea); Löbl & Löbl, 2015: 1007 (Palaearctic catalog; Jilin, Liaoning; Japan, South Korea).	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFF8B244FF6AFAA83396FAAD.taxon	materials_examined	Type material examined: female, with labels as follows: “ Holo-, Typus ” / “ D., japonicus Came, TYPE ” / “ HOLOTYPE, Diochus, japonicus, cameron, 1930, det. R. G. Booth 2013 ” / “ M. Cameron., Bequest., B. M. 1955 – 147. ” / “ JAPAN, Kobe, 27 - 11 - 30 ” (BMNH).	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFF8B244FF6AFAA83396FAAD.taxon	description	Description. Measurement. BL = 5.95 mm, FL = 2.51 mm, HL = 0.67 mm, HW = 0.56 mm, PL = 0.94 mm, PW = 0.74 mm, EL = 0.82 mm, EW = 1.01 mm. Body nearly fusiform shaped, large sized (Fig. 3 A). Body brown, except abdominal intersegmental membrane yellowish brown. Legs yellow brown. Antennae brown, basal two antennomeres smoother and lighter than remaining antennomeres. Maxillary and labial palpi yellow brown. Head. Oval in shape (HL to HW ratio 1.2), tempora (behind eyes) dilated and arcuate posteriad, widest at basal 1 / 3, posterior angles rounded. Dorsal integument shiny, extensively bearing micropunctures and microstriae, also with 5 – 7 large, scattered punctures around each eye, and numerous smaller punctures on deflexed portion of tempus and near the posterior margin. Each side of cranium with large characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three ocular punctures near dorsal margin of eye, temporal puncture at the basal 1 / 5, also with occipital puncture at the basal 1 / 4. Frontal furrows and anteocular furrows absent. Eye medium sized, slightly longer than half the length of tempora (eye: tempora = 0.20: 0.38 mm), distinctly protruding laterad. Epistoma not flattened, abruptly oblique, anterior margin emarginate medially. Distance between antennal insertions 0.16 mm being distinctly longer than distance from antenna to eye (0.11 mm). Antennae. Total length 1.69 mm. Scape rod-shaped, slightly thickened apically, 0.26 mm, shorter than two subsequent antennomeres combined; antennomere II elongate, 0.15 mm; III 0.19 mm, distinctly longer than II; IV 0.11 mm, IV to X gradually shortened, IV slightly longer than X; XI 0.11 mm, slightly longer than X. Neck. Surface shiny, only bearing shallow microsculpture, in trapezoidal shape, width of 0.21 mm, slightly wider than 1 / 3 width of head. Pronotum. Elliptical (PL to PW ratio 1.3), distinctly longer than head. Anterior region near anterior angles deflexed and narrowed, lateral margins widened posteriad to basal 1 / 3 (with the widest at apical 1 / 3), then convergent backwards, anterior and posterior angles broadly rounded. Dorsal surface glossy extensively with micropunctures, but without any microsculpture. Each side of the midline with symmetrical five large lateral punctures, with additional 13 – 15 punctures near each margins (including anterior, lateral and posterior margins). Mesoscutellum. Triangular and small. Surface shiny, bearing some light microstriae, but without any punctures. Elytra. Transverse and short (EL to EW ratio 0.81), distinctly wider and shorter than pronotum. Humeri well developed, lateral margins divergent posteriorly, hind margin not rounded, but obliquely truncate. Dorsal integument shiny, flattened, without microsculpture; each elytron with a row of 5 punctures along suture, a row of 5 punctures in median, also with 4 – 5 rows of punctures on deflexed portion. Legs. First four segments of protarsi stout, wider than those of meso- and metatarsi slender. Last segment of protarsi longer than II – IV together; that of mesotarsi slightly longer than II and III together; that of meta- longer than II and III together. Abdomen. Broadest at segment V. Tergites III – VII covered with dense and obvious transverse microstriae; each tergite with dense yellow pubescence, also with a row of darker and coarser setae at posterior margin of III – VI. Tergites III – VII with a basal impression near anterior margin, respectively, each one only possessing one basal transverse carina; surface shiny, bearing polygonal reticulum, but without any punctures. The intersegmental membrane bearing transparent yellow fenestra. All abdominal sternites shiny, with microstriae and punctures as those on tergites. Male. Unknown.	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFF8B244FF6AFAA83396FAAD.taxon	distribution	Distribution. China (Jilin, Liaoning, Taiwan); Japan; South Korea.	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFF8B244FF6AFAA83396FAAD.taxon	discussion	Remarks. This species possesses some similar characters to D. bicornutus sp. nov., like the shape of the antennomeres, pronotum shape and body color, but the body size is large (ca. 6.0 mm), and the head is shorter, length 1.2 times longer than width. However, up to now, only one female specimen has been found, this problem is thus left open for further study.	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFF9B241FF6AFA7C366EF9BF.taxon	description	(Fig. 3 G; Fig. 10 A – N; Fig. 10 - 1 A – H) Cameron, 1918: 84 (Type locality: Singapore: Sembawang); Cameron, 1921: 404 (Singapore); Scheerpeltz, 1933: 1324 (catalog); Herman, 2001: 2448 (catalog); Smetana, 2004 (Palaearctic catalog; Hongkong); Löbl & Löbl, 2015: 1007 (Palaearctic catalog; Hongkong).	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFF9B241FF6AFA7C366EF9BF.taxon	materials_examined	Type material examined: male, with labels as follows: “ Holo-, Type ” / “ TYPE ” / “ HOLOTYTYPE, Diochus, pulchellus, Cameron, 1918, det. R. G. Booth 2013 ” / “ Sembawang, Singapore., Dr. Cameron. ” / “ Debris ” / “ M. Cameron., Bequest., B. M. 1955 – 147. ” (BMNH).	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFF9B241FF6AFA7C366EF9BF.taxon	description	Description. Measurement. BL = 2.94 mm, FL = 1.61 mm, HL = 0.48 mm, HW = 0.40 mm, PL = 0.52 mm, PW = 0.53 mm, EL = 0.57 mm, EW = 0.70 mm. Body nearly fusiform shaped, small sized (Fig. 3 G). Body yellow brown; head, basal half of elytra and abdomen darker. Legs, antennae, maxillary and labial palpi yellow brown. Head (Fig. 10 A, D). Rounded triangular in shape (HL to HW ratio 1.2), tempora (behind eyes) gradually dilated and arcuate posteriad, widest at basal 1 / 5, posterior angles rounded. Dorsal integument shiny, extensively bearing microstriae, but with some polygonal reticulum on frontal region, also with 3 – 5 small, scattered punctures behind each eye, and numerous smaller punctures on deflexed portion of tempus and near the posterior margin. Each side of cranium with large characteristic punctures: one frontal puncture on frontal region, two anterolateral punctures near antennal insertion, three ocular punctures near dorsal margin of eye, temporal puncture at the basal 1 / 6, also with occipital puncture near hind margin. Frontal furrows and anteocular furrows absent, with big and round elevation between eyes. Eye large sized, longer than half the length of tempora (eye: tempora = 0.13: 0.20 mm), distinctly protruding laterad. Epistoma not flattened, abruptly oblique, anterior margin emarginate medially. Distance between antennal insertions 0.15 mm being distinctly longer than distance from antenna to eye (0.06 mm). Ventral surface (Fig. 10 D) with scattered punctures, interspaces between them ca. 5 punctures’ diameter. Gular sutures deep, confluent at base 1 / 3. Antennae (Fig. 10 F). Total length 1.00 mm. Scape rod-shaped, slightly thickened apically, 0.19 mm, shorter than two subsequent antennomeres combined; antennomere II elongate, 0.08 mm; III 0.09 mm, slightly longer than II; IV 0.08 mm, longer than V; VI – X subequal in length of 0.06 mm; XI 0.11 mm, longer than X. Mouthparts (Fig. 10 G – I). Labrum semi-circle shaped (Fig. 10 G), lateral margin slightly waved. Mandibles falciform, left one with one or two indistinct teeth on inner edge, right one with one sharper distinct tooth (Fig. 10 I – J). Neck (Fig. 10 J – K). Surface shiny, only bearing shallow microsculpture, in trapezoidal shape, width of 0.15 mm, slightly wider than 1 / 3 width of head, with a shallow bisinuate basal furrow near posterior margin (Fig. 4 J). Ventral surface with observable gular sutures, not forming obvious triangular region, which missing transverse median furrow (Fig. 4 K). Prothorax (Fig. 10 B, E). Pronotum elliptical (PL to PW ratio 0.98), with subequal length to head, but distinctly wider. Anterior region near anterior angles not deflexed or not narrowed, lateral margins parallel, anterior and posterior angles broadly rounded. Dorsal surface glossy extensively with micropunctures, but without any microsculpture. Each side of the midline with symmetrical five large lateral punctures, with additional 13 – 15 punctures near each margins (including anterior, lateral and posterior margins). Prosternum bearing observable transverse ridge between basisternum and furcasternum, which angularly backwards (Fig. 10 E). Pterothorax. Mesoscutellum triangular and small (Fig. 10 M). Surface shiny, bearing some light microstriae, but without any punctures. The transverse ridge on mesoventrites in wave shape; the paired oblique furrows short but deep; the carina (limiting mesocoxal cavity) slightly arcuate. The discrimen region on metaventrite normally elevated. Elytra (Fig. 10 C). Transverse and short (EL to EW ratio 0.81), longer than pronotum, and distinctly wider. Humeri well developed, lateral margins divergent posteriorly, hind margin not rounded, but obliquely truncate. Dorsal integument shiny, flattened, without microsculpture; each elytron with a row of 5 punctures along suture, a row of 5 punctures in median, also with 4 – 5 rows of punctures on deflexed portion. Legs (Fig. 10 L). First four segments of protarsi not dilated, as wide as those of meso- and metatarsi. Last segment of protarsi as long as II and III together; that of mesotarsi slightly longer than II and III together; that of meta- longer than II and III together. Abdomen (Fig. 10 N). Broadest at segment V. Tergites III – VII covered with dense and obvious transverse microstriae; each tergite with dense yellow pubescence, also with a row of darker and coarser setae at posterior margin of III – VI. Tergites III – VII with a basal impression near anterior margin, respectively, each one only possessing one basal transverse carina; surface shiny, bearing polygonal reticulum, but without any punctures. The intersegmental membrane bearing transparent yellow fenestra. All abdominal sternites shiny, with microstriae and punctures as those on tergites. Male (Fig. 10 - 1 A – H). Posterior margin of tergite VIII arcuate (Fig. 10 - 1 A), that of sternite VIII bisinuate (Fig. 10 - 1 B). Tergite IX symmetrical, connected mediobasally (Fig. 10 - 1 C), sharpened apically. Sternite IX symmetrical, widest near the middle, without longitudinal furrow near each lateral margin, and basal margin concaved (Fig. 10 - 1 D). Tergite X symmetrical, large with arcuate posterior margin (Fig. 10 - 1 C). Aedeagus pearlike shaped and symmetrical (Fig. 10 - 1 E – G), small sized, ca. 0.31 mm long. Parameres long and symmetrical, slightly curved apically, 4 / 5 times the length of basal bulb. Internal sac possessing inner sclerites, sperm pump coiled once, not thinned toward apex (Fig. 10 - 1 H). Female. Unknown.	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFF9B241FF6AFA7C366EF9BF.taxon	distribution	Distribution. China (Hongkong); Singapore.	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
7C5887A3FFF9B241FF6AFA7C366EF9BF.taxon	discussion	Remarks. This species is distinguished from its congeners by the combination of the following characters: 1) the rounded labrum; 2) the blunt teeth on left mandible; 3) emarginated posterior margin of abdominal sternite VIII; 4) slender and elongate parameres and apical broad sclerites of aedeagus.	en	Zhou, Yu-Lingzi, Zhou, Hong-Zhang (2016): Taxonomy of the genus Diochus Erichson, 1839 (Coleoptera: Staphylinidae, Staphylininae, Diochini) in China with descriptions of four new species. Zootaxa 4127 (1): 1-30, DOI: 10.11646/zootaxa.4127.1.1
