taxonID	type	description	language	source
832E87FA1518EE52E7EDFC0ACF79A34A.taxon	description	Of the 5 genera belonging to this subtribe (Chant & McMurtry, 2004 b), only Phytoscutus Muma is known to occur in sub-Saharan Africa.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1518EE52E7EDFAF8CF8EA1AF.taxon	description	Five species of this genus are reported in this paper. They have female idiosomal setal pattern 9 A: 6 E / JV- 3: ZV- 3, except Phytoscutus eugenus (Ueckermann & Loots), whose setal pattern is 9 A: 6 E / JV- 3,4: ZV- 3; all setae smooth, except for P. glomus (Pritchard & Baker) in which Z 5 is very lightly barbed; J 2, S 2 and S 4 absent; r 3 and R 1 inserted on interscutal membrane; Jv 5 inserted on the soft cuticle; chaetotaxy of genua II and III 2 – 2 / 0, 2 / 0 – 1 and 1 – 2 / 1, 2 / 0 – 1, respectively.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1519EE53E7EDFF08CF77A5FC.taxon	description	(Fig. 1)	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1519EE53E7EDFF08CF77A5FC.taxon	description	Dorsum. Dorsal shield smooth, 441 long and 302 wide. Setae j 1 [28], j 3 [72], j 4 [9], j 6 [9], J 5 [9], z 2 [82], z 4 [25], z 5 [9], Z 1 [19], Z 4 [186], Z 5 [227], s 4 [173], S 5 [9], r 3 [13], R 1 [13]. Venter. Sternal shield smooth, posterior margin concave; distances between St 1 - St 3 [54], St 2 - St 2 [66]. Genital shield striate; distance between St 5 - St 5 [95]. Ventrianal shield reticulate, pentagonal, anterior margin straight, [176] long, [183] wide at level of Zv 2 and [151] wide at level of anus, with 3 pairs of preanal setae (Jv 1, Jv 2 and Zv 2; only one member of Jv 4 present); Zv 1 on soft cuticle; Jv 5 long [189]; preanal pores posteromesad to Jv 2. With 1 pair of metapodal shields. Chelicera. Movable digit [28]; fixed digit [25] (because of the position, it is not possible to count the number of teeth). Spermatheca. Calyx cup-shaped, [6] long. Legs. Sharp-tipped macrosetae; Sge I [50], Sge II [32], Sge III [63], Sti III [50], Sge IV [101], Sti IV [88], St IV [95]. Specimens examined. Angola: Salazar, holotype, on Eugenia uniflora, 18 - IV- 1972, M. K. P. Smith Meyer. World distribution. Angola.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA151EEE54E7EDFF08C94CA59C.taxon	description	(Fig. 2)	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA151EEE54E7EDFF08C94CA59C.taxon	description	Dorsum. Dorsal shield smooth, [371] long and [285] wide. Setae j 1 [32], j 3 broken, j 4 broken, j 6 [more than 13], J 5 [10], z 2 broken, z 4 [18], z 5 [5], Z 1 [44], Z 4 broken, Z 5 [136], s 4 [145], S 5 [6], r 3 [8], R 1 [6]. Venter. Sternal shield reticulate laterally; posterior margin concave; distances between St 1 - St 3 [54], St 2 - St 2 [72]. Genital shield reticulate; distance between St 5 - St 5 95. Ventrianal shield reticulate, pentagonal, anterior margin straight, [172] long, [180] wide at level of Zv 2 and [168] wide at level of anus; with 3 pairs of preanal setae (Jv 1, Jv 2 and Zv 2); Jv 4 and Zv 1 on soft cuticle; Jv 5 short [21]; preanal pores posteromesad to Jv 2. With 1 pair of metapodal shields. Chelicera. Not possible to measure movable and fixed digits and to count their teeth. Spermatheca. Calyx funnel-shaped, [10] long. Legs. Sharp-tipped macrosetae; Sge II [22], Sge III [36], Sti III [31], Sge IV [64], Sti IV [48], St IV [41]. Specimens examined. Democratic Republic of Congo (Belgian Congo): Lwiro, holotype, on an ornamental tree, 18 - V- 1955, E. W. Baker. Remarks. No additional specimens of this species were collected in this study. World distribution. Democratic Republic of Congo, Kenya and Zimbabwe.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA151FEE56E7EDFF08CF4BA4FA.taxon	description	(Fig. 3)	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA151FEE56E7EDFF08CF4BA4FA.taxon	description	Dorsum. Dorsal shield mostly smooth, with few undulating transverse striae between setae Z 4, 399 (368 – 456) [379] long and 332 (304 – 368) [321] wide. Setae j 1 28 (24 – 32) [30], j 3 61 (53 – 69) [60], j 4 7 (6 – 8) [4], j 6 16 (11 – 19) [14], J 5 9 (8 – 11) [12], z 2 9 (8 – 11) [10], z 4 18 (14 – 22) [17], z 5 8 (6 – 10) [4], Z 1 37 (30 – 43) [30], Z 4 174 (168 – 187) [166], Z 5 203 (192 – 216) [197], s 4 158 (149 – 171) [167], S 5 13 (10 – 16) [17], r 3 9 (5 – 11) [8], R 1 6 [6]. Venter. Sternal shield smooth posterior margin concave; distances between St 1 - St 3 54 (50 – 58) [54], St 2 - St 2 65 (61 – 69) [62]. Genital shield reticulate; distance between St 5 - St 5 104 (101 – 110) [95]. Ventrianal shield reticulate, anterior margin concave, lateral and posterior margins forewing a semicircle, 165 (142 – 184) [158] long, 239 (208 – 256) [223] wide at the level of Zv 2 and 136 (122 – 158) wide at level of anus, with 5 pairs of preanal setae (Jv 1, Jv 2, Jv 4, Zv 1 and Zv 2); Jv 5 short (24); preanal pores mesad and slightly anterior to Jv 2. With 1 pair of metapodal shields, each with a protuberance on the inner margin, suggesting the fusion of a large, laterad and a small, mesad shield. Chelicera. Movable digit 27 (26 – 28) [27] long, with 3 [3] teeth; fixed digit 19 (18 – 19) long, with 8 [apparently 7] teeth and a pilus dentilis. Spermatheca. Calyx funnel-shaped, 15 (11 – 19) long. Legs. Sharp-tipped macrosetae; Sge I 30 (26 – 35), Sge III 37 (32 – 40) [36], Sti III 27 (24 – 32), Sge IV 69 (62 – 78) [66], Sti IV 48 (42 – 58) [54], St IV 48 (40 – 54) [48]. Specimens examined. Cameroon: 1 km S Yangamo, on Cassia sp., 02 - II- 1991, L. Louis. Democratic Republic of Congo (Belgian Congo): Stanleyville, on Vitex congolensis, 18 - IV- 1955, E. W. Baker. Ghana: Senchi, on unknown plant, 13 - XI- 1989, G. J. de Moraes. Nigeria: 4 km S Elele, on Alchornea cordifolia, 15 - IV- 1992, A. Onzo. Uganda: Rachai, 34 km S Masaka, on Coffea sp., 05 - X- 1990, J. S. Yaninek. Zambia: 84 km S Kasama, on Brachystegia boehmii, 16 - VI- 1992, J. S. Yaninek. Remarks. Differently from the holotype, the specimens collected in this study have macrosetae on genu I and tibia III. World distribution. Cameroon, Democratic Republic of Congo, Ghana, Kenya, Madagascar, Nigeria, Sierra-Leone, Uganda and Zambia.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA151CEE56E7EDFC88CF5AA19C.taxon	description	(Fig. 4) Diagnosis. This species is characterized by having seta z 2 more than twice as long as z 4; seta j 4 at about the same level as z 2; sternal shield totally reticulate; Zv 1, Jv 4 and Jv 5 on integument. Female. (2 specimens measured) Dorsum. Dorsal shield mostly smooth, with few undulating transverse striae between setae Z 4, 435 long and 315 wide. Setae j 1 36 (34 – 38), j 3 65 (62 – 67), j 4 8, j 6 13, J 5 11, z 2 79 (77 – 82), z 4 30 (29 – 32), z 5 8, Z 1 29, Z 4 186 (182 – 189), Z 5 236 (235 – 237), s 4 194 (173 – 216), S 5 8, r 3 8, R 1 8. Venter. Sternal shield reticulate; posterior margin concave; distances between St 1 - St 3 54, St 2 - St 2 72. Genital shield reticulate, distance between St 5 - St 5 91. Ventrianal shield reticulate, anterior margin straight, lateral and posterior margins rounded, 176 long, 177 (173 – 181) wide at widest level and 158 wide at level of anus, with 3 pairs of preanal setae (Jv 1, Jv 2, Zv 2); Jv 4 and Zv 1 on soft cuticle; Jv 5 long (104); preanal pores mesad and slightly posterior to Jv 2. With 1 pair of metapodal shields. Chelicera. Movable digit 27 long, with 2 teeth; fixed digit 18 long, with 7 teeth and a pilus dentilis. Spermatheca. Calyx funnel-shaped, 14 long. Legs. Sharp-tipped macrosetae; Sge I 53, Sge III 62, Sti III 61, Sge IV 104 (98 – 110), Sti IV 94 (91 – 96), St IV 77. Locality and type material. Holotype female from Mangifera indica, 5 km N Muyuka, Cameroon, 09 - II- 1991, L. Louis, deposited at ESALQ-USP. One paratype female from unknown plant, 12 km N Obala, Cameroon, 05 - II- 1991, L. Louis, deposited at ESALQ-USP. Etymology. This new species is named after Liénou Louis, a Cameroonian acarologist involved in the collections of most of the phytoseiids from Cameroon treated in this paper. Remarks. This species falls into the gongylus species group. Indeed, its characters fit those of this species group.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA151DEE58E7EDFADACE7FA572.taxon	description	(Fig. 5)	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA151DEE58E7EDFADACE7FA572.taxon	description	Dorsum. Dorsal shield mostly smooth, with some undulating transverse striae between Z 1 and Z 4, 425 (408 – 443) [403] long and 362 (352 – 384) [302] wide. Setae j 1 28 (26 – 34) [32], j 3 57 (37 – 77) [69], j 4 7 (6 – 8) [9], j 6 19 (18 – 21) [13], J 5 10 (8 – 11) [13], z 2 10 (10 – 11) [9], z 4 13 (10 – 16) [16], z 5 9 (8 – 11) [9], Z 1 86 (58 – 139) [139], Z 4 251 (243 – 259) [239], Z 5 319 (302 – 331) [296] s 4 236 (216 – 259) [230], S 5 15 (11 – 19) [25], r 3 8 (5 – 11) [9], R 1 10 (8 – 13) [9]. Venter. Sternal shield mostly smooth, with some rare lateral striae; posterior margin concave; distances between St 1 - St 3 59 (56 – 62) [54], St 2 - St 2 73 (72 – 74) [63]. Genital shield reticulate; distance between St 5 - St 5 116 (110 – 122) [101]. Ventrianal shield reticulate, lateral and posterior margins rounded, 203 (200 – 206) [192] long, 295 (269 – 344) [290] wide at the level of Zv 2 and 218 (213 – 225) [214] wide at level of anus, with 5 pairs of preanal setae (Jv 1, Jv 2, Jv 4, Zv 1 and Zv 2); JV 5 short (31 long); preanal pores mesad and about in line with setae Jv 2. With 2 pairs of metapodal shields, 1 of which is much smaller than the other. Chelicera. Movable digit 27 (27 – 28) [25] long, with 3 – 4 [4] teeth; fixed digit 19 (18 – 20) [22] long, with 8 [7] teeth and a pilus dentilis. Spermatheca. Calyx funnel-shaped, 17 (14 – 19) [13] long. Legs. Sharp-tipped macrosetae; Sge I 56 [57], Sge III 77 [73], Sti III 43 [44], Sge IV 112 [126], Sti IV 86 [91], St IV 56 [69]. Specimens examined. Angola: Salazar, from Eugenia uniflora, 18 - IV- 1972, M. K. P. Smith Meyer. Uganda: 34 km S Masaka, Rakai District, Coffea sp., 05 - X- 1990, J. S. Yaninek; Nsambya, Rakai District, Mangifera indica, 06 - X- 1990, B. Odongo; 1 km N Muzizi River, Hoima, Artocarpus heterophyllus, 10 - X- 1990, J. S. Yaninek. Remarks. The holotype of this species has a single pair of metapodal shields, instead of 2 as in the specimens from Uganda, collected in this study; in the latter specimens, those shields are distinctly separate in some specimens, but partially fused in others. Considerable variation in setal lengths was observed between specimens collected in Uganda within and between sites / hosts, and even between right and left sides of one of the specimens. Specimens from 1 km N Muzizi River, Hoima, have setae j 3 and Z 1 much shorter than those of the holotype and of the specimens from the other sites however it was not considered as a new species because of the high variation in setal lengths observed in some specimens of P. wiesei in Uganda. World distribution. Angola, Liberia and Uganda.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1513EE59E7EDFC45CF5BA0B7.taxon	description	Nine species of this genus are reported in this paper. They have tan to brown color and setae r 3 and R 1 inserted on interscutal membrane.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1513EE5AE7EDF8CACEF5A262.taxon	description	(Fig. 6) Diagnosis. This new species is characterized by having z 2 shorter than the distance between its base and the base of z 4; z 4 longer than the distance between its base and the base of s 4; s 4 as long as Z 4, and both longer than Z 5; sternal and genital shields smooth; Jv 5 smooth; and by lacking S 4. Female. (1 specimen measured). Dorsum. Dorsal shield smooth, 389 long and 233 wide. Setae j 1 21, j 3 35, j 4 2, j 5 2, j 6 2, J 5 8, z 2 13, z 4 74, z 5 2, Z 1 10, Z 4 86, Z 5 64, s 4 88, S 2 11, S 5 11, r 3 14, R 1 11. Setae smooth, except Z 4 and Z 5 which are serrate; S 4 absent. Venter. Sternal shield mostly smooth, with some rare lateral striae; posterior margin concave; distances between St 1 - St 3 53, St 2 - St 2 80. Genital shield smooth, with 3 pairs of distinct, elongate muscle scars; distance between St 5 - St 5 106. Ventrianal shield pentagonal, with transverse striae and 2 pairs of distinct, roundish muscle scars, anterior margin slightly concave, 115 long, 110 wide at level of Zv 2 and 106 wide at level of anus, preanal pores small and posteromesad to Jv 2. Two pairs of metapodal shields. Ventral setae smooth. Chelicera. Movable digit 33 long, apparently with 1 tooth; fixed digit 29 long, apparently with 5 teeth and a pilus dentilis. Spermatheca. Calyx cup-shaped 10 long; atrium distinct. Legs. Sharp-tipped macrosetae; Sge III 26, Sge IV 53, Sti IV 40, St IV 72. Chaetotaxy: genu II: 2 – 2 / 1,2 / 0 – 1: genu III: 1 – 2 / 1,2 / 0 – 1. Male. Unknown. Locality and type material. Holotype female from Citrus sp., Luwero, Kyankonwa, Uganda, 05 - X- 1990, J. S. Yaninek, deposited at ESALQ-USP. Etymology. The epithet atrichos refers to the absence of seta S 4 on the dorsal shield of the type of this new species. Remarks. The characters of this species fit those of the genus Proprioseiopsis, except the absence of S 4 that separate this species from the other Proprioseiopsis species.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1511EE5CE7EDFF08CC73A402.taxon	description	(Fig. 7) Diagnosis. Species characterized by having dorsal shield smooth; seta Z 5 slightly longer than setae z 4, Z 4 and s 4; setae z 4, Z 4, Z 5, s 4 and Jv 5 smooth; z 2 and z 4 about as long as distances between their bases and the bases of the setae next behind; sternal shield smooth; ventrianal shield pentagonal, smooth, longer than wide at level of Zv 2, fixed digit not more than half the length of those in other Proprioseiospsis. Female. (2 specimens measured). Dorsum. Dorsal shield smooth, 334 (326 – 341) long and 238 (218 – 259) wide. Setae j 1 21, j 3 54 (50 – 59), j 4 6, j 5 6, j 6 12 (11 – 13), J 5 8, z 2 46 (43 – 50), z 4 64 (59 – 69), z 5 6, Z 1 10, Z 4 68 (64 – 72), Z 5 81 (74 – 88), s 4 81 (80 – 82), S 2 29 (21 – 37), S 4 22 (14 – 30), S 5 26 (22 – 30), r 3 37 (34 – 40), R 1 6 (5 – 6). All setae smooth. Venter. Sternal shield smooth, posterior margin concave; distances between St 1 - St 3 54 (53 – 56), St 2 - St 2 64 (62 – 66). Genital shield smooth; distance between St 5 - St 5 70 (67 – 72). Ventrianal shield pentagonal, smooth, anterior margin slightly convex, 109 (106 – 112) long, 97 (96 – 98) wide at level of Zv 2 and 83 (78 – 88) wide at level of anus, preanal pores well posterior and slightly mesad to Jv 2. Ventral setae smooth. Chelicera. Movable digit short, 16 (15 – 17) long, with 1 tooth; fixed, very short, 13 long, with 2 teeth and a pilus dentilis. Spermatheca. Calyx cup-shaped 7 (5 – 8) long 9 (7 – 10) in diameter at its distal extreme; atrium appressed to the base of the calyx. Legs. Sharp-tipped macrosetae; Sge I 29 (27 – 30), Sge II 28 (22 – 34), Sge III 25 (24 – 26), Sti III 20 (18 – 23), Sge IV 34 (32 – 35), Sti IV 34 (32 – 35), St IV 43 (42 – 45). Chaetotaxy: genu II: 2 – 2 / 0,2 / 0 – 1; genu III: 1 – 2 / 0,2 / 0 – 1. Male. Unknown. Locality and type material. Holotype female from Citrus sp., Luwero, Kyankonwa, Uganda, 05 - X- 1990, J. S. Yaninek, deposited at ESALQ-USP. One paratype female from Persea americana, 14 km SW Mvieng, Cameroon, 07 - II- 1991, L. Louis, deposited at IITAIM. Etymology. The epithet brachydactylus refers to the short fixed cheliceral digit of the types of this new species. Remarks. The new species fits the original description of the genus. However, it is different from all known Proprioseiopsis species by having fixed cheliceral digit at least twice shorter.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1516EE5DE7EDFD00CCBFA262.taxon	description	(Fig. 8) Diagnosis. Species characterized by having dorsal shield mostly smooth, with few anterolateral striae; seta Z 5 slightly shorter than setae z 4, Z 4 and s 4; setae z 4, Z 4, Z 5, s 4 and Jv 5 serrate; z 2 and z 4 about as long as distances between their bases and the bases of the following setae; sternal shield with few lateral striae; ventrianal shield subtriangular, reticulate, about as long as wide at level of Zv 2; fixed cheliceral digit with few teeth (apparently 3 teeth); calyx of spermatheca cup-shaped. face; C. Spermatheca; D. Leg IV. Female. (1 specimen measured). Dorsum. Dorsal shield smooth, with few anterolateral striae, 341 long and 240 wide. Setae j 1 24, j 3 41, j 4 5, j 5 5, j 6 5, J 5 10, z 2 29, z 4 70, z 5 5, Z 1 17, Z 4 74, Z 5 62, s 4 74, S 2 34, S 4 24, S 5 14, r 3 17, R 1 12. Setae z 4, Z 4, Z 5 and s 4 serrate, other setae smooth. Venter. Sternal shield mostly smooth, few anterior and lateral striae; posterior margin concave; distances between St 1 - St 3 50, St 2 - St 2 67. Genital shield smooth; distance between St 5 - St 5 91. Ventrianal shield subtriangular, reticulate, anterior margin very slightly concave, 118 long, 127 wide at level of Zv 2, elliptical preanal pores mesad and slightly posterior to Jv 2; Jv 5 serrate and other ventral setae smooth. Chelicera. Movable digit 30 long, with 1 tooth; fixed digit 27 long, apparently with 3 teeth and a pilus dentilis. Spermatheca. Calyx cup-shaped 7 long, 10 in diameter at its distal extreme; atrium small. Legs. Sharp-tipped macrosetae; Sge III 19, Sge IV 34, Sti IV 19, St IV 65. Chaetotaxy: genu II: 2 – 2 / 0,2 / 0 – 1; genu III: 1 – 2 / 1,2 / 0 – 1. Male. Unknown. Locality and type material. Holotype female from Capsicum sp., 3 km N Brema, Ghana, 05 - IV- 1990, B. Haruna and K. Antwi, deposited at ESALQ-USP. Etymology. The epithet bremaensis refers to the location where the type of this new species was collected. Remarks. Proprioseiopsis bregetovae differs from this new species by having the posterior region of the dorsal shield reticulate and because its z 4 and Jv 5 are smooth. Proprioseiopsis okanagensis differs from this new species by having Z 5 longer than z 4, Z 4 and s 4; in addition, it has all dorsal setae as well as Jv 5 smooth, and fixed cheliceral digit multidentate. Both of those species differ from the new species by having calyx of spermatheca bell-shaped.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1517EE5DE7EDFBE0CF36A1B4.taxon	description	(Fig. 9)	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1517EE5DE7EDFBE0CF36A1B4.taxon	description	Dorsum. Dorsal shield smooth, 328 [356] long and 230 [236] wide. Setae j 1 19 [22], j 3 41, j 4 6, j 5 6, j 6 9, J 5 9, z 2 22, z 4 22, z 5 6, Z 1 9, Z 4 60 [63], Z 5 72 [76], s 4 57 [59], S 2 9, S 4 6, S 5 8, r 3 19, R 1 13. Seta z 4 rather closely associated with z 2 and mediolateral to z 2 - s 4. Setae smooth, except Z 4 and Z 5 which are serrate. Venter. Sternal shield smooth, posterior margin concave; distances between St 1 - St 3 54 [57], St 2 - St 2 55 [60]. Genital shield smooth; distance between, St 5 - St 5 67 [74]. Ventrianal shield pentagonal, reticulate, anterior margin straight, 113 long, 111 [117] wide at level of Zv 2 and 95 [98] wide at level of anus, small preanal pores posteromesad to Jv 2. Ventral setae smooth. Chelicera. Movable digit 28 long, with 1 tooth; fixed digit 28 long, with 3 teeth and a pilus dentilis. Spermatheca. Calyx saccular, 13 [16] long; atrium small, bifid. Legs. Sharp-tipped macrosetae; Sge IV 44 [45], St IV 62 [63]. Chaetotaxy: Sge II: 2 – 2 / 1, 2 / 0 – 1; Sge III: 1 - 2 / 1, 2 / 0 - 1. Specimens examined. South Africa: Potchefstroom, North-West Province, on grass, 20 - X- 1954, P. A. J. Ryke. Remarks. Compared to the original description of the genus (Chant & McMurtry, 2005 a), the macroseta on tibia IV of the types of this species is not much longer than the other setae on the segment. World distribution. South Africa.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1514EE5FE7EDFB5ECF36A712.taxon	description	(Fig. 10)	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1514EE5FE7EDFB5ECF36A712.taxon	description	Dorsum. Dorsal shield smooth, 333 (315 – 343) [343] long and 246 (224 – 261) [261] wide. Setae j 1 27 (25 – 28) [28], j 3 54 (41 – 60) [60], j 4 5 (4 – 5) [5], j 5 5 (4 – 5) [5], j 6 6 (5 – 6) [5], J 5 8 (6 – 9), z 2 25, z 4 16 (13 – 19), z 5 5 (4 – 5) [5], Z 1 6 (5 – 6) [6], Z 4 81 (54 – 91) [91], Z 5 76 (63 – 88) [88], s 4 77 (67 – 82) [82], S 2 6, S 4 8 (6 – 9) [8], S 5 10 (9 – 13) [9], r 3 16 (13 – 18) [18], R 1 9. Setae smooth, except Z 4 and Z 5 which are serrate. Venter. Sternal shield smooth, posterior margin concave; distances between St 1 - St 3 51 (50 – 52) [52], St 2 - St 2 65 (63 – 66) [60]. Genital shield smooth; distance between St 5 - St 5 76 (69 – 79) [69]. Ventrianal shield subtriangular, reticulate, anterior margin slightly concave, 107 (101 – 120) [120] long, 105 (95 – 115) [115] wide at level of Zv 2 and 80 (72 – 85) [85] wide at level of anus, small preanal pores posteromesad to Jv 2. Ventral setae smooth. Chelicera. Movable digit 28 long, with 1 tooth; fixed digit 28 long, with 4 teeth and a pilus dentilis. Spermatheca. Calyx saccular, 17 (16 – 19) long; atrium appressed to the base of the calyx. Legs. Sharp-tipped macrosetae; Sge II 27 (25 – 28) [25], Sge III 32 (28 – 35) [28], Sge IV 59 (57 – 63) [63], Sti IV 43 (35 – 50) [50], St IV 58 (50 – 66) [66]. Chaetotaxy: Sge II: 2 - 2 / 1, 2 / 0 - 1; Sge III: 1 - 2 / 1, 2 / 0 - 1. Specimens examined. South Africa: Potchefstroom, North-West Province, holotype and paratype, on grass and Medicago sativa, 20 - X- 1954 and April 1956, P. A. J. Ryke; Charters Creek, near Lake St. Lucia, Kwazulu / Natal Province, on grass, 24 - I- 1966, L. J. Erasmus; Groblersdal, Mpumalanga Province, on Lantana rugosa, 5 - XII- 1979, M. K. P. Smith Meyer; Gladdespruit, near Badplaas, Mpumalanga Province, on grass, 26 - IV- 1972, K. N. de Kock. World distribution. South Africa.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1515EE40E7EDFAA2CE69A402.taxon	description	(Fig. 11)	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1515EE40E7EDFAA2CE69A402.taxon	description	Dorsum. Dorsal shield smooth, 386 (384 – 387) long and 248 (244 – 252) wide. Setae j 1 32, j 3 46 (41 – 50), j 4 4, j 5 4, j 6 9 (8 – 9), J 5 13, z 2 35, z 4 17, z 5 5, Z 1 7 (6 – 8), Z 4 115 (107 – 123), Z 5 169 (154 – 183), s 4 82 (76 – 88), S 2 9, S 4 9, S 5 18 (16 – 19), r 3 22 (21 – 22), R 1 13. Setae smooth, except Z 4 and Z 5 which are serrate. Venter. Sternal shield mostly smooth, with lateral striae, posterior margin concave; distances between St 1 - St 3 61 (59 – 62), St 2 - St 2 77 (67 – 82). Genital shield smooth; distance between St 5 - St 5 82 (81 – 82). Ventrianal shield pentagonal, reticulate, anterior margin very slightly concave, 122 (117 – 126) long, 112 (110 – 113) wide at level of Zv 2 and 91 wide at level of anus, small preanal pores posterior to Jv 2 (see remarks bellow). Ventral setae smooth. Chelicera. Movable digit 29 (28 – 30) long, with 1 tooth; fixed digit 28 (27 – 28) long, with 4 teeth and a pilus dentilis. Spermatheca. Calyx saccular, 15 (13 – 16) long; atrium appressed to base of the calyx. Legs. Sharp-tipped macrosetae; Sge II 30 (28 – 32), Sge III 40 (38 – 41), Sge IV 79, Sti IV 63 (60 – 66), St IV 69. Chaetotaxy: Sge II: 2 - 2 / 1, 2 / 0 - 1; Sge III: 1 - 2 / 1, 2 / 0 - 1. Specimens examined: South Africa: Stellenbosch, Western Cape Province, from Populus sp., 12 - I- 1955, P. A. J. Ryke. Remarks. In both specimens, one member of preanal pores is posterior and slightly mesad to Jv 2, whereas the other is well posterior and laterad to JV 2, next to the margin of the ventrianal shield. World distribution. Algeria, Armenia, Australia, Azerbaijan, France, Gaza, Georgia, Germany, Greece, Israel, Italy, Morocco, New Zealand and South Africa.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA150AEE42E7EDF930CCCDA76A.taxon	description	(Fig. 12)	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA150AEE42E7EDF930CCCDA76A.taxon	description	Dorsum. Dorsal shield mostly smooth, with some rare lateral striae, 340 (336 - 344) [355] long and 223 (214 – 230) [216] wide. Setae j 1 17 (16 – 18) [22], j 3 28 (26 – 29) [28], j 4 5 (5 – 6) [6], j 5 4 (3 – 5) [5], j 6 6 (5 – 6) [7], J 5 9 (8 – 10) [7], z 2 12 (8 – 16) [14], z 4 9 (8 – 11) [10], z 5 4 (3 – 5) [5], Z 1 8 [6], Z 4 69 (62 – 74) [73], Z 5 102 (94 – 112) [98], s 4 54 (53 – 56) [56], S 2 9 (8 – 10) [8], S 4 9 (8 – 10) [8], S 5 10 (10 – 11) [8], r 3 15 (14 – 16) [13], R 1 8 (6 – 10) [7]. Setae smooth, except Z 4 and Z 5 which are serrate. Venter. Sternal shield reticulate, posterior margin very slightly concave; distances between St 1 - St 3 60 (59 – 61), St 2 - St 2 64. Genital shield smooth; distance between St 5 - St 5 65 (61 – 69). Ventrianal shield pentagonal, reticulate, anterior margin straight, 113 (109 – 115) [112] long, 97 (91 – 101) [96] wide at level of Zv 2 and 77 (72 – 80) wide at level of anus, preanal pores mesad and slightly posterior to Jv 2. Ventral setae smooth. Chelicera. Movable digit 34 (33 – 34) long, with 1 tooth; fixed digit 29 (28 – 29) long, with 8 teeth and a pilus dentilis. Spermatheca. Calyx shallow cup-shaped, 3 (3 – 4) long, 9 (9 – 10) in diameter; atrium appressed to the base of the calyx. Legs. Sharp-tipped macrosetae; Sge II 23 (22 – 24) [25], Sge III 23 (22 – 24) [25], Sge IV 49 (45 – 51) [54], Sti IV 27 [32], St IV 58 [62]. Chaetotaxy: genu II: 2 - 2 / 0,2 / 0 - 1; genu III: 1 – 2 / 1,2 / 0 – 1. Specimens examined. Ghana: Atimpoku, Adomi Bridge, Eastern Region, on Centrosema pubescens, 13 - XI- 1989, G. J. de Moraes. Kenya: 6 km N Mtwapa, Coastal Province, on Synedrella nodiflora, 30 - XI- 1989, J. S. Yaninek; 3 km E Kisi, on unknown plant, 3 - XII- 1989; J. S. Yaninek. Mexico: Holotype female intercepted at Brownsville, Texas, USA, on Zinnia sp. 27 - VI- 1937, G. F. Callaghan. Remarks. This is the first record of this species in Africa. World distribution. Australia, Brazil, Colombia, Cuba, Ghana, Guadeloupe, Hawaii, Kenya, Mexico, New Zealand, Panama, USA.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1508EE43E7EDFE18C95BA76A.taxon	description	(Fig. 13)	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1508EE43E7EDFE18C95BA76A.taxon	description	Dorsum. Dorsal shield smooth, 348 (324 – 381) long and 270 (227 – 290) wide. Setae j 1 29 (28 – 32), j 3 64 (57 – 69), j 4 6 (4 – 8), j 5 6 (5 – 8), j 6 9 (8 – 10), J 5 8 (6 – 10), z 2 39 (32 – 44), z 4 25 (19 – 34), z 5 6 (5 – 8), Z 1 21 (14 – 24), Z 4 108 (88 – 120), Z 5 88 (67 – 107), s 4 99 (91 – 106), S 2 20 (14 – 26), S 4 14 (9 – 16), S 5 10 (9 – 11), r 3 19 (16 – 21), R 1 10 (9 – 11). Setae smooth, except Z 4 and Z 5 which are serrate. Venter. Sternal shield reticulate along lateral margins, posterior margin concave; distances between St 1 - St 3 53 (50 – 56), St 2 - St 2 70 (63 – 77). Genital shield mostly smooth, with lateral striae; distance between St 5 - St 5 89 (82 – 96). Ventrianal shield pentagonal, reticulate, anterior margin straight, 109 (98 – 122) long, 108 (104 – 115) wide at level of Zv 2 and 90 (84 – 104) wide at level of anus, preanal pores mesad and slightly posterior to Jv 2. Ventral setae smooth. Chelicera. Movable digit 33 (32 – 34) long, with 1 tooth; fixed digit 30 (28 – 32) long, with 5 teeth and a pilus dentilis. Spermatheca. Calyx saccular, 16 (13 – 19) long; atrium small, nodular. Legs. Sharp-tipped macrosetae; Sge III 28 (25 – 32), Sge IV 58 (46 – 66), Sti IV 41 (34 – 47), St IV 84 (77 – 91). Chaetotaxy: Sge II: 2 - 2 / 0, 2 / 0 - 1; Sge III: 1 – 2 / 1, 2 / 0 – 1. Specimens examined. Ghana: 10 km E Obuasi, Ashanti Region, on unknown plant, 10 - XI- 1989, J. S. Yaninek; 1 km W River Pora, Beposo, Central Region, on unknown plant, 12 - XI- 1989, J. S. Yaninek; Atimpoku, Adomi Bridge, Eastern Region, on unknown plant, 13 - XI- 1989, G. J. de Moraes; 23 km N Otumi, Chromolaena odorata, 28 - III- 1990, J. Gyemenah. Kenya: 27.9 km S Mombasa, Coastal Province on Zea mays, 28 - XI- 1989; J. S. Yaninek; 17 km N Lunga Lunga, Coastal Province on Physalis angula, 28 - XI- 1989, J. S. Yaninek. Mozambique: 5.5 km SE Mitili, Zambezia Province, on Tridax procumbens, 25 - IV- 01, B. Agboton. Sierra-Leone: Njala University, on Lantana camara, 12 - XII- 1991, J. S. Yaninek. USA: Holotype female from Winter Haven, Florida, on Canna sp .. South Africa: Tzaneen, Limpopo Province, from Vernonia stipulacea, 19 - II- 1965, M. K. P. Smith Meyer (holotype and paratype of P. peltatus); Machadodorp, Mpumalanga Province, from Pennisetum clandestinum, 17 - I- 1977, E. A. Ueckermann; Zimbabwe: Triangle, from Cynodon dactylon, 03 - XI- 1969, M. K. P. Smith Meyer. Remarks. Proprioseiopsis peltatus was synonymized with P. ovatus by Tseng (1983). The measurements of specimens from South Africa and other sub-Saharan African countries examined in this study are intermediate to those given by Moraes & McMurtry (1983) for the holotypes of Proprioseiopsis cannaensis (Muma, 1962) and P. ovatus. The spermathecae of the specimens considered in this study also seem intermediate to those shown by Moraes & McMurtry (1983) for the holotypes of those species. It seems that P. ovatus could also be a senior synonym of P. cannaensis as admitted by Muma & Denmark (1969). World distribution. Brazil, Costa Rica, Cuba, Ecuador, Egypt, Ghana, Hawaii, Honduras, Kenya, Philippines, Puerto Rico, Sierra Leone, South Africa, Taiwan, USA and Zimbabwe.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1509EE44E7EDFA9ECF36A42A.taxon	description	(Fig. 14)	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1509EE44E7EDFA9ECF36A42A.taxon	description	Dorsum. Dorsal shield smooth, 293 [318] long and 208 [235] wide. Setae j 1 25, j 3 35 [38], j 4 5 [6], j 5 5 [6], j 6 6, J 5 8, z 2 16 [19], z 4 9 [13], z 5 5 [6], Z 1 6, Z 4 79 [91], Z 5 72 [85], s 4 63 [72], S 2 9, S 4 9, S 5 9, r 3 19 [22], R 1 13 [16]. Setae smooth, except Z 4 and Z 5 which are serrate. Venter. Sternal shield smooth, posterior margin concave; distances between St 1 - St 3 54 [57], St 2 - St 2 63 [65]. Genital shield smooth; distance between St 5 - St 5 63 [66]. Ventrianal shield pentagonal, with transverse striation, anterior margin slightly concave, 101 [113] long, 95 [101] wide at level of Zv 2 and 82 [85] wide at level of anus, preanal pores mesad and slightly posterior to Jv 2. Ventral setae smooth. Chelicera. Movable digit 28 long, with 2 teeth; fixed digit 28 long, with 8 teeth and a pilus dentilis. Spermatheca. Calyx trumpet-shaped, 25 long; atrium nodular. Legs. Sharp-tipped macrosetae; Sge IV 57, Sti IV 38 [41], St IV 57. Chaetotaxy: Sge II: 2 - 2 / 0, 2 / 0 - 1; Sge III: 1 - 2 / 1, 2 / 0 - 1. MALE. (Specimen measured. South Africa: 1 paratype). Dorsum. Dorsal shield pattern as in female, 252 long and 161 wide. Setae j 1 19, j 3 25, j 4 6, j 5 6, j 6 6, J 5 6, z 2 13, z 4 9, z 5 5, Z 1 6, Z 4 51, Z 5 50, s 4 44, S 2 6, S 4 6, S 5 6, r 3 16, R 1 13. Setae smooth except Z 4 and Z 5 which are serrate. Venter. Ventrianal shield subtriangular and faintly reticulate; 104 long and 139 wide at anterior corners, with 3 pairs of preanal setae, 1 pair of preanal pores, additional pores not visible. Spermatodactyl. Shaft 14 long. Legs. Macrosetae sharp-tipped; Sge IV 32, Sti IV 22, St IV 44. Chaetotaxy of genua II and III as in female. Specimens examined. South Africa: Potchefstroom, North-West Province, from soil covered with Eragrostis curvula, 1963, L. J. Erasmus (holotype female and paratype male); Buffelspoort near Brits, North-West Province, from Hyparrhenia hirta, 15 - III- 1977, E. A. Ueckermann. World distribution. South Africa.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA150FEE45E7EDFCA0CFF7A36C.taxon	description	Of the 8 genera belonging to this subtribe (Chant & McMurtry, 2005 b), only Afrodromips Chant & McMurtry, Aristadromips Chant & McMurtry, Scapulaseius Karg & Oomen-Kalsbeek and Typhlodromips De Leon are known from sub-Saharan Africa.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA150FEE46E7EDF8AACF62A742.taxon	discussion	Remarks. This species was described from specimens collected in Sanya, Tanzania, where it was associated with the ant Crematogaster nigriceps, on an unknown plant. No additional specimens were collected in this study. World distribution. Tanzania.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA150CEE46E7EDFEC0CEF0A494.taxon	description	Two species of this genus were found in sub-Saharan Africa.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA150CEE47E7EDFCADCE04A232.taxon	description	(Fig. 15) Diagnosis. This new species is distinct by having dorsal shield mostly smooth, with lateral striae between j 1 and S 2; z 2 and z 4 approximately as long as distances between their bases and the bases of the setae next behind; ventrianal shield of the female constricted at level of Jv 2, with transverse striae anteriorly to preanal pores and with reticulation between preanal pores and anterior margin of anus; male with 6 pairs of preanal setae. Female. (5 specimens measured). Dorsum. Dorsal shield mostly smooth, with lateral striae in region anterior to S 2, 352 (344 – 362) long and 206 (200 – 211) wide. Setae j 1 26 (24 – 29), j 3 39 (35 – 40), j 4 21 (19 – 24), j 5 15 (14 – 16), j 6 18 (16 – 19), J 2 19 (18 – 19), J 5 10 (10 – 11), z 2 29 (27 – 30), z 4 36 (30 – 37), z 5 12 (11 – 13), Z 1 28 (27 – 30), Z 4 55 (53 – 58) Z 5 76 (74 – 80), s 4 51 (50 – 53), S 2 47 (45 – 50), S 4 28 (26 – 30), S 5 27 (24 – 30), r 3 35 (32 – 37), R 1 22 (21 – 24). Setae smooth, except Z 4 and Z 5 which are lightly serrate; r 3 and R 1 inserted on interscutal membrane. Venter. Sternal shield smooth, posterior margin straight; distances between St 1 - St 3 66 (62 – 67), St 2 - St 2 63 (62 – 64). Genital shield smooth; distance between St 5 - St 5 64 (61 – 69). Ventrianal shield pentagonal, constricted laterally at level of Jv 2, with transverse striae anteriorly to preanal pores and with reticulation between preanal pores and anterior margin of anus; anterior margin about straight, 120 (117 – 125) long, 82 (80 – 85) wide at level of Zv 2 and 78 (75 – 80) wide at level of anus, elliptical preanal pores posteromesad to Jv 2. Two pairs of metapodal shields. Ventral setae smooth. Chelicera. Movable digit 33 (33 – 34) long, apparently with 3 teeth; fixed digit 29 (28 – 29) long, apparently with 12 teeth and a pilus dentilis. Spermatheca. Calyx bell-shaped, 18 (16 – 24) long; atrium appressed to the base of the calyx. Legs. Sharp-tipped macrosetae; Sge II 25 (22 – 29), Sge III 22 (19 – 24), Sti III 20 (18 – 21), Sge IV 41 (40 – 43), Sti IV 31 (27 – 34), St IV 66 (64 – 72). Chaetotaxy: genu II: 2 - 2 / 0,2 / 0 - 1; genu III: 1 – 2 / 1,2 / 0 – 1. Male. (1 specimen measured). Dorsum. Dorsal shield pattern as in female, 275 long and 197 wide. Setae j 1 21, j 3 32, j 4 21, j 5 16, j 6 18, J 2 16, J 5 10, z 2 19, z 4 29, z 5 14, Z 1 19, Z 4 40, Z 5 51, s 4 42, S 2 38, S 4 24, S 5 22, r 3 26, R 1 24. Setae smooth, except Z 4 and Z 5 which are lightly serrate. Venter. Ventrianal shield subtriangular, reticulate; 118 long, 147 wide at the anterior corners; with 6 pairs of preanal setae, 4 pairs of lyrifissures (anteromesad to Zv 1, posterolateral to Zv 1, anterolateral to Zv 3 and between Zv 2 and Zv 3), elliptical preanal pores in line with Jv 2. Spermatodactyl. Shaft distally bifurcate, 30 long. Legs. Sharp-tipped macrosetae; Sge II 18, Sge III 18, Sti III 16, Sge IV 27, Sti IV 24 and St IV 51. Chaetotaxy of genua II and III as in female. Locality and type material. Holotype female, 7 paratype females and 1 allotype male from Erynoglossum sp., Aberdares, Kenya, 26 - XII- 1989, H. Van der Berg, deposited at ESALQ-USP. Etymology. The epithet aberdaresensis refers to the location where the types of this species were collected. Remarks. This species fits the description of Aristadromips except for the larger value of the sum of the ratios “ length of dorsal shield / length of S 4 ” and “ length of dorsal shield / length of S 5 ” (ca. 25.6) and for having z 2 and z 4 aproximately as long as the distances between their bases and the bases of the following setae. Such characteristics are intermediate between those of Aristadromips and Typhlodromips De Leon. Within the latter genus, the shape of the spermatheca would place this species closer to those of the ariri species group of Chant & McMurtry (2005 b). The number of male preanal setae, however, indicates that it should be placed in the genus Aristadromips.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1502EE48E7EDFF08CF36A46C.taxon	discussion	Remarks. This species was described from specimens collected in Golden Gate Highland National Park, Orange Free State, South Africa, on Cydonia oblonga. No additional specimens were collected in the present study. Chant and McMurtry did not list this species in any genus in their revision of the subfamily Amblyseiinae. However, it fits the description of Aristadromips (tribe Typhlodromipsini). World distribution. South Africa.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1502EE48E7EDFAA8CDAEA164.taxon	discussion	Remarks. This species was described from specimens collected in Bogor, Java, Indonesia, from an unspecified substrate. It was also collected in Angola (Carmona 1968). No additional specimens were collected in this study. World distribution. Angola, China, Hong-Kong, India, Indonesia, Malaysia, Philippines, Singapore, Thailand.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1502EE48E7EDFDE0CE8DA2A4.taxon	description	markwelli species group of Amblyseius, Schicha, 1987: 25. japonicus species group of Amblyseius, Schicha, 1987: 26. oguroi species group of Amblyseius, Wu & Ou, 1999: 103.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1502EE48E7EDFDE0CE8DA2A4.taxon	description	A single species of this genus was reported from sub-Saharan Africa.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1502EE49E7EDF8E8CEE4A45A.taxon	description	Eight species of this genus were found in sub-Saharan Africa.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1503EE49E7EDFDE8CF36A1B4.taxon	description	(Fig. 16)	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1503EE49E7EDFDE8CF36A1B4.taxon	description	Dorsum. Dorsal shield mostly reticulate, but smooth in the central region anterior to J 2 and in the region posterior to the pores immediately anterior and mesad to Z 4, 334 long and 208 wide. Setae j 1 19, j 3 22, j 4 9, j 5 9, j 6 13, J 2 16, J 5 9, z 2 13, z 4 13 [14], z 5 9, Z 1 13 [15], Z 4 38 [39], Z 5 69 [74], s 4 30 [32], S 2 13, S 4 9, S 5 9, r 3 16, R 1 13. Venter. Sternal shield smooth, posterior margin almost straight; distances between St 1 - St 3 47, St 2 - St 2 52 [54]. Genital shield smooth; distance between St 5 - St 5 60 [62]. Ventrianal shield pentagonal, smooth, anterior margin straight, 110 [110] long, 82 [85] wide at level of Zv 2 and 79 [82] wide at level of anus, preanal pores mesad and slightly posterior to Jv 2. Two pairs of metapodal shields. Ventral setae smooth. Chelicera. Movable digit 25 [26] long, with 3 teeth; fixed digit 22 [25] long, with 11 – 12 teeth and a pilus dentilis. Spermatheca. Calyx dish-shaped, 2 [3], with a tubular section adjacent to the atrium; the latter kidneyshaped. Legs. Sharp-tipped macrosetae; Sge I 30 [32], Sge II 25 [28], Sge III 28, Sti III 22 [26], Sge IV 49 [47], Sti IV 41, and St IV 63. Chaetotaxy: Sge II: 2 - 2 / 0, 2 / 0 - 1; Sge III: 1 - 2 / 1, 2 / 0 - 1. Male. (Specimen measured. South Africa: 1). Dorsum. Dorsal shield pattern as in female, 268 long and 183 wide. Setae j 1 19, j 3 25, j 4 9, j 5 9, j 6 9, J 2 13, J 5 9, z 2 13, z 4 13, z 5 9, Z 1 9, Z 4 38, Z 5 54, s 4 25, S 2 9, S 4 9, S 5 9, r 3 16, R 1 11. Setae smooth, except Z 4 and Z 5 which are serrate. Venter. Ventrianal shield subtriangular and faintly reticulate; 113 long and 151 wide at anterior corners; with 3 pairs of preanal setae, preanal pores mesad and slightly posterior to Jv 2, additional pores not visible. Spermatodactyl. Shaft 19 long. Legs. Sharp-tipped macrosetae; Sge I 24, Sge II 22, Sge III 22, Sti III 25, Sge IV 35, Sti IV 32, and St IV 54. Chaetotaxy of genua II and III as in female. Specimens examined. South Africa: Munster, Kwazulu / Natal Province, from an unidentified plant, 15 - IV- 1955, P. A. J. Ryke (holotype and paratype); Agricultural Research Station, East London, Eastern Cape Province, from Achyranthes sicula, 2 - XII- 1977, M. K. P. Smith Meyer. Remarks. This species seems very similar, if not identical to Typhlodromips mangleae De Leon, 1967, a species reported from the Caribbean area and South America. World distribution. South Africa.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1500EE4AE7EDFBC8CC85A334.taxon	discussion	Remarks. This species was described from specimens collected in Takon, Oueme Province, Benin, on Morinda lucida. No additional specimens were collected in this study. World distribution. Benin.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1500EE4BE7EDFA4DC9BEA5EA.taxon	description	(Fig. 17)	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1500EE4BE7EDFA4DC9BEA5EA.taxon	description	Dorsum. Dorsal shield reticulate, with reticulation more evident laterally, 322 (302 – 339) long and 210 (207 – 226) wide. Setae j 1 17 (13 – 21), j 3 20 (18 – 24), j 4 10 (8 – 11), j 5 10 (8 – 11), j 6 11 (11 – 13), J 2 13 (11 – 14), J 5 9 (8 – 13), z 2 11 (10 – 13), z 4 12 (11 – 13), z 5 9 (8 – 11), Z 1 13 (11 – 14), Z 4 36 (32 – 42), Z 5 73 (66 – 80), s 4 28 (24 – 34), S 2 11 (10 – 13), S 4 9 (8 – 11), S 5 8 (8 – 11), r 3 16 (14 – 18), R 1 12 (10 – 14). Setae smooth, except Z 4 and Z 5 which are serrate. Venter. Sternal shield mostly smooth, except for few lateral striae, posterior margin slightly concave; distances between St 1 - St 3 53 (48 – 56), St 2 - St 2 60 (56 – 64). Genital shield smooth; distance between St 5 - St 5 64 (59 – 67). Ventrianal shield pentagonal, reticulate anteriorly to preanal pores, 107 (96 – 117) long, 89 (80 – 96) wide at level of Zv 2 and 80 (72 – 89) wide at level of anus, elliptical preanal pores mesad and slightly posterior to Jv 2. Two pairs of metapodal shields. Ventral setae smooth. Chelicera. Movable digit 28 (27 – 28) long, with 3 teeth; fixed digit 24 (23 – 25) long, with 8 teeth and a pilus dentilis. Spermatheca. Calyx dome-shaped, 9 (8 – 10) long; atrium kidney-shaped. Legs. Tip of macrosetae of genua I – IV and of tibia and tarsus IV with tiny knobs; tip of other macrosetae blunt; Sge I: 28 (26 – 32), Sge II 25 (24 – 29), Sge III 30 (26 – 32), Sti III 23 (21 – 26), Sge IV 48 (43 – 53), Sti IV 37 (32 – 42), St IV 61 (54 – 67). Chaetotaxy: genu II: 2 - 2 / 0,2 / 0 - 1; genu III: 1 – 2 / 1,2 / 0 – 1. Specimens examined. Benin: International Institute of Tropical Agriculture (IITA) Cotonou Research Station, on unknown plant, 02 - XI- 1989, I. D. Zannou; Ikpinle, on Byrsocarpus coccineus, 11 - XI- 1991, A. Onzo. Cameroon: 1 km SW Nkoteng, Central Province, on unknown plant, 15 - II- 1991, A. Onzo. Ghana: Kotoku, Accra, on Tridax procumbens, 07 - XI- 1989, G. J. de Moraes; Nsutan, on Manihot esculenta, 08 - XI- 1989, J. S. Yaninek; 18 km N Kumassi, Ashanti Region, on unknown plant, 09 - XI- 1989, G. J. de Moraes; 18 km N Kumassi, Ashanti Region, on Centrosema pubescens, 09 - XI- 1989, J. S. Yaninek; 15 km S Abuassi, Ashanti Region, on unknown plant, 10 - XI- 1989, J. S. Yaninek; 2 km W Jakobu, Ashanti Region, on Vitis vinifera, 10 - XI- 1989, J. S. Yaninek; Mankesim, Central Region, on unknown plant, 12 - XI- 1989, J. S. Yaninek. Malawi: 2.5 SE Mzuzu, Northern Province, on Zea mays, 04 - V- 2000, B. Eklou. Rwanda: 5 km N Ruhengeri, on Ipomoea sp. 08 - XII- 1989, J. S. Yaninek. Sierra-Leone: Port Loko, on Z. mays, 16 - XII- 1991, J. S. Yaninek. World distribution. Benin, Cameroon, Ghana, Malawi, Nigeria, Rwanda, Sierra-Leone.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1506EE4DE7EDFF08CFB7A5EA.taxon	description	(Fig. 18) Diagnosis. This species is characterized by having dorsal shield reticulate, ventrianal shield lightly striate anteriorly to preanal pores and reticulate posteriorly to those, ca. 90 % as wide as long, fixed cheliceral digit with 8 teeth and calyx of spermatheca dish-shaped, without a distinct tubular section next to atrium. Female. (2 specimens measured). Dorsum. Dorsal shield reticulate anteriorly to Z 4, and mostly smooth, with few transverse striae posteriorly, 366 (355 – 376) long and 226 (226 – 227) wide. Setae j 1 18, j 3 22, j 4 14, j 5 14, j 6 16, J 2 19, J 5 9 (8 – 10), z 2 18 (18 – 19), z 4 19, z 5 15 (14 – 16), Z 1 18 (18 – 19), Z 4 37, Z 5 72, s 4 31 (30 – 32), S 2 16, S 4 12 (11 – 13), S 5 10 (10 – 11), r 3 20 (19 – 21), R 1 17 (16 – 18). Setae smooth, except Z 4 and Z 5 which are serrate. Venter. Sternal shield smooth, posterior margin slightly concave; distances between St 1 - St 3 54, St 2 - St 2 64. Genital shield smooth; distance between, St 5 - St 5 64 (62 – 66). Ventrianal shield pentagonal, lightly striate anteriorly to preanal pores and reticulate posteriorly to those; with or without lateral constriction, anterior margin convex, 118 (117 – 118) long, 108 (104 – 112) wide at level of Zv 2 and 93 wide at level of anus, elliptical preanal pores slightly posteromesad to Jv 2. Two pairs of metapodal shields. Ventral setae smooth. Chelicera. Movable digit 30 long, with 3 teeth; fixed digit 27 long, with 8 teeth and a pilus dentilis. Spermatheca. Calyx dish-shaped, 2 long, 15 in diameter; atrium kidney-shaped. Legs. Tip of macrosetae of genua I – IV and of tarsus IV with tiny knobs; tip of other macrosetae blunt; Sge I 27, Sge II 24, Sge III 27 (26 – 29), Sti III 22 (21 – 22), Sge IV 42, Sti IV 32, St IV 54 (53 – 56). Chaetotaxy: genu: II 2 - 2 / 0,2 / 0 - 1; genu III: 1 – 2 / 1,2 / 0 – 1. Male. (1 specimen measured). Dorsum. Dorsal shield pattern as in female, 272 long and 197 wide. Setae j 1 16, j 3 22, j 4 11, j 5 11, j 6 13, J 2 16, J 5 5, z 2 13, z 4 16, z 5 10, Z 1 14, Z 4 32, Z 5 54, s 4 26, S 2 11, S 4 8, S 5 8, r 3 18, R 1 11. Setae smooth, except Z 4 and Z 5 which are serrate. Venter. Ventrianal shield subtriangular, reticulate anteriorly to preanal pores, 130 long, 160 wide at the anterior corners, with 3 pairs of preanal setae, 5 pairs of lyrifissures (1 directly anterior to Jv 1, 2 anterolaterad to Jv 1, 1 anterolaterad to Zv 2 and 1 posterolaterad to Zv 2), elliptical preanal pores mesad and slightly posterior to Jv 2. Spermatodactyl. Shaft 20 long. Legs. Tip of macrosetae of genua I – IV and of tarsus IV with tiny knobs; tip of other macrosetae blunt; Sge I 16, Sge II 19, Sge III 19, Sti III 16, Sge IV 29, Sti IV 24 and St IV 40. Chaetotaxy of genua II and III as in female. Locality and type material. Holotype female, 1 paratype female and 1 allotype male from Citrus sp., Kigali, ISAR-Karama Station, Rwanda, 09 - XII- 1989, J. S. Yaninek, deposited at ESALQ-USP. Etymology. The epithet neoshi refers to the similarity between this species and Typhlodromips shi (Pritchard & Baker). Remarks. Typhlodromips shi (Pritchard & Baker, 1962) has generally shorter dorsal setae, ventrianal shield narrower and smooth or with few and light striae anteriorly to preanal pores. Typhlodromips ibadanensis (Ueckermann & Loots, 1988) differs from this new species by having ventrianal shield reticulate anteriorly to preanal pores compared to lightly striate anteriorly to preanal pores and reticulate posteriorly to those in T. neoshi, and calyx of spermatheca dome-shaped.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1507EE4EE7EDFB98CCE8A76A.taxon	description	(Fig. 19) Diagnosis. This species is characterized by having reticulate dorsal shield; all dorsal setae smooth, except Z 5, which is serrate; sternal shield mostly smooth, with few lateral striae; ventrianal shield reticulate; 1 pair of metapodal shields; calyx of spermatheca cup-shaped; Sge IV about as long as Sti IV and both ca. 50 % as long as St IV. Female. (1 specimen measured). Dorsum. Dorsal shield with faint reticulation on anterior half and with distinct reticulation on posterior half, 424 long and 238 wide. Setae j 1 29, j 3 32, j 4 14, j 5 11, j 6 broken, J 2 19, J 5 11, z 2 19, z 4 16, z 5 13, Z 1 19, Z 4 46, Z 5 77, s 4 42, S 2 26, S 4 22, S 5 16, r 3 21, R 1 22. All setae smooth, except Z 5 which is serrate. Venter. Sternal shield mostly smooth, with few lateral striae; posterior margin irregular and slightly concave; distances between St 1 - St 3 83, St 2 - St 2 85. Genital shield smooth; distance between St 5 - St 5 85. Ventrianal shield pentagonal, reticulate; anterior margin slightly convex, 128 long, 118 wide at level of Zv 2 and 105 wide at level of anus, preanal pores posterior to and approximately vertically aligned with Jv 2. A single pair of distinct metapodal shields. Ventral setae smooth. Chelicera. Movable digit 45 long, with 2 teeth; fixed digit 40 long, with 7 teeth and a pilus dentilis. Spermatheca. Calyx cup-shaped, 9 long; atrium small. Legs. Sharp-tipped macrosetae; Sge II 29, Sge III 29, Sti III 27, Sge IV 46, Sti IV 45, St IV 91. Chaetotaxy: genu II: 2 – 2 / 0,2 / 1 – 1; genu III: 1 – 2 / 0,2 / 1 – 1. Male. Unknown. Locality and type material. Holotype female from unknown plant, 39 km W Nakuru, Valley Province, Kenya, 04 - XII- 1989, J. S. Yaninek, deposited at ESALQ-USP. Etymology. The epithet neoswellendamensis refers to the similarity between this new species and Typhlodromips swellendamensis (Ueckermann & Loots). Remarks. Typhlodromips swellendamensis (Ueckermann & Loots, 1988) and Typhlodromips tee (Schicha, 1983) differ from this new species by having dorsal shield centrally smooth, sternal and ventrianal shields smooth and 2 pairs of metapodal shields. In addition, T. tee has movable and fixed cheliceral digits with 3 and 10 teeth, respectively.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA1504EE70E7EDF987C8FAA7E2.taxon	description	(Fig. 20) Diagnosis. This new species is characterized by having dorsal shield mostly smooth and without distinct constriction at level of seta R 1; ventrianal shield vase-shaped and striate on anterior half; calyx of spermatheca funnel-shaped and stout; atrium long and vacuolate; movable and fixed cheliceral digits with 4 and 10 teeth, respectively. Female. (5 specimens measured). Dorsum. Dorsal shield mostly smooth, except for few anterolateral striae, without distinct constriction at level of seta R 1, 356 (339 – 384) long and 257 (245 – 264) wide. Setae j 1 22 (19 – 26), j 3 28 (26 – 30), j 4 10 (10 – 11), j 5 9 (8 – 10), j 6 10 (8 – 11), J 2 14 (13 – 14), J 5 10, z 2 11 (10 – 13), z 4 12 (10 – 14), z 5 10 (8 – 11), Z 1 17 (14 – 19), Z 4 47 (43 – 53), Z 5 94 (86 – 98), s 4 37 (34 – 40), S 2 15 (14 – 16), S 4 14 (13 – 14), S 5 11 (10 – 13), r 3 17 (16 – 19), R 1 11 (10 – 13). All setae smooth, except Z 5 which is serrate. Venter. Sternal shield mostly smooth, except for few lateral striae; posterior margin straight; distances between St 1 - St 3 57 (53 – 62), St 2 - St 2 68 (66 – 70). Genital shield smooth; distance between St 5 - St 5 72 (67 – 75). Ventrianal shield vase-shaped, striate on anterior half, anterior margin convex, 114 (109 – 120) long, 69 (61 – 82) wide at level of Zv 2 and 76 (74 – 82) wide at level of anus, elliptical preanal pores slightly posteromesad to Jv 2. Two pairs of metapodal shields. Ventral setae smooth. Chelicera. Movable digit 31 long, with 4 teeth; fixed digit 27 long, with 10 teeth and a pilus dentilis. Spermatheca. Calyx funnel-shaped, stout 16 (15 – 16) long; atrium vacuolate. Legs. Sharp-tipped macrosetae; Sge I 32 (29 – 34), Sge II 35 (32 – 37), Sge III 50 (45 – 53), Sti III 33 (32 – 35), Sge IV 74 (70 – 77), Sti IV 33 (32 – 35), St IV 62 (61 – 64). Chaetotaxy: genu: II 2 - 2 / 0,2 / 0 - 1; genu III: 1 - 2 / 1,2 / 0 - 1. Male. Unknown. Locality and type material. Holotype female from unknown plant, 28 km S Dunkwa, Western Region, Ghana, 11 - XI- 1989, J. S. Yaninek, deposited at ESALQ-USP. One paratype female from Elaeis guineensis, 28 km S Dunkwa, Western Region, Ghana, 11 - XI- 1989, J. S. Yaninek, deposited at IITAIM. One paratype female from Securinega virosa, Agbotagon, Atlantic Province, Benin, 10 - X- 1991, B. Eklou, deposited at IITAIM. One paratype female from Anacardium occidentale, Bo China Farm, Sierra-Leone, 14 - XII- 1991, B. Kristensen, deposited at IITAIM. One paratype female from Coffea sp., 34 km S Masaka, Uganda, 05 - X- 1990, J. S. Yaninek, deposited at ESALQ-USP. Etymology. The epithet robusticalyx refers to the shape of the calyx of the types of this species. Remarks. Similarly to this new species, Typhlodromips lambatinus (Schicha & Corpuz-Raros, 1992), T. mitassini (Chant & Hansell, 1971) and T. sinensis Denmark & Muma, 1972 also have ventrianal shield vaseshaped, which is uncommon for species in this genus. However, in all of those species a small but distinct constriction is observed at level of seta R 1, and the ventrianal shield is smooth. In addition, T. lambatinus has an extra dorsal shield seta, on one side, regarded as a “ freak feature ” and its dorsal shield is reticulate; T. mistassini has spermatheca very small and poorly sclerotized and it has seta z 2 almost as long as the distance between its base and the base of s 4. Typhlodromips sinensis does not have stout calyx and vacuolate atrium.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA153AEE70E7EDFE60CC16A1A4.taxon	description	(Fig. 21)	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA153AEE70E7EDFE60CC16A1A4.taxon	description	Dorsum. Dorsal shield reticulate anteriorly to Z 4, and mostly smooth, with scanty striae posteriorly, 332 (310 – 347) [330] long and 217 (200 – 226) [204] wide. Setae j 1 16 (13 – 21) [16], j 3 19 (16 – 24) [19], j 4 10 (8 – 13) [11], j 5 10 (8 – 11) [10], j 6 12 (10 – 13) [11], J 2 13 (11 – 14) [12], J 5 8 (8 – 10) [8], z 2 12 (11 – 13) [13], z 4 12 (10 – 13) [13], z 5 10 (8 – 11) [9], Z 1 13 (13 – 14) [14], Z 4 31 (22 – 37) [31], Z 5 67 (53 – 74) [67], s 4 21 (16 – 26) [18], S 2 11 (8 – 13) [12], S 4 10 (8 – 13) [11], S 5 9 (8 – 10) [9], r 3 14 (11 – 16) [14], R 1 11 (8 – 13) [10]. Setae smooth, except Z 4 and Z 5 which are serrate. Venter. Sternal shield mostly smooth, with few lateral striae; posterior margin slightly concave, distances between St 1 - St 3 54 (50 – 58) [53], St 2 - St 2 63 (59 – 66) [63]. Genital shield smooth, distance between St 5 - St 5 65 (59 – 70) [66]. Ventrianal shield pentagonal, smooth or with few transverse striae anteriorly to preanal pores, 108 (94 – 115) [110] long, 84 (74 – 94) [82] wide at level of Zv 2 and 79 (72 – 86) [78] wide at level of anus, elliptical preanal pores mesad and about in line with Jv 2. Two pairs of metapodal shields. Ventral setae smooth. Chelicera. Movable digit 27 (27 – 28) long, with 3 teeth; fixed digit 23 (23 – 24) long, with 8 teeth and a pilus dentilis. Spermatheca. Calyx dish-shaped, 6 (5 – 8) long, 13 (13 – 14) in diameter; atrium kidney-shaped. Legs. Tip of macrosetae of genua I – IV and of tarsus IV with tiny knobs; tip of other macrosetae blunt; Sge I: 26 (22 – 30) [27], Sge II 23 (19 – 29) [24], Sge III 29 (24 – 32) [28], Sti III 23 (19 – 24) [24], Sge IV 42 (35 – 48) [41], Sti IV 33 (26 – 40) [34], St IV 52 (42 – 67) [46]. Chaetotaxy: genu II: 2 – 2 / 0,2 / 0 – 1; genu III: 1 – 2 / 1,2 / 0 – 1. Specimens examined. Benin: International Institute of Tropical Agriculture (IITA) Cotonou Research Station, on Chromolaena odorata, 02 - XI- 1989, I. D. Zannou; Agbotagon, on Macrosphyra longistyla, 12 - VIII- 1991, B. Eklou. Cameroon: 52 km N Kribi, on Elaeis guineensis, 08 - II- 1991, L. Louis. Democratic Republic of Congo (Belgian Congo): Stanleyville, Eastern Province, Zaire, on Afromomum sp., 20 - IV- 1955, E. W. Baker. Ghana: Nsutam, on Manihot esculenta, 08 - XI- 1989, J. S. Yaninek; 2 km W Jakobu, Ashanti Region, on Ageratum conyzoides, 10 - XI- 1989, J. S. Yaninek; Somanya, on Chromolaena odorata, 13 - XI- 1989, J. S. Yaninek; Somanya, on unknown plants, 13 - XI- 1989, G. J. de Moraes. Kenya: 3 km N Lunga Lunga, Coastal Province, on Musa sapientum, 29 - XI- 1989; J. S. Yaninek; 8 km N Kinango, on Cassia sp. 29 - XI- 1989, J. S. Yaninek. Mozambique: 8 km S Movia, on A. conyzoides, 29 - IV- 2001, B. Agboton. Sierra-Leone: Port Loko, on Zea mays, 16 - XII- 1991, J. S. Yaninek. World distribution. Angola, Benin, Democratic Republic of Congo, Ghana, Kenya, Mozambique, Nigeria and Sierra-Leone.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA153BEE72E7EDFA08CF36A51A.taxon	description	(Fig. 22)	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
832E87FA153BEE72E7EDFA08CF36A51A.taxon	description	Dorsum. Dorsal shield striate-reticulate laterally and posteriorly, smooth centrally, 383 (353 – 413) [413] long and 213 (198 – 225) [225] wide. Setae j 1 32 (32 – 35) [35], j 3 41 (38 – 44) [44], j 4 16 [16], j 5 16 (13 – 16) [16], j 6 17 (16 – 19) [16], J 2 17 (16 – 19), J 5 12 (11 – 13) [12], z 2 23 (22 – 25) [23], z 4 22 [22], z 5 15 (13 – 16) [15], Z 1 23 (22 – 25) [23], Z 4 52 (47 – 55) [55], Z 5 72 (69 – 76) [76], s 4 46 (42 – 54) [49], S 2 38 (35 – 41) [40], S 4 34 (32 – 38) [35], S 5 21 (16 – 25) [23], r 3 22 (22 – 25) [22], R 1 27 (25 – 32) [32]. Setae smooth, except Z 5 which is serrate. Venter. Sternal shield mostly smooth, except for few lateral striae, posterior margin straight, distances between St 1 - St 3 69 (66 – 74) [74], St 2 - St 2 74 (69 – 79) [79]. Genital shield smooth; distance between St 5 - St 5 73 (66 – 82). Ventrianal shield pentagonal, smooth, anterior margin slightly convex, 130 (120 – 139) long, 105 (98 – 113) wide at level of Zv 2 and 90 (79 – 101) [101] wide at level of anus, small preanal pores posterolaterad to Jv 2. Two pairs of metapodal shields. Ventral setae smooth. Chelicera. Movable digit 43 (41 – 46) [41] long, with 2 teeth; fixed digit 37 (33 – 41) [41] long, with 5 – 6 teeth and a pilus dentilis. Spermatheca. Calyx cup-shaped, 10 (8 – 9) [9] long; atrium appressed to the base of the calyx. Legs. Sharp-tipped macrosetae; Sge II 31 (28 – 32) [31], Sge III 31 (28 – 32) [31], Sti III 28 (25 – 30) [28], Sge IV 46 (44 – 49) [49], Sti IV 41 (38 – 44) [44], St IV 80 (76 – 88) [88]. Chaetotaxy: Sge II: 2 – 2 / 0, 2 / 0 – 1; Sge: III 1 – 2 / 1, 2 / 0 – 1. Specimens examined. South Africa: Swellendam, Western Cape Province, from Citrus sp., 2 - VIII- 1976, S. Kambarov (holotype and 2 paratypes); Silver mine Nature Reserve, Cape Town, Western Cape Province, from Pinus radiata, 08 - III- 1983, D. P. Keetch; 20 km from Machadodorp, Mpumalanga Province, from an unknown plant, 25 - VIII- 1981, E. A. Ueckermann; Ebenezer Dam near Tzaneen, Limpopo Province, from Castanea sativa, 24 - V- 1983, E. A. Ueckermann; Ebenezer Dam near Tzaneen, Limpopo Province, from Spiraea cantoniensis, 24 - V- 1983, E. A. Ueckermann. Remarks. Although in the original description it was mentioned that 1 and 4 teeth were found on the movable and fixed digits, respectively, a re-examination of the types and of additional specimens collected showed that the correct numbers were 2 and 5 – 6 teeth, respectively. World distribution. South Africa.	en	De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid, Yaninek, John S. (2007): Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa 1448 (1): 1-39, DOI: 10.11646/zootaxa.1448.1.1, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1
