taxonID	type	description	language	source
D5C92A25F65D5116B1B13813B178B06F.taxon	distribution	Distribution. Eastern Palearctic and Oriental (Fig. 1).	en	Kolcsar, Levente-Peter, Kato, Daichi, Gamboa, Maribet, Watanabe, Kozo (2020): Revision of Japanese species of Nipponomyia Alexander, 1924 (Diptera, Pediciidae). ZooKeys 1000: 71-105, DOI: http://dx.doi.org/10.3897/zookeys.1000.55021, URL: http://dx.doi.org/10.3897/zookeys.1000.55021
D5C92A25F65D5116B1B13813B178B06F.taxon	biology_ecology	Biology. Adults swarm in the air close to the ground or above the vegetation, in shadow and windless conditions. They rest on ventral surfaces of substrates like leaves, spreading their wings horizontally, even during copulation. Nipponomyia kuwanai and N. trispinosa males walk fast on the vegetation and fly short distances to find females. Nipponomyia kuwanai females were observed ovipositing in muddy, wet soil, near mosses on a mountain lakeshore. A N. trispinosa female was observed searching for oviposition sites around wet soil, rich of organic matter next to a waterfall, but the oviposition has not yet been observed. Sometimes N. kuwanai, N. trispinosa, and N. pentacantha inhabit the same habitat.	en	Kolcsar, Levente-Peter, Kato, Daichi, Gamboa, Maribet, Watanabe, Kozo (2020): Revision of Japanese species of Nipponomyia Alexander, 1924 (Diptera, Pediciidae). ZooKeys 1000: 71-105, DOI: http://dx.doi.org/10.3897/zookeys.1000.55021, URL: http://dx.doi.org/10.3897/zookeys.1000.55021
ADCBFAB26FCD570D97A6E3C442AC858C.taxon	materials_examined	Material examined. Non-types: Japan: [Honshu] • 1 ♂; Aomori, Hirosaki, Ichinowatari-washinosu; alt. 205 m; 40 ° 31.15 ' N, 140 ° 26.33 ' E; 17 Jun. 2013; D. Kato leg. (pinned, BLKU) • 1 ♂; Aomori, Nishimeya, Shirakami Nature observation garden, Kawaratai; alt. 255 m; 40 ° 31.13 ' N, 140 ° 12.89 ' E; 21 Jun. 2013; D. Kato leg. (pinned, BLKU) • 1 ♀; Aomori, Hirosaki, Inekari River, Koguriyama; alt. 170 m; 40 ° 32.19 ' N, 140 ° 29.22 ' E; 26 Jun. 2013; D. Kato leg. (pinned, BLKU) • 1 ♂; same data as previous except 25 Jul. 2013; D. Kato leg. (pinned, BLKU) • 3 ♂, 2 ♀; Aomori, Towada, Sakura Spa, Okuse; alt. 854 m; 40 ° 37.64 ' N, 140 ° 54.59 ' E; 5 Aug. 2013; D. Kato leg. (pinned, BLKU) • 1 ♂, 1 ♀; Aomori, Towada, Tsutanuma Path, Okuse; alt. 468 m; 40 ° 35.45 ' N, 140 ° 57.42 ' E; 10 Jun. 2014; D. Kato leg. (pinned, BLKU) • 1 ♂; same data as previous except 21 Jun. 2014; D. Kato leg. (pinned, BLKU). [Hokkaido]: • 5 ♂, 1 ♀ (♂ GenBank # MT 874511); Hokkaido, Higashikawa, Asahidake, River Yukomabetsu; alt. 1120 m; 43 ° 39.14 ' N, 142 ° 48.14 ' E; 23 Jul. 2019; L. - P. Kolcsar leg. (pinned or in ethanol, CKLP) • 2 ♀; Hokkaido, Murayama, Kijihiki Highland, muddy area; alt. 565 m; 41 ° 57.13 ' N, 140 ° 36.57 ' E; 30 Jul. 2019; L. - P. Kolcsar leg. (pinned or in ethanol, CKLP).	en	Kolcsar, Levente-Peter, Kato, Daichi, Gamboa, Maribet, Watanabe, Kozo (2020): Revision of Japanese species of Nipponomyia Alexander, 1924 (Diptera, Pediciidae). ZooKeys 1000: 71-105, DOI: http://dx.doi.org/10.3897/zookeys.1000.55021, URL: http://dx.doi.org/10.3897/zookeys.1000.55021
ADCBFAB26FCD570D97A6E3C442AC858C.taxon	description	Diagnostic characters. Yellowish species with 11 - 13 dark spots on thorax (7 dark spots in N. okinawensis Kolcsar & Kato, sp. nov., 11 dark spots in N. pentacantha). Wing with transverse dark lines in costal cell. Brown marking extending from base of R 2 + 3 to base of M 1, often not reaching M 1 (brown marking extending from R 2 + 3 to base of M 1 in N. okinawensis Kolcsar & Kato, sp. nov. and to m-m in N. pentacantha). Second sternite with black marking at corner of membranous area (without this marking in N. pentacantha and N. okinawensis Kolcsar & Kato, sp. nov.), and without other line (a diffuse line positioned same level as line on sternite 3 in N. pentacantha). Gonostylus with 11 - 14 spines (4 or 5 spines in N. pentacantha). Aedeagus short, as long as wide in lateral view, tip rounded (twice as long as wide in N. pentacantha). Cercus curved upward (straight in N. okinawensis Kolcsar & Kato, sp. nov.). Genital opening Y-shaped (T-shaped in N. okinawensis Kolcsar & Kato, sp. nov.), lateral sclerite less than 1 / 5 of length of genital fork (1 / 3 of length of genital fork in N. pentacantha and less than 1 / 6 - 1 / 7 of length of genital fork in N. okinawensis Kolcsar & Kato, sp. nov.), genital fork spoon-shaped (cross-shaped in N. pentacantha and N. okinawensis Kolcsar & Kato, sp. nov.).	en	Kolcsar, Levente-Peter, Kato, Daichi, Gamboa, Maribet, Watanabe, Kozo (2020): Revision of Japanese species of Nipponomyia Alexander, 1924 (Diptera, Pediciidae). ZooKeys 1000: 71-105, DOI: http://dx.doi.org/10.3897/zookeys.1000.55021, URL: http://dx.doi.org/10.3897/zookeys.1000.55021
ADCBFAB26FCD570D97A6E3C442AC858C.taxon	distribution	Distribution. Japan: Honshu (Nakamura 2014; Oosterbroek 2020), first records from Hokkaido (Fig. 13).	en	Kolcsar, Levente-Peter, Kato, Daichi, Gamboa, Maribet, Watanabe, Kozo (2020): Revision of Japanese species of Nipponomyia Alexander, 1924 (Diptera, Pediciidae). ZooKeys 1000: 71-105, DOI: http://dx.doi.org/10.3897/zookeys.1000.55021, URL: http://dx.doi.org/10.3897/zookeys.1000.55021
2E7C04B0F37957B19F2D4ED19A78A5E3.taxon	description	Diagnostic characters. Anterior part of thorax dark brown to black, posterior part yellowish brown (yellow in N. pentacantha and N. kuwanai), abdomen yellow. Thorax with 7 darker patches (11 in N. pentacantha and 11 - 13 in N. kuwanai), 2 in presutural area of scutum. Wing with transverse dark lines in costal cell. Brown marking extending from R 2 + 3 to base of M 1 (brown marking usually not extending to base of M 1 in N. kuwanai and extending to crossvein m-m in N. pentacantha). Second sternite without dark line (with black marking at corner of membranous area in N. kuwanai and with a diffuse line positioned same level as line on sternite 3 in N. pentacantha). Cercus straight (curved upward in N. kuwanai and N. pentacantha). Genital opening T-shaped (Y-shaped in N. kuwanai and N. pentacantha), lateral sclerite very small, less than 1 / 6 - 1 / 7 of length of genital fork (1 / 3 of length of genital fork in N. pentacantha and less than 1 / 5 of length of genital fork in N. kuwanai). Genital fork cross-shaped, lateral branch curved caudally (spoon-shaped in N. kuwanai and cross-shaped in N. pentacantha but lateral branch almost straight).	en	Kolcsar, Levente-Peter, Kato, Daichi, Gamboa, Maribet, Watanabe, Kozo (2020): Revision of Japanese species of Nipponomyia Alexander, 1924 (Diptera, Pediciidae). ZooKeys 1000: 71-105, DOI: http://dx.doi.org/10.3897/zookeys.1000.55021, URL: http://dx.doi.org/10.3897/zookeys.1000.55021
2E7C04B0F37957B19F2D4ED19A78A5E3.taxon	description	Description. Body length: female: 12 mm. Wing length: female 10 mm. Head: General coloration brown (Fig. 14 B, C). Palpus dark brown, 5 - segmented, segments 2 - 4 almost same in length, last segment 1.5 - 1.6 x longer than segment 4. Tip of last flagellomere darker than other part of palpus. Antenna 1.5 x longer than head. Flagellum 13 - segmented, scape and pedicel brown, flagellum gradually lightening from base to tip. Thorax: Apical half of thorax dark brown, almost black, partly due to decay inside, posterior part yellowish brown (Fig. 14). Pattern of thorax hardly recognizable, only 2 lateral large spots on presutural area of scutum distinct. Postsutural area of scutum with 5 spots, 1 triangular black spot at middle of suture, other 2 spots at anterior corners of transverse suture, and 2 small spots at posterior corners of scutum (parascutum) (Fig. 14 B, C). Legs: General coloration yellow, covered with yellowish setae. Femora without apical dark area, tip of tibiae with a narrow darker ring. Apical ends of tarsomeres 1 to 4 each with narrow dark yellow to light brown ring. Tarsomeres 4 and 5 yellowish (Fig. 14 A). Tarsomeres each with 2 spurs, black, easily discernible. Wings: As in Fig. 4 E. Wing with transverse dark lines in costal cell. Crossvein m-m present. Narrow band on R 2 + 3 not extending to crossvein m-m. Small yellowish brown area around connection of m-cu to Cu. Abdomen: Yellow to light brown, relatively short setae dark on tergites and pale on sternites. Tergites 2 - 6, each with longitudinal narrow black line on lateral side, 1 / 4 - 1 / 3 length of tergite length, less prominent compared to other species. Sternite 2 without dark mark. Sternites 3 - 5 each with narrow brown line, not continuous in sternite 3 (Fig. 14 A, B). The abdomen removed in specimen for DNA extraction. Female terminalia, ovipositor: General coloration dark yellow (Fig. 14 A). Cercus almost straight (Fig. 12 C). Genital fork cross-shaped, widening at posterior 1 / 3, lateral branches directed caudally. Lateral sclerite of genital plate very small, indistinct, less than 1 / 6 - 1 / 7 of length of genital fork. Genital opening T-shaped, two darker areas between genital fork and genital opening diffuse, twice longer than lateral sclerite (Fig. 12 D). Male: Unknown. Larva: Unknown. Pupa: Unknown.	en	Kolcsar, Levente-Peter, Kato, Daichi, Gamboa, Maribet, Watanabe, Kozo (2020): Revision of Japanese species of Nipponomyia Alexander, 1924 (Diptera, Pediciidae). ZooKeys 1000: 71-105, DOI: http://dx.doi.org/10.3897/zookeys.1000.55021, URL: http://dx.doi.org/10.3897/zookeys.1000.55021
2E7C04B0F37957B19F2D4ED19A78A5E3.taxon	distribution	Distribution. Japan: Ryukyu Islands: Okinawa Island (Fig. 9). Oriental region.	en	Kolcsar, Levente-Peter, Kato, Daichi, Gamboa, Maribet, Watanabe, Kozo (2020): Revision of Japanese species of Nipponomyia Alexander, 1924 (Diptera, Pediciidae). ZooKeys 1000: 71-105, DOI: http://dx.doi.org/10.3897/zookeys.1000.55021, URL: http://dx.doi.org/10.3897/zookeys.1000.55021
9D563E1DCB635234AC883B59B2DF1F92.taxon	materials_examined	Material examined. Non-types: Japan: [Honshu] • 2 ♂; Aomori, Nishimeya, Shirakami Nature observation garden, Kawaratai; alt. 255 m; 40 ° 31.13 ' N, 140 ° 12.89 ' E; 4 Jul. 2013; leg. D. Kato (pinned, BLKU) • 1 ♀ (GenBank # MT 874514); same data as previous except 6 Jul. 2013; D. Kato leg. (pinned, BLKU) • 1 ♂; Aomori, Nishimeya, Okawa Path, Kawaratai; alt. 300 m; 40 ° 30.04 ' N, 140 ° 12.24 ' E; 15 Jul. 2013; D. Kato leg. (pinned, BLKU) • 1 ♀; Aomori, Hirosaki, Inekari River, Koguriyama; alt. 170 m; 40 ° 32.19 ' N, 140 ° 29.22 ' E; 25 Jul. 2013; D. Kato leg. (pinned, BLKU) • 1 ♂; Aomori, Towada, Tsutanuma Path, Okuse; alt. 468 m; 40 ° 35.45 ' N, 140 ° 57.42 ' E; 21 Jun. 2014; D. Kato leg. (pinned, BLKU) • 3 ♂; Fukushima, Hinoemata, Hiuchigatake; alt. 1530 m; 36 ° 59.4 ' N, 139 ° 16.82 ' E; 16 Jul. 2019; D. Kato leg. (pinned, BLKU) • 1 ♂; Niigata, Tokamachi, Matsunoyama-Amamizukoshi, Mt Amamizu; alt. 920 m; 37 ° 1.47 ' N, 138 ° 33.77 ' E; 3 Jul. 2019; D. Kato leg. (pinned, BLKU) • 4 ♂, 1 ♀; Niigata, Tokamachi, Matsunoyama, Kyororo; alt. 310 m; 37 ° 5.97 ' N, 138 ° 36.98 ' E; 21 Jul. 2019; D. Kato leg. (pinned, BLKU).	en	Kolcsar, Levente-Peter, Kato, Daichi, Gamboa, Maribet, Watanabe, Kozo (2020): Revision of Japanese species of Nipponomyia Alexander, 1924 (Diptera, Pediciidae). ZooKeys 1000: 71-105, DOI: http://dx.doi.org/10.3897/zookeys.1000.55021, URL: http://dx.doi.org/10.3897/zookeys.1000.55021
9D563E1DCB635234AC883B59B2DF1F92.taxon	description	Diagnostic characters. Yellowish species with 11 dark spots on thorax (7 dark spots in N. okinawensis Kolcsar & Kato, sp. nov., 11 - 13 dark spots in N. kuwanai). Wing with transverse dark lines in costal cell. Brown marking extending from R 2 + 3 to crossvein m-m (brown marking extending from R 2 + 3 to maximum to base of M 1 in N. kuwanai and N. okinawensis Kolcsar & Kato, sp. nov.). Sternite 2 without black marking at corner of membranous area (with black marking at corner of membranous area in N. kuwanai), a diffuse line positioned same level as line on sternite 3 (N. kuwanai without this line, N. okinawensis Kolcsar & Kato, sp. nov. with any line and dark marking on sternite 2). Gonostylus with 4 or 5 spines (11 - 14 spines in N. kuwanai). Aedeagus short, twice as long as wide, rounded (as long as wide in N. kuwanai). Cercus curved upward (straight in N. okinawensis Kolcsar & Kato, sp. nov.). Female genital opening Y-shaped (T-shaped in N. okinawensis Kolcsar & Kato, sp. nov.), lateral sclerite 1 / 3 of length of genital fork (less than 1 / 5 of length of genital fork in N. kuwanai and less than 1 / 6 - 1 / 7 of length of genital fork in N. okinawensis Kolcsar & Kato, sp. nov.), genital fork cross-shaped (spoon-shaped in N. kuwanai, cross-shaped in N. okinawensis Kolcsar & Kato, sp. nov. but lateral branch curved caudally).	en	Kolcsar, Levente-Peter, Kato, Daichi, Gamboa, Maribet, Watanabe, Kozo (2020): Revision of Japanese species of Nipponomyia Alexander, 1924 (Diptera, Pediciidae). ZooKeys 1000: 71-105, DOI: http://dx.doi.org/10.3897/zookeys.1000.55021, URL: http://dx.doi.org/10.3897/zookeys.1000.55021
9D563E1DCB635234AC883B59B2DF1F92.taxon	distribution	Distribution. Japan: Honshu Island (Oosterbroek 2020, Nakamura 2014) (Fig. 9).	en	Kolcsar, Levente-Peter, Kato, Daichi, Gamboa, Maribet, Watanabe, Kozo (2020): Revision of Japanese species of Nipponomyia Alexander, 1924 (Diptera, Pediciidae). ZooKeys 1000: 71-105, DOI: http://dx.doi.org/10.3897/zookeys.1000.55021, URL: http://dx.doi.org/10.3897/zookeys.1000.55021
5CCE30F02A6756B88CFC0795527F093B.taxon	materials_examined	Material examined. Non-types: Japan: [Hokkaido] • 2 ♂, (GenBank # MT 874512); Hokkaido, Sobetsu, River Benkei; alt. 238 m; 42 ° 33.52 ' N, 140 ° 59.29 ' E; 29 Jul. 2019; L. - P. Kolcsar leg. (pinned or in ethanol, CKLP). [Honshu] • 3 ♂, 1 ♀; Aomori, Hirosaki, Ichinowatari-washinosu; alt. 205 m; 40 ° 31.15 ' N, 140 ° 26.33 ' E; 5 Sep. 2013; D. Kato leg. (pinned, BLKU) • 1 ♂; Aomori, Hirosaki, Inekari River, Koguriyama; alt. 170 m; 40 ° 32.19 ' N, 140 ° 29.22 ' E; 10 Sep. 2013; D. Kato leg. (pinned, BLKU) • 3 ♂; Aomori, Towada, Tsutanuma Path, Okuse; alt. 468 m; 40 ° 35.45 ' N, 140 ° 57.42 ' E; 30 Aug. 2014; D. Kato leg. (pinned, BLKU) • 2 ♂; Aomori, Towada, Sakura Spa, Okuse; alt. 854 m; 40 ° 37.64 ' N, 140 ° 54.59 ' E; 3 Aug. 2015; D. Kato leg. (pinned, BLKU) • 1 ♂; Gifu, Nakatsugawa, Nishimata-dani Valley, Kashimo; alt. 800 m; 35 ° 44.51 ' N, 137 ° 25.57 ' E; 7 Aug. 2015; D. Kato leg. (pinned, BLKU) • 1 ♂, 1 ♀; Hiroshima, Hatsukaichi, Nakatsudani-gawa River, Yoshiwa; alt. 900 m; 43 ° 0.83 ' N, 141 ° 20.01 ' E; 2 Sep. 2015; D. Kato leg. (pinned, BLKU) • 2 ♂; Hiroshima, Hatsukaichi, Mt Misaka-yama, Yoshiwa; alt. 1070 m; 34 ° 30.68 ' N, 132 ° 2.81 ' E; 2 Sep. 2015; D. Kato leg. (pinned, BLKU) • 2 ♂; Nagano, Ueda, Daimyozin stream, Sugadaira MRC; alt. 1315 m; 36 ° 31.2 ' N, 138 ° 21.24 ' E; 20 Aug. 2013; D. Kato leg. (pinned, BLKU) • 1 ♂; Tochigi, Nikko; alt. 675 m; 36 ° 44.44 ' N, 139 ° 37.1 ' E; 8 Sep. 2011; D. Kato leg. (pinned, BLKU) • 2 ♂; Tottori, Yazu, Mt Ogino-sen; alt. 905 m; 35 ° 25.85 ' N, 134 ° 25.57 ' E; 17 Sep. 2014; D. Kato leg. (pinned, BLKU) • 1 ♂; Yamagata, Sakata, Yunodai Spa, Kusatsu; alt. 475 m; 39 ° 1.57 ' N, 140 ° 1.56 ' E; 18 Sep. 2014; D. Kato leg. (pinned, BLKU). [Kyushu] • 2 ♂; Fukuoka, Mt Sefuri, Itaya, Sawara-ku; alt. 970 m; 33 ° 26.29 ' N, 130 ° 22 ' E; 5 Sep. 2015; D. Kato leg. (pinned, BLKU) • 2 ♀; Fukuoka, Miyako, Notoge Pass, Saigawa-Hobashira; alt. 740 m; 33 ° 29.74 ' N, 130 ° 57.69 ' E; 21 Sep. 2015; D. Kato leg. (pinned, BLKU) • 1 ♂; Fukuoka, Fukuoka, Katae, Mt Abura; alt. 225 m; 33 ° 31.83 ' N, 130 ° 21.96 ' E; 20 Oct. 2015; D. Kato leg. (pinned, BLKU) • 1 ♀; Oita, Yufu, Shonai-cho-asono; alt. 870 m; 33 ° 9.65 ' N, 131 ° 20.21 ' E; 10 Sep. 2016; D. Kato leg. (pinned, BLKU) • 1 ♂; Saga, Saga-shi, Kasa River near Hokuza Dam, Fujimachi-sekiya; alt. 330 m; 33 ° 25.99 ' N, 130 ° 13.93 ' E; 15 Oct. 2015; D. Kato leg. (pinned, BLKU). [Shikoku] • 1 ♂; Ehime, Wakayama, small waterfall and stream; alt. 1305 m; 33 ° 44.71 ' N, 133 ° 8.23 ' E; 10 Sep. 2019; L. - P. Kolcsar leg. (pinned or in ethanol, CKLP) • 3 ♂, 1 ♀, (GenBank # MT 874513); Ehime, Toon, Shiraino waterfall; alt. 685 m; 33 ° 45.54 ' N, 132 ° 58.16 ' E; 16 Sep. 2019; L. - P. Kolcsar leg. (pinned or in ethanol, CKLP) • 2 ♂; Tokushima, Miyoshi, Ochiai Pass, Higashiiya-Ochiai; alt. 1460 m; 33 ° 55.33 ' N, 133 ° 56.88 ' E; 15 May 2015; D. Kato leg. (pinned, BLKU).	en	Kolcsar, Levente-Peter, Kato, Daichi, Gamboa, Maribet, Watanabe, Kozo (2020): Revision of Japanese species of Nipponomyia Alexander, 1924 (Diptera, Pediciidae). ZooKeys 1000: 71-105, DOI: http://dx.doi.org/10.3897/zookeys.1000.55021, URL: http://dx.doi.org/10.3897/zookeys.1000.55021
5CCE30F02A6756B88CFC0795527F093B.taxon	description	Diagnostic characters. Yellowish species with 11 darker spots on thorax (N. yakushimensis Kolcsar & Kato, sp. nov. dark yellow species with 11 large dark spots, N. gracilis without dark spots on thorax). Wing without transverse dark line in costal cell. Brown band running from base of R 2 + 3 to tip of M 4 and to m-cu (brown band not reaching wing margin in N. yakushimensis Kolcsar & Kato, sp. nov.). Dark band along crossveins r-m and m-cu conspicuous. Second sternite with black marking at corner of membranous area, but without other line. Gonostylus with 3 spines (2 spines in N. yakushimensis Kolcsar & Kato, sp. nov. and N. gracilis), aedeagus short, triangular, and acute at tip in lateral view (aedeagus long, rod-shaped in N. yakushimensis Kolcsar & Kato, sp. nov. and N. gracilis). Cercus long and straight, just gently curved upward. Genital opening T-shaped, genital fork cross-shaped, lateral sclerite large, half as long as genital fork.	en	Kolcsar, Levente-Peter, Kato, Daichi, Gamboa, Maribet, Watanabe, Kozo (2020): Revision of Japanese species of Nipponomyia Alexander, 1924 (Diptera, Pediciidae). ZooKeys 1000: 71-105, DOI: http://dx.doi.org/10.3897/zookeys.1000.55021, URL: http://dx.doi.org/10.3897/zookeys.1000.55021
5CCE30F02A6756B88CFC0795527F093B.taxon	distribution	Distribution. Russia: Kuril Islands. Japan: Honshu, Shikoku (Nakamura 2014; Oosterbroek 2020), first records from Hokkaido and Kyushu (Fig. 17).	en	Kolcsar, Levente-Peter, Kato, Daichi, Gamboa, Maribet, Watanabe, Kozo (2020): Revision of Japanese species of Nipponomyia Alexander, 1924 (Diptera, Pediciidae). ZooKeys 1000: 71-105, DOI: http://dx.doi.org/10.3897/zookeys.1000.55021, URL: http://dx.doi.org/10.3897/zookeys.1000.55021
79092BEEC9E35D3CB9D0E918F2C73F3A.taxon	description	Diagnostic characters. Dark yellow species with 11 large darker spots on thorax (N. trispinosa light yellowish species with 11 smaller dark spots, N. gracilis brownish species without any dark spots on thorax). Wing without transverse dark line on costal cell. Brown band running from base of R 2 + 3 to crossvein m-m, but not reaching wing margin (reaching the wing margin in N. trispinosa). Brown band along crossveins r-m and m-cu conspicuous. Second sternite with black marking at corner of membranous area, but without other line. Gonostylus with 2 spines (3 spines in N. trispinosa), aedeagus long, rod-shaped and acute at tip (aedeagus short, triangular in N. trispinosa).	en	Kolcsar, Levente-Peter, Kato, Daichi, Gamboa, Maribet, Watanabe, Kozo (2020): Revision of Japanese species of Nipponomyia Alexander, 1924 (Diptera, Pediciidae). ZooKeys 1000: 71-105, DOI: http://dx.doi.org/10.3897/zookeys.1000.55021, URL: http://dx.doi.org/10.3897/zookeys.1000.55021
79092BEEC9E35D3CB9D0E918F2C73F3A.taxon	description	Description. Body length: male 8 - 8.5 mm. Wing length: male 8 - 8.5 mm. Head: Light brown to brown with grayish pruinosity (Fig. 18 B, C). Palpi dark brown, 5 - segmented, segments 2 - 4 subequal in length, last segment twice as long as palpomere 4. Antenna short, just a little longer than head. Antenna brown, flagellomeres darker than scape and pedicel. Flagellum 13 - segmented, flagellomeres gradually narrowing to apical end. Thorax: General coloration dark yellow, dorsal parts somewhat darker (Fig. 18 A). Presutural area of scutum with 4 large spots, very conspicuous (Fig. 18 B, C) and 7 spots on postsutural area of scutum also distinct (Fig. 18 C). Setae on thorax relatively long and dark. Legs: General coloration yellow, covered with yellowish setae. Femora without clear apical dark area, but with some darker setae. Apical part of tibiae light brown, with a few darker setae. Apical ends of tarsomeres narrowly dark yellow to light brown (Fig. 18 A). Tarsomeres with spurs very small, hardly discernible. Wing: As in Fig. 5 G. Yellow pattern less intensive compared to other Japanese species. Spots around yellow costal region brown, not blackish as in N. trispinosa. Brown band running from base of R 2 + 3 to crossvein m-m, but not reaching wing margin. Brown band along crossveins r-m and m-cu conspicuous. In paratype, wing with Rs divided to R 2 + 3 + 4 and R 5 (Fig. 5 G), in holotype as usual in genus, divided to R 2 + 3 and R 4 + 5. Abdomen: Abdomen covered with relative long dark setae. Tergites 2 - 6 each with a longitudinal narrow black line on lateral side, its length ranging from 1 / 2 - 3 / 4 of tergite length. Sternite 2 with a short black line at corner of membranous fold. Sternites 3 - 7 each with a broad brown patch, covering anterior half of segment (Fig. 18 A, B). The abdomen of paratype removed for DNA extraction. Male terminalia: Dark yellow to light brown (Fig. 18 A). Median part of tergite 9 with posterior margin convex with two small obtuse peaks laterally (Fig. 19 A, B). Gonocoxite with apical lobe 2.2 x longer than wide (in the middle) and 1.6 x longer than tergite 9 (Fig. 19 E, F). Apical lobe of gonocoxite squarish in dorsal and ventral views (Fig. 19 A-D), as long as width of gonocoxite at middle in lateral view (Fig. 19 G, H). Basal lobe of gonocoxite prominent, both in ventral and lateral views, triangular in ventral view (Fig. 19 C, D). Outer part of gonostylus slender in inner view (Fig. 19 G, H), with 2 black spines, inner part of gonostylus triangular (Fig. 19 A-D). Interbase elongated, gradually widening to tip, widest part twice wider than basal part in dorsal view (Fig. 19 A, B), interbase curved dorsally in lateral view (Fig. 19 G, H). Aedeagus rod-shaped, extending beyond interbase, tip acute, curved dorsally (Fig. 19 I, J). Female: Unknown. Larva: Unknown. Pupa: Unknown.	en	Kolcsar, Levente-Peter, Kato, Daichi, Gamboa, Maribet, Watanabe, Kozo (2020): Revision of Japanese species of Nipponomyia Alexander, 1924 (Diptera, Pediciidae). ZooKeys 1000: 71-105, DOI: http://dx.doi.org/10.3897/zookeys.1000.55021, URL: http://dx.doi.org/10.3897/zookeys.1000.55021
79092BEEC9E35D3CB9D0E918F2C73F3A.taxon	distribution	Distribution. Japan: Ryukyu Islands: Yakushima Island (Fig. 9).	en	Kolcsar, Levente-Peter, Kato, Daichi, Gamboa, Maribet, Watanabe, Kozo (2020): Revision of Japanese species of Nipponomyia Alexander, 1924 (Diptera, Pediciidae). ZooKeys 1000: 71-105, DOI: http://dx.doi.org/10.3897/zookeys.1000.55021, URL: http://dx.doi.org/10.3897/zookeys.1000.55021
