taxonID	type	description	language	source
73FEB24B8930A04DB39EFB223FCBED8A.taxon	materials_examined	Material examined. Holotype. ♂, THAILAND, Surat Thani Province, Phanom District, near Khlong Phanom National Park, Pha Daeng, 8 ° 53 ' 41 " N, 98 ° 33 ' 12 " E, ca. 67 m a. s. l., 7 Aug. 2016, ASRU members leg. (CUMZ-pxDGT 00205). Paratypes. 17 ♂♂, 24 ♀♀, same data as for holotype (CUMZ- pxDGT 00206); 1 ♂, 1 ♀, same data as for holotype (ZMUC 00040249); 1 ♂, 1 ♀, same data as for holotype (NHMW 9423). Further specimens, not paratypes. 5 ♂♂, 3 ♀♀, THAILAND, Surat Thani Province, Phanom District, near Khlong Phanom National Park, Pha Daeng, 8 ° 53 ' 41 " N, 98 ° 33 ' 12 " E, ca. 67 m a. s. l., 6 Aug. 2015, ASRU members leg. (CUMZ).	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
73FEB24B8930A04DB39EFB223FCBED8A.taxon	etymology	Etymology. The species name is an adjective, refers to the two additional process-like lobes on the solenophore (one on lamina lateralis and one on lamina medialis).	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
73FEB24B8930A04DB39EFB223FCBED8A.taxon	diagnosis	Diagnosis. Differs from other species by having: metaterga 5 - 19 with 2 + 2 cones in anterior row and 3 + 3 cones in posterior row; sternal lobe between male coxae 4 subtrapeziform; male femora 8 and 9 with apophyses distally; lamina lateralis with an additional process-like protruding lobe; lamina medialis basally with an additional protruding process-like lobe.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
73FEB24B8930A04DB39EFB223FCBED8A.taxon	description	Description. SIZE. Length 15 - 17 mm (male), 16 - 18 mm (female); width of midbody metazona 1.0 - 1.2 mm (male), 1.3 - 1.5 mm (female). Width of rings 2 = 3 <4 <collum <5 <head = 6 - 17, thereafter body gradually tapering towards telson. Colour (Figure 6 A, B). Specimens in life brown / pale brown; paraterga brownish white; head, antennae (except whitish distal part of antennomeres 7 and 8) and collum brown; prozona, metaterga (except white spines in posterior row) and surface below paraterga brown / pale brown; sterna pale brown / whitish brown; epiproct and legs whitish brown; a few basal podomeres whitish brown / white. Antennae. Reaching to body ring 7 or 8 (male) and 6 (female) when stretched dorsally. Collum (Figure 7 A). With three transverse rows of setiferous cones, 4 + 4 in anterior row, 1 + 1 in intermediate row and 2 + 2 in posterior row; with one inconspicuous setiferous notch at lateral margin; paraterga wing-like, quite short, tip blunt, elevated at ca. 15 ° - 20 ° (male) 10 ° - 20 ° (female), directed almost caudad. Tegument. Quite shining; collum coarsely microgranulate; metaterga and surface below paraterga finely microgranulate. Metaterga (Figure 7 A, C, E). With two transverse rows of setiferous cones; metaterga 2 - 4 with 2 + 2 cones in anterior row and 2 (3) + 2 (3) cones in posterior row; metaterga 5 - 19 with 2 + 2 cones in anterior row and 3 + 3 cones in posterior row; all cones subequal in length and size. An additional cone-like denticle at base of paraterga near anterior row of cones. Paraterga (Figure 7 A-D, H). Quite short; directed dorsocaudad on body rings 3 - 17, elevated at ca. 60 ° - 70 ° (male) 55 ° - 70 ° (female), directed more caudad on body ring II and increasingly so on rings 18 and 19. Denticle of paraterga located at base of paraterga and very close to anterior row of cones on metaterga. Ozopore visible in lateral view. Telson (Figure 7 E, I, H). Epiproct quite long; tip subemarginate; lateral setiferous tubercles conspicuous; apical tubercles conspicuous. Hypoproct subtrapeziform, wide; caudal margin round (in some specimens subtruncate), with inconspicuous setiferous tubercles. Sterna (Figs 4 A; 7 F, G). Sternal lobe between male coxae 4 subtrapeziform, broad, and thin, tips subtruncate, in situ directed ventroanteriad; posterior surface of sternal lobe with two pores borne on swollen and short lobe. Legs (Figs 4 B, C; 7 J). Male femora 8 and 9 with apophyses distally. Gonopods (Figs 5 A, 8). Coxa shorter than femur. Prefemoral part ca. half as long as femur. Femur not enlarged distally. Postfemoral part broad. Mesal sulcus wide; lateral sulcus narrow. Solenophore subequal in size to postfemoral part: lamina lateralis small and short; with a protruding lobe, process-like, directed mesad; apically round: lamina medialis long; base enlarged and stout, slightly attenuated near the tip, basally with a protruding lobe, process-like, directed mesad; tip curving down, bifurcating into two small spines. Solenomere curving down, compressed in transverse section, tip directed posteriad.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
73FEB24B8930A04DB39EFB223FCBED8A.taxon	distribution	Distribution and habitat (Figure 6 C). Known only from the type locality which is a small isolated limestone mountain between Khao Sok and Khlong Phanom National Parks. The new species blended perfectly with the humid rock walls, and most specimens were found inside rock holes / crevices. S. biloba sp. n. co-occurs with Desmoxytes cervina (Pocock, 1895) (Srisonchai et al. 2018 a) in the same habitat. Several attempts (2017 - 2018) have been made to find further specimens near the type locality, but none were found. As the new species has only been found at the type locality only, we regard S. biloba sp. n. as endemic to Thailand.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
E8FFC23ACFAEB1C82C1AEAAE10208984.taxon	materials_examined	Material examined. Holotype. ♂, MYANMAR, Tanintharyi Region, Myeik, 20 km northeast of Monoron, Lenya National Park, limestone mountain near Ngawun Chaung River, 11 ° 40 ' 20 " N, 99 ° 13 ' 30 " E, ca. 64 m a. s. l., 9 Jun. 2015, FFI staff and ASRU members leg. (CUMZ-pxDGT 00207). Paratypes. 20 ♂♂, 25 ♀♀, same data as for holotype (CUMZ-pxDGT 00208); 1 ♂, 1 ♀, same data as for holotype (ZMUC 00040250); 1 ♂, 1 ♀, same data as for holotype (ZMUM); 1 ♂, 1 ♀, same data as for holotype (NHMW 9422); 1 ♂, 1 ♀, same data as for holotype (NHMUK).	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
E8FFC23ACFAEB1C82C1AEAAE10208984.taxon	etymology	Etymology. The name is an artificially constructed homophone (efefi = FFI) honouring FFI (Fauna and Flora International, Myanmar), an organization for biodiversity conservation; in recognition of their hard work to protect wildlife including invertebrates.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
E8FFC23ACFAEB1C82C1AEAAE10208984.taxon	diagnosis	Diagnosis. Sternal lobe between male coxae 4 not bilobed and male femora without modification. Similar in this respect to S. hasta sp. n., but differs by having: collum with 5 (4) + 5 (4) tubercles in anterior row, 2 + 2 tubercles in intermediate row and 3 + 3 tubercles in posterior row; metaterga 2 - 8 with 3 + 3 cones in anterior row and 3 + 3 cones in posterior row; metaterga 9 - 18 with 3 + 3 cones in anterior row and 4 + 4 cones in posterior row; metatergum 19 with 3 + 3 tubercles / cones in anterior row and 4 + 4 tubercles / cones in posterior row; postfemoral part of gonopod with a triangular process and a triangular ridge.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
E8FFC23ACFAEB1C82C1AEAAE10208984.taxon	description	Description. SIZE. Length 26 - 30 mm (male), 30 - 32 mm (female); width of midbody metazona 2.1 - 2.2 mm (male), 2.7 - 2.9 mm (female). Width of collum = ring 2 = 3 = 4 <head = 5 - 17, thereafter body gradually tapering towards telson. Colour (Figure 9 A, B). Specimens in life with body brown / yellowish brown; paraterga yellow; antennae (except whitish distal part of antennomeres 7 and 8), head and prozona brown / blackish brown; collum, metaterga and surface below paraterga brown / yellowish brown; sterna, epiproct and legs brown; a few basal podomeres pale brown / whitish brown. Antennae. Reaching to body ring 9 or 10 (male) and 7 or 8 (female) when stretched dorsally. Collum (Figure 10 A). With three transverse rows of setiferous tubercles, 5 (4) + 5 (4) tubercles in anterior row, 2 + 2 (1) tubercles in intermediate row and 3 + 3 tubercles in posterior row; with two inconspicuous setiferous notches at lateral margin; paraterga wing-like, quite short and small, tip obtuse, elevated at ca. 15 ° - 25 ° (male) 10 ° - 15 ° (female), directed caudolaterad. Tegument. Quite dull; collum, metaterga (posterior part) and surface below paraterga coarsely microgranulate; metaterga (anterior part) smooth. Metaterga (Figure 10 A, C, E). With two transverse rows of setiferous cones; metaterga 2 - 8 with 3 + 3 cones in anterior row and 3 + 3 cones in posterior row; metaterga 9 - 19 with 3 (4) + 3 (4) cones in anterior row and 4 (5) + 4 (5) cones in posterior row; all cones subequal in length and size. Paraterga (Figure 10 A-E, H). Very long; directed almost dorsad on body rings 2 - 16, elevated at ca. 65 ° - 80 ° (male) 60 ° - 70 ° (female); directed dorsocaudad on ring 17; directed increasingly caudad on body rings 18 and 19. Ozopore visible in lateral view. Telson (Figure 10 E, H, I). Epiproct quite long; tip subtruncate; lateral setiferous tubercles conspicuous; apical tubercles inconspicuous. Hypoproct subsemicircular; caudal margin round (in some specimens angular), with conspicuous setiferous tubercles. Sterna (Figs 4 D; 10 F, G). Sternal lobe between male coxae 4 erect, subrectangular, very long; tips emarginate, in situ directed ventrad; posterior surface bearing one pore near tip. Legs (Figs 4 E, 10 J). Male femora without modification. Gonopods (Figs 5 B, 11). Coxa subequal in length to femur. Prefemoral part ca. 2 / 3 as long as femur. Femur not enlarged distally, ventrally swollen in middle part. Postfemoral part broad; mesally with a long triangular process (directed mesoanteriad) and a long triangular ridge, between process and ridge with a wide furrow. Mesal sulcus and lateral sulcus wide. Solenophore bigger and longer than postfemoral part; basally very broad: lamina lateralis long and slender, curved, tip round: lamina medialis long and slender; with two ridges in middle portion; slightly attenuated near tip; tip in situ resting very close to solenomere, terminating in small spines. Solenomere flat, curving down; tip terminating in three sharp spines, directed mesoventrad.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
E8FFC23ACFAEB1C82C1AEAAE10208984.taxon	distribution	Distribution and habitat (Figure 9 C). Known only from the type locality. The specimens were found exclusively on rock walls or in caves. We have tried to find this species in other places near the type locality, but no further specimens have been collected. Given the finding only at the type locality, the new species is considered to be endemic to southern Myanmar.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
90F95CA9F7EBA1A853B70F51883E00D8.taxon	materials_examined	Material examined. Holotype. ♂, THAILAND, Chumphon Province, Thung Tako District, Khao Ma Ngaen, 10 ° 05 ' 27 " N, 99 ° 04 ' 25 " E, ca. 28 m a. s. l., 23 Oct. 2016, ASRU members leg. (CUMZ-pxDGT 00209). Paratypes. 5 ♂♂, 6 ♀♀, same data as for holotype (CUMZ-pxDGT 00210); 1 ♂, 1 ♀, same data as for holotype (ZMUC 00040251). Further specimens, not paratypes, all from THAILAND, Chumphon Province. Mueang Chumphon District: 8 ♂♂, 1 ♀, Wat Tham Sanook, 10 ° 28 ' 52 " N, 99 ° 04 ' 29 " E, ca. 54 m a. s. l., 3 Jul. 2017, ASRU members leg. (CUMZ). Lang Suan District: 2 ♀♀, Wat Ratcha Burana School, 9 ° 56 ' 21 " N, 99 ° 02 ' 26 " E, ca. 34 m a. s. l., 10 Sep. 2016, ASRU members leg. (CUMZ); 1 ♂, 5 ♀♀, Wat Tham Khao Kriap (Khao Kriap Cave), 9 ° 49 ' 08 " N, 99 ° 02 ' 22 " E, ca. 102 m a. s. l., 5 Jun. 2009, ASRU members leg. (CUMZ). Sawi District: 8 ♂♂, 3 ♀♀, Wat Nam Cha, 10 ° 17 ' 54 " N, 99 ° 01 ' 57 " E, ca. 95 m a. s. l., 5 Jun. 2009, ASRU members leg. (CUMZ).	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
90F95CA9F7EBA1A853B70F51883E00D8.taxon	etymology	Etymology. The name is a Latin noun in apposition meaning spear, referring to the shape of the sternal lobe between male coxae 4 which is somewhat similar to a spear.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
90F95CA9F7EBA1A853B70F51883E00D8.taxon	diagnosis	Diagnosis. Sternal lobe between male coxae 4 not bilobed, not bifurcate; male femora without modification. Similar in this respect to S. efefi sp. n., but differs by having: collum with 4 + 4 tubercles in anterior row, 1 + 1 tubercles in intermediate row and 2 + 2 tubercles in posterior row; metaterga 2 - 8 with 2 + 2 cones in anterior row and 2 + 2 cones in posterior row; metaterga 9 - 18 with 2 + 2 cones in anterior row and 2 + 2 cones in posterior row; metatergum 19 with 2 + 2 tubercles / cones in anterior row and 2 + 2 tubercles / cones in posterior row; lamina medialis (lm) with a large lobe in middle part.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
90F95CA9F7EBA1A853B70F51883E00D8.taxon	description	Description. SIZE. Length 23 - 33 mm (male), 26 - 33 mm (female); width of midbody metazona 1.7 - 2.2 mm (male), 2.1 - 2.8 mm (female). Width of collum = ring 2 = 3 = 4 <head = 5 - 16, thereafter body gradually tapering towards telson. Colour (Figure 12 A-C). Specimens in life with body black / brownish black; paraterga white / yellowish white / whitish yellow; antennae (except whitish distal part of antennomeres 7 and 8) and metaterga (posterior part) brown / brownish black; head and collum brown / blackish brown; prozona and metaterga (anterior part) black; surface below paraterga black / brownish black; sterna and epiproct brown; legs brown / pale brown; a few basal podomeres pale whitish brown. Antennae (Figure 13 M). Reaching to body ring 9 or 10 (male) and 7 (female) when stretched dorsally. Collum (Figure 13 A). With three transverse rows of setiferous tubercles, 4 + 4 tubercles in anterior row, 1 (0) + 1 tubercles in intermediate row and 2 + 2 tubercles in posterior row; with two inconspicuous setiferous notches at lateral margin; paraterga wing-like, quite short and broad, tip sharp, elevated at ca. 10 ° - 15 ° (male) 10 ° - 15 ° (female), directed caudolaterad. Tegument. Quite dull; collum, metaterga and surface below paraterga finely microgranulate. Metaterga (Figure 13 A, C, E). With two transverse rows of setiferous tubercles / cones and spines; metaterga 2 - 8 with 2 + 2 tubercles / cones in anterior row and 2 + 2 spines in posterior row; metaterga 2 - 18 with 2 + 2 tubercles / cones in anterior row and 2 + 2 spines in posterior row; metatergum 19 with 2 + 2 tubercles in anterior row and 2 + 2 tubercles in posterior row; lateral spines of posterior row bigger and longer than mesal ones, gradually reduced in length and size on the following rings. Paraterga (Figure 13 A-E, H). Very long; directed dorsolaterad on body rings 4 - 16, elevated at ca. 60 ° - 70 ° (male) 50 ° - 60 ° (female); directed caudolaterad on rings 2, 3 and 17; directed increasingly caudad on body rings 18 and 19. Ozopore visible in lateral view. Telson (Figure 13 E, H, I). Epiproct long; tip subtruncate; lateral setiferous tubercles conspicuous (in some specimens inconspicuous); apical tubercles inconspicuous. Hypoproct subtrapeziform (in some specimens subsemicircular); caudal margin round (in some specimens angular), with inconspicuous setiferous tubercles. Sterna (Figs 4 F; 13 F, G). Sternal lobe between male coxae 4 coniform, long, spear-like; base stout; tips sharp, in situ directed almost ventrad; posterior surface bearing one pore. Legs (Figs 4 G, 13 J). Male femora without modification. Gonopods (Figs 5 C, 14). Coxa shorter than femur. Prefemoral part ca. 2 / 3 as long as femur. Femur not enlarged distally, basally indented. Postfemoral part narrow. Mesal sulcus and lateral sulcus wide. Solenophore bigger and longer than postfemoral part: lamina lateralis broad and long, flattened laterally: lamina medialis long; base enlarged, slightly attenuated near the tip; middle part with a large lobe; tip a bit curving up, terminating in several small spines. Solenomere circular in transverse section, curving up, tip directed anteriad.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
90F95CA9F7EBA1A853B70F51883E00D8.taxon	distribution	Distribution and habitat (Figure 12 D). The specimens were found on rocks or walls with plants, and some were found in a small cave. S. hasta sp. n. is distributed only in Chumphon Province, and we regard the new species as endemic for the Thai fauna. At Wat Nam Cha the new species coexists with Desmoxytes cervina.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
334D026F96583BC5FA1AD12AC8AAC23A.taxon	materials_examined	Material examined. Holotype. ♂, THAILAND, Krabi Province, Mueang Krabi District, Wat Tham Sue (Tiger Cave), valley behind Tiger Cave, 8 ° 07 ' 38 " N, 98 ° 55 ' 26 " E, ca. 87 m a. s. l., 9 Jul. 2017, ASRU members leg. (CUMZ-pxDGT 00211). Paratypes. 5 ♂♂, 9 ♀♀, 1 juvenile, same data as for holotype (CUMZ-pxDGT 00212); 1 ♂, 1 ♀, same data as for holotype (ZMUC 00040252). Further specimens, not paratypes, all from THAILAND, Krabi Province. Ao Luek District: 2 ♂♂, 1 ♀, Than Bok Khorani, 8 ° 23 ' 28 " N, 98 ° 44 ' 07 " E, ca. 46 m a. s. l., 14 Jan. 2013, ASRU members leg. (CUMZ); 5 ♂♂, 1 ♀, Than Bok Khorani, 8 ° 23 ' 28 " N, 98 ° 44 ' 07 " E, ca. 46 m a. s. l., 23 Aug. 2014, ASRU members leg. (CUMZ); 12 ♂♂, 7 ♀♀, 1 juvenile, Than Bok Khorani, 8 ° 23 ' 28 " N, 98 ° 44 ' 07 " E, ca. 46 m a. s. l., 30 Aug. 2015, ASRU members leg. (CUMZ); 3 ♂♂, Than Bok Khorani, 8 ° 23 ' 28 " N, 98 ° 44 ' 07 " E, ca. 46 m a. s. l., Jan. 2016, ASRU members leg. (CUMZ); 1 ♂, 5 ♀♀, P. N. Mountain Resort, 8 ° 24 ' 09 " N, 98 ° 44 ' 18 " E, ca. 46 m a. s. l., 30 Aug. 2015, ASRU members leg. (CUMZ); 1 ♂, 1 broken ♂, Tham Sa Yuan Thong (Sa Yuan Thong Cave), 8 ° 23 ' 29 " N, 98 ° 46 ' 17 " E, ca. 7 m a. s. l., 9 Oct. 2006, ASRU members leg. (CUMZ). Muaeng Krabi District: 1 broken ♂, Wat Tham Sue (Tiger Cave), valley behind Tiger Cave, 8 ° 07 ' 38 " N, 98 ° 55 ' 26 " E, ca. 87 m a. s. l., 25 Oct. 2007, ASRU members leg. (CUMZ); 1 broken ♂, 1 ♀, Wat Tham Sue (Tiger Cave), valley behind Tiger Cave, 8 ° 07 ' 38 " N, 98 ° 55 ' 26 " E, ca. 87 m a. s. l., 7 Oct. 2009, ASRU members leg. (CUMZ); 1 ♂, Wat Tham Sue (Tiger Cave), valley behind Tiger Cave, 8 ° 07 ' 38 " N, 98 ° 55 ' 26 " E, ca. 87 m a. s. l., 24 Aug. 2014, ASRU members leg. (CUMZ); 1 ♀, Wat Tham Sue (Tiger Cave), valley behind Tiger Cave, 8 ° 07 ' 38 " N, 98 ° 55 ' 26 " E, ca. 87 m a. s. l., 30 Aug. 2015, P. Pimvichai, P. Prasankok and N. Natarat leg. (CUMZ); 2 ♂♂, 1 ♀, 1 broken ♀, Wat Tham Sue (Tiger Cave), valley behind Tiger Cave, 8 ° 07 ' 38 " N, 98 ° 55 ' 26 " E, ca. 87 m a. s. l., 25 Jul. 2017, ASRU members leg. (CUMZ).	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
334D026F96583BC5FA1AD12AC8AAC23A.taxon	etymology	Etymology. The new species is named after the province where the type locality lies.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
334D026F96583BC5FA1AD12AC8AAC23A.taxon	diagnosis	Diagnosis. Male femora 6 and 7 humped distally. Similar in this respect to S. macaca sp. n., but differs by having: paraterga orange, longer; male femora 6 smaller; tip of lamina lateralis round, not protuding as digitiform; tip of lamina medialis terminating in two lobes.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
334D026F96583BC5FA1AD12AC8AAC23A.taxon	description	Description. SIZE. Length 28 - 31 mm (male), 30 - 33 mm (female); width of midbody metazona 1.8 - 2.0 mm (male), 2.2 - 2.5 mm (female). Width of rings 2 = 3 = 4 <collum <head = 5 - 16, thereafter body gradually tapering towards telson. Colour (Figure 15 A-D). Specimens in life with body black / brownish black; paraterga orange; head, antennae (except whitish distal part of antennomeres 7 and 8), collum, prozona and epiproct black; metaterga and surface below paraterga black / brownish black; sterna brown; legs brown / blackish brown; a few basal podomeres whitish brown. Antennae (Figure 16 M). Reaching to body ring 8 (male) and 6 (female) when stretched dorsally. Collum (Figure 16 A). With three transverse rows of setiferous tubercles / cones, 4 + 4 in anterior row, 1 + 1 in intermediate row and 2 + 2 in posterior row; with one conspicuous setiferous notch at lateral margin; paraterga spiniform, long, tip sharp, elevated at ca. 20 ° - 30 ° in both male and female, directed caudolaterad. Tegument. Quite shining; collum coarsely microgranulate; metaterga and surface below paraterga smooth. Metaterga (Figure 16 A, C, E). With two transverse rows of setiferous tubercles and setiferous cones / spines; metaterga 2 - 7 with 2 + 2 tubercles in anterior row and 2 + 2 spines in posterior row; 8 - 19 with 2 + 2 tubercles in anterior row and 2 + 2 cones in posterior row; lateral cones / spines of posterior row bigger and longer than mesal ones, gradually reduced in size and length on the following rings. Paraterga (Figure 16 A-E, H). Extremely long; directed dorsolaterad on body rings 2 - 16, elevated at ca. 45 ° - 60 ° (male) 40 ° - 50 ° (female), directed dorsocaudad on ring 17, directed increasingly caudad on body rings 18 and 19. Ozopore visible in subdorsal view. Telson (Figure 16 E, H, I). Epiproct long; tip subtruncate; lateral setiferous tubercles mostly inconspicuous (in some specimens conspicuous); apical tubercles inconspicuous. Hypoproct subtrapeziform; caudal margin round, with inconspicuous setiferous tubercles. Sterna (Figs 4 H; 16 F, G). Sternal lobe between male coxae 4 bifurcate, long; tips sharp, in situ directed ventroanteriad; posterior surface bearing one pore. Legs (Figs 4 I, J; 16 J). Male femora 6 a bit humped; male femora 7 strongly humped. Gonopods (Figs 3, 5 D, 17). Coxa subequal in length to femur. Prefemoral part ca. half as long as femur. Femur obviously enlarged distally. Postfemoral part narrow. Mesal sulcus and lateral sulcus wide. Solenophore bigger than postfemoral part: lamina lateralis small, compact, tip round: lamina medialis long; basally enlarged and slightly attenuated near the tip; tip a bit curved, terminating in two lobes. Solenomere curved and twisted, compressed in transverse section, tip directed lateroposteriad.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
334D026F96583BC5FA1AD12AC8AAC23A.taxon	distribution	Distribution and habitat (Figure 15 E). S. krabiensis sp. n. inhabits Krabi Province. Considering its narrow distribution, we regard this species as endemic for the Thai fauna. It is syntopic with Desmoxytes delfae (Jeekel, 1964), Desmoxytes cervina and Gigaxytes gigas (Golovatch & Enghoff, 1994), which were collected from the same location (Than Bok Khorani and Wat Tham Sue (Tiger Cave )), but the new species was encountered living on rock walls or in small caves while the others were usually found on leaf litter or on tree branches.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
1C33B2F6B1A8B6ADCFC80C5E77F1E993.taxon	materials_examined	Material examined. Holotype. ♂, THAILAND, Phang Nga Province, Takua Thung District, Wat Suwan Khuha (Monkey Cave), 8 ° 25 ' 42 " N, 98 ° 28 ' 22 " E, ca. 27 m a. s. l., 8 Aug. 2016, ASRU members leg. (CUMZ-pxDGT 00213). Paratypes. 7 ♂♂, 2 ♀♀, same data as for holotype (CUMZ-pxDGT 00214); 1 ♂, 1 ♀, same data as for holotype (ZMUC 00040253); 1 ♂ same data as for holotype (ZMUM).	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
1C33B2F6B1A8B6ADCFC80C5E77F1E993.taxon	etymology	Etymology. The species is named after the monkey, long-tailed macaque (Macaca fascicularis) living at the type locality (Monkey Cave).	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
1C33B2F6B1A8B6ADCFC80C5E77F1E993.taxon	diagnosis	Diagnosis. Male femora 6 and 7 humped distally. Similar in this respect to S. krabiensis sp. n., but differs from it by having: paraterga brownish white, shorter; male femora 6 larger; tip of lamina lateralis (ll) protruding as a small lobe, digitiform; tip of lamina medialis (lm) bent, terminating in one lobe.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
1C33B2F6B1A8B6ADCFC80C5E77F1E993.taxon	description	Description. SIZE. Length 27 - 29 mm (male), 29 - 32 mm (female); width of midbody metazona 1.8 - 1.9 mm (male), 2.0 - 2.3 mm (female). Width of collum = 2 = 3 <4 <head = 5 - 16, thereafter body gradually tapering towards telson. Colour (Figure 18 A, B). Specimens in life with body black; paraterga brownish white; head, antennae (except whitish distal part of antennomeres 7 and 8), collum, metaterga, prozona and surface below paraterga black; sterna brown; epiproct black / brownish black; legs blackish brown; a few basal podomeres pale brown / whitish brown. Antennae (Figure 19 M). Reaching to body ring 8 or 9 (male) and 7 (female) when stretched dorsally. Collum (Figure 19 A). With three transverse rows of setiferous tubercles / cones, 4 + 4 tubercles / cones in anterior row, 1 + 1 tubercles / cones in intermediate row and 2 + 2 tubercles / cones in posterior row; with one conspicuous setiferous notch at lateral margin; paraterga spiniform, long, tip sharp, elevated at ca. 15 ° - 20 ° in both male and female, directed caudolaterad. Tegument. Quite shining; collum, metaterga and surface below paraterga smooth. Metaterga (Figure 19 A, C, E). With two transverse rows of setiferous tubercles and cones / spines; metaterga 2 - 19 with 2 + 2 tubercles in anterior row and 2 + 2 cones / spines in posterior row; lateral cones / spines of posterior row bigger and longer than mesal ones, gradually reduced in length and size on the following rings. Paraterga (Figure 19 A-E, H). Long; directed dorsolaterad on body rings 2 - 16, elevated at ca. 60 ° - 70 ° (male) 50 ° - 60 ° (female), directed dorsocaudad on ring 17, directed increasingly caudad on body rings 18 and 19. Ozopore visible in dorsolateral view. Telson (Figure 19 E, H, I). Epiproct quite short; tip subtruncate; lateral setiferous tubercles conspicuous; apical tubercles inconspicuous. Hypoproct subtrapeziform; caudal margin round, with conspicuous setiferous tubercles. Sterna (Figs 4 K; 19 F, G). Sternal lobe between male coxae 4 bifurcate, long; base stout; tips very sharp, in situ directed ventroanteriad; posterior surface bearing 1 pore. Legs (Figs 4 L, M; 19 J). Male femora 6 and 7 humped ventrally in distal part. Gonopods (Figs 5 E, 20). Coxa shorter than femur. Prefemoral part ca. half as long as femur. Femur quite enlarged distally. Postfemoral part short and narrow. Mesal sulcus and lateral sulcus wide. Solenophore longer than postfemoral part: lamina lateralis small, compact; apically protruding as a small lobe, directed mesoventrad: lamina medialis long; basally enlarged and slightly attenuated near the tip; tip bent, sharp and curving up. Solenomere curved and twisted, metazona in transverse section, tip directed posteriad.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
1C33B2F6B1A8B6ADCFC80C5E77F1E993.taxon	distribution	Distribution and habitat (Figure 18 C). All specimens were collected in small caves near the big Monkey Cave, crawling on rock walls. It is difficult to see the new species without using a flashlight / torch as the black body colour blends in with dark rocks. This species can be found in syntopy with Desmoxytes cervina. For the time being, S. macaca sp. n. is known only from the type locality and we regard it as endemic to Thailand.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
095D147E436CDE04BA4757B82F9B36C8.taxon	materials_examined	Material examined. Holotype. ♂, THAILAND, Surat Thani Province, Phanom District, Wat Tham Wararam, 8 ° 53 ' 07 " N, 98 ° 40 ' 01 " E, ca. 51 m a. s. l., 6 Aug. 2016, ASRU members leg. (CUMZ-pxDGT 00215). Paratypes. 23 ♂♂, 22 ♀♀, same data as for holotype (CUMZ-pxDGT 00216); 1 ♂, 1 ♀, same data as for holotype (ZMUC 00040254); 1 ♂, 1 ♀, same data as for holotype (ZMUM); 1 ♂, 1 ♀, same data as for holotype (NHMW 9425); 1 ♂, 1 ♀, same data as for holotype (NHMUK). Further specimens, not paratypes. THAILAND: 1 ♂, 2 ♀♀, Surat Thani Province, Phanom District, Wat Tham Wararam, 8 ° 53 ' 07 " N, 98 ° 40 ' 01 " E, ca. 51 m a. s. l., 5 Aug. 2014, ASRU members leg. (CUMZ).	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
095D147E436CDE04BA4757B82F9B36C8.taxon	etymology	Etymology. The species name is a Latin adjective, referring to the tip of lamina medialis which is somewhat hand-shaped.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
095D147E436CDE04BA4757B82F9B36C8.taxon	diagnosis	Diagnosis. Male femora without modification. Similar in this respect to S. efefi sp. n., S. hasta sp. n., S. sutchariti sp. n. and S. tortioverpa sp. n., but differs from them by having: anterior part of sternal lobe between male coxae 4 bifurcate, fork-like; tip of lamina medialis expanded, hand-shaped.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
095D147E436CDE04BA4757B82F9B36C8.taxon	description	Description. SIZE. Length 26 - 30 mm (male), 27 - 32 mm (female); width of midbody metazona 1.9 - 2.2 mm (male), 2.0 - 2.4 mm (female). Width of collum = 2 = 3 = 4 <head <5 - 16, thereafter body gradually tapering towards telson. Colour (Figure 21 A-C). Specimens in life with body black; paraterga orange; head, antennae (except whitish distal part of antennomeres 7 and 8), collum, prozona, metaterga (except white spines in posterior row) and surface below paraterga black; sterna and legs brown; epiproct pale brown; a few basal podomeres whitish brown. Antennae (Figure 22 M). Reaching to body ring 8 (male) and 6 or 7 (female) when stretched dorsally. Collum (Figure 22 A). With three transverse rows of setiferous tubercles / cones, 4 + 4 in anterior row, 1 (0) + 1 (0) in intermediate row and 2 (1) + 2 (1) in posterior row; with one conspicuous setiferous notch at lateral margin; paraterga spiniform, long, tip sharp, elevated at ca. 15 ° - 20 ° (male) 10 ° - 15 ° (female), directed almost laterad. Tegument. Very shining; collum coarsely microgranulate; metaterga and surface below paraterga smooth. Metaterga (Figure 22 A, C, E). With two transverse rows of setiferous cones and setiferous spines; metaterga 2 - 19 with 2 + 2 cones in anterior row and 2 + 2 spines in posterior row; lateral cones / spines of posterior row bigger and longer than mesal ones, gradually reduced in size and length on the following rings. Paraterga (Figure 22 A-E, H). Very long; directed dorsolaterad on body rings 2 - 17, elevated at ca. 50 ° - 60 ° (male) 45 ° - 60 ° (female), directed increasingly caudad on body rings 18 and 19. Ozopore visible in dorsolateral view. Telson (Figure 22 E, H, I). Epiproct quite short; tip subtruncate; lateral setiferous tubercles conspicuous; apical tubercles inconspicuous. Hypoproct subtrapeziform (in some specimens subtriangular); caudal margin round (in some specimens angular), with inconspicuous setiferous tubercles. Sterna (Figs 4 N; 22 F, G). Sternal lobe between male coxae 4 with two parts; anterior part bifurcate, tuning-fork-like, long, tips sharp, in situ directed ventroanteriad; posterior margin of anterior part bearing 1 pore; posterior part swollen, short. Legs (Figs 4 O, 22 J). Male femora without modification. Gonopods (Figs 5 F, 23). Coxa shorter than femur. Prefemoral part ca. half as long as femur. Femur not enlarged distally. Postfemoral part broad. Mesal sulcus and lateral sulcus wide. Solenophore a bit bigger than postfemoral part: lamina lateralis small, oval, tip round: lamina medialis long; basally enlarged and slightly attenuated near the tip; tip fringed, hand-shaped; tip curving down, in situ resting close to solenomere. Solenomere curved and twisted, compressed in transverse section, tip directed posteriad.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
095D147E436CDE04BA4757B82F9B36C8.taxon	distribution	Distribution and habitat (Figure 21 D). S. palmata sp. n. is known only from the type locality. We regard this species as endemic for the Thai fauna. The new species can be found in syntopy with Desmoxytes corythosaurus Srisonchai, Enghoff & Panha, 2018, crawling on humid rock walls.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
72414FE8601811D2439AF3809C336220.taxon	diagnosis	Diagnosis. The genus Spinaxytes gen. n. is characterized by: 1. Paraterga spiniform. 2. Metaterga with two rows of tubercles / cones / spines. 3. Postfemoral part of gonopod conspicuous, demarcated from femur by deep mesal and lateral sulci. 4. Lamina lateralis distinctly demarcated from lamina medialis. 5. Lamina medialis very long, curved, larger and longer than lamina lateralis.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
72414FE8601811D2439AF3809C336220.taxon	etymology	Etymology. The name is a noun in apposition; from the Latin ' spina', referring to the spine-like paraterga of all constituent species; ' - xytes' ensures harmony with Desmoxytes (and its synonym ' Pteroxytes ').	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
72414FE8601811D2439AF3809C336220.taxon	distribution	Distribution and habitat. All species of the new genus are allopatric. However, some of the new species can be found in syntopy with some species of Desmoxytes. The nine species described here are confined to limestone habitats and have narrow distributions; we therefore regard them as endemic. All species blend perfectly with habitat environment, mostly living on humid rock walls and / or inside small caves. According to the current knowledge, Spinaxytes gen. n. is distributed only in the Malay Peninsula (south Myanmar, south Thailand, and north Malaysia): Myanmar: Thanintharyi Region; Thailand: Chumphon Province (Thung Tako, Mueng Chumphon, Lang Suan and Sawi Districts), Surat Thani Province (Phanom District), Phang Nga Province (Mueng Phang Nga and Takua Thung Districts), Krabi Province (Ao Luek and Muaeng Krabi Districts); Malaysia: Perak State (Figure 30).	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
F9DF34622ABDAE14E4695A8643751306.taxon	materials_examined	Material examined. Holotype. ♂, THAILAND, Krabi Province, Muang Krabi District, Tham Na Mee (Na Mee Cave), 8 ° 08 ' 12 " N, 98 ° 48 ' 23 " E, ca. 70 m a. s. l., 31 Aug. 2015, C. Sutcharit leg. (CUMZ-pxDGT 00217). Paratypes. 7 ♂♂, 4 ♀♀, same data as for holotype (CUMZ-pxDGT 00218).	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
F9DF34622ABDAE14E4695A8643751306.taxon	etymology	Etymology. The name honours associate professor Dr. Chirasak Sutcharit, malacologist of ASRU (CUMZ), collector of this new species and numerous other dragon millipedes.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
F9DF34622ABDAE14E4695A8643751306.taxon	diagnosis	Diagnosis. Male femora without modification, sternal lobe between male coxae 4 incompletely bilobed. Similar in these respects to S. palmata sp. n., but differs by having: a large and round lamina lateralis; tip of lamina medialis terminating in two spines; distal part of solenomere circular in tranverse section.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
F9DF34622ABDAE14E4695A8643751306.taxon	description	Description. SIZE. Length 20 - 25 mm (male), 23 - 27 mm (female); width of midbody metazona 1.5 - 1.8 mm (male), 1.9 - 2.2 mm (female). Width of collum = 2 = 3 = 4 <head = 5 - 16, thereafter body gradually tapering towards telson. Colour (Figure 24 A-C). Specimens in life with body black; paraterga brownish white; head, antennae (except whitish distal part of antennomeres 7 and 8), collum, prozona, metaterga and epiproct black; surface below paraterga black / brownish black; sterna brown; legs brown / blackish brown; a few basal podomeres whitish brown. Antennae. Reaching to body ring 8 or 9 (male) and 6 (female) when stretched dorsally. Collum. With three transverse rows of setiferous tubercles, 4 + 4 tubercles in anterior row, 1 + 1 tubercles in intermediate row and 2 + 2 tubercles in posterior row; with one inconspicuous setiferous notch at lateral margin; paraterga spiniform, quite short, tip sharp, elevated at ca. 15 ° - 20 ° in both male and female, directed caudolaterad. Tegument. Quite dull; collum and metaterga (posterior part) coarsely microgranulate; metaterga (anterior part) and surface below paraterga smooth. Metaterga. With two transverse rows of setiferous tubercles and setiferous spines; metaterga 2 - 19 with 2 + 2 tubercles in anterior row and 2 + 2 spines in posterior row; lateral spines of posterior row bigger and longer than mesal ones, subequal in size and length on all body rings. Paraterga. Long; directed dorsolaterad on body rings 2 - 16, elevated at ca. 45 ° - 50 ° (male) 40 ° - 50 ° (female), directed dorsocaudad on ring 17, directed increasingly caudad on body rings 18 and 19. Ozopore visible in lateral view. Telson. Epiproct quite long; tip subtruncate; lateral setiferous tubercles conspicuous; apical tubercles inconspicuous. Hypoproct subtrapeziform (in some specimens subtriangular); caudal margin round (in some specimens angular), with inconspicuous setiferous tubercles. Sterna (Figure 4 P). Sternal lobe between male coxae 4 incompletely bilobed; tips sharp, in situ directed laterad; posterior surface bearing 2 pores. Legs (Figure 4 Q). Male femora without modification. Gonopods (Figs 5 G, 25). Coxa subequal in length to femur. Prefemoral part ca. 2 / 3 as long as femur. Femur not enlarged distally. Postfemoral part narrow. Mesal sulcus and lateral sulcus wide. Solenophore bigger and longer than postfemoral part: lamina lateralis oval, large, long, tip round: lamina medialis long and slender; base enlarged, slightly attenuated near the tip; tip curving down, with two sharp spines (one smaller, one bigger). Solenomere curving up, circular in transverse section, tip directed anteriad.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
F9DF34622ABDAE14E4695A8643751306.taxon	distribution	Distribution and habitat (Figure 24 D). S. sutchariti sp. n. is known only from the type locality, and we regard it as endemic to Thailand. The new species can be found in the same area as Gigaxytes gigas, but we assume that they live in different microhabitats: G. gigas was collected from the ground in leaf litter, whereas the new species was found on humid rock walls.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
D10F18D5012CE22BE1CEF631C05ADFE6.taxon	materials_examined	Material examined. Holotype. ♂, MALAYSIA, Perak State, Ipoh City, Gua Tempurung, limestone mountain, 4 ° 24 ' 58 " N, 101 ° 11 ' 16 " E, ca. 92 m a. s. l., 27 Sep. 2007, B. W. Ng and ASRU members leg. (CUMZ-pxDGT 00219). Paratypes. 1 ♂, 1 ♂ gonopods missing, 1 ♀, same data as for holotype (CUMZ-pxDGT 00220).	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
D10F18D5012CE22BE1CEF631C05ADFE6.taxon	etymology	Etymology. The name is a noun in apposition, from the Latin tortio meaning torsion and verpa for penis, refers to the distal part of gonopod (postfemoral part, solenophore and solenomere) which makes a 90 degrees torsion relative to the femoral part.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
D10F18D5012CE22BE1CEF631C05ADFE6.taxon	diagnosis	Diagnosis. Male femora without modification; collum with 4 + 4 tubercles in anterior row, 1 + 1 tubercles in intermediate row and 2 + 2 tubercles in posterior row. Similar in these respects to S. hasta sp. n. and S. palmata sp. n., but differs from them by having: a completely bilobed sternal lobe between male coxae 4; postfemoral part, solenophore and solenomere angled 90 degrees with femoral part; lamina lateralis with two lobes - the lateral one spine-like, the mesal one shorter and ridge-like; solenomere very long, longer than lamina medialis.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
D10F18D5012CE22BE1CEF631C05ADFE6.taxon	description	Description. SIZE. Length 28 - 30 mm (male), 30 - 33 mm (female); width of midbody metazona 2.0 mm (male), 2.8 mm (female). Width of rings 2 = 3 = 4 <head = 5 - 16, thereafter body gradually tapering towards telson. Colour. Specimens in life with body black / brownish black. Colour in alcohol: after 10 years changed to brown; paraterga brownish white; antennae brown (except whitish distal part of antennomeres 7 and 8); head, collum, metaterga and prozona blackish brown; surface below paraterga brown / blackish brown; sterna, epiproct and legs pale brown; a few basal podomeres whitish brown. Antennae. Reaching to body ring 8 or 9 (male) and 6 or 7 (female) when stretched dorsally. Collum. With three transverse rows of setiferous tubercles, 4 + 4 tubercles in anterior row, 1 + 1 tubercles in intermediate row and 2 + 2 tubercles in posterior row; anterior margin truncate; with one inconspicuous setiferous notch at lateral margin; paraterga wing-like, long and broad, tip sharp, elevated at ca. 15 ° - 20 ° in both male and female, directed caudolaterad. Tegument. Quite dull; collum, metaterga and surface below paraterga finely microgranulate. Metaterga. With two transverse rows of setiferous tubercles / cones and spines; metaterga 2 - 8 with 2 + 2 cones in anterior row and 2 + 2 spines in posterior row; metaterga 9 - 19 with 2 + 2 tubercles / cones in anterior row and 2 + 2 spines in posterior row; mesal spines of posterior row bigger and longer than lateral ones, gradually reduced in length and size on posterior rings. Paraterga. Very long; directed dorsolaterad on body rings 2 - 17, elevated at ca. 65 ° - 70 ° (male) 60 ° - 70 ° (female), directed increasingly caudad on body rings 18 and 19. Ozopore visible in dorsolateral view. Telson. Epiproct quite short; tip subtruncate; lateral setiferous tubercles inconspicuous; apical tubercles inconspicuous. Hypoproct subsemicircular; caudal margin round, with inconspicuous setiferous tubercles. Sterna (Figure 4 R). Sternal lobe between male coxae 4 completely divided into two lobes, long, spine-like; tips in situ directed ventrad; posterior surface a bit swollen, bearing 2 pores. Legs (Figure 4 S). Male femora without modification. Gonopods (Figs 5 H, 26). Coxa subequal in length to femur. Prefemoral part about 2 / 3 as long as femur. Femur quite enlarged distally. Postfemoral part large, broad and wide; angled 90 degrees with femur. Mesal sulcus and lateral sulcus wide. Solenophore smaller than postfemoral part: lamina lateralis apparently with two lobes demarcated from each other; lateral lobe very long, process-like, its tip in situ directed ventrad; mesal lobe short and wide, supporting solenomere: lamina medialis long, base not enlarged, tip directed mesad. Solenomere obviously longer than lamina medialis, circular in transverse section, curving down, tip directed laterad.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
D10F18D5012CE22BE1CEF631C05ADFE6.taxon	distribution	Distribution and habitat. Known only from the type locality which is currently a tourist attraction (cave). We regard this species as endemic to Malaysia.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
9513175F6C72A291882334650F6D26DC.taxon	materials_examined	Material examined. Holotype. ♂, THAILAND, Phang Nga Province, Mueang Phang Nga District, Phung Chang Cave, 8 ° 26 ' 34 " N, 98 ° 30 ' 59 " E, ca. 24 m a. s. l., 8 Aug. 2016, ASRU members leg. (CUMZ-pxDGT 00221). Paratypes. 15 ♂♂, 16 ♀♀, same data as for holotype (CUMZ-pxDGT 00222); 1 ♂, 1 ♀, same data as for holotype (ZMUC 00040255); 1 ♂, 1 ♀, same data as for holotype (ZMUM); 1 ♂, 1 ♀, same data as for holotype (NHMW 9424). Further specimens, not paratypes, all from THAILAND, Phang Nga Province. Muaeng Phang Nga District: 4 ♂♂, 2 ♀♀, Phung Chang Cave, 8 ° 26 ' 34 " N, 98 ° 30 ' 59 " E, ca. 24 m a. s. l., 6 Aug. 2014, ASRU members leg. (CUMZ); 4 ♂♂, Phung Chang Cave, 8 ° 26 ' 34 " N, 98 ° 30 ' 59 " E, ca. 24 m a. s. l., 5 Aug. 2015, ASRU members leg. (CUMZ); 2 ♂♂, 4 broken ♀♀, Tham Nam Pud, 8 ° 27 ' 50 " N, 98 ° 32 ' 36 " E, ca. 58 m a. s. l., 8 Oct. 2006, ASRU members leg. (CUMZ); 3 ♂♂, 2 ♀♀, Tham Nam Pud, 8 ° 27 ' 50 " N, 98 ° 32 ' 36 " E, ca. 58 m a. s. l., 5 Aug. 2015, ASRU members leg. (CUMZ); 2 ♂♂, 1 ♀, Wat Tham Bang Toei, 8 ° 27 ' 52 " N, 98 ° 34 ' 10 " E, ca. 24 m a. s. l., 10 Jul. 2017, ASRU members leg. (CUMZ); 8 ♂♂, 2 ♀♀, Tham Pha Phueng Bureau of Monks, 8 ° 28 ' 24 " N, 98 ° 32 ' 15 " E, ca. 78 m a. s. l., 10 Jul. 2017, ASRU members leg. (CUMZ). Thap Put District: 11 ♂♂, 3 ♀♀, Wat Kerewong (Tham Koab), 8 ° 31 ' 52 " N, 98 ° 34 ' 39 " E, ca. 76 m a. s. l., 9 Jul. 2017, ASRU members leg. (CUMZ).	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
9513175F6C72A291882334650F6D26DC.taxon	etymology	Etymology. The name is a Latin noun in apposition (uncus), meaning hook, and refers to the hook-like lamina medialis of gonopod.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
9513175F6C72A291882334650F6D26DC.taxon	diagnosis	Diagnosis. Differs from other species by having only male femora 7 strongly humped distally, in combination with the distal part of lamina medialis hook-like, tip long and sharp.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
9513175F6C72A291882334650F6D26DC.taxon	description	Description. SIZE. Length 20 - 27 mm (male), 25 - 29 mm (female); width of midbody metazona 1.4 - 1.6 mm (male), 2.1 - 2.3 mm (female). Width of collum <2 = 3 <4 <head = 5 - 16, thereafter body gradually tapering towards telson. Colour (Figure 27 A-E). Specimens in life with body black; paraterga yellow / whitish yellow; head, antennae (except whitish distal part of antennomeres 7 and 8) and prozona black; collum, metaterga and surface below paraterga black / brownish black; sterna and epiproct brown; legs yellow; a few basal podomeres white. Antennae (Figure 28 M). Reaching to body ring 8 or 9 (male) and 6 or 7 (female) when stretched dorsally. Collum (Figure 28 A). With three transverse rows of setiferous cones, 4 + 4 in anterior row, 1 + 1 in intermediate row and 2 + 2 in posterior row; with two conspicuous setiferous notches at lateral margin (first notch located at base of paraterga very close to cones of anterior row; paraterga spiniform, long, tip sharp, elevated at ca. 20 ° - 30 ° in both male and female, directed almost laterad. Tegument. Quite shining; collum, metaterga (posterior part) and surface below paraterga finely microgranulate; metaterga (anterior part) coarsely microgranulate. Metaterga (Figure 28 A, C, E). With two transverse rows of setiferous cones and setiferous spines; metaterga 2 - 19 with 2 + 2 cones in anterior row and 2 + 2 spines in posterior row; lateral spines of posterior row very long, bigger and longer than mesal ones. Paraterga (Figure 28 A-E, H). Very long; directed dorsolaterad on body rings 2 - 16, elevated at ca. 40 ° - 50 ° (male) 40 ° - 45 ° (female), directed dorsocaudad on ring 17, directed increasingly caudad on body rings 18 and 19. Ozopore visible in dorsolateral view. Telson (Figure 28 E, I, H). Epiproct short; tip subtruncate; lateral setiferous tubercles inconspicuous; apical tubercles inconspicuous. Hypoproct subtrapeziform (in some specimens subsemicircular); caudal margin round, with conspicuous setiferous tubercles (in some specimens inconspicuous). Sterna (Figs 4 T; 28 F, G). Sternal lobe between male coxae 4 bifurcate, long; tips sharp, in situ directed ventroanteriad; posterior surface bearing one pore. Legs (Figs 5 U, V; 28 J). Male femora 7 strongly humped ventrally in distal part. Gonopods (Figs 5 I, 29). Coxa subequal in length to femur. Prefemoral part almost half as long as femur. Femur obviously enlarged distally. Postfemoral part small, narrow. Mesal sulcus wide; lateral sulcus narrow. Solenophore bigger and longer than postfemoral part: lamina lateralis very small, compact, tip round: lamina medialis long; basally enlarged and slightly attenuated near the tip; apically sharp, long, hook-like; tip curving down, in situ resting close to solenomere. Solenomere curved and twisted, compressed in transverse section, tip directed posteriad.	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
9513175F6C72A291882334650F6D26DC.taxon	distribution	Distribution and habitat (Figure 27 F). Known only from Phang Nga Province; we regard S. uncus sp. n. as endemic for the Thai fauna. Most specimens were found on rock walls near the cave, some were seen crawling on leaf litter on the rock. The new species has been encountered in syntopy with Desmoxytes cervina at Phung Chang Cave, Tham Nam Pud and Wat Kerewong (Tham Koab).	en	Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai, Panha, Somsak (2018): A revision of dragon millipedes IV: the new genus Spinaxytes, with the description of nine new species (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 797: 19-69, DOI: http://dx.doi.org/10.3897/zookeys.797.29510, URL: http://dx.doi.org/10.3897/zookeys.797.29510
