taxonID	type	description	language	source
8619E9769459D915EA132F0220CDFDFB.taxon	description	(Figures 1 – 6, 9, 12 – 13; Tables 1 – 4) Holotype: UW 112244, 157.0 mm, female, 52.3405 ºN, 172.7455 ºW, 441 m depth, 31 May 2002, F / V Morning Star, cruise 2002 ­ 01, haul 32, J. W. Orr. Paratypes: UW 112243, 3 (145.0 – 190.0 mm), 52.3732 ºN, 171.3548 ºW, 324 m depth, 10 August 2002, F / V Sea Storm, cruise 2002 ­ 01, haul 225, R. C. Harrison; UW 112235, 1 (162.0 mm), 52.3796 ºN, 170.6611 ºW, 278 m depth, 31 May 2000, F / V Vesteraalen, cruise 2000 ­ 01, haul 51, W. C. Flerx; UW 112241, 3 (108.0 – 167.0 mm), 52.331 ºN, 172.7470 ºW, 441 m depth, 23 July 2002, F / V Sea Storm, cruise 2002 ­ 01, haul 156, J. W. Orr; UW 112237, 1 (152.9 mm), 52.2383 ºN, 172.1247 ºW, 357 m depth, 29 July 2002, F / V Sea Storm, cruise 2002 ­ 01, haul 167, benthic bag, R. C. Harrison; UW 112236, 2 (129.0 – 133.0 mm; 129.0 mm cleared and stained), 52.4877 ºN, 172.5677 ºW, 419 m depth, 26 June 1997, M / V Dominator, cruise 1997 ­ 01, haul 71, R. C. Harrison; UW 112242, 3 (139.5 – 176.0 mm), 52.3217 ºN, 172.7447 ºW, 458 m depth, 16 June 1994, F / V Pacific Knight, cruise 1994 ­ 01, haul 54; UW 112278, 8 (109.8 – 157.0 mm), 52.3405 ºN, 172.7455 ºW, 441 m depth, 31 May 2002, F / V Morning Star, cruise 2002 ­ 01, haul 32, J. W. Orr; UW 112239, 1 (161.0 mm), 52.3282 ºN, 172.7468 ºW, 444 m depth, F / V Vesteraalen, cruise 2000 ­ 01, haul 67, W. C. Flerx; UW 112238, 2 (110.8 – 119.5 mm), 52.3282 ºN, 172.7468 ºW, 444 m depth, 4 June 2000, F / V Vesteraalen, cruise 2000 ­ 01, haul 67, benthic bag, W. C. Flerx; UW 113691, 16 (72.0 – 170.0 mm), 52.3255 ºN, 172.7466 ºW, 450 m depth, 19 June 2004, F / V Sea Storm, cruise 2004 ­ 01, haul 60, J. W. Orr; USNM 385683, 2 (113.0 – 165.0 mm), 52.3255 ºN, 172.7466 ºW, 450 m depth, 19 June 2004, F / V Sea Storm, cruise 2004 ­ 01, haul 60, benthic bag, J. W. Orr; USNM 385684, 1 (156.0 mm), 52.4888 ºN, 172.5564 ºW, 397 m depth, 20 June 2004, F / V Sea Storm, cruise 2004 ­ 01, haul 61, J. W. Orr; CAS 223481, 1 (146.0 mm), 52.5251 ºN, 172.0747 ºW, 364 m depth, 21 June 2004, F / V Sea Storm, cruise 2004 ­ 01, haul 66, J. W. Orr; CAS 223482, 1 (195.0 mm), 52.2102 ºN, 172.2060 ºW, 341 m depth, 23 June 2004, F / V Sea Storm, cruise 2004 ­ 01, haul 73, J. W. Orr.	en	Orr, James Wilder, Busby, Morgan Scott (2006): Revision of the snailfish genus Allocareproctus Pitruk & Fedorov (Teleostei: Liparidae), with descriptions of four new species from the Aleutian Islands. Zootaxa 1173: 1-37, DOI: 10.5281/zenodo.2645716
8619E9769459D915EA132F0220CDFDFB.taxon	diagnosis	Diagnosis Teeth simple (Fig. 2 C); nasal pore 1 with pigmented papilla; peritoneum black; orobuccal valve with 4 – 6 finger­like projections; orobuccal cavity dark; gill rakers 14 – 18, stout, with strong spines at tip; interorbital papilla absent; body blotchy red, darker gray anteriorly; iris silver gray.	en	Orr, James Wilder, Busby, Morgan Scott (2006): Revision of the snailfish genus Allocareproctus Pitruk & Fedorov (Teleostei: Liparidae), with descriptions of four new species from the Aleutian Islands. Zootaxa 1173: 1-37, DOI: 10.5281/zenodo.2645716
8619E9769459D915EA132F0220CDFDFB.taxon	description	Description Body depth at pectoral­fin base 11.8 – 18.6 (15.4) %, at center of pelvic disk 13.8 – 21.2 (18.0) %, at anal­fin origin 14.0 – 22.0 (18.8) %. Predorsal length 22.7 – 28.6 (25.2) %. Preanal length 38.8 – 47.0 (43.4) %. Head large, width 12.4 – 19.6 (18.4) %, length 21.7 – 24.3 (22.4) %. Interorbital width 4.3 – 7.7 (5.1) %. Snout 5.5 – 8.8 (6.9) %. Mouth small, maxilla 7.5 – 11.3 (11.3) %, extending to anterior portion of orbit. Teeth simple, recurved, rarely with weak shoulders (Fig. 2 C), in a narrow band of 4 – 12 oblique rows of 5 – 10 teeth per row in all except the posterior outer row; outer row of 34 teeth forming comb­like series. In all except outer row, middle teeth of each row largest; in outer row, first 20 teeth equally sized, succeeding teeth gradually smaller posteriorly. Orbit large, diameter 5.3 – 7.3 (6.1) %. Papillae present on many pores: present on nasal pore 1, absent from nasal pore 2; present on maxillary pores 2 – 6, rarely present on pore 1 (2 – 6); present on preoperculomandibular pores 5 – 7, often present also on pore 4 (5 – 7); present on suprabranchial pores 1 – 2 (Fig. 4 D). Papillae with slight pigment or unpigmented. Interorbital papilla absent. Cephalic free neuromasts large and profuse to nearly indiscernible over the interorbit and nape. Gill opening small, 4.6 – 6.7 (5.6) %, entirely above pectoral fin. Gill rakers on anterior arch 14 – 18 (16), stout with strong spines at tip. Orobuccal valve with 4 – 6 (4) finger­like lobes, central lobe largest, other lobes successively smaller (Fig. 12 B). Dorsal­fin rays 42 – 45 (44), tips of anterior 4 – 6 rays projecting from fin membrane, anteriormost rays about 30 % free from membrane, succeeding rays less so; posteriormost ray attached membranously to dorsalmost caudal­fin ray for 2.0 – 4.4 (2.7) %. Anal fin with 33 – 37 (36) rays, posteriormost ray membranously attached to ventralmost caudal­fin ray for 2.6 – 5.1 (3.4) %. One to three (2) anal­fin pterygiophores and associated rays anterior to first haemal spine. Pectoral­fin rays 36 – 40 (37) in two lobes separated by shallow notch, 9 – 12 (10) rays in lower lobe. Dorsalmost ray at level just below ventral rim of orbit. Upper lobe rounded, extending to anal­fin origin, length 15.8 – 20.1 (17.6) %, with rays 6 – 7 (6) longest; length of shortest notch ray 4.3 – 9.5 (6.7) %; length of lower lobe 13.3 – 20.2 (15.0) %, extending between posterior margin of pelvic disk and anus, with ray 3 longest, rays 9 – 12 shortening ventrally. Pelvic disk large, length 8.0 – 9.7 (8.5) %, width 7.1 – 8.8 (7.8) %. Distance from disk to anus 2.7 – 9.4 (6.2) %, about 28 – 105 (73.7) % DL, distance from anus to anal­fin origin 11.6 – 18.9 (16.6) %, about 125 – 225 (196) % DL. Urogenital papilla conical, short, about 10 – 25 (11.2) % DL when protruded, unpigmented. Pyloric caeca about 22 – 27 (22), on right side of body, finger­like, long, 25 – 50 (48.4) % HL. Caudal fin slightly rounded, 12.8 – 17.1 (14.3) %, depth at hypural plate 2.7 – 4.6 (2.9) %, with principal rays 11 – 14 (12), dorsal principal rays 5 – 7 (5), ventral principal rays 6 – 7 (7). Dorsal procurrent rays 2 – 3 (2), borne on epural and posteriormost neural spine; ventral procurrent ray 1 – 2 (1), borne on expanded posteriormost haemal spine. Vertebrae 45 – 49 (49), abdominal vertebrae 12, caudal vertebrae 33 – 37 (37). Color in life light red, with darker red blotches scattered over body, dark speckling at origin of dorsal fin, anterior part of body blue­gray, posterior part of body blotchy red; iris silver gray. Color in alcohol dark gray anteriorly, becoming pale posteriorly, with pigment on cephalic papillae and at origin of dorsal fin. Peritoneum black; orobranchial cavity dark. Largest specimen examined a 195 mm male (CAS 223482). Smallest female with yolked eggs 162 mm; smallest male with enlarged, swollen testes 124 mm. Range Allocareproctus kallaion has been collected only in the east­central Aleutian Islands, from Seguam Pass to Yunaska Island at depths of 278 – 458 m (Fig. 9).	en	Orr, James Wilder, Busby, Morgan Scott (2006): Revision of the snailfish genus Allocareproctus Pitruk & Fedorov (Teleostei: Liparidae), with descriptions of four new species from the Aleutian Islands. Zootaxa 1173: 1-37, DOI: 10.5281/zenodo.2645716
8619E9769459D915EA132F0220CDFDFB.taxon	etymology	Etymology The specific epithet is derived from the Greek “ ” for “ comb ”, an allusion to the multiple projections on the orobuccal valve and the single row of teeth on the posterior part of the dentary.	en	Orr, James Wilder, Busby, Morgan Scott (2006): Revision of the snailfish genus Allocareproctus Pitruk & Fedorov (Teleostei: Liparidae), with descriptions of four new species from the Aleutian Islands. Zootaxa 1173: 1-37, DOI: 10.5281/zenodo.2645716
8619E9769459D915EA132F0220CDFDFB.taxon	discussion	Remarks Unlike other species of Allocareproctus, larger specimens of A. kallaion were often collected with greatly distended stomachs filled with water. This mechanism of inflation in A. kallaion is similar to that of puffer fishes (Tetraodontiformes; Brainerd 1994, Wainwright & Turingan 1997) and frogfishes (Antennariidae; Pietsch & Grobecker 1987). Members of the putative sister family of the Liparidae, the Cyclopteridae, are well known for their ability to expand when disturbed, although the mechanism of inflation has not been examined (Mecklenburg et al. 2002). Comparisons	en	Orr, James Wilder, Busby, Morgan Scott (2006): Revision of the snailfish genus Allocareproctus Pitruk & Fedorov (Teleostei: Liparidae), with descriptions of four new species from the Aleutian Islands. Zootaxa 1173: 1-37, DOI: 10.5281/zenodo.2645716
8619E9769459D915EA132F0220CDFDFB.taxon	description	Allocareproctus kallaion is easily distinguished from all other species of Allocareproctus by the following characters. Its body in life is a blotchy red unlike the uniform red to peach coloration of other species. Both in life and when preserved, the anterior part of the body and orobuccal cavity are darker, unlike the pale color of other species. It differs from all other species in having multiple lobes on the orobuccal valve (Fig. 12 B), high numbers (14 – 18 vs. 7 – 12 in all other species) of larger gill rakers with stronger spines on the tips, invariably 12 abdominal vertebrae, and a comb­like uniserial row of teeth on the dentary. Its simple teeth further distinguish it from A. tanix, A. unangas, and A. ungak, which all possess moderately to strongly trilobed teeth (Fig. 2). Several morphometric characters differ significantly between A. kallaion and all other species of Allocareproctus. Head length, body depth, snout length, suborbital depths to oral cleft and mandible, predorsal length, snout to anus length, pelvic­disk length and width, and lengths of dorsal­ and anal­fin connections to the caudal fin are less, while pelvic disk to anus length is greater than in all other species.	en	Orr, James Wilder, Busby, Morgan Scott (2006): Revision of the snailfish genus Allocareproctus Pitruk & Fedorov (Teleostei: Liparidae), with descriptions of four new species from the Aleutian Islands. Zootaxa 1173: 1-37, DOI: 10.5281/zenodo.2645716
8619E9769455D92FEA13289020D2FA8B.taxon	description	(Figures 1 – 2, 6, 10, 13; Tables 1 – 4) Holotype: UW 112308, 82.5 mm, 53.1274 ºN, 169.9643 ºW, 387 m depth, 1 June 2002, F / V Morning Star, cruise 2002 ­ 01, haul 35, J. W. Orr / D. E. Stevenson. Paratypes: UW 112300, 1 (129.7 mm), female, 53.2067 ºN, 169.8452 ºW, 399 m depth, 4 June 2002, F / V Morning Star, cruise 2002 ­ 01, haul 52, J. W. Orr; UW 112313, 1 (115.2 mm), 53.0363 ºN, 170.2986 ºW, 210 m depth, 27 May 2000, F / V Vesteraalen, cruise 2000 ­ 0 1, haul 39, W. C. Flerx; UW 112303, 2 (79 – 105.8 mm), 51.5636 ºN, 178.3331 ºE, 465 m depth, 30 June 2000, M / V Dominator, cruise 2000 ­ 01, haul 159, K. P. Maslenikov; UW 112306, 2 (40.3 – 64.4 mm), 53.1899 ºN, 169.8622 ºW, 398 m depth, 19 July 2002, F / V Vesteraalen, cruise 2002 ­ 01, haul 195, R. N. Clark; UW 112305, 1 (104.2 mm), 52.2762 ºN, 170.5968 ºW, 231 m depth, 30 May 2000, M / V Dominator, cruise 2000 ­ 01, haul 49, J. W. Orr; UW 112304, 1 (126.2 mm), 52.40236 ºN, 171.8333 ºW, 270 m depth, 8 August 2002, F / V Sea Storm, cruise 2002 ­ 01, haul 213, R. C. Harrison; UW 112087, 1 (83 mm), 53.1303 ºN, 169.9617 ºW, 433 m depth, 26 May 2000, F / V Vesteraalen, cruise 2000 ­ 01, haul 35, W. C. Flerx; UW 112089, 2 (73.5 – 78.6 mm), 52.4212 ºN, 170.2785 ºW, 211 m depth, 29 May 2000, M / V Dominator, cruise 2000 ­ 01, haul 43, benthic bag, J. W. Orr; UW 112301, 1 (117.7 mm), 52.3896 ºN, 171.36 ºW, 234 m depth, 11 June 2000, M / V Dominator, cruise 2000 ­ 01, haul 82, K. P. Maslenikov; UW 112302, 2 (76.7 – 98.2 mm), 52.3675 ºN, 171.3377 ºW, 318 m depth, 11 June 2000, M / V Dominator, cruise 2000 ­ 01, haul 83, benthic bag, K. P. Maslenikov; UW 112085, 1 (78 mm), 51.9056 ºN, 176.6003 ºE, 264 m depth, 5 July 2000, F / V Vesteraalen, cruise 2000 ­ 0 1, haul 173, E. S. Brown; UW 112086, 1 (101.6 mm), 52.3679 ºN, 171.2465 ºW, 316 m depth, 18 June 1997, F / V Vesteraalen, cruise 1997 ­ 01, haul 42, W. C. Flerx; UW 112088, 2 (84.2 – 102 mm; 84.2 mm cleared and stained), 52.5303 ºN, 172.1104 ºW, 364 m depth, 26 June 1997, F / V Vesteraalen, cruise 1997 ­ 01, haul 78, R. C. Harrison; UW 112307, 1 (101.7 mm), 52.2383 ºN, 172.1247 ºW, 357 m depth, 29 July 2002, F / V Sea Storm, cruise 2002 ­ 01, haul 167, R. C. Harrison; USNM 385685, 4 (68.3 – 96.1 mm), 52.3687 ºN, 171.2406 ºW, 323 m depth, 5 May 2000, M / V Dominator, cruise 2000 ­ 01, haul 51, benthic bag, J. W. Orr; USNM 385686, 1 (118 mm), 52.5080 ºN, 172.2650 ºW, 215 m depth, 21 June 2004, F / V Sea Storm, cruise 2004 ­ 01, haul 65, J. W. Orr; USNM 385687, 2 (95 – 115 mm), 52.9580 ºN, 169.4346 ºW, 429 m depth, 12 June 2004, F / V Gladiator, cruise 2004 ­ 01, haul 26, K. P. Maslenikov; CAS 223483, 1 (92 mm), 52.8965 ºN, 169.4597 ºW, 341 m depth, 13 June 2004, F / V Gladiator, cruise 2004 ­ 01, haul 29, benthic bag, K. P. Maslenikov; CAS 223484, 1 (113 mm), 53.2200 ºN, 169.7387 ºW, 323 m depth, 13 June 2004, F / V Gladiator, cruise 2004 ­ 01, haul 27, K. P. Maslenikov; CAS 223485, 1 (105 mm), 53.0341 ºN, 170.2061 ºW, 176 m depth, 14 June 2004, F / V Gladiator, cruise 2004 ­ 01, haul 32, benthic bag, K. P. Maslenikov; UW 28351, 2 (82.2 – 128.7 mm), no data.	en	Orr, James Wilder, Busby, Morgan Scott (2006): Revision of the snailfish genus Allocareproctus Pitruk & Fedorov (Teleostei: Liparidae), with descriptions of four new species from the Aleutian Islands. Zootaxa 1173: 1-37, DOI: 10.5281/zenodo.2645716
8619E9769455D92FEA13289020D2FA8B.taxon	diagnosis	Diagnosis Teeth strongly trilobed (Fig. 2 D); nasal papillae absent; papillae associated with few pores, invariably present only on maxillary pore 6 and suprabranchial pores 1 – 2; peritoneum dark; orobuccal valve with 1 finger­like projection; orobuccal cavity pale; gill rakers 8 – 12, short, blunt; interorbital papilla present; pyloric caeca on left side, short; body color light red; iris golden.	en	Orr, James Wilder, Busby, Morgan Scott (2006): Revision of the snailfish genus Allocareproctus Pitruk & Fedorov (Teleostei: Liparidae), with descriptions of four new species from the Aleutian Islands. Zootaxa 1173: 1-37, DOI: 10.5281/zenodo.2645716
8619E9769455D92FEA13289020D2FA8B.taxon	description	Description Body depth at pectoral­fin base 14.2 – 22.6 (20.1) %, at center of pelvic disk 18.9 – 24.6 (20.4) %, at anal­fin origin 18.8 – 26.5 (25.5) %. Predorsal length 26.5 – 33.3 (29.7) %. Preanal length 37.4 – 46.4 (41.9) %. Head large, width 12.7 – 18.8 (15.3) %, length 25.1 – 28.2 (25.7) %. Interorbital width 5.1 – 8.2 (5.1) %, about equal to orbit length. Snout 6.5 – 9.2 (8.8) %. Mouth small, maxilla 8.4 – 12.0 (11.1) %, extending to anterior portion of orbit. Teeth strongly trilobed (Fig. 2 D), in a band of 8 – 12 oblique rows of 5 – 9 teeth per row. Orbit large, diameter 5.0 – 7.8 (5.0) %. Papillae small, present on few pores: absent from nasal pores 1 – 2; present on maxillary pore 6, often present on pore 5 (5 – 6); often present on preoperculomandibular pores 6 – 7, rarely absent (6 – 7); present on suprabranchial pores 1 – 2 (Fig. 4 E). Interorbital papilla present. Papillae unpigmented. Cephalic free neuromasts reduced or nearly indiscernible over the interorbit and nape. Gill opening small, 4.7 – 7.1 (5.9) %, entirely above pectoral fin or extending to pectoral­fin ray 3 (entirely above in the holotype). Gill rakers on anterior arch 8 – 12 (9), blunt and stout. Central projection of orobuccal valve a moderately elongate single lobe. Dorsal­fin rays 42 – 45 (44), tips of anterior 4 – 6 rays projecting from fin membrane, anteriormost rays about 30 % free from membrane, succeeding rays less so; posteriormost ray attached membranously to dorsalmost caudal­fin ray for 2.5 – 5.0 (3.6) %. Anal fin with 36 – 39 (38) rays, posteriormost ray membranously attached to ventralmost caudal­fin ray for 3.0 – 5.3 (3.6) %. One to three (2) anal­fin pterygiophores and associated rays anterior to first haemal spine. Pectoral­fin rays 38 – 42 (39) in two lobes separated by shallow notch, 9 – 10 (10) rays in lower lobe. Dorsalmost ray at level of ventral rim of orbit. Upper lobe rounded, extending to or just past anal­fin origin, length 15.4 – 20.1 (17.0) %, with ray 6 longest; length of shortest notch ray 4.9 – 10.9 (10.9) %; length of lower lobe 13.4 – 19.5 (15.7) %, extending to anus, with ray 2 longest, rays 3 – 10 shortening ventrally. Pelvic disk large, length 7.8 – 11.4 (10.3) %, width 7.4 – 10.7 (9.9) %. Distance from disk to anus 1.7 – 8.3 (5.6) %, about 20 – 90 (54.5) % DL, distance from anus to anal­fin origin 5.7 – 15.8 (13.5) %, about 50 – 150 (130.6) % DL. Urogenital papilla conical, short, 9.2 – 30.5 (30.5) % DL, unpigmented. Pyloric caeca about 22, on left side of body, fingerlike, short, 17.2 – 28.5 (24.0) % HL. Caudal fin slightly rounded, 11.7 – 17.0 (13.5) %, depth at hypural plate 2.4 – 5.2 (3.4) %, with principal rays 11 – 12 (12), dorsal principal rays 5, ventral principal rays 6 – 7 (7). Dorsal procurrent rays 1 – 3 (2), borne on epural and posteriormost neural spine; ventral procurrent ray 1 – 2 (1), borne on expanded posteriormost haemal spine. Vertebrae 46 – 50 (48), abdominal vertebrae 10 – 11 (10), caudal vertebrae 36 – 40 (38). Body color in life uniform light red; iris golden. Color in alcohol pale, with pigment absent from cephalic papillae and absent from origin of dorsal fin. Peritoneum black; orobranchial cavity pale. Largest specimen examined a 129.7 mm female (UW 112300). Smallest female with yolked eggs 91 mm; smallest male with enlarged, swollen testes 76.7 mm. Range Allocareproctus unangas has been collected only in the Aleutian Islands, from Buldir Pass to the Islands of Four Mountains, at depths of 210 to 465 m (Fig. 10).	en	Orr, James Wilder, Busby, Morgan Scott (2006): Revision of the snailfish genus Allocareproctus Pitruk & Fedorov (Teleostei: Liparidae), with descriptions of four new species from the Aleutian Islands. Zootaxa 1173: 1-37, DOI: 10.5281/zenodo.2645716
8619E9769455D92FEA13289020D2FA8B.taxon	etymology	Etymology Named in honor of the people of the Aleutian Islands, the specific epithet “ unangas ” is the autonym of the Aleuts of Atka Island, a major island near the center of the species distribution. It is to be treated as a noun in apposition.	en	Orr, James Wilder, Busby, Morgan Scott (2006): Revision of the snailfish genus Allocareproctus Pitruk & Fedorov (Teleostei: Liparidae), with descriptions of four new species from the Aleutian Islands. Zootaxa 1173: 1-37, DOI: 10.5281/zenodo.2645716
8619E9769455D92FEA13289020D2FA8B.taxon	discussion	Remarks Eggs of A. unangas were collected from the octocoral Primnoa sp. from Seguam Pass, with eggs of another liparid (Busby et al., in press). These are the first records of spawning in nature outside of lithodid crabs for liparids other than species of Liparis. See description and discussion of Busby et al. (in press). Comparisons	en	Orr, James Wilder, Busby, Morgan Scott (2006): Revision of the snailfish genus Allocareproctus Pitruk & Fedorov (Teleostei: Liparidae), with descriptions of four new species from the Aleutian Islands. Zootaxa 1173: 1-37, DOI: 10.5281/zenodo.2645716
8619E9769455D92FEA13289020D2FA8B.taxon	description	Allocareproctus unangas is most similar to A. ungak, which also has trilobed teeth (Fig. 2 D – E) and a uniform red body. It is distinguished from A. ungak by its golden iris (vs. silver gray in A. ungak), papilla on nasal pore 1, fewer pores with associated papillae (Fig. 4), and pyloric caeca shorter and on the left side of the body. The two species also exhibit significant modal differences in several meristic characters (Tables 1 – 3), including dorsal­fin rays (39 – 42 in A. ungak vs. 42 – 45 in A. unangas), anal­fin rays (33 – 35 vs. 36 – 39), pectoral­fin rays (35 – 40 vs. 38 – 42), and caudal vertebrae (34 – 36 vs. 36 – 40). Morphometric differences include a greater head length, body depths at anal­fin origin and pelvic disk, predorsal length, and snout to anus length in A. unangas than in A. ungak. See other comparisons under species accounts above. Allocareproctus ungak new species Whiskered Snailfish (Figures 1 – 2, 4 – 6, 11, 13; Tables 1 – 4)	en	Orr, James Wilder, Busby, Morgan Scott (2006): Revision of the snailfish genus Allocareproctus Pitruk & Fedorov (Teleostei: Liparidae), with descriptions of four new species from the Aleutian Islands. Zootaxa 1173: 1-37, DOI: 10.5281/zenodo.2645716
8619E9769455D92FEA13289020D2FA8B.taxon	materials_examined	Holotype: UW 111933, 91.0 mm, female, 52.3687 ºN, 171.2406 ºW, 323 m depth, 31 May 2000, M / V Dominator, cruise 2000 ­ 01, haul 51, benthic bag, J. W. Orr. Paratypes: UW 112084, 4 (55.0 – 106.9 mm), 52.3282 ºN, 172.7468 ºW, 444 m depth, 4 June 2000, F / V Vesteraalen, cruise 2000 ­ 01, haul 67, benthic bag, W. C. Flerx; UW 111934, 1 (106 mm), 52.3282 ºN, 172.7468 ºW, 444 m depth, 4 June 2000, F / V Vesteraalen, cruise 2000 ­ 01, haul 67, W. C. Flerx; UW 111935, 2 (97.3 – 98.4 mm), 52.3131 ºN, 171.5057 ºW, 445 m depth, 31 May 2000, M / V Dominator, cruise 2000 ­ 01, haul 54, benthic bag, J. W. Orr; UW 111929, 1 (105.5 mm), 52.1920 ºN, 171.6730 ºW, 461 m depth, 6 August 2002, F / V Sea Storm, cruise 2002 ­ 01, haul 210, R. C. Harrison; UW 45239, 17 (81.5 – 101.2 mm; 84.5 mm cleared and stained), 52.3184 ºN, 172.7453 ºW, 455 m depth, 22 June 1997, F / V Vesteraalen, cruise 1997 ­ 01, haul 58, W. C. Flerx; UW 111937, 1 (87 mm), 52.5517 ºN, 169.4788 ºW, 330 m depth, 25 May 2003, F / V Northwest Explorer, cruise 2003 ­ 01, haul 6, J. W. Orr; UW 111933, 1 (91 mm, in 95 % ethanol), 52.3687 ºN, 171.2406 ºW, 323 m depth, 31 May 2000, M / V Dominator, cruise 2000 ­ 01, haul 51, J. W. Orr; UW 111938, 1 (112 mm), 52.3687 ºN, 171.2406 ºW, 323 m depth, 31 May 2000, M / V Dominator, cruise 2000 ­ 01, haul 51, J. W. Orr; UW 45238, 3 (72.1 – 91.7 mm), 52.3310 ºN, 172.7470 ºW, 441 m depth, 23 July 2002, F / V Sea Storm, cruise 2002 ­ 0 1, haul 156, J. W. Orr; UW 111928, 2 (85.8 – 93.2 mm), 52.3687 ºN, 171.2406 ºW, 323 m depth, 31 May 2000, M / V Dominator, cruise 2000 ­ 01, haul 51, benthic bag, J. W. Orr; UW 111927, 1 (94.3 mm), 52.3732 ºN, 171.3548 ºW, 324 m depth, 10 August 2002, F / V Sea Storm, cruise 2002 ­ 01, haul 225, R. C. Harrison; USNM 385688, 3 (97 – 115 mm), 52.5251 ºN, 172.0747 ºW, 364 m depth, 21 June 2004, F / V Sea Storm, cruise 2004 ­ 01, haul 66, J. W. Orr; USNM 385689, 3 (62 – 78 mm), 52.3255 ºN, 172.7466 ºW, 450 m depth, 19 June 2004, F / V Sea Storm, cruise 2004 ­ 01, haul 60, benthic bag, J. W. Orr; CAS 223486, 7 (91 – 107 mm), 52.3728 ºN, 171.3587 ºW, 318 m depth, 21 June 2004, F / V Gladiator, cruise 2004 ­ 01, haul 65, K. P. Maslenikov; UW 113675, 1 (94.5 mm), 52.1932 ºN, 171.6718 ºW, 461 m depth, 20 June 2004, F / V Gladiator, cruise 2004 ­ 01, haul 58, K. P. Maslenikov; UW 113676, 1 (129.3 mm), 52.9580 ºN, 169.4346 ºW, 429 m depth, 12 June 2004, F / V Gladiator, cruise 2004 ­ 01, haul 26, K. P. Maslenikov.	en	Orr, James Wilder, Busby, Morgan Scott (2006): Revision of the snailfish genus Allocareproctus Pitruk & Fedorov (Teleostei: Liparidae), with descriptions of four new species from the Aleutian Islands. Zootaxa 1173: 1-37, DOI: 10.5281/zenodo.2645716
8619E9769455D92FEA13289020D2FA8B.taxon	diagnosis	Diagnosis Teeth strongly trilobed (Fig. 2 E); nasal pore 1 with unpigmented papilla; peritoneum black; orobuccal valve with 1 finger­like projection; orobuccal cavity pale; gill rakers 7 – 11, short, blunt; interorbital papilla present or absent; pyloric caeca on right or left side; body red; iris silver gray.	en	Orr, James Wilder, Busby, Morgan Scott (2006): Revision of the snailfish genus Allocareproctus Pitruk & Fedorov (Teleostei: Liparidae), with descriptions of four new species from the Aleutian Islands. Zootaxa 1173: 1-37, DOI: 10.5281/zenodo.2645716
8619E9769455D92FEA13289020D2FA8B.taxon	description	Description Body depth at pectoral­fin base 15.0 – 21.0 (19.5) %, at center of pelvic disk 16.6 – 22.9 (21.9) %, at anal­fin origin 18.2 – 25.2 (22.0) %. Predorsal length 25.9 – 29.7 (27.3) %. Preanal length 38.4 – 47.3 (43.1) %. Head large, width 11.8 – 20.4 (14.1) %, length 23.9 – 27.3 (24.7) %. Interorbital width 4.9 – 8.0 (5.7) %, about equal to orbit length. Snout 6.8 – 9.3 (7.9) %. Mouth small, maxilla 9.0 – 12.1 (10.1) %, extending to midorbit. Teeth strongly trilobed (Fig. 2 E), in a broad band of 8 – 12 oblique rows of 5 – 16 teeth per row, band narrowing posteriorly to 2 – 3 rows on premaxilla and to a short uniserial row of 6 teeth on dentary. Orbit large, diameter 5.2 – 7.0 (6.2) %. Papillae strong, present on many pores: present on nasal pore 1, absent from nasal pore 2; present on maxillary pores 4 – 6, often present on pores 2 – 3 (4 – 6); present on preoperculomandibular pores 5 – 6, often present also on pore 7 (5 – 7); present on suprabranchial pores 1 – 2 (Fig. 4 F). Interorbital papilla present or absent (present in holotype). Papillae unpigmented. Cephalic free neuromasts reduced and nearly indiscernible over the interorbit and nape. Gill opening small, 4.9 – 7.8 (7.6) %, entirely above pectoral fin or extending to pectoral­fin ray 3 (1). Gill rakers on anterior arch 7 – 11 (10), short and blunt. Central projection of orobuccal valve a moderately elongate single lobe. Dorsal­fin rays 39 – 42 (41), tips of anterior 4 – 6 rays projecting from fin membrane, anteriormost rays about 30 % free from membrane, succeeding rays less so; posteriormost ray attached membranously to dorsalmost caudal­fin ray for 2.3 – 5.8 (4.0) %. Anal fin with 33 – 35 (34) rays, posteriormost ray membranously attached to ventralmost caudal­fin ray for 3.0 – 5.9 (3.6) %. One or two (2) anal­fin pterygiophores and associated rays anterior to first haemal spine. Pectoral­fin rays 35 – 40 (37) in two lobes, separated by shallow notch, 8 – 10 (9) rays in lower lobe. Dorsalmost ray at level of ventral rim of orbit. Upper lobe rounded, extending to anal­fin origin, length 16.0 – 20.0 (17.1) %, with rays 5 – 6 (5) longest; length of shortest notch ray 5.8 – 10.5 (6.0) %; length of lower lobe 14.4 – 19.8 (15.9) %, extending between posterior margin of pelvic disk and anus, with ray 3 longest, rays 4 – 10 shortening ventrally. Pelvic disk large, length 8.6 – 11.8 (8.8) %, width 7.1 – 11.1 (8.2) %. Distance from disk to anus 1.5 – 5.5 (4.8) %, about 13 – 55 (55) % DL, distance from anus to anal­fin origin 6.7 – 18.8 (13.4) %, about 50 – 200 (152.5) % DL. Urogenital papilla conical, relatively long, 4 – 96.5 (6.0) % DL, unpigmented. Pyloric caeca about 20, primarily on right side of body, finger­like, long, about 25 – 50 (40) % HL. Caudal fin slightly rounded, 13.0 – 17.1 (14.4) %, depth at hypural plate 3.3 – 5.2 (4.2) %, with principal rays 11 – 13 (12), dorsal principal rays 5 – 6 (5); ventral principal rays 7 – 8 (7). Dorsal procurrent rays 2, borne on epural and posteriormost neural spine; ventral procurrent ray 1, borne on expanded posteriormost haemal spine. Vertebrae 45 – 47 (46), abdominal vertebrae 10 – 11 (11), caudal vertebrae 34 – 36 (35). Body in life uniform light red, scattered speckling rarely present at origin of dorsal fin; iris silver gray. Color in alcohol pale, with pigment absent from cephalic papillae and rarely present at origin of dorsal fin. Peritoneum black; orobranchial cavity pale. Largest specimen examined a 129.3 mm male (UW 113676). Smallest female with yolked eggs 85 mm; smallest male with enlarged, swollen testes 95 mm. Range Allocareproctus ungak has been collected only in the Aleutian Islands, from Seguam Pass to the Islands of Four Mountains, at depths of 318 to 461 m (Fig. 11).	en	Orr, James Wilder, Busby, Morgan Scott (2006): Revision of the snailfish genus Allocareproctus Pitruk & Fedorov (Teleostei: Liparidae), with descriptions of four new species from the Aleutian Islands. Zootaxa 1173: 1-37, DOI: 10.5281/zenodo.2645716
8619E9769455D92FEA13289020D2FA8B.taxon	etymology	Etymology The specific epithet “ ungak ” is taken from the Alutiiq word for “ whiskers ” an allusion to the many papillae associated with cephalic pores. It is to be treated as a noun in apposition. Comparisons	en	Orr, James Wilder, Busby, Morgan Scott (2006): Revision of the snailfish genus Allocareproctus Pitruk & Fedorov (Teleostei: Liparidae), with descriptions of four new species from the Aleutian Islands. Zootaxa 1173: 1-37, DOI: 10.5281/zenodo.2645716
8619E9769455D92FEA13289020D2FA8B.taxon	description	Allocareproctus ungak is most similar to A. unangas, which also has trilobed teeth (Fig. 2 D – E) and a uniform red body. It is distinguished from A. unangas by the presence of a papilla on nasal pore 1 and higher number of pores with associated papillae (Fig. 4). Pyloric caeca are longer and typically on the right side of the body, unlike A. unangas in which they are shorter and on the left. For meristic and morphometric differences from other species of Allocareproctus, see species accounts above.	en	Orr, James Wilder, Busby, Morgan Scott (2006): Revision of the snailfish genus Allocareproctus Pitruk & Fedorov (Teleostei: Liparidae), with descriptions of four new species from the Aleutian Islands. Zootaxa 1173: 1-37, DOI: 10.5281/zenodo.2645716
