taxonID	type	description	language	source
862DE85F73594F37DCBE7117FB60FD21.taxon	type_taxon	Type species: Compressalges nipponiae Dubinin, 1950, by original designation.	en	Waki, Tsukasa, Shimano, Satoshi (2022): Redescription of two parasitic feather mites sampled from the last two Crested Ibises, Nipponia nippon (Temminck, 1835) (Pelecaniformes: Threskiornithidae) lived in Japan. Zootaxa 5116 (1): 136-150, DOI: 10.11646/zootaxa.5116.1.7
862DE85F73594F37DCBE7117FB60FD21.taxon	discussion	This genus includes only the type species. Within the family Caudiferidae, this genus clearly differs from two other genera, Caudifera Gaud & Mouchet, 1959 and Semicaudifera Gaud & Atyeo, 1996, in having the following features. In males of Compressalges, the opisthosoma is strongly narrowed posteriorly and with a pair of elongated opisthosomal lobes, the ventral side of opisthosoma has long epimerites IVa and adanal apodemes, and legs IV are hypertrophied; in females, the external copulatory tube is absent and the copulatory opening is situated terminally between setae ps 1 (Dubinin 1950; Gaud & Atyeo 1996).	en	Waki, Tsukasa, Shimano, Satoshi (2022): Redescription of two parasitic feather mites sampled from the last two Crested Ibises, Nipponia nippon (Temminck, 1835) (Pelecaniformes: Threskiornithidae) lived in Japan. Zootaxa 5116 (1): 136-150, DOI: 10.11646/zootaxa.5116.1.7
862DE85F73594F33DCBE72E1FA0DF81F.taxon	description	(Figs. 2 – 6)	en	Waki, Tsukasa, Shimano, Satoshi (2022): Redescription of two parasitic feather mites sampled from the last two Crested Ibises, Nipponia nippon (Temminck, 1835) (Pelecaniformes: Threskiornithidae) lived in Japan. Zootaxa 5116 (1): 136-150, DOI: 10.11646/zootaxa.5116.1.7
862DE85F73594F33DCBE72E1FA0DF81F.taxon	vernacular_names	[Japanese name: Toki-umoudani]	en	Waki, Tsukasa, Shimano, Satoshi (2022): Redescription of two parasitic feather mites sampled from the last two Crested Ibises, Nipponia nippon (Temminck, 1835) (Pelecaniformes: Threskiornithidae) lived in Japan. Zootaxa 5116 (1): 136-150, DOI: 10.11646/zootaxa.5116.1.7
862DE85F73594F33DCBE72E1FA0DF81F.taxon	materials_examined	Material examined: 4 females and 4 males (MPM Coll. No. 21820) from feathers of Nipponia nippon (Temminck, 1835) (Threskiornithidae), JAPAN, Niigata Prefecture, Sado Island, Ministry of the Environment, Sado Japanese Crested Ibis Conservation Center, 21, August 1993 – 4, October 1994, feather coll. Dr. Yoshinori Kaneko.	en	Waki, Tsukasa, Shimano, Satoshi (2022): Redescription of two parasitic feather mites sampled from the last two Crested Ibises, Nipponia nippon (Temminck, 1835) (Pelecaniformes: Threskiornithidae) lived in Japan. Zootaxa 5116 (1): 136-150, DOI: 10.11646/zootaxa.5116.1.7
862DE85F73594F33DCBE72E1FA0DF81F.taxon	description	Description. FEMALE (Figs. 2 A – B, 4 A – D, 5 A – B, 6 A – B, D). Dorsal idiosoma. Gnathosoma: subcapitulum nearly square-shaped, length 97 – 126, greatest width 79 – 84. Body length including gnathosoma 506 – 558 long and 247 – 272 wide. Idiosoma 439 – 497 long, hysterosoma 349 – 377 long. Prodorsal shield strongly enlarged posteriorly, shaped as inverted mushroom, covering most part of prodorsum, anterior part with network pattern, posterior part with transverse striation, length 110 – 117, width of posterior part 117 – 124 (Figs. 2 B, 5 A). Setae ve rudimentary, situated on lateral margins of prodorsal shield, setae si and se near posterior margin of prodosal shield, setae se separated by 94 – 109. Hysteronotal shield enlarged in anterior part and gradually attenuate posteriorly, anterior margin with striations and shaped as trapezoid, close to prodorsal shield, length 329 – 374, greatest width 216 – 242, and surface with numerous small circular lacunae (Figs. 2 B, 5 B). Setae cp 153 – 187 long. Setae c 2 thin spiculiform, 37 – 43 long. Setae c 3 spiculiform, 34 – 39 long. Setae h 3 spiculiform 78 – 94 long. Hysteronotal gland openings gl between levels of d 2 and e 2, cupules im between d 2 and gl. Distances between setae and hysteronotal gland opening: se: se 87 – 92, si: si 51 – 55, d 2: e 2 56 – 76, d 2: gl 31 – 44, e 1: gl 63 – 75, h 1: h 1 23 – 29, h 2: h 2 57 – 63, h 3: h 3 45 – 52. Length of other dorsal setae: vi 8 – 11, se 94 – 109, c 1 7 – 10, d 1 9 – 13, d 2 12 – 23, e 1 13 – 16, e 2 11 – 13, f 2 17 – 22, h 1 10 – 13, h 2 210 – 252. Ventral idiosoma. Anus close to posterior margin of idiosoma. Oviporus shaped as an inverted Y, situated at level of sejugal furrow (Fig. 2 A, 6 A). Epigynum bow-shaped, 46 – 66 long and 41 – 62 wide. Cupules ih near lateral edges of opisthosoma at level of ps 3. Length of ventral setae: 1 a 38 – 56, 3 a 49 – 52, 4 a 47 – 57, 4 b 20 – 28, ps 2 10 – 11, ps 3 9 – 13. Copulatory opening situated terminally, near posterior margin between setae ps 1 (Fig. 2 A, 6 B). Legs (Figs. 4 A – D, Table 1). Length of tarsi, I – IV: 49 – 57, 46 – 54, 50 – 56, 58 – 69. Setation of legs I – IV: tarsi 8 – 7 – 6 – 6; tibiae 1 – 1 – 1 – 1; genua 2 – 2 – 0 – 0; femora 1 – 1 – 0 – 0; trochanters 1 – 1 – 1 – 0; coxae I – IV 1 – 0 – 1 – 0. Solenidiotaxy of legs I – IV: tarsi 2 – 1 – 0 – 0, tibiae I 1 – 1 – 1 – 1, genua 2 – 1 – 1 – 0. Famulus epsilon ɛ short spine-like (Figs. 4 A, 6 D), close to solenidion ω 1; setae ba of tarsi I, II distant from bases of solenidia ω 1. MALE. (Figs. 3 A – B, 4 E, 5 C – D, 6 C). Dorsal idiosoma. Gnathosoma similar to that of female, length 88 – 101, width 66 – 83. Body length, including gnathosoma, 471 – 494, greatest width 230 – 253. Idiosoma length 400 – 442, hysterosoma length 316 – 329. Prodorsal shield as in female, length 103 – 111, width of posterior part 107 – 113. Setae si and se in posterior part of prodosal shield, setae se separated by 79 – 84. Hysteronotal shield strongly enlarged in anterior part and gradually attenuate posteriorly, anterior margin trapezoid-shaped, slightly concave in middle part, length 266 – 300, greatest width 186 – 212, surface with small circular lacunae in central area (Fig. 5 C), and with large net-like ornamentation in posterior part (Fig. 5 D). Setae c 2 and c 3 spiculiform. Opisthosomal lobes small, elongate, almost parallel-sided, length 42 – 47, greatest width 26 – 27. Terminal cleft U-shaped, greatest width 9 – 14. Outer margins of opisthosomal lobes with lateral membranes spreading from bases to posterior one third. Setae h 1 narrowly lanceolate and situated on inner margins of opisthosomal lobes. Setae f 2 spiculiform, situated near outer margins of opisthosomal lobes. Setae ps 2 narrowly lanceolate, situated on lateral margins of opisthosomal lobes. Macrosetae h 2 and h 3 situated on posterolateral angles of opisthosomal lobes. Setae ps 1 wide fan-shaped with radial striation, situated on terminus of opisthosomal lobes. Cupules im and hysteronotal gland openings gl between d 2 and e 2. Distances between setae and hysteronotal gland opening: se: se 79 – 86, si: si 46 – 54, d 2: e 2 77 – 94, d 2: gl 59 – 76, e 1: gl 30 – 38, h 1: h 1 11 – 15, h 2: h 2 60 – 68 h 3: h 3 45 – 52. Length of dorsal setae: vi 5 – 9, si 4 – 7, se 39 – 49, cp 141 – 205, c 1 5 – 8, c 2 28 – 38, c 3 25 – 33, d 1 5 – 8, d 2 6 – 8, e 1 4 – 7, e 2 6, f 2 18 – 25, h 1 22 – 25, h 2 197 – 241, h 3 183 – 229. Ventral idiosoma. Genital apparatus between trochanters IV, flanked laterally with setae g and ps 3. Setae ps 3 occasionally duplicated, additional pair smaller and situated anterior to original ps 3 (Fig. 6 C). Genital papillae posterior to setae 4 a. Aedeagus stylet-like, 30 – 34 in length. Adanal suckers barrel-shaped, retracted in opisthosoma, 20 – 24 in length and 16 – 25 in width. Anus near anterior end of terminal cleft. Epimerites IVa long; adanal apodemes well developed, situated immediately posterior and parallel to epimerites IVa. Cupules ih near posterior ends of opisthosomal lobes. Length of ventral setae: 1 a 32 – 39, 3 a 31 – 42, 4 a 28 – 38, 4 b 23 – 28, ps 1 31 – 34, ps 2 23 – 29, ps 3 13 – 17, g 21 – 27. Legs (Fig. 4 E, Table 1): Legs IV hypertrophied. Length of tarsi I – IV: 46 – 51, 42 – 49, 45 – 50, 67 – 70. Leg setation as in female. Seta e of tarsus IV filiform, seta d small spiculiform and situated at level of distal 1 / 4 of this segment.	en	Waki, Tsukasa, Shimano, Satoshi (2022): Redescription of two parasitic feather mites sampled from the last two Crested Ibises, Nipponia nippon (Temminck, 1835) (Pelecaniformes: Threskiornithidae) lived in Japan. Zootaxa 5116 (1): 136-150, DOI: 10.11646/zootaxa.5116.1.7
862DE85F73594F33DCBE72E1FA0DF81F.taxon	discussion	Remarks. In the original description of Compressalges nipponiae and in the subsequent work, Dubinin (1950, 1956) erroneously interpreted the hysteronotal glands, which are dark-colored and have longitudinal striation, as some ventral setae modified in “ large funnel-shaped suckers ”. Gaud & Atyeo (1996) provided correct illustrations of this mite, but their drawing were too small-sized and did not show some tiny details of the genital apparatus, adanal suckers and opisthosoma of male.	en	Waki, Tsukasa, Shimano, Satoshi (2022): Redescription of two parasitic feather mites sampled from the last two Crested Ibises, Nipponia nippon (Temminck, 1835) (Pelecaniformes: Threskiornithidae) lived in Japan. Zootaxa 5116 (1): 136-150, DOI: 10.11646/zootaxa.5116.1.7
862DE85F73534F3DDCBE70ECF95DFDD4.taxon	type_taxon	Type species: Freyanopterolichus nipponiae Dubinin, 1953, by original designation.	en	Waki, Tsukasa, Shimano, Satoshi (2022): Redescription of two parasitic feather mites sampled from the last two Crested Ibises, Nipponia nippon (Temminck, 1835) (Pelecaniformes: Threskiornithidae) lived in Japan. Zootaxa 5116 (1): 136-150, DOI: 10.11646/zootaxa.5116.1.7
862DE85F73534F3DDCBE70ECF95DFDD4.taxon	discussion	The genus Freyanopterolichus currently includes 10 species occurring on ibises (Threskiornithidae), storks (Ciconiidae), hamerkops (Scopidae) and cranes (Gruidae) (Trouessart & Mégnin, 1885; Dubinin 1953; Gaud & Mouchet, 1959; Gaud 1982 a, 1982 b). Within the family, this genus is clearly characterized by the following combination of characters. In both sexes, two internal vertical setae vi are present, epimerites I are free, the prodorsal shield is wide triangular and encompasses both pairs of scapular setae, the ambulacral discs of legs I are larger than those of legs II; in females, macrosetae h 2 and h 3 are long whip-like and usually dilated in basal part, the epigynum is situated between the extremities of epimerites I, and the oviporus is moved anterior and situated between levels of trochanters II and III.	en	Waki, Tsukasa, Shimano, Satoshi (2022): Redescription of two parasitic feather mites sampled from the last two Crested Ibises, Nipponia nippon (Temminck, 1835) (Pelecaniformes: Threskiornithidae) lived in Japan. Zootaxa 5116 (1): 136-150, DOI: 10.11646/zootaxa.5116.1.7
862DE85F73534F3FDCBE72F1FDF2F9B2.taxon	vernacular_names	[Japanese name: Toki-enban-umoudani]	en	Waki, Tsukasa, Shimano, Satoshi (2022): Redescription of two parasitic feather mites sampled from the last two Crested Ibises, Nipponia nippon (Temminck, 1835) (Pelecaniformes: Threskiornithidae) lived in Japan. Zootaxa 5116 (1): 136-150, DOI: 10.11646/zootaxa.5116.1.7
862DE85F73534F3FDCBE72F1FDF2F9B2.taxon	description	(Figs. 7 – 10)	en	Waki, Tsukasa, Shimano, Satoshi (2022): Redescription of two parasitic feather mites sampled from the last two Crested Ibises, Nipponia nippon (Temminck, 1835) (Pelecaniformes: Threskiornithidae) lived in Japan. Zootaxa 5116 (1): 136-150, DOI: 10.11646/zootaxa.5116.1.7
862DE85F73534F3FDCBE72F1FDF2F9B2.taxon	materials_examined	Material examined. 4 females and 4 males (MPM Coll. No. 21821) from feathers of Nipponia nippon (Temminck, 1835) (Threskiornithidae), Japan, Niigata Prefecture Sado Island, Ministry of the Environment, Sado Japanese Crested Ibis Conservation Center, 21, August 1993 – 4, October 1994, feather coll. Dr. Yoshinori Kaneko.	en	Waki, Tsukasa, Shimano, Satoshi (2022): Redescription of two parasitic feather mites sampled from the last two Crested Ibises, Nipponia nippon (Temminck, 1835) (Pelecaniformes: Threskiornithidae) lived in Japan. Zootaxa 5116 (1): 136-150, DOI: 10.11646/zootaxa.5116.1.7
862DE85F73534F3FDCBE72F1FDF2F9B2.taxon	description	Description. FEMALE (Figs. 7 A – B, 9 A – D, 10 A – B, E, G). Dorsal idiosoma. Subcapitulum trapezoidal, length 77 – 120, width at base 91 – 139. Body length including gnathosoma 438 – 490. Idiosoma transversally ovate, slightly wider than long, strongly flattened dorsoventrally, length 386 – 445 and greatest width 386 – 445, width including lateral membranes 467 – 509. Prodorsal shield large, occupying entire surface of prodorsum, roughly triangular, with two pairs of angular lateral incisions, length 79 – 97, greatest width 228 – 249, (Fig. 7 A). Setae vi spiculiform, situated near anterior end of prodorsal shield. Setae si and se situated at posterior margin of this shield, setae si narrowly lanceolate, and setae se long sword-shaped, separated by 104 – 108. Scapular shields represented by small curved sclerites. Lateral margins of hysterosoma with narrow membranes stretching from sejugal furrow to bases of setae f 2; length of hysterosoma 310 – 330. Hysteronotal shield covering most part of hysterosoma, wider than long, anterior margin straight, length 278 – 290, greatest width 318 – 457, most surface with polygonal net-work pattern (Fig. 10 A), ovate median area near posterior margin with small pit-like lacunae (Fig. 10 B). Seta c 2 narrowly lanceolate, situated on soft tegument near lateral margins of propodosoma. Seta cp long whip-like, situated marginally slightly posterior to bases of setae c 2. Setae d 2, e 2, f 2, h 2, h 3, ps 1, ps 2 situated near lateral margins of hysterosoma. Setae d 2 filiform. Setae e 2, f 2, ps 1 dilated, narrowly lanceolate, macrosetae h 2 and h 3 dilated along all their length, gradually attenuate to apices. Seta ps 2 short filiform. Hysteronotal gland opening gl posteromesal from setae e 2. Cupules ip close tosetae f 2, cupules im indistinct. Distance between setae and hysteronotal gland openings: si: si 53 – 64, se: se 104 – 118, c 1: d 1 78 – 89, d 1: gl 65 – 95, d 2: gl 93 – 120, e 1: gl 56 – 68, e 1: h 1 89 – 101, h 1: h 1 114 – 154, h 2: h 2 219 – 247, h 3: h 3 148 – 160, ps 1: ps 1 103 – 122. Length of dosal setae: vi 11 – 15, si 78 – 113, se 156 – 158, c 1 12 – 23, c 2 70 – 85, cp 126 – 156, d 1 12 – 19, d 2 20 – 24, e 1 16 – 24, e 2 53 – 80, f 2 57 – 82, h 1 17 – 24, h 2 266 – 353, h 3 241 – 357, ps 1 63 – 84. Ventral idiosoma. Epimerite I free. Seta c 3 filiform, situated posterior to humeral shields. Epigynum almost semicircular, situated between tips of epimerites I. Oviporus long, shaped as an inverted Y, apodemes of oviporus extending to level of trochanters III (Figs. 7 A, 10 E). Anus close to posterior margin of idiosoma. Length of ventral setae: 1 a 35 – 50, 3 a 14 – 25, 4 a 40 – 52, 4 b 38, c 3 20 – 23, ps 2 33 – 49, ps 3 17 – 22, g 20 – 29. Distance between setae: 1 a: 4 b 64 – 78, 3 a: 4 b 46 – 68, 4 b: g 18 – 24, 4 a: g 55 – 68, 4 a: ps 3 149 – 169. Legs (Figs. 9 A – D, 10 G, Table 1). Length of tarsi I – IV: 47 – 53, 60 – 63, 58 – 66 and 61 – 87, respectively. Length of tarsus I and II ca. 2 times of each greatest width. Setation of legs I – IV (excluding proral setae p and q): tarsi 8 – 8 – 4 – 6; tibiae 1 – 1 – 1 – 1; genua I – IV 2 – 2 – 0 – 0; femora I – IV 1 – 1 – 0 – 0; trochanters I – IV 0 – 1 – 1 – 0; coxae I – IV 1 – 0 – 1 – 0. Solenidiotaxy of legs I – II – III – IV: tarsi 2 – 1 – 0 – 0, tibiae 1 – 1 – 1 – 1, genua 1 – 1 – 1 – 0. Ambulacral discs I with smooth margin, discs of leg II – IV with ca. 10 denticles on distal edges. MALE. Dorsal idiosoma (Figs. 8 A – B, 9 E, 10 C – D). Gnathosoma as in male, length 71 – 81, width 86 – 101. Body length including gnathosoma 425 – 450. Idiosoma transversally ovate, slightly wider than long, length 340 – 394 and greatest width 423 – 443, width including lateral membranes 437 – 448. Length of hysterosoma 281 – 297. Prodorsal shield as in female, length 75 – 92 and width 213 – 228. Setae vi spiculiform, situated near anterior end of prodorsal shield. Setae si narrowly lanceolate, and setae se long sword-shaped, separated by 94 – 105. Scapular shields represented by small bow-shaped sclerites. Lateral margins of hysterosoma with narrow membranes stretching from sejugal furrow to bases of setae f 2. Hysteronotal shield: roughly ovate, wider than long, anterior margin slightly convex, length 245 – 273 and greatest width 388 – 401, most surface with numerous polygonal network pattern, area near posterior end additionally with small pits (Figs. 10 C, D). Setae cp and c 3 as in female. Setae d 2, e 2, f 2, h 2, h 3, ps 1, ps 2 situated near lateral margins of hysterosoma. Setae d 2 filiform. Setae e 2, f 2, ps 2 and ps 1 dilated, daggershaped; macrosetae h 2 and h 3 long dilated along all their length, gradually attenuate to apices. Distance between setae and hysteronotal gland openings: si: si 37 – 46, se: se 94 – 105, c 1: d 1 77 – 90, d 1: gl 74 – 96, d 2: gl 99 – 108, e 1: gl 54 – 67, e 1: h 1 80 – 88, h 1: h 1 125 – 131, h 2: h 2 209 – 244, h 3: h 3 140 – 171, ps 1: ps 1 81 – 120. Length of dorsal setae: vi 13 – 18, si 95 – 107, se 148 – 165, c 1 18 – 23, c 2 69 – 91, d 1 12 – 18, d 2 22 – 30, e 1 12 – 15, e 2 77 – 87, f 2 61 – 74, h 1 14 – 17, h 2 186 – 342, h 3 223 – 294, ps 1 67 – 82. Ventral idiosoma. Epimerites I free, closer to each other than in female. Genital apparatus at level of trochanters IV. Aedeagus hook-shaped (Fig. 10 F), 43 – 46 long. Both pairs of genital papillae and setae g approximately at midlength of genital arch. Anus close to posterior margin of opisthosoma. Adanal suckers situated at level of anus, disc-shaped, 23 – 26 in diameter, with radial patterns. Cupules ip close to f 2. Length of ventral setae: 1 a 57 – 74, 3 a 43 – 53, 4 a 26 – 40, 4 b 28 – 49, c 3 28 – 43, ps 2 65 – 70, ps 3 25 – 30, g 30 – 34. Legs (Fig. 9 E, Table 1). Shape and setation as in female except tarsus IV. Length of tarsi I – IV 45 – 66, 45 – 65, 53 – 73, 61 – 81. Setae d and e of tarsus IV small spine-like, situated in distal 1 / 3 of segment.	en	Waki, Tsukasa, Shimano, Satoshi (2022): Redescription of two parasitic feather mites sampled from the last two Crested Ibises, Nipponia nippon (Temminck, 1835) (Pelecaniformes: Threskiornithidae) lived in Japan. Zootaxa 5116 (1): 136-150, DOI: 10.11646/zootaxa.5116.1.7
862DE85F73534F3FDCBE72F1FDF2F9B2.taxon	discussion	Remarks. Among four species of the genus Freyanopterolichus associated with ibises (Mégnin & Trouessart 1884; Trouessart & Mégnin 1885; Gaud & Mouchet 1959; Gaud 1982 b), F. nipponiae is most close to F. chorioptoides (Mégnin & Trouessart, 1884) from Bostrychia carunculata (Threskiornithidae) in having: in both sexes, the idiosoma almost circular in shape, macrosetae h 2 and h 3 dilated, and idiosomal setae f 2, and ps 1 dagger-shaped; and in females, setae ps 2 dagger-shaped. Freyanopterolichus nipponiae can be distinguished from the latter in the following combination of characters. In both sexes, the idiosoma is transversely ovate and slightly wider than long, tarsi I, II are approximately 2 times longer than greatest wide, macrosetae h 2 and h 3 are dilated along all their length and gradually attenuate to apices; in males, the terminal cleft is absent; in females, setae ps 1 are as long as the distance between their bases. In both sexes of F. chorioptoides, the idiosoma is approximately as long as wide, tarsi I, II are approximately 3 times longer than their greatest wide, dilatations of macrosetae h 2 and h 3 are developed only in the basal part and have a noticeable angular extension on the inner margin; in males, the terminal cleft is shaped as a wide and shallow concavity; in females, setae ps 1 are nearly two times as long as the distance between their bases.	en	Waki, Tsukasa, Shimano, Satoshi (2022): Redescription of two parasitic feather mites sampled from the last two Crested Ibises, Nipponia nippon (Temminck, 1835) (Pelecaniformes: Threskiornithidae) lived in Japan. Zootaxa 5116 (1): 136-150, DOI: 10.11646/zootaxa.5116.1.7
