taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
8569BD6CFF96126031A91C47FDC2F972.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4416665/files/figure.png	https://doi.org/10.5281/zenodo.4416665	FIGURE 2. Heth pivari n. sp., female. A) Habitus, sublateral view, right side. B) Anterior end, sublateral view. C) Part of anterior region, lateral view showing neck retractor muscles; rp, ramified projection of the stoma entrance lumen (Spiridonov & Yushin 2000). D) Collar, lateral view; box shows dimensions of shield. E) Oviduct-uterus region with migrating sperm. Scale bars: A, 230 µm; B, C: 25 µm; D, 40 µm; E, 115 µm.	FIGURE 2. Heth pivari n. sp., female. A) Habitus, sublateral view, right side. B) Anterior end, sublateral view. C) Part of anterior region, lateral view showing neck retractor muscles; rp, ramified projection of the stoma entrance lumen (Spiridonov & Yushin 2000). D) Collar, lateral view; box shows dimensions of shield. E) Oviduct-uterus region with migrating sperm. Scale bars: A, 230 µm; B, C: 25 µm; D, 40 µm; E, 115 µm.	2020-10-20	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.		Zenodo	biologists	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.			
8569BD6CFF96126031A91C47FDC2F972.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4416668/files/figure.png	https://doi.org/10.5281/zenodo.4416668	FIGURE 3. SEM micrographs of Heth pivari n. sp. females. A) Anterior region, sublateral view. B) Anterior region, lateral view. C) Head, lateral view. D) Shield region. Arrows indicate button-like papillae. Scale bars: A, B, 50 µm; C, D, 20 µm.	FIGURE 3. SEM micrographs of Heth pivari n. sp. females. A) Anterior region, sublateral view. B) Anterior region, lateral view. C) Head, lateral view. D) Shield region. Arrows indicate button-like papillae. Scale bars: A, B, 50 µm; C, D, 20 µm.	2020-10-20	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.		Zenodo	biologists	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.			
8569BD6CFF96126031A91C47FDC2F972.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4416670/files/figure.png	https://doi.org/10.5281/zenodo.4416670	FIGURE 4. Heth pivari n. sp., male. A) Habitus, lateral view, right side. B) Anterior region, lateral view. C) Anterior region, subdorsal view. D) Posterior region, lateral view, right side. E) Gubernaculum and fused spicules, dorsal view. Scale bars: A, 250 µm; B–E, 50 µm.	FIGURE 4. Heth pivari n. sp., male. A) Habitus, lateral view, right side. B) Anterior region, lateral view. C) Anterior region, subdorsal view. D) Posterior region, lateral view, right side. E) Gubernaculum and fused spicules, dorsal view. Scale bars: A, 250 µm; B–E, 50 µm.	2020-10-20	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.		Zenodo	biologists	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.			
8569BD6CFF96126031A91C47FDC2F972.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4416674/files/figure.png	https://doi.org/10.5281/zenodo.4416674	FIGURE 5. SEM micrographs of Heth pivari n. sp. males. A) Anterior end, stoma slit-like. B) Anterior end, stoma open. C) Closeup of stoma showing striae and minute amphid aperture (a). D) Posterior region showing ventral sucker (S), papillae 1,2 and 4, and tail. E) Posterior region showing arrangement of posterior papillae 2–7. F) Closeup of cloacal region. Scale bars: A, B, E, 30 µm; C, F, 20 µm; D, 200 µm.	FIGURE 5. SEM micrographs of Heth pivari n. sp. males. A) Anterior end, stoma slit-like. B) Anterior end, stoma open. C) Closeup of stoma showing striae and minute amphid aperture (a). D) Posterior region showing ventral sucker (S), papillae 1,2 and 4, and tail. E) Posterior region showing arrangement of posterior papillae 2–7. F) Closeup of cloacal region. Scale bars: A, B, E, 30 µm; C, F, 20 µm; D, 200 µm.	2020-10-20	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.		Zenodo	biologists	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.			
8569BD6CFF96126031A91C47FDC2F972.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4416676/files/figure.png	https://doi.org/10.5281/zenodo.4416676	FIGURE 6. DIC micrographs of Heth pivari n. sp., female and male structures. A) Lateral ornamentation in anterior region of female. B) Male, lateral somal extensions of posterior arcade surrounding anterior esophagus. C) Female, secretory-excretory pore (arrow). D. Female, somatic papillae anterior to vulva. E) Fusiform to comet-shaped sperm in male. All scale bars 20 µm.	FIGURE 6. DIC micrographs of Heth pivari n. sp., female and male structures. A) Lateral ornamentation in anterior region of female. B) Male, lateral somal extensions of posterior arcade surrounding anterior esophagus. C) Female, secretory-excretory pore (arrow). D. Female, somatic papillae anterior to vulva. E) Fusiform to comet-shaped sperm in male. All scale bars 20 µm.	2020-10-20	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.		Zenodo	biologists	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.			
8569BD6CFF96126031A91C47FDC2F972.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4416680/files/figure.png	https://doi.org/10.5281/zenodo.4416680	FIGURE 7. Heth pivari n. sp. A) Live female with single egg. B–E) Female enlargement and increasing numbers of eggs. The uterus is capacious but the vagina holds just one egg at a time (B, D). F) Female, dense trichomycete attachment in anterior region. G) Male, arthropod seta lodged in anteriormost region of intestine. H) Male, lateral chord (arrows). Scale bars: A, 500 µm; B–F, 250 µm; G, 100 µm; H, 50 µm.	FIGURE 7. Heth pivari n. sp. A) Live female with single egg. B–E) Female enlargement and increasing numbers of eggs. The uterus is capacious but the vagina holds just one egg at a time (B, D). F) Female, dense trichomycete attachment in anterior region. G) Male, arthropod seta lodged in anteriormost region of intestine. H) Male, lateral chord (arrows). Scale bars: A, 500 µm; B–F, 250 µm; G, 100 µm; H, 50 µm.	2020-10-20	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.		Zenodo	biologists	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.			
8569BD6CFF96126031A91C47FDC2F972.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4416663/files/figure.png	https://doi.org/10.5281/zenodo.4416663	FIGURE 1. Heth mauriesi female, terminology used to describe the anterior ornamentation of Heth spp. A) Drawings of H. mauriesi from Adamson (1982), courtesy of Dr. Martin Adamson. B) SEM of H. mauriesi, previously unpublished image courtesy of Dr. Ramon Carreno. Scale bars 50 µm.	FIGURE 1. Heth mauriesi female, terminology used to describe the anterior ornamentation of Heth spp. A) Drawings of H. mauriesi from Adamson (1982), courtesy of Dr. Martin Adamson. B) SEM of H. mauriesi, previously unpublished image courtesy of Dr. Ramon Carreno. Scale bars 50 µm.	2020-10-20	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.		Zenodo	biologists	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.			
8569BD6CFF96126031A91C47FDC2F972.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4416693/files/figure.png	https://doi.org/10.5281/zenodo.4416693	FIGURE 12. Worldwide Heth spp. Group 2 (continued). A) H. tonkinensis (after Malysheva & Spiridonov 2010). B) H. tuxtlensis (after Mejía-Madrid 2014). C) H. vietnamensis (after Malysheva & Spiridonov 2010). D) H. xaniophora (after Hunt 1994), en face, lateral and dorso-ventral views.	FIGURE 12. Worldwide Heth spp. Group 2 (continued). A) H. tonkinensis (after Malysheva & Spiridonov 2010). B) H. tuxtlensis (after Mejía-Madrid 2014). C) H. vietnamensis (after Malysheva & Spiridonov 2010). D) H. xaniophora (after Hunt 1994), en face, lateral and dorso-ventral views.	2020-10-20	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.		Zenodo	biologists	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.			
8569BD6CFF96126031A91C47FDC2F972.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4416684/files/figure.png	https://doi.org/10.5281/zenodo.4416684	FIGURE 8. Molecular relationships among Heth spp. and other Rhigonematomorpha and Oxyuridomorpha using partial sequences of the D2-D3 expansion segment of 28S LSU rDNA. Bootstrap and posterior probability scores are placed above and below, respectively, corresponding nodes. XXXXXXXDNA sequence of H.mauriesi unpublished.	FIGURE 8. Molecular relationships among Heth spp. and other Rhigonematomorpha and Oxyuridomorpha using partial sequences of the D2-D3 expansion segment of 28S LSU rDNA. Bootstrap and posterior probability scores are placed above and below, respectively, corresponding nodes. XXXXXXXDNA sequence of H.mauriesi unpublished.	2020-10-20	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.		Zenodo	biologists	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.			
8569BD6CFF84127831A91E78FA5EF95A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4416686/files/figure.png	https://doi.org/10.5281/zenodo.4416686	FIGURE 9. Worldwide Heth spp., continuous collar and no shield (Group 1). A) H. albertoi (after García & Fontana 2004). B) H. amazonensis (after Kloss 1965). C) H. bifidispiculum (after Adamson 1982). D) H. clunyi (after Adamson 1985). E) H. coyi (after García et al. 1997). F) H.gordae (after Mejía-Madrid 2014). G) H.hexaspinosum (after Chitwood 1935).H) H. hispaniolae (after García et al. 2001). I) H. konoplevi (after Malysheva & Sumaya 2017). J) H. maicuru (after Adamson 1983).	FIGURE 9. Worldwide Heth spp., continuous collar and no shield (Group 1). A) H. albertoi (after García & Fontana 2004). B) H. amazonensis (after Kloss 1965). C) H. bifidispiculum (after Adamson 1982). D) H. clunyi (after Adamson 1985). E) H. coyi (after García et al. 1997). F) H.gordae (after Mejía-Madrid 2014). G) H.hexaspinosum (after Chitwood 1935).H) H. hispaniolae (after García et al. 2001). I) H. konoplevi (after Malysheva & Sumaya 2017). J) H. maicuru (after Adamson 1983).	2020-10-20	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.		Zenodo	biologists	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.			
8569BD6CFF84127831A91E78FA5EF95A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4416688/files/figure.png	https://doi.org/10.5281/zenodo.4416688	FIGURE 10. Worldwide Heth spp. Group 1 (continued). A) H. mauriesi (after Adamson 1982). B) H. perarmatum (after Dollfus 1952). C, D) H. pinnatum, head end and neck region (after García & Coy 1995). E) H. spinosum (after Adamson 1983). F) H. xarochae (after Mejía-Madrid 2014). G) H. zeuglocantha (after Hunt 1994).	FIGURE 10. Worldwide Heth spp. Group 1 (continued). A) H. mauriesi (after Adamson 1982). B) H. perarmatum (after Dollfus 1952). C, D) H. pinnatum, head end and neck region (after García & Coy 1995). E) H. spinosum (after Adamson 1983). F) H. xarochae (after Mejía-Madrid 2014). G) H. zeuglocantha (after Hunt 1994).	2020-10-20	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.		Zenodo	biologists	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.			
8569BD6CFF84127831A91E78FA5EF95A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4416691/files/figure.png	https://doi.org/10.5281/zenodo.4416691	FIGURE 11. Worldwide Heth spp., continuous collar with cervical shield (Group 2). A) H. baudini (after Malysheva & Cribb 2012). B) H. costata (after Hunt 1994). C. H. dimorphum (after Chitwood 1935). D) H. hamatus (after Bowie 1985). E) H. impalutiensis (after Malysheva et al. 2012). F) H. insularis (after Kloss 1965). G) H. initiensis (after Malysheva & Sumaya 2017). H) H. juli (after Cobb 1898). I) H. orthopori (after Adamson 1987). J) H. ortonwilliamsi (after Hunt 1994). K) H. sutherlandi (after Hunt 1994). L) H. taybaci (after Malysheva & Spiridonov 2010). M) H. taynguyeni (after Malysheva & Spiridonov 2010).	FIGURE 11. Worldwide Heth spp., continuous collar with cervical shield (Group 2). A) H. baudini (after Malysheva & Cribb 2012). B) H. costata (after Hunt 1994). C. H. dimorphum (after Chitwood 1935). D) H. hamatus (after Bowie 1985). E) H. impalutiensis (after Malysheva et al. 2012). F) H. insularis (after Kloss 1965). G) H. initiensis (after Malysheva & Sumaya 2017). H) H. juli (after Cobb 1898). I) H. orthopori (after Adamson 1987). J) H. ortonwilliamsi (after Hunt 1994). K) H. sutherlandi (after Hunt 1994). L) H. taybaci (after Malysheva & Spiridonov 2010). M) H. taynguyeni (after Malysheva & Spiridonov 2010).	2020-10-20	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.		Zenodo	biologists	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.			
8569BD6CFF84127831A91E78FA5EF95A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4416695/files/figure.png	https://doi.org/10.5281/zenodo.4416695	FIGURE 13. Worldwide Heth spp., discontinuous collar, no shield (Group 3). A) H. artigasi (after Dollfus 1952). B) H. artigasi (after Adamson 1983). C) H. baracoa (after Spiridonov 1989). D) H. duvidosum (after Artigas 1929). E) H. imias (after Spridonov 1989). F) H. macrocephala (after Kloss 1965). G) H. multiplus (after Kloss 1965). H) H. parartigasi (after Adamson 1985).	FIGURE 13. Worldwide Heth spp., discontinuous collar, no shield (Group 3). A) H. artigasi (after Dollfus 1952). B) H. artigasi (after Adamson 1983). C) H. baracoa (after Spiridonov 1989). D) H. duvidosum (after Artigas 1929). E) H. imias (after Spridonov 1989). F) H. macrocephala (after Kloss 1965). G) H. multiplus (after Kloss 1965). H) H. parartigasi (after Adamson 1985).	2020-10-20	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.		Zenodo	biologists	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.			
8569BD6CFF84127831A91E78FA5EF95A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4416697/files/figure.png	https://doi.org/10.5281/zenodo.4416697	FIGURE 14. Worldwide Heth spp. Group 3 (continued). A) H. sinediscus (after Kloss 1965). B) H. spinalatum (after Kloss 1965). C) H. spoliatus (after García & Coy 1995). D) H. travassosi (Dollfus 1952). E) H. travfilhoi (after Adamson 1983). F) H. tuzetae (after Adamson 1983).	FIGURE 14. Worldwide Heth spp. Group 3 (continued). A) H. sinediscus (after Kloss 1965). B) H. spinalatum (after Kloss 1965). C) H. spoliatus (after García & Coy 1995). D) H. travassosi (Dollfus 1952). E) H. travfilhoi (after Adamson 1983). F) H. tuzetae (after Adamson 1983).	2020-10-20	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.		Zenodo	biologists	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.			
8569BD6CFF84127831A91E78FA5EF95A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4416699/files/figure.png	https://doi.org/10.5281/zenodo.4416699	FIGURE 15. Worldwide Heth spp. not conforming to any other group (Group 4). A) H. adolphi (after Sanchez-Velazquez 1979). B) H. josephinae (after Sanchez-Velazquez 1979). C) H. lamothei (after Sanchez-Velazquez 1979). D) H. poeyi (after Otero et al. 1993).	FIGURE 15. Worldwide Heth spp. not conforming to any other group (Group 4). A) H. adolphi (after Sanchez-Velazquez 1979). B) H. josephinae (after Sanchez-Velazquez 1979). C) H. lamothei (after Sanchez-Velazquez 1979). D) H. poeyi (after Otero et al. 1993).	2020-10-20	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.		Zenodo	biologists	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.			
8569BD6CFF84127931A91DDAFA5EFB35.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4416686/files/figure.png	https://doi.org/10.5281/zenodo.4416686	FIGURE 9. Worldwide Heth spp., continuous collar and no shield (Group 1). A) H. albertoi (after García & Fontana 2004). B) H. amazonensis (after Kloss 1965). C) H. bifidispiculum (after Adamson 1982). D) H. clunyi (after Adamson 1985). E) H. coyi (after García et al. 1997). F) H.gordae (after Mejía-Madrid 2014). G) H.hexaspinosum (after Chitwood 1935).H) H. hispaniolae (after García et al. 2001). I) H. konoplevi (after Malysheva & Sumaya 2017). J) H. maicuru (after Adamson 1983).	FIGURE 9. Worldwide Heth spp., continuous collar and no shield (Group 1). A) H. albertoi (after García & Fontana 2004). B) H. amazonensis (after Kloss 1965). C) H. bifidispiculum (after Adamson 1982). D) H. clunyi (after Adamson 1985). E) H. coyi (after García et al. 1997). F) H.gordae (after Mejía-Madrid 2014). G) H.hexaspinosum (after Chitwood 1935).H) H. hispaniolae (after García et al. 2001). I) H. konoplevi (after Malysheva & Sumaya 2017). J) H. maicuru (after Adamson 1983).	2020-10-20	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.		Zenodo	biologists	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.			
8569BD6CFF84127931A91DDAFA5EFB35.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4416688/files/figure.png	https://doi.org/10.5281/zenodo.4416688	FIGURE 10. Worldwide Heth spp. Group 1 (continued). A) H. mauriesi (after Adamson 1982). B) H. perarmatum (after Dollfus 1952). C, D) H. pinnatum, head end and neck region (after García & Coy 1995). E) H. spinosum (after Adamson 1983). F) H. xarochae (after Mejía-Madrid 2014). G) H. zeuglocantha (after Hunt 1994).	FIGURE 10. Worldwide Heth spp. Group 1 (continued). A) H. mauriesi (after Adamson 1982). B) H. perarmatum (after Dollfus 1952). C, D) H. pinnatum, head end and neck region (after García & Coy 1995). E) H. spinosum (after Adamson 1983). F) H. xarochae (after Mejía-Madrid 2014). G) H. zeuglocantha (after Hunt 1994).	2020-10-20	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.		Zenodo	biologists	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.			
8569BD6CFF84127931A91DDAFA5EFB35.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4416691/files/figure.png	https://doi.org/10.5281/zenodo.4416691	FIGURE 11. Worldwide Heth spp., continuous collar with cervical shield (Group 2). A) H. baudini (after Malysheva & Cribb 2012). B) H. costata (after Hunt 1994). C. H. dimorphum (after Chitwood 1935). D) H. hamatus (after Bowie 1985). E) H. impalutiensis (after Malysheva et al. 2012). F) H. insularis (after Kloss 1965). G) H. initiensis (after Malysheva & Sumaya 2017). H) H. juli (after Cobb 1898). I) H. orthopori (after Adamson 1987). J) H. ortonwilliamsi (after Hunt 1994). K) H. sutherlandi (after Hunt 1994). L) H. taybaci (after Malysheva & Spiridonov 2010). M) H. taynguyeni (after Malysheva & Spiridonov 2010).	FIGURE 11. Worldwide Heth spp., continuous collar with cervical shield (Group 2). A) H. baudini (after Malysheva & Cribb 2012). B) H. costata (after Hunt 1994). C. H. dimorphum (after Chitwood 1935). D) H. hamatus (after Bowie 1985). E) H. impalutiensis (after Malysheva et al. 2012). F) H. insularis (after Kloss 1965). G) H. initiensis (after Malysheva & Sumaya 2017). H) H. juli (after Cobb 1898). I) H. orthopori (after Adamson 1987). J) H. ortonwilliamsi (after Hunt 1994). K) H. sutherlandi (after Hunt 1994). L) H. taybaci (after Malysheva & Spiridonov 2010). M) H. taynguyeni (after Malysheva & Spiridonov 2010).	2020-10-20	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.		Zenodo	biologists	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.			
8569BD6CFF84127931A91DDAFA5EFB35.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4416663/files/figure.png	https://doi.org/10.5281/zenodo.4416663	FIGURE 1. Heth mauriesi female, terminology used to describe the anterior ornamentation of Heth spp. A) Drawings of H. mauriesi from Adamson (1982), courtesy of Dr. Martin Adamson. B) SEM of H. mauriesi, previously unpublished image courtesy of Dr. Ramon Carreno. Scale bars 50 µm.	FIGURE 1. Heth mauriesi female, terminology used to describe the anterior ornamentation of Heth spp. A) Drawings of H. mauriesi from Adamson (1982), courtesy of Dr. Martin Adamson. B) SEM of H. mauriesi, previously unpublished image courtesy of Dr. Ramon Carreno. Scale bars 50 µm.	2020-10-20	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.		Zenodo	biologists	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.			
8569BD6CFF85127A31A91E3FFA5EFD5D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4416691/files/figure.png	https://doi.org/10.5281/zenodo.4416691	FIGURE 11. Worldwide Heth spp., continuous collar with cervical shield (Group 2). A) H. baudini (after Malysheva & Cribb 2012). B) H. costata (after Hunt 1994). C. H. dimorphum (after Chitwood 1935). D) H. hamatus (after Bowie 1985). E) H. impalutiensis (after Malysheva et al. 2012). F) H. insularis (after Kloss 1965). G) H. initiensis (after Malysheva & Sumaya 2017). H) H. juli (after Cobb 1898). I) H. orthopori (after Adamson 1987). J) H. ortonwilliamsi (after Hunt 1994). K) H. sutherlandi (after Hunt 1994). L) H. taybaci (after Malysheva & Spiridonov 2010). M) H. taynguyeni (after Malysheva & Spiridonov 2010).	FIGURE 11. Worldwide Heth spp., continuous collar with cervical shield (Group 2). A) H. baudini (after Malysheva & Cribb 2012). B) H. costata (after Hunt 1994). C. H. dimorphum (after Chitwood 1935). D) H. hamatus (after Bowie 1985). E) H. impalutiensis (after Malysheva et al. 2012). F) H. insularis (after Kloss 1965). G) H. initiensis (after Malysheva & Sumaya 2017). H) H. juli (after Cobb 1898). I) H. orthopori (after Adamson 1987). J) H. ortonwilliamsi (after Hunt 1994). K) H. sutherlandi (after Hunt 1994). L) H. taybaci (after Malysheva & Spiridonov 2010). M) H. taynguyeni (after Malysheva & Spiridonov 2010).	2020-10-20	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.		Zenodo	biologists	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.			
8569BD6CFF85127A31A91E3FFA5EFD5D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4416693/files/figure.png	https://doi.org/10.5281/zenodo.4416693	FIGURE 12. Worldwide Heth spp. Group 2 (continued). A) H. tonkinensis (after Malysheva & Spiridonov 2010). B) H. tuxtlensis (after Mejía-Madrid 2014). C) H. vietnamensis (after Malysheva & Spiridonov 2010). D) H. xaniophora (after Hunt 1994), en face, lateral and dorso-ventral views.	FIGURE 12. Worldwide Heth spp. Group 2 (continued). A) H. tonkinensis (after Malysheva & Spiridonov 2010). B) H. tuxtlensis (after Mejía-Madrid 2014). C) H. vietnamensis (after Malysheva & Spiridonov 2010). D) H. xaniophora (after Hunt 1994), en face, lateral and dorso-ventral views.	2020-10-20	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.		Zenodo	biologists	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.			
8569BD6CFF85127A31A91E3FFA5EFD5D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4416665/files/figure.png	https://doi.org/10.5281/zenodo.4416665	FIGURE 2. Heth pivari n. sp., female. A) Habitus, sublateral view, right side. B) Anterior end, sublateral view. C) Part of anterior region, lateral view showing neck retractor muscles; rp, ramified projection of the stoma entrance lumen (Spiridonov & Yushin 2000). D) Collar, lateral view; box shows dimensions of shield. E) Oviduct-uterus region with migrating sperm. Scale bars: A, 230 µm; B, C: 25 µm; D, 40 µm; E, 115 µm.	FIGURE 2. Heth pivari n. sp., female. A) Habitus, sublateral view, right side. B) Anterior end, sublateral view. C) Part of anterior region, lateral view showing neck retractor muscles; rp, ramified projection of the stoma entrance lumen (Spiridonov & Yushin 2000). D) Collar, lateral view; box shows dimensions of shield. E) Oviduct-uterus region with migrating sperm. Scale bars: A, 230 µm; B, C: 25 µm; D, 40 µm; E, 115 µm.	2020-10-20	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.		Zenodo	biologists	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.			
8569BD6CFF85127A31A91E3FFA5EFD5D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4416676/files/figure.png	https://doi.org/10.5281/zenodo.4416676	FIGURE 6. DIC micrographs of Heth pivari n. sp., female and male structures. A) Lateral ornamentation in anterior region of female. B) Male, lateral somal extensions of posterior arcade surrounding anterior esophagus. C) Female, secretory-excretory pore (arrow). D. Female, somatic papillae anterior to vulva. E) Fusiform to comet-shaped sperm in male. All scale bars 20 µm.	FIGURE 6. DIC micrographs of Heth pivari n. sp., female and male structures. A) Lateral ornamentation in anterior region of female. B) Male, lateral somal extensions of posterior arcade surrounding anterior esophagus. C) Female, secretory-excretory pore (arrow). D. Female, somatic papillae anterior to vulva. E) Fusiform to comet-shaped sperm in male. All scale bars 20 µm.	2020-10-20	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.		Zenodo	biologists	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.			
8569BD6CFF87127431A91F37FA5FFEA5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4416697/files/figure.png	https://doi.org/10.5281/zenodo.4416697	FIGURE 14. Worldwide Heth spp. Group 3 (continued). A) H. sinediscus (after Kloss 1965). B) H. spinalatum (after Kloss 1965). C) H. spoliatus (after García & Coy 1995). D) H. travassosi (Dollfus 1952). E) H. travfilhoi (after Adamson 1983). F) H. tuzetae (after Adamson 1983).	FIGURE 14. Worldwide Heth spp. Group 3 (continued). A) H. sinediscus (after Kloss 1965). B) H. spinalatum (after Kloss 1965). C) H. spoliatus (after García & Coy 1995). D) H. travassosi (Dollfus 1952). E) H. travfilhoi (after Adamson 1983). F) H. tuzetae (after Adamson 1983).	2020-10-20	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.		Zenodo	biologists	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.			
8569BD6CFF87127431A91F37FA5FFEA5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4416695/files/figure.png	https://doi.org/10.5281/zenodo.4416695	FIGURE 13. Worldwide Heth spp., discontinuous collar, no shield (Group 3). A) H. artigasi (after Dollfus 1952). B) H. artigasi (after Adamson 1983). C) H. baracoa (after Spiridonov 1989). D) H. duvidosum (after Artigas 1929). E) H. imias (after Spridonov 1989). F) H. macrocephala (after Kloss 1965). G) H. multiplus (after Kloss 1965). H) H. parartigasi (after Adamson 1985).	FIGURE 13. Worldwide Heth spp., discontinuous collar, no shield (Group 3). A) H. artigasi (after Dollfus 1952). B) H. artigasi (after Adamson 1983). C) H. baracoa (after Spiridonov 1989). D) H. duvidosum (after Artigas 1929). E) H. imias (after Spridonov 1989). F) H. macrocephala (after Kloss 1965). G) H. multiplus (after Kloss 1965). H) H. parartigasi (after Adamson 1985).	2020-10-20	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.		Zenodo	biologists	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.			
8569BD6CFF88127431A91EDBFA5EF8E4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/4416699/files/figure.png	https://doi.org/10.5281/zenodo.4416699	FIGURE 15. Worldwide Heth spp. not conforming to any other group (Group 4). A) H. adolphi (after Sanchez-Velazquez 1979). B) H. josephinae (after Sanchez-Velazquez 1979). C) H. lamothei (after Sanchez-Velazquez 1979). D) H. poeyi (after Otero et al. 1993).	FIGURE 15. Worldwide Heth spp. not conforming to any other group (Group 4). A) H. adolphi (after Sanchez-Velazquez 1979). B) H. josephinae (after Sanchez-Velazquez 1979). C) H. lamothei (after Sanchez-Velazquez 1979). D) H. poeyi (after Otero et al. 1993).	2020-10-20	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.		Zenodo	biologists	Phillips, Gary;Moulton, John K.;Bernard, Ernest C.			
