taxonID	type	description	language	source
0001EEF23D1A56B58F16857DCEC2258D.taxon	etymology	Etymology. The specific name “ nujiang ” refers to the location of type specimens, the area around the Nujiang River. As common name we suggest “ Nujiang green pit-viper ” in English and “ Nù Jiāng Zhú Yè Qīng (怒江竹叶青) ” in Chinese.	en	Liang, Ya-Ting, Ding, Li, Vogel, Gernot, Chen, Ze-Ning, Wu, Zheng-Jun (2025): A new species of the genus Trimeresurus Lacépède, 1804 (Reptilia, Squamata, Viperidae) from Southwestern China. Zoosystematics and Evolution 101 (4): 1653-1668, DOI: 10.3897/zse.101.162424
0001EEF23D1A56B58F16857DCEC2258D.taxon	diagnosis	Diagnosis. A species of the genus Trimeresurus has a combination of the following characters: (1) Dorsal body olive drab or grass green, without bands or markings; interstitial skin greyish-black; ventral body yellow green. (2) Tail mostly reddish brown with dark brown tail end. (3) Dorsum of the head has the same color as the body, the upper labials are light green. (4) Moderate body size, with the maximum total length exceeding 804 mm. (5) White (above) and dark red (below) ventrolateral stripe present on the first row of dorsal scales, and the ventrolateral stripe continuous on the tail in males; faint white ventrolateral stripe in females. (6) Postocular streak absent in both sexes. (7) Iris golden-yellow in both sexes in life. (8) First supralabial completely separated from nasal scale. (9) Head scales not keeled; dorsal scale row 19 (rarely 21 or 22) – 19 – 15 (rarely 13) (N = 11), feebly keeled except the outermost rows. (10) Internasals not in contact, most usually separated by 1 – 2 scales. (11) Supraoculars large but elongate, separated by 10 – 11 smooth cephalic scales. (12) Ventral scales 164 – 173 in males (N = 6), 165 – 168 in females (N = 5). (13) Subcaudal scales 59 – 68 in males (N = 4), 57 – 60 in females (N = 5). (14) Total number of VEN + SC 226 – 241 in males (N = 4), 222 – 226 in females (N = 5). (15) Tail relatively short, and tail to total length ratio (TaL / TL) 0.147 – 0.163 in males (N = 4), 0.144 – 0.152 in females (N = 5). (16) Hemipenes short and spinose, bilobed at 6 th / 7 th plate when unextruded, tips reaching SC 10, sulcus spermaticus shallow, visible, divides at the base of the organ.	en	Liang, Ya-Ting, Ding, Li, Vogel, Gernot, Chen, Ze-Ning, Wu, Zheng-Jun (2025): A new species of the genus Trimeresurus Lacépède, 1804 (Reptilia, Squamata, Viperidae) from Southwestern China. Zoosystematics and Evolution 101 (4): 1653-1668, DOI: 10.3897/zse.101.162424
0001EEF23D1A56B58F16857DCEC2258D.taxon	description	Description of holotype male DL R 353. (Figs 3, 4). Morphology. Body cylindrical, long and thin, SVL 694 mm; TaL 110 + mm, the tail a bit missing; TL 804 + mm, ratio of tail length to total length (TaL / TL) over 13.7 %. Head triangular, elongate, clearly distinct from the neck, head length 35.2 mm (HL / SVL 0.051); head width 22.6 mm (HW / HL 0.642). Distance between tip of snout and anterior border of eye 11.1 mm on both sides. Eyes large, eye diameter 4.4 mm (ED / HL 0.125); pupil vertically elliptic. Body scalation. Dorsal scales in 21 – 19 – 13 rows; dorsal scales rhomboid, feebly keeled except the first row of scales, which are smooth; 169 ventral scales (plus 1 preventrals); 41 + / 42 + subcaudals, paired; anal shield single and entire. Head scalation. Rostral slightly visible when viewed from above, subtriangular; a large pair of enlarged and subrectangular internasals bordered by one scale; nostril completely enclosed in nasal scale; nasal scale complete, completely separated from 1 st supralabial, with only a trace of a suture, sub-pentagonal; three elongated preoculars on each side of the head, two lower preoculars and 2 nd supralabial encompass the loreal pit; one elongate and narrow supraocular; one long, thin, crescent-like subocular scale; 2 nd supralabials completely contact the anterior margin of the pit; two small scales on the left side and one small scale on the right side between the nasal and the foveal; 10 small and irregular cephalic scales between the supraoculars; temporal and occipital scales feebly keeled. 9 / 9 supralabials; 1 st supralabial triangular, small; 2 nd as high as the 1 st supralabial, nearly of the same width throughout; 3 rd supralabial largest, wider than high, and in contact with the subocular; 4 th and 5 th supralabials separated from the subocular by a single row of smooth scales; 6 th supralabials separated from subocular by 2 / 2 scales. 12 / 12 infralabials; the first pair of infralabials are in contact with each other behind the mental; the first three pairs of infralabials are in contact with the anterior chin shields. Coloration in preservation (Figs 3, 4). Description based on holotype DL R 353 fixed in formalin and later stored in 75 % ethanol for approximately three years. Dorsal head and body olive drab, ventral color faded to whitish green. Lateral head olive above lower margin of eye, upper labials light green with bluish tint. The ventral surfaces of the head are creamy white with a bluish tint. Ventral surface of tail whitish green anteriorly, becoming paler bluish-green at the middle and end of tail. Ventrolateral stripe present, with white (above) and dark brown (below) on first dorsal scale rows. Coloration in life (Fig. 5). Dorsal body generally olive drab in males and grass green in females, with greyish-black interstitial skin. Lateral head jungle-green above lower margin of eyes, and light green below, without postocular stripes in both genders. Lateral body deep green above and gradually become light green below. A white (above) and dark red (below) ventrolateral stripe present in males, a thin and light white ventrolateral stripe present in females, which ventrolateral stripe present on each scale of the first dorsal scale. Venter uniform yellowish-green. Tail mostly reddish-brown, dark brown at the end of the tail. Iris golden-yellow in both males and females, pupils edged with lighter color. Hemipenis. Description based on two specimens, namely the holotype DL R 353 and paratype DL 19. The hemipenis is relatively short, papilloma, reaching only 10 th subcaudal, and bifurcates at 6 th – 7 th subcaudal when unextruded. Sulcus spermaticus shallow, visible, divides at the base of the organ. Intraspecific morphological variation. The ten paratypes agree with the description of the holotype in most aspects except for the following difference: (1) ventrolateral stripe always present, males have obvious white or bicolored stripes, while females have thin and light white stripes; (2) dorsal scales in 19 (rarely 21 or 22) – 19 – 15 (rarely 13) rows; (3) ventral scales 164 – 173 in males (N = 6), while 165 – 168 in females (N = 5); (4) subcaudal scales 59 – 68 in males (N = 4), while 57 – 60 in females (N = 5); (5) total number of VEN + SC 226 – 241 in males (N = 4), while 222 – 226 in females (N = 5). (6) TaL / TL 0.147 – 0.163 in males (N = 4), 0.144 – 0.152 in females (N = 5); supralabial scales vary between 9 or 10 (N = 11); (7) infralabial scales vary from 11 to 13 (N = 11); (8) cephalic scales vary from 10 to 11 (N = 11) (See details in Table 2).	en	Liang, Ya-Ting, Ding, Li, Vogel, Gernot, Chen, Ze-Ning, Wu, Zheng-Jun (2025): A new species of the genus Trimeresurus Lacépède, 1804 (Reptilia, Squamata, Viperidae) from Southwestern China. Zoosystematics and Evolution 101 (4): 1653-1668, DOI: 10.3897/zse.101.162424
0001EEF23D1A56B58F16857DCEC2258D.taxon	distribution	Distribution and habitat. Currently, Trimeresurus nujiang sp. nov. is known only from the type locality in Gongshan County, Nujiang Lisu Autonomous Prefecture, Yunnan Province, China (Fig. 6). The collected individuals were encountered at night, perched on tree branches near the Grand Canyon of the Nujiang. It is currently known to be found in subtropical broad-leaved evergreen forests at elevations reaching approximately 1500 – 1700 m in Gongshan, Yunnan, China (Fig. 7). Gongshan County is located in the Nujiang Lisu Autonomous Prefecture in the northwestern part of Yunnan Province, bordering Myanmar to the west. Nujiang River originates on the Tibetan Plateau and flows through the Nujiang Lisu Autonomous Prefecture in Yunnan Province, where it exits and flows into Myanmar. It is assumed that the new species may also be distributed in Myanmar.	en	Liang, Ya-Ting, Ding, Li, Vogel, Gernot, Chen, Ze-Ning, Wu, Zheng-Jun (2025): A new species of the genus Trimeresurus Lacépède, 1804 (Reptilia, Squamata, Viperidae) from Southwestern China. Zoosystematics and Evolution 101 (4): 1653-1668, DOI: 10.3897/zse.101.162424
