identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
842B6D7DFFFDFFD8FF679C005399B9EC.text	842B6D7DFFFDFFD8FF679C005399B9EC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Genyocerus Motschulsky	<div><p>Genyocerus Motschulsky</p><p>Genyocerus Motschulsky, 1858: 68 . Type species, by monotypy, G. albipennis Motschulsky, 1858 .</p><p>Diacavus Schedl, 1939: 363 . Type species, by original designation, Diapus frontalis Strohmeyer, 1914 . Synonymy: Wood 1969: 118.</p><p>Length 1.8–5.4 mm. 3.0–4.2 times as long as wide. Colour reddish brown to black, underside and appendages usually paler, elytral disc sometimes partly testaceous. Vestiture sparse except on female frons. Head with a very short rostrum, frons angularly separated from vertex, flattened, sometimes with a longitudinal keel or keels in the male; young females with one to several brushes or pencils of long hairs on frons and sometimes also on epistoma, often largely concealing frontal surface but lost later during gallery construction. Eyes oval or egg–shaped, not extending onto ventral surface of head. Antennae inserted close to lower margin of eyes in males, and in females of some species, near upper margin of eyes in females of species with long hair brushes on upper part of frons. Antennal scape short, club–shaped in male, slightly longer and scarcely widened distally in female, funicle four–segmented, club oval, flattened, anterior and posterior surfaces uniformly pubescent to base, a narrow testaceous strip lacking sensory setae along the posterior margin. Maxillae with separate galea and lacinia. Submentum separated from margin of oral opening by a groove on each side. Pronotum usually longer than wide, sides shallowly constricted in anterior half, widest about one–third from base, femoral grooves without carinae anteriorly or posteriorly, disc weakly convex, usually with one to many large mycangial pores on each side of median line in basal third. Mesonotum without a median carina, scutellum slightly medially impressed, flush with elytral surface posteriorly. Elytra elongate, horizontal, disc weakly striate–punctate, declivity very short or absent, male elytra with spines or teeth on some interstriae at discal apex, female elytral apex simply rounded. Metasternal–metepisternal depression of male unarmed. Abdomen with fifth ventrite concave and subvertical in male, convex in female, apical margin of fourth ventrite sometimes extended as a hyaline membrane of fused hairs in male. Procoxae very large, longer than femur, widely separated by a trapezoidal intercoxal plate, mesocoxae globular, less widely separated, metacoxae transverse, extending more than halfway across first abdominal ventrite, narrowly separated medially. Protibiae narrow, granulate or with rasp–like teeth in both sexes, the apex with a single recurved spine.</p><p>The genus is distinguished from other platypodine genera except Diapus by the very large, widely separated procoxae. The species of Genyocerus can be distinguished from those of Diapus by their wider, more obvious scutellum, which is flush with the elytral surface posteriorly. The mycangial pores of Genyocerus are usually larger than those of Diapus and never fused to form a transverse or crescentic bar on each side of the midline. The antennal club of Genyocerus never has a median, testaceous strip lacking sensillae on the outer surface. The males of Diapus never possess a membranous extension of the apical margin of the fourth abdominal ventrite. The females of Genyocerus never possess dehiscent mandibular appendages. It may be noted that the larvae, so far as known (Browne 1972), show little evidence of generic distinctions. The two genera appear closely related, and more detailed studies, including DNA data and cladistic analyses are needed to test this phylogenetic hypothesis.</p><p>Sexual dimorphism</p><p>The males of Genyocerus are characterised by the presence of teeth or spines at the apex of the elytral disc; in the females, the elytral apices are subtruncate or transversely rounded. In the males of some species, the apical margin of the fourth ventrite extends as a hyaline membranous structure projecting over the basal part of the fifth ventrite (Fig. 12). This probably helps to make the concave fifth ventrite a more effective 'shovel' as the male collects and expels frass from the gallery system. The fifth ventrite of the female is essentially convex, and there is never an extension of the fourth ventrite. The young females bear brushes of hairs on the frons. The arrangement of these is species-specific, and they are probably used in courtship (Browne 1961a, Roberts 1993) but lost after mating when the female takes over the extension of the gallery system from the male. In the male, the antennae are always inserted below the middle of the frons. In the females, the position of the antennae is associated with the presence or absence of large hair brushes on the upper part of the frons. When these are absent, the antennae are inserted in the same position as in the male (Fig. 7); when present, the antennae are inserted on the upper part of the frons at or above the level of the upper margin of the eyes (e.g. Figs 17, 34). The number of mycangial pores on the pronotum is variable but is almost always greater in the female than in the male (see below).</p><p>Arrangement of teeth on male elytral apex</p><p>Schedl (e.g. 1966) states that the first and second major teeth at the apex of the male elytra are formed from extensions of interstriae 1 and 3, whereas Browne (1962) writes that they are formed from extensions of interstriae 2 and 4. In fact, the first and second teeth are borne on interstriae 3 and 5. The interstriae in Genyocerus are often impunctate and can only be distinguished by the lines of strial punctures between them. Even the arrangement of the striae on the elytra is often difficult to determine in the smaller species, because the punctures are often small and obsolescent, but it can be seen in larger species such as G. biporus (Fig. 1). The first interstriae are normally developed along the suture. The apex may form a small secondary tooth at the inner side of the larger primary tooth on interstriae 3. Striae 1 are very short and usually terminate not more than one-third along the elytra. Thus the second interstriae are very short and taper to a point about onethird from the base of the elytra. Interstriae 3 and 5 are broadened distally and extend over the declivity as teeth. The teeth are usually flattened and blunt or with bifid tips (Figs 11, 15), but sometimes are more slen- der, tapering and spine-like (Figs 5, 30). Interstriae 4 and 6 are narrowed posteriorly and may not reach the end of the elytra. On the posterolateral part of the elytra, striae 6 - 8 end about 1/3 from the elytral apex, and there is a smooth, shining, punctureless area extending across interstriae 7 - 8 and sometimes interstriae 9 (Fig. 9). One or both interstriae 7 and 8 and interstriae 9 may bear apical teeth, or interstriae 7 - 9 may be fused apically to form an acute edge bearing several small teeth (Fig. 27). In a few species (e.g. G.tenellus, G. exilis), a tooth is also present on the elytral declivity.</p><p>Intraspecific variation</p><p>It has become clear during the course of this study that the authors who described the majority of species in the genus Genyocerus underestimated the intraspecific variation in the number of mycangial pores on the pronotum. Both Schedl and Browne in many of their papers (see Wood &amp; Bright 1992 for references) noted that the number of pores differs between the sexes, with the female usually having a greater number (although the reverse is the case in Genyocerus puer (Schedl), Roberts 1993). In addition, the number of pores can differ between the two sides of the same individual, and there can also be considerable variation between individuals from the same collection site, and from different sites, independently of the differences between males and females. Similarly, in the males of some species, the shape and size of the teeth at the end of the elytral disc can be quite variable intraspecifically. Such variations have sometimes been described as separate species. However, the gradual accumulation of more specimens from a wider range of collection sites has indicated that intermediates exist. The resulting new synonyms are reported below.</p><p>Biology</p><p>The species of Genyocerus normally breed only in host trees in the family Dipterocarpaceae . The only partial exception to this is G. pendleburyi, which may occasionally attack trees in the family Fagaceae in areas in which dipterocarps are scarce (Browne 1977). A few other records from non-dipterocarp trees may or may not involve actual breeding. By contrast, species of Diapus, like most platypodines, are generally polyphagous, attacking any trees in which the symbiotic ambrosia fungi will grow. Genyocerus species usually attack dying or felled trees, but Browne (1961a) noted that attacks may occur on stressed trees after drought.</p><p>The general features of the life history are given by Beeson (1961) and Browne (1961a). The gallery system is started by the male, but only a short vertical gallery is made. It is probable that the female is attracted by a male-produced pheromone (Beaver 2000b), although no studies have been made on Genyocerus or Diapus . After mating, the female continues to extend the gallery into the wood. The male remains in the entrance, keeping out potential predators and parasites and removing the frass that accumulates from the boring activities of the female. The gallery system branches mainly in one transverse plane and may extend deeply into the tree. The mycangia on the pronotum transport an ambrosial fungus (or fungi) that is released into the gallery, grows in the wood and provides the only food for both adults and larvae, which feed on the hyphae growing on the gallery walls. The ambrosia fungi associated with Genyocerus and Diapus have not been investigated. Mature larvae construct pupal cells above and below the gallery at the ends of the branches. The adults of the new generation emerge through the parental entrance hole.</p></div>	https://treatment.plazi.org/id/842B6D7DFFFDFFD8FF679C005399B9EC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beaver, R. A.;L. - Y	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF4FFD3FF679D185242B982.text	842B6D7DFFF4FFD3FF679D185242B982.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Genyocerus albipennis Motschulsky 1858	<div><p>Genyocerus albipennis Motschulsky</p><p>Genyocerus albipennis Motschulsky, 1858: 68 .</p><p>Diacavus irregularis Browne, 1970: 582 . Synonymy: Wood, 1981: 121.</p><p>Taxonomy: The type material of G. albipennis was long thought to be lost, but Wood (1969) found a single female in the Motschulsky collection in Moscow and clarified the status of the genus and the species. Later, Wood (1981) placed Diacavus irregularis in synonymy. We have examined the type series of Diacavus irregularis (BMNH) . The male is closely related to, but distinct from, G. diaphanus (Schedl) (see key to males above), but we are unable to distinguish the female from G. diaphanus (see key to females above). The species was referred to by Beeson (1961) as Diacavus zeylanicus, but this name is a nomen nudum (Browne 1977).</p><p>Distribution: Sri Lanka only. Records from other countries result from confusion with Diapus albipennis Strohmeyer, which is in fact a synonym of G. pendleburyi (Schedl) (see below) and not a synonym of G. albipennis Motschulsky, as indicated by Schedl (1972a).</p><p>Biology: The only identified host is Dipterocarpus zeylanicus (Dipterocarpaceae) (Browne 1977).</p></div>	https://treatment.plazi.org/id/842B6D7DFFF4FFD3FF679D185242B982	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beaver, R. A.;L. - Y	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF4FFD3FF679BD85233BADA.text	842B6D7DFFF4FFD3FF679BD85233BADA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Genyocerus biporus (Schedl)	<div><p>Genyocerus biporus (Schedl)</p><p>(Figs 1–4)</p><p>Diacavus biporus Schedl, 1942: 217 .</p><p>Genyocerus biporus (Schedl): Browne, 1977: 97.</p><p>Diacavus intermedius Browne, 1968: 134 . n. syn.</p><p>Taxonomy: We have compared the male lectotype of D. biporus (NMW) and a further male determined by Schedl with a long series of both sexes in RAB's collection, which had earlier been compared to the male holotype and male and female paratypes of D. intermedius (BMNH) . Only a single species is represented. Browne (1968) suggested that the two species were distinguishable by the smaller size of G. biporus and differences in the shape of the elytral teeth of the male. However, it seems possible that he had not examined the lectotype of G. biporus, because his description of the elytral teeth of G. biporus as 'of subequal breadth and truncate' is incorrect and apparently based on a misidentified specimen in BMNH. The elytral teeth have the same form in the male types of both species.</p><p>Distribution: The species is distributed from northeastern India (Assam) through Southeast Asia to Indonesia and the Philippines. It has often been intercepted in Japan in timber from the region but is not known to be established there.</p><p>Biology: Recorded only from genera belonging to the family Dipterocarpaceae ( Anisoptera, Dipterocarpus, Dryobalanops, Hopea, Parashorea, Shorea). It usually attacks freshly cut logs (Browne 1961a).</p></div>	https://treatment.plazi.org/id/842B6D7DFFF4FFD3FF679BD85233BADA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beaver, R. A.;L. - Y	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF5FFD2FF679FAD55DFBB64.text	842B6D7DFFF5FFD2FF679FAD55DFBB64.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Genyocerus borneensis (Browne)	<div><p>Genyocerus borneensis (Browne)</p><p>(Figs 5–7)</p><p>Diapus borneensis Browne, 1980b: 500 .</p><p>Genyocerus borneensis (Browne): Beaver, 1998: 185.</p><p>Taxonomy: We have examined the male holotype (BMNH) and a series of specimens collected by S. Adebratt in East Malaysia (Sabah) (ZMLU), which includes both sexes. The species was transferred from Diapus to Genyocerus by Beaver (1998). The female is described here for the first time. It is characterised by the presence of two narrow, forwardly directed, diverging pencils of hairs on the lower part of the frons and by a patch of small mycangial pores in addition to the one or two large pores on each side of the pronotal midline.</p><p>Female: Generally resembling male, 2.7 mm long (excluding frontal hair brushes and part of abdomen extending beyond elytra), about 4.1 times as long as wide. Frons weakly convex above epistoma, shallowly, transversely impressed above antennal insertions, just above epistoma and close to midline on each side, a pencil–like brush of long, pale yellow hairs, directed anteriad and somewhat ventrad, each brush inserted on a slightly raised base, cylindrical near base, tapering towards tip, brushes parallel close to base, then divergent, between their insertions on lower part of frons in midline a small tuft of much shorter, ventrally directed hairs (Fig. 7). Antennae inserted at level of lower margin of eyes, club oval, with a few longer hairs, pubescent to base except for a narrow corneous strip along posterior margin, as in male. Pronotum about 1.3 times as long as wide, slightly more elongate than in male, one or two large mycangial pores on each side of midline close to base, and lateral to these a group of about 25–40 much smaller pores. (In the male, a patch of about 12–15 small pores laterally to the pair of large mycangial pores was not mentioned by Browne (1980b) in his description.) Elytra 1.7–1.8 times as long as pronotum, 2.1–2.2 times longer than wide, carina at base of elytra reduced and restricted to humeral region, not extending to scutellum as in male, a row of long semi– recumbent hairs on interstriae 3 (as in male), lacking spines at apex of elytral disc, elytral apices separately rounded. Fifth ventrite plano–concave, without a distinct median groove, lateral margins indistinct, vestiture a little shorter and less dense than in male, largely confined to marginal areas.</p><p>Female specimens examined: MALAYSIA: Sabah, Sipitang, Mendolong, 26.xi.1987, 27.xi.1987, various dates from 13.iv.–11.v.1988, 19.ii.1989 (S. Adebratt) (16 ♀) (ZMLU, RAB) .</p><p>Distribution: The species was previously known only from specimens intercepted in Japan in timber shipped from East Malaysia (Sabah and Sarawak) (Browne 1980b, Ohno 1990b). The occurrence of the species in Sabah can now be confirmed, and it is recorded for the first time from Brunei Darussalam.</p><p>New Records: BRUNEI, 115 o 7'E, 4 o 34'N, Kuala Belalong FSC, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.11667&amp;materialsCitation.latitude=4.5666666" title="Search Plazi for locations around (long 115.11667/lat 4.5666666)">Dipterocarp forest</a>, site 16–2, 520m alt., Fog 28, 18.iii.1992 (N. Mawdsley) (1 ♂) (BMNH); MALAYSIA: Sabah, Sipitang, Mendolong, 26.xi.1987, 27.xi.1987, various dates from 13.iv.–13.v.1988, 19.ii.1989, 19.iii.1989 (S. Adebratt) (23 ♂, 16 ♀) (ZMLU, RAB) .</p><p>Biology: Recorded from four genera of Dipterocarpaceae ( Anisoptera, Dipterocarpus, Dryobalanops, Shorea) and from ' Sapotaceae sp.' (Ohno 1990b).</p></div>	https://treatment.plazi.org/id/842B6D7DFFF5FFD2FF679FAD55DFBB64	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beaver, R. A.;L. - Y	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF5FFD1FF6799FA5294B88C.text	842B6D7DFFF5FFD1FF6799FA5294B88C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Genyocerus diaphanus (Schedl)	<div><p>Genyocerus diaphanus (Schedl)</p><p>(Fig. 8)</p><p>Diacavus diaphanus Schedl, 1939: 384 .</p><p>Genyocerus diaphanus (Schedl): Wood 1969: 118.</p><p>Diacavus assamensis Browne 1964: 760 . Synonymy: Schedl 1972b: 159.</p><p>Diacavus abdominalis Schedl, 1942: 218 . n. syn. (male only)</p><p>Diacavus philippinensis Schedl, 1942: 216 . n. syn.</p><p>Diacavus sexporus Schedl, 1942: 217 . n. syn.</p><p>Taxonomy: G. diaphanus was described by Schedl from the female only. The male was described by Browne (1962) from a specimen taken in association with the female. Schedl (1962) independently also described the male. We have compared the male holotype of D. abdominalis (NMW), the male holotype and a female described by Schedl (1966) of D. philippinensis (NMW) and the male lectotype of D. sexporus (NMW) with a series of males and females in RAB's collection from East Malaysia (Sabah) and Brunei, which had earlier been compared with a paralectotype female and the male described by Browne (1962) of D. diaphanus (BMNH) . Only a single, somewhat variable species is represented, the males of which vary from 2.5–3.2 mm in length. The characters used by Browne (1964) and Schedl (1942) to distinguish the species intergrade in longer series. The number of mycangial pores present on the pronotum of the male varies from none to 2 or 3 on each side. In the female, the number of mycangial pores varies from 4–10 on each side of the midline. The two sides usually differ by no more than a single pore, but there is much greater variation between individuals. There is also variation in the shape of the male elytral teeth. The inner four teeth on each side may be parallelsided or tapering, and bifid to almost truncate at the tip. The fifth, outer tooth varies from acutely pointed to obliquely truncate. All females that we have seen, with the exception of the supposed female of D. abdominalis, have the same type of frontal ornamentation and are essentially indistinguishable. Schedl (1942) noted that the single female of D. abdominalis, badly damaged and collected at a locality over 1800 km from that of the male holotype, 'könnte ... das zugehörige Geschlecht sein' and labelled this specimen as the allotype of the species. However, the frontal ornamentation is not of the type found in G. diaphanus, and Schedl's 'allotype' is in fact a female of G. pendleburyi .</p><p>Distribution: G. diaphanus is widespread, occurring from India through Southeast Asia to Borneo and the Philippines. It has frequently been intercepted in dipterocarp timber imported to Japan (e.g. Schedl 1966, Ohno 1990b) and Korea (Choo &amp; Woo 1983). It is newly recorded here from northern Thailand.</p><p>New Records: THAILAND, Chiang Mai, C[hiang] M[ai].Univ., 300m, ex EtOH trap, various dates from 10.i.2005 – 4.iv.2005, (W. Puranasakul) (1 ♂, 3 ♀) (CMU, RAB) .</p><p>Biology: The species, like its congeners, breeds primarily in trees of the family Dipterocarpaceae, and has been recorded from the following genera ( Anisoptera, Balanocarpus, Dipterocarpus, Hopea, Parashorea, Pentacme, Shorea, Upuna, Vatica). There are a few records from trees of other families: Gonystylus (Thymeleaceae) (Browne 1961a), Gluta or Melanorrhoea, Mangifera (Anacardiaceae), Durio (Bombacaceae), Palaquium (Sapotaceae), Pterocymbium (Sterculiaceae) (Ohno 1990b), but these involve small numbers of specimens. Ohno (1990b, as Genyocerus abdominalis) records a total of 9851 (6938 ♂, 2913 ♀) specimens bred from imported logs of seven genera of Dipterocarpaceae, but only 24 (21 ♂, 3 ♀) obtained from imported logs of six non-dipterocarp genera. This suggests that breeding was very rare, if it occurred at all, in the latter.</p></div>	https://treatment.plazi.org/id/842B6D7DFFF5FFD1FF6799FA5294B88C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beaver, R. A.;L. - Y	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF6FFD0FF679AEF5461BCCA.text	842B6D7DFFF6FFD0FF679AEF5461BCCA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Genyocerus exilis (Schedl 1942)	<div><p>Genyocerus exilis (Schedl)</p><p>(Figs 9–10)</p><p>Diacavus exilis Schedl, 1942: 218 (male); Browne, 1962: 219 (female).</p><p>Genyocerus exilis (Schedl): Browne, 1980a: 371.</p><p>Taxonomy: We have examined the male and female specimens taken in association by F. G. Browne and used by him to describe the female (BMNH) and have compared them with specimens of both sexes collected in Brunei (BMNH) and Sabah (ZMLU, RAB). The males are characterised by the presence of a large mycangial pore close to the midline on each side and lateral to these a group of 5–13 smaller pores. In the females, there are usually two large pores in addition to the group of smaller pores. In both sexes, occasional individuals lack the large pores.</p><p>Distribution: East Malaysia (Sarawak), Indonesia (Maluku, Bacan I.), West Malaysia. It is newly recorded here from Brunei Darussalam and Sabah. It has been intercepted in Japan in timber from Borneo and Maluku.</p><p>New Records: BRUNEI: 4 o 34'N, 115 o 7'E, Kuala Belalong FSC, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.11667&amp;materialsCitation.latitude=4.5666666" title="Search Plazi for locations around (long 115.11667/lat 4.5666666)">Dipterocarp forest</a>, 220 – 275 m. alt, fogging, malaise trap, flight intercept trap, various dates from 16.vi.1991 – 6.viii.1991, 8.ii.1992, 10.iii.1992 (N. Mawdsley) (8 ♂, 10 ♀) (BMNH); MALAYSIA: Sabah, Sipitang, Mendolong, 8.xii.1987, various dates from 10.iv.1988 – 13.v.1988, 10 – 31.iii.1989 (S. Adebratt) (20 ♂, 32 ♀) (ZMLU, RAB); Sabah, Poring Spring, lower montane, mixed dipterocarp forest,&gt; 650m. , fogging Aporusa sp., 22.iii.1993 (A. Floren) (1 ♂) (BZUW) .</p><p>Biology: Recorded from the following genera of Dipterocarpaceae: Anisoptera, Dipterocarpus, Dryobalanops, Shorea, Upuna, Vatica (Browne 1961a, Ohno 1990b)..</p></div>	https://treatment.plazi.org/id/842B6D7DFFF6FFD0FF679AEF5461BCCA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beaver, R. A.;L. - Y	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF7FFD7FF679D10520CBCC9.text	842B6D7DFFF7FFD7FF679D10520CBCC9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Genyocerus frontalis (Strohmeyer)	<div><p>Genyocerus frontalis (Strohmeyer)</p><p>(Figs 11–14)</p><p>Diapus frontalis Strohmeyer, 1914b: 5 .</p><p>Diacavus frontalis (Strohmeyer): Schedl, 1939: 364.</p><p>Genyocerus frontalis (Strohmeyer): Wood, 1969: 118.</p><p>Diacavus haticollis Browne, 1962: 217 .</p><p>Diacavus laticollis Browne, 1964: 753 (justified emendation). n. syn.</p><p>Taxonomy: Diapus frontalis was described from a single male collected by G. Doria in Borneo (Sarawak) in 1865-66 and deposited in the Stettin Museum. Wood &amp; Bright (1992) state that the type is presumed lost; however, Wegrzynowicz &amp; Mokrycki (1996) list a 'syntype’ in IZW. We have examined this specimen. There can be no doubt that it is the holotype of the species. It was evidently seen by Chapuis, who indicates on one of the eight labels below the specimen that it is a new species, but did not describe it. We have compared the holotype to a series of specimens from East Malaysia (Sabah) (ZMLU), which had earlier been compared to the holotype male of Diacavus laticollis Browne from West Malaysia (BMNH). Only a single species is represented. The previously unknown female is described below.</p><p>Female: Generally resembling male, 3.7–3.8 mm long (excluding frontal hair brushes and part of abdomen extending beyond elytra), 2.9–3.0 times as long as wide. Frons with antennae inserted on upper part of frons close to upper margin of eyes, brushes of long, golden–yellow hairs present in young females both above epistoma and on upper part of frons (Figs 13–14), a large teardrop–shaped, shallow, shining, impunctate impression on each side, extending from vertex to lower quarter of frons, the impressions separated by a narrow, median carina, upper part of frons lateral to impressions flat, bearing on each side a large brush of hairs, extending dorsally well above antennal insertion and nearly meeting medially just below vertex, the majority of hairs directed forward, downcurved at tips, each brush extending ventrally in a narrow band close to inner margin of eye nearly to its lower margin, the lowest group of hairs longer and incurved towards midline; epistomal brushes divided into two parts on each side, the long hairs upcurved below those of the upper brushes, almost reaching vertex, lowest part of frons below impressions finely punctured, the punctures with much shorter erect hairs, normally concealed below epistomal brushes. Pronotum 1.05–1.1 times longer than wide, similar to that of male but with 9–12 large mycangial pores on each side of midline close to base (Fig. 13), rather than 0–2 as in male. Elytra wider than and about 1.6 times as long as pronotum, 1.5 times longer than wide, carina on base of elytra reduced and more rounded, striae less strongly impressed, becoming obsolete in posterior third of elytra, interstriae with sparse, semi–recumbent hairs, elytral apices lacking teeth, separately rounded, with more abundant short hairs.</p><p>Female specimens examined: MALAYSIA: Sabah, Sipitang, Mendolong, 28.xi.1987, 6.v.1988, 13.v.1988 (S. Adebratt) (5 ♀) (ZMLU, RAB) .</p><p>Distribution: The species is recorded only from East Malaysia (Sarawak) and West Malaysia. It is newly recorded here from Brunei Darussalam and Sabah.</p><p>New records: BRUNEI: 4 o 34'N, 115 o 7'E, Kuala Belalong FSC, Dipterocarp forest, site 1, 260m alt., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.11667&amp;materialsCitation.latitude=4.5666666" title="Search Plazi for locations around (long 115.11667/lat 4.5666666)">Fog</a> 8, 9.viii.1991 (N. Mawdsley) (1 ♂) (BMNH) ; Temburong, nr. K. Belalong Field Stud. Centre, 4 o 33'N, 115 o 09'E, 150m, 1.iii.1992 (R. A. Beaver) (3 ♂); as previous except: 200m, ex Shorea sp., 10.iii.1992 (2 ♂) (all RAB) ; [MALAYSIA (Sabah)] North Borneo (SE) , Forest camp, 19 km. N. of Kalabakan, 60m., 13.x.1962 (Y. Hirashima) (1 ♂); as previous except: 21.x.1962 (1 ♂); as previous except: 19.x.1962 (K. J. Kuncheria) (1 ♂) (all BPBM) ; MALAYSIA: Sabah, Sipitang, Mendolong, 28.xi.1987, 6.v.1988, 13.v.1988 (S. Adebratt) (1 ♂, 5 ♀) (ZMLU, RAB) .</p><p>Biology: The only host record is from the dipterocarp tree Shorea leprosula (Browne 1962) .</p></div>	https://treatment.plazi.org/id/842B6D7DFFF7FFD7FF679D10520CBCC9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beaver, R. A.;L. - Y	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF0FFD7FF679D175503B9B6.text	842B6D7DFFF0FFD7FF679D175503B9B6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Genyocerus multiporus (Schedl)	<div><p>Genyocerus multiporus (Schedl)</p><p>(Figs 15–18)</p><p>Diapus multiporus Schedl, 1936b: 67 .</p><p>Diacavus multiporus (Schedl): Schedl, 1939: 364.</p><p>Genyocerus multiporus (Schedl): Wood, 1969: 118.</p><p>Diacavus decemspinatus Schedl, 1942: 216 (as D. 10- spinatus). n. syn.</p><p>Taxonomy: We have compared the female holotype of D. multiporus (NMW) and the male lectotype of D. decemspinatus (NMW) directly with a long series of male and female specimens collected by S. Adebratt in East Malaysia (Sabah) (ZMLU, RAB), in which the sexes can be reliably associated. Schedl (1942) already suggested that the male of D. multiporus might be D. decemspinatus, and this can now be confirmed. The species is closely related to G. papuanus .</p><p>Distribution: The species occurs in East and West Malaysia and in the Philippines and has been imported in timber to Japan. Its occurrence in Vietnam (Schedl 1972a) needs to be substantiated.</p><p>Biology: Recorded hosts are all in the family Dipterocarpaceae ( Anisoptera, Shorea, Vatica) (Browne 1961a, 1985; Ohno et al. 1988).</p></div>	https://treatment.plazi.org/id/842B6D7DFFF0FFD7FF679D175503B9B6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beaver, R. A.;L. - Y	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF0FFD6FF679A2A53BEBC35.text	842B6D7DFFF0FFD6FF679A2A53BEBC35.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Genyocerus papuanus Roberts 1993	<div><p>Genyocerus papuanus Roberts</p><p>(Figs 19 –22)</p><p>Genyocerus papuanus Roberts, 1993: 27 .</p><p>Taxonomy: We have examined the holotype male, allotype female and a female paratype (BPBM) from Papua New Guinea and compared them with a series of males collected in Indonesia: Papua Barat (formerly Irian Jaya) (NKME). The available material suggests that the species occurs in two size classes. The holotype male from Papua New Guinea (BPBM) and nine males (NKME) from Indonesia: Papua Barat are 4.5–4.8 mm long; two further males (NKME) from a second locality in Papua Barat (about 100 km distant from the first, but in a different river basin) are 5.5 and 5.7 mm long. The smaller males have a total of 8–9 mycangial pores; the larger males have 14–17. We are unable to find any other significant differences between the two groups of males. Although the size difference suggests reproductive isolation, it would be premature to describe the larger males as a distinct species, because intermediates may yet be found (as in G.quadriporus (see below)). Roberts (1993) suggests that the male can be distinguished from G.biporus by the hair ornamentation of the lower part of the frons. However, the basic arrangement of hairs in four longitudinal rows is the same in both species. Alternative distinguishing characters are given in the keys above.</p><p>Distribution: Known only from the island of New Guinea (Indonesia: Papua Barat, and Papua New Guinea).</p><p>New Records: INDONESIA, Irian Jaya, Nabire area, road Nabire-Ilaga, km 62, 03 o 30'936''S,</p><p>135 o 41'945''E, 250m NN, LEK, X.1997 (M. Balke) (9 ♂) (NKME, RAB) (all smaller form); Wapoga River, E</p><p>Asori, km 64, Kwadewa camp, 2 o 49'S, 136 o 28'E, 10.i.1999 (A. Weigel) (2 ♂) (NKME, RAB) (larger form). Biology: The only recorded host is Anisoptera sp. (Dipterocarpaceae) (Roberts 1993).</p></div>	https://treatment.plazi.org/id/842B6D7DFFF0FFD6FF679A2A53BEBC35	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beaver, R. A.;L. - Y	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF1FFD6FF679EAA53D7B83B.text	842B6D7DFFF1FFD6FF679EAA53D7B83B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Genyocerus pendleburyi (Schedl)	<div><p>Genyocerus pendleburyi (Schedl)</p><p>Diapus pendleburyi Schedl, 1936a: 18 .</p><p>Genyocerus pendleburyi (Schedl) . Beaver, 1998: 185.</p><p>Diapus albipennis Strohmeyer, 1942: 284 . n. syn.</p><p>Genyocerus strohmeyeri Wood, 1992: 78 . (Replacement name for G. albipennis (Strohmeyer 1942 nec Motschulsky 1858) n. syn.</p><p>Taxonomy: We have compared four male syntypes of D. albipennis Strohmeyer (DEI) with a male of G. pendleburyi in RAB's collection, which had previously been compared with a paralectotype male in BMNH. Strohmeyer (1942) notes that D. albipennis is very close to D. pendleburyi, and the supposed difference in the pronotal pores which he gives to distinguish the species does not exist. Following the transfer of D. albipennis to the genus Genyocerus by Wood (1992), a new name for the species was required because of homonymy with G. albipennis Motschulsky, the type species of the genus. Wood (1992) renamed the species G. strohmeyeri . However, Wood's name must also fall as a synonym of D. pendleburyi . Schedl (1942) briefly described the female of G. pendleburyi but evidently based his description on a specimen in which the brushes of hairs on the frons had been abraded. These brushes are described and figured by Strohmeyer (1942) in his description of the female of D. albipennis . As noted above, the supposed 'allotype' of Diacavus abdominalis is a female of this species.</p><p>Distribution: The species is distributed from Northeastern India (Assam) through Southeast Asia to Borneo and the Philippines. It has been intercepted in timber from Indonesia and the Philippines imported into Japan and Korea.</p><p>Biology: It usually breeds in trees of the family Dipterocarpaceae ( Anisoptera, Dipterocarpus, Dryobalanops, Pentacme, Shorea), but has also been recorded from Fagaceae (Quercus) . Browne (1977) notes that the record from Quercus sp. came from a site at high altitude where dipterocarps were scarce.</p></div>	https://treatment.plazi.org/id/842B6D7DFFF1FFD6FF679EAA53D7B83B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beaver, R. A.;L. - Y	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF1FFD6FF679AA156BFBA7B.text	842B6D7DFFF1FFD6FF679AA156BFBA7B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Genyocerus plumatus (Schedl)	<div><p>Genyocerus plumatus (Schedl)</p><p>Diapus plumatus Schedl, 1936b: 66 .</p><p>Diacavus plumatus (Schedl) . Schedl, 1972a: 268.</p><p>Genyocerus plumatus (Schedl) . Wood &amp; Bright, 1992: 1095.</p><p>Taxonomy: This species was previously known only from the female holotype (NMW), collected in the Philippines. The female is easily distinguished from all other species by the single median brush of hairs, downcurved at the tips, on the lower part of the frons. The male has not been described. It is not one of the two species ( G.shoreae (Browne), G. spinatus (Browne)) of which the females are unknown.</p><p>Distribution: Philippines (Luzon?). A single female is recorded here from East Malaysia.</p><p>New Record: MALAYSIA, Sabah, Sipitang, Mendolong, 17.ii.1989 (S. Adebratt) (1 ♀) (RAB) .</p><p>Biology: Unknown.</p></div>	https://treatment.plazi.org/id/842B6D7DFFF1FFD6FF679AA156BFBA7B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beaver, R. A.;L. - Y	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF2FFD5FF679FAD55BDBEF1.text	842B6D7DFFF2FFD5FF679FAD55BDBEF1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Genyocerus puer (Schedl)	<div><p>Genyocerus puer (Schedl)</p><p>(Figs 23–25)</p><p>Diapus puer Schedl, 1970: 223 .</p><p>Genyocerus puer (Schedl) . Wood &amp; Bright, 1992: 1095.</p><p>Diacavus trispinatus Schedl, 1974: 471 . Synonymy: Roberts, 1993: 28.</p><p>Genyocerus puer trispinatus (Browne, 1985: 197 nec Schedl, 1974). Given subspecific status by Roberts, 1993: 29.</p><p>Taxonomy: Somewhat confusingly, Roberts (1993) divided this species into two subspecies, G. puer puer (Schedl), with the male of Diacavus trispinatus Schedl as synonym, and G. puer trispinatus (Browne) based on females described by Browne (1985) as Genyocerus trispinatus Schedl. Roberts (1993) described for the first time the female of G. puer puer and the male of G. puer trispinatus . Genyocerus puer fergussonus Roberts (Wood &amp; Bright 1992) is a nomen nudum based on specimens labelled by Roberts as Genyocerus fergussonus Roberts but later included by him (Roberts 1993) under the name G. puer trispinatus . We have examined a male paratype of G. puer (NMW) and two females from the series described by Roberts (1993) as G. puer puer (BPBM), the female lectotype of G. puer trispinatus (BMNH) and two males and two females of ' G. fergussonus' (BPBM). Given the extent of variation found in other species of Genyocerus, we believe that only a single species is represented and that recognition of G. puer trispinatus as a subspecies is not valid. The species is unique in the genus in that the male has many more mycangial pores than the female. The male has 30–40 large pores on each side of the midline; some females have 8–10 pores but others have none.</p><p>Distribution: Papua New Guinea only.</p><p>Biology: Specimens have been collected from sticky traps placed on felled trees of Anisoptera sp. and Hopea sp. (Dipterocarpaceae) (Roberts 1993).</p></div>	https://treatment.plazi.org/id/842B6D7DFFF2FFD5FF679FAD55BDBEF1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beaver, R. A.;L. - Y	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF2FFD5FF679B7D56EFBAE9.text	842B6D7DFFF2FFD5FF679B7D56EFBAE9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Genyocerus quadriporus (Schedl)	<div><p>Genyocerus quadriporus (Schedl)</p><p>(Figs 27 –29)</p><p>Diacavus quadriporus Schedl, 1942: 217 .</p><p>Diacavus quadridens Browne, 1962: 218 . n. syn.</p><p>Diacavus compactus Schedl, 1966: 40 . n. syn.</p><p>Diacavus compactus dubiosus Schedl, 1966: 41 . n. syn.</p><p>Diacavus quadrifoveolatus Schedl, 1970: 369 . n. syn.</p><p>Taxonomy: We have compared the male holotype, female allotype and a female paratype of D. quadriporus (NMW), a male paratype of D. quadrifoveolatus (NMW), 2 male and 4 female paratypes of D. compactus (NMW) and the male lectotype and a female labelled as 'allotype’ of D. compactus ssp. dubiosus (NMW) with a series of specimens of both sexes in RAB's collection from Malaysia and Thailand, which had earlier been compared with paratypes of D. quadridens (BMNH) . The species form a series that is continuously variable in size but without specifically distinct characters in either sex. Males vary from 2.5–3.2 mm long, females from 2.7–3.4 mm long. The subspecies dubiosus is simply a small form of the species and not subspecifically distinct. The form described as D. quadrifoveolatus represents the other extreme in size. Schedl (1963) synonymised D. quadridens with D. philippinensis, stating that the species agree in all particulars. This is incorrect. As indicated above, D. philippinensis is conspecific with D. diaphanus, a species that is not very closely related to D. quadridens .</p><p>Distribution: The species is recorded from West Malaysia, East Malaysia (Sabah, Sarawak), Sumatra, Thailand and the Philippines. It has been intercepted in Japan and South Korea in timber exported from the region.</p><p>Biology: All recorded host trees are members of the family Dipterocarpaceae ( Anisoptera, Dipterocarpus, Dryobalanops, Shorea).</p></div>	https://treatment.plazi.org/id/842B6D7DFFF2FFD5FF679B7D56EFBAE9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beaver, R. A.;L. - Y	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF3FFD4FF679FAD57F0BF9C.text	842B6D7DFFF3FFD4FF679FAD57F0BF9C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Genyocerus quadrispinosus (Schedl)	<div><p>Genyocerus quadrispinosus (Schedl)</p><p>(Figs 30–31)</p><p>Diapus quadrispinosus Schedl, 1969: 218 .</p><p>Genyocerus quadrispinosus (Schedl) . Beaver, 2000a: 256.</p><p>Taxonomy: We have examined two male and two female paratypes (NMW). The species is related to G. pendleburyi, G. spinatus and G. borneensis (Beaver 2000a), but can be distinguished by the characters given in the keys.</p><p>Distribution: The species is known only from the Philippines, from which it was intercepted in timber imported to Japan. The record by Schedl (1973) from Nepal must be regarded as dubious and is possibly the result of misidentification of G. pendleburyi, which occurs in the area.</p><p>Biology: The species was originally taken from 'lauan' wood, probably a species of Shorea (Dipterocarpaceae) .</p></div>	https://treatment.plazi.org/id/842B6D7DFFF3FFD4FF679FAD57F0BF9C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beaver, R. A.;L. - Y	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF3FFD4FF679DD7526CB8C4.text	842B6D7DFFF3FFD4FF679DD7526CB8C4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Genyocerus serratus (Schedl 1936)	<div><p>Genyocerus serratus (Schedl)</p><p>(Figs 32–34)</p><p>Diapus serratus Schedl, 1936: 17 .</p><p>Diacavus serratus (Schedl): Schedl, 1939: 364.</p><p>Genyocerus serratus (Schedl): Beaver &amp; Browne, 1979: 618.</p><p>Taxonomy: The species was described in the genus Diapus but transferred to Diacavus by Schedl (1939). It resembles G.talurae but can be distinguished by the characters given in the keys.</p><p>Distribution: East Malaysia (Sarawak), West Malaysia. The species is newly recorded here from Brunei Darussalam and East Malaysia (Sabah).</p><p>New Records: BRUNEI, 115 o 7'E, 4 o 34'N, Kuala Belalong FSC, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.11667&amp;materialsCitation.latitude=4.5666666" title="Search Plazi for locations around (long 115.11667/lat 4.5666666)">Dipterocarp forest</a>, 270m, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=115.11667&amp;materialsCitation.latitude=4.5666666" title="Search Plazi for locations around (long 115.11667/lat 4.5666666)">Aerial</a> malaise, 16.vi.1991 (N. Mawdsley) (1 ♀) ; as previous except: Ground F[light] I[ntercept] T[rap], 17.vi.1991 (1 ♀) ; 220m, Aerial FIT, 16.vii.1991 (1 ♀ ; 210m, Ground malaise, 8.ii.1992 (1 ♂)(all BMNH) . MALAYSIA, Sabah, Sipitang, Mendolong, 1.xii.1987 (S. Adebratt) (9 ♂, 6 ♀) ; as previous except various dates from 29.iv.1988 – 11.v.1988 (10 ♂, 6 ♀) (ZMLU, RAB).</p><p>Biology: The species has been recorded from species of Dipterocarpus, Dryobalanops, Hopea, Parashorea, Shorea and Vatica (all Dipterocarpaceae) (Browne 1961a, b; Beaver &amp; Browne 1979). The gallery system is described by Browne (1961a), who suggests that the life cycle in West Malaysia spans 5–6 weeks.</p></div>	https://treatment.plazi.org/id/842B6D7DFFF3FFD4FF679DD7526CB8C4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beaver, R. A.;L. - Y	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF3FFD4FF67991A531FBA9A.text	842B6D7DFFF3FFD4FF67991A531FBA9A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Genyocerus shoreae Browne 1981	<div><p>Genyocerus shoreae Browne</p><p>Genyocerus shoreae Browne, 1981: 605 .</p><p>Taxonomy: The holotype male (BMNH) has been examined. This is the largest species of Genyocerus, 5.4 mm long, and characterised by the combination of long, acutely pointed elytral teeth and the presence of a hyaline extension of the fourth ventrite. The species is known only from two males imported from the Philippines to Japan, intercepted at Nagoya port .</p><p>Distribution: Philippines (Mindanao).</p><p>Biology: Recorded from Shorea sp. (Dipterocarpaceae) (Browne 1981).</p></div>	https://treatment.plazi.org/id/842B6D7DFFF3FFD4FF67991A531FBA9A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beaver, R. A.;L. - Y	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFECFFCBFF679FAD563DBFFC.text	842B6D7DFFECFFCBFF679FAD563DBFFC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Genyocerus spinatus Browne	<div><p>Genyocerus spinatus Browne</p><p>Diapus spinatus Browne, 1980: 499 .</p><p>Genyocerus spinatus (Browne): Beaver, 1998: 185.</p><p>Taxonomy. We have examined the male holotype (BMNH). The female has not been described. The species is closely related to G. pendleburyi and distinguishable by little more than its greater size and the relative lengths of the spines (see key to males). It is possible that further collections will show that it overlaps in these characters with G. pendleburyi . The female is expected to be very similar to G. pendleburyi and will probably run to the same couplet in the key to females.</p><p>Distribution: Intercepted in Japan in timber imported from 'various shipment localities' in Borneo (Ohno 1990b) and from Indonesia (Sumatera) (Ohno et al. 1986).</p><p>Biology: Recorded from four genera of Dipterocarpaceae ( Anisoptera, Dipterocarpus, Dryobalanops, Shorea), with nearly 3,000 specimens obtained from Dipterocarpus sp., and as single specimens from Dyera sp. (Apocyanaceae) and Tarrietia sp. (Sterculiaceae) (Ohno 1990b). The latter two genera are unlikely to be normal hosts.</p></div>	https://treatment.plazi.org/id/842B6D7DFFECFFCBFF679FAD563DBFFC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beaver, R. A.;L. - Y	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFECFFCAFF679C6D54CABDDC.text	842B6D7DFFECFFCAFF679C6D54CABDDC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Genyocerus talurae (Stebbing)	<div><p>Genyocerus talurae (Stebbing)</p><p>(Figs 35–36)</p><p>Diapus talurae Stebbing, 1906: 418 .</p><p>Genyocerus talurae (Stebbing) . Wood &amp; Bright, 1992: 1096.</p><p>Diapus furtivus Sampson, 1913: 450 . Synonymy: Beeson, 1921: 22.</p><p>Diapus mirus Sampson, 1913: 452 . Synonymy: Sampson in Stebbing, 1914: 633.</p><p>Genyocerus dipterocarpi Browne, 1977: 98 . n. syn.</p><p>Taxonomy: We have compared male and female paratypes of G. dipterocarpi (BMNH) with specimens in RAB's collection that had earlier been compared with syntypes of D. furtivus (BMNH) . The synonymy of D. furtivus with G.talurae, suggested by Beeson (1921), was confirmed by Wood after examination of type material (Wood &amp; Bright 1992). The synonymy of D. mirus with D. furtivus was already suggested by Sampson in a personal communication to Stebbing (1914) and has been noted by Schedl (1972a) and Wood &amp; Bright (1992). Apart from a small difference in size (3.1–3.4 mm length for G. dipterocarpi, 3.5–3.7 mm for G.talurae), the only real difference between the males of the two species is the smaller number of mycangial pores on the pronotum of G. dipterocarpi (one or rarely two pores per side in G. dipterocarpi, three to five pores per side in G. talurae). As noted above, the number of pronotal pores can be quite variable and must be used with caution as a character at specific level. The females of G. dipterocarpi are smaller (3.1–3.2 mm long, excluding the hair brushes, and the part of the abdomen extending beyond the elytra, relative to 3.5–3.6 mm for G. talurae), but otherwise agree. G. talurae ranges from India to Vietnam, whereas G. dipterocarpi is known only from the Andaman Islands. However, in the absence of any information on genetic distinctiveness, we do not believe that G. dipterocarpi is morphologically sufficiently distinct to be given subspecific rank.</p><p>Distribution: India (including Andaman Is.), Myanmar, Pakistan, Thailand, Vietnam. It has been intercepted in timber imported to Japan from East Malaysia (Sabah, Sarawak) (Browne 1986, Ohno 1990b) and Indonesia (Sumatera) (Ohno et al. 1986), but its presence in these countries should be confirmed.</p><p>Biology: Recorded from the following genera of Dipterocarpaceae: Anisoptera, Dipterocarpus, Shorea, Vateria, Vatica . Other recorded hosts are: Canarium euphyllum (Burseraceae), Lannea coromandelica (= Odina wodier) ( Anacardiaceae) and Quercus semicarpifolia (Fagaceae) (Browne 1977, Wood &amp; Bright 1992). It is unlikely that the three latter species are regular hosts. The life history, gallery system and economic importance of the species are described by Beeson (1917, 1961). Of particular interest is the observa- tion by Beeson (1917) that the male secretes wax from minute pores on the posterior part of the elytra. This wax is used to form a short tube at the entrance to the gallery system.</p></div>	https://treatment.plazi.org/id/842B6D7DFFECFFCAFF679C6D54CABDDC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beaver, R. A.;L. - Y	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFEDFFCAFF679E0D520EBFCA.text	842B6D7DFFEDFFCAFF679E0D520EBFCA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Genyocerus tenellus (Schedl)	<div><p>Genyocerus tenellus (Schedl)</p><p>(Figs 37–39)</p><p>Diacavus tenellus Schedl, 1966: 42 .</p><p>Genyocerus tenellus (Schedl): Ohno et al., 1987: 90.</p><p>Taxonomy: We have examined two male and two female paratypes from the Schedl collection (NMW). This small species is closely related to G. quadriporus but can be distinguished by the characters given in the keys .</p><p>Distribution: Philippines (Luzon). It is known only from specimens intercepted in Japan in timber imported from Luzon (Schedl 1966, Ohno et al. 1987).</p><p>Biology: Recorded from Shorea sp. and 'lauan' (presumably Shorea sp.) ( Dipterocarpaceae).</p></div>	https://treatment.plazi.org/id/842B6D7DFFEDFFCAFF679E0D520EBFCA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beaver, R. A.;L. - Y	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
