taxonID	type	description	language	source
842B6D7DFFFDFFD8FF679C005399B9EC.taxon	description	The genus is distinguished from other platypodine genera except Diapus by the very large, widely separated procoxae. The species of Genyocerus can be distinguished from those of Diapus by their wider, more obvious scutellum, which is flush with the elytral surface posteriorly. The mycangial pores of Genyocerus are usually larger than those of Diapus and never fused to form a transverse or crescentic bar on each side of the midline. The antennal club of Genyocerus never has a median, testaceous strip lacking sensillae on the outer surface. The males of Diapus never possess a membranous extension of the apical margin of the fourth abdominal ventrite. The females of Genyocerus never possess dehiscent mandibular appendages. It may be noted that the larvae, so far as known (Browne 1972), show little evidence of generic distinctions. The two genera appear closely related, and more detailed studies, including DNA data and cladistic analyses are needed to test this phylogenetic hypothesis. Sexual dimorphism The males of Genyocerus are characterised by the presence of teeth or spines at the apex of the elytral disc; in the females, the elytral apices are subtruncate or transversely rounded. In the males of some species, the apical margin of the fourth ventrite extends as a hyaline membranous structure projecting over the basal part of the fifth ventrite (Fig. 12). This probably helps to make the concave fifth ventrite a more effective ' shovel' as the male collects and expels frass from the gallery system. The fifth ventrite of the female is essentially convex, and there is never an extension of the fourth ventrite. The young females bear brushes of hairs on the frons. The arrangement of these is species-specific, and they are probably used in courtship (Browne 1961 a, Roberts 1993) but lost after mating when the female takes over the extension of the gallery system from the male. In the male, the antennae are always inserted below the middle of the frons. In the females, the position of the antennae is associated with the presence or absence of large hair brushes on the upper part of the frons. When these are absent, the antennae are inserted in the same position as in the male (Fig. 7); when present, the antennae are inserted on the upper part of the frons at or above the level of the upper margin of the eyes (e. g. Figs 17, 34). The number of mycangial pores on the pronotum is variable but is almost always greater in the female than in the male (see below). Arrangement of teeth on male elytral apex Schedl (e. g. 1966) states that the first and second major teeth at the apex of the male elytra are formed from extensions of interstriae 1 and 3, whereas Browne (1962) writes that they are formed from extensions of interstriae 2 and 4. In fact, the first and second teeth are borne on interstriae 3 and 5. The interstriae in Genyocerus are often impunctate and can only be distinguished by the lines of strial punctures between them. Even the arrangement of the striae on the elytra is often difficult to determine in the smaller species, because the punctures are often small and obsolescent, but it can be seen in larger species such as G. biporus (Fig. 1). The first interstriae are normally developed along the suture. The apex may form a small secondary tooth at the inner side of the larger primary tooth on interstriae 3. Striae 1 are very short and usually terminate not more than one-third along the elytra. Thus the second interstriae are very short and taper to a point about onethird from the base of the elytra. Interstriae 3 and 5 are broadened distally and extend over the declivity as teeth. The teeth are usually flattened and blunt or with bifid tips (Figs 11, 15), but sometimes are more slen- der, tapering and spine-like (Figs 5, 30). Interstriae 4 and 6 are narrowed posteriorly and may not reach the end of the elytra. On the posterolateral part of the elytra, striae 6 - 8 end about 1 / 3 from the elytral apex, and there is a smooth, shining, punctureless area extending across interstriae 7 - 8 and sometimes interstriae 9 (Fig. 9). One or both interstriae 7 and 8 and interstriae 9 may bear apical teeth, or interstriae 7 - 9 may be fused apically to form an acute edge bearing several small teeth (Fig. 27). In a few species (e. g. G. tenellus, G. exilis), a tooth is also present on the elytral declivity. Intraspecific variation It has become clear during the course of this study that the authors who described the majority of species in the genus Genyocerus underestimated the intraspecific variation in the number of mycangial pores on the pronotum. Both Schedl and Browne in many of their papers (see Wood & Bright 1992 for references) noted that the number of pores differs between the sexes, with the female usually having a greater number (although the reverse is the case in Genyocerus puer (Schedl), Roberts 1993). In addition, the number of pores can differ between the two sides of the same individual, and there can also be considerable variation between individuals from the same collection site, and from different sites, independently of the differences between males and females. Similarly, in the males of some species, the shape and size of the teeth at the end of the elytral disc can be quite variable intraspecifically. Such variations have sometimes been described as separate species. However, the gradual accumulation of more specimens from a wider range of collection sites has indicated that intermediates exist. The resulting new synonyms are reported below. Biology The species of Genyocerus normally breed only in host trees in the family Dipterocarpaceae. The only partial exception to this is G. pendleburyi, which may occasionally attack trees in the family Fagaceae in areas in which dipterocarps are scarce (Browne 1977). A few other records from non-dipterocarp trees may or may not involve actual breeding. By contrast, species of Diapus, like most platypodines, are generally polyphagous, attacking any trees in which the symbiotic ambrosia fungi will grow. Genyocerus species usually attack dying or felled trees, but Browne (1961 a) noted that attacks may occur on stressed trees after drought. The general features of the life history are given by Beeson (1961) and Browne (1961 a). The gallery system is started by the male, but only a short vertical gallery is made. It is probable that the female is attracted by a male-produced pheromone (Beaver 2000 b), although no studies have been made on Genyocerus or Diapus. After mating, the female continues to extend the gallery into the wood. The male remains in the entrance, keeping out potential predators and parasites and removing the frass that accumulates from the boring activities of the female. The gallery system branches mainly in one transverse plane and may extend deeply into the tree. The mycangia on the pronotum transport an ambrosial fungus (or fungi) that is released into the gallery, grows in the wood and provides the only food for both adults and larvae, which feed on the hyphae growing on the gallery walls. The ambrosia fungi associated with Genyocerus and Diapus have not been investigated. Mature larvae construct pupal cells above and below the gallery at the ends of the branches. The adults of the new generation emerge through the parental entrance hole.	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF4FFD3FF679D185242B982.taxon	materials_examined	Taxonomy: The type material of G. albipennis was long thought to be lost, but Wood (1969) found a single female in the Motschulsky collection in Moscow and clarified the status of the genus and the species. Later, Wood (1981) placed Diacavus irregularis in synonymy. We have examined the type series of Diacavus irregularis (BMNH). The male is closely related to, but distinct from, G. diaphanus (Schedl) (see key to males above), but we are unable to distinguish the female from G. diaphanus (see key to females above). The species was referred to by Beeson (1961) as Diacavus zeylanicus, but this name is a nomen nudum (Browne 1977). Distribution: Sri Lanka only. Records from other countries result from confusion with Diapus albipennis Strohmeyer, which is in fact a synonym of G. pendleburyi (Schedl) (see below) and not a synonym of G. albipennis Motschulsky, as indicated by Schedl (1972 a). Biology: The only identified host is Dipterocarpus zeylanicus (Dipterocarpaceae) (Browne 1977).	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF4FFD3FF679BD85233BADA.taxon	description	(Figs 1 – 4)	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF4FFD3FF679BD85233BADA.taxon	distribution	Distribution: The species is distributed from northeastern India (Assam) through Southeast Asia to Indonesia and the Philippines. It has often been intercepted in Japan in timber from the region but is not known to be established there. Biology: Recorded only from genera belonging to the family Dipterocarpaceae (Anisoptera, Dipterocarpus, Dryobalanops, Hopea, Parashorea, Shorea). It usually attacks freshly cut logs (Browne 1961 a).	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF5FFD2FF679FAD55DFBB64.taxon	description	(Figs 5 – 7)	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF5FFD2FF679FAD55DFBB64.taxon	description	Female: Generally resembling male, 2.7 mm long (excluding frontal hair brushes and part of abdomen extending beyond elytra), about 4.1 times as long as wide. Frons weakly convex above epistoma, shallowly, transversely impressed above antennal insertions, just above epistoma and close to midline on each side, a pencil – like brush of long, pale yellow hairs, directed anteriad and somewhat ventrad, each brush inserted on a slightly raised base, cylindrical near base, tapering towards tip, brushes parallel close to base, then divergent, between their insertions on lower part of frons in midline a small tuft of much shorter, ventrally directed hairs (Fig. 7). Antennae inserted at level of lower margin of eyes, club oval, with a few longer hairs, pubescent to base except for a narrow corneous strip along posterior margin, as in male. Pronotum about 1.3 times as long as wide, slightly more elongate than in male, one or two large mycangial pores on each side of midline close to base, and lateral to these a group of about 25 – 40 much smaller pores. (In the male, a patch of about 12 – 15 small pores laterally to the pair of large mycangial pores was not mentioned by Browne (1980 b) in his description.) Elytra 1.7 – 1.8 times as long as pronotum, 2.1 – 2.2 times longer than wide, carina at base of elytra reduced and restricted to humeral region, not extending to scutellum as in male, a row of long semi – recumbent hairs on interstriae 3 (as in male), lacking spines at apex of elytral disc, elytral apices separately rounded. Fifth ventrite plano – concave, without a distinct median groove, lateral margins indistinct, vestiture a little shorter and less dense than in male, largely confined to marginal areas. Female specimens examined: MALAYSIA: Sabah, Sipitang, Mendolong, 26. xi. 1987, 27. xi. 1987, various dates from 13. iv. – 11. v. 1988, 19. ii. 1989 (S. Adebratt) (16 ♀) (ZMLU, RAB). Distribution: The species was previously known only from specimens intercepted in Japan in timber shipped from East Malaysia (Sabah and Sarawak) (Browne 1980 b, Ohno 1990 b). The occurrence of the species in Sabah can now be confirmed, and it is recorded for the first time from Brunei Darussalam. New Records: BRUNEI, 115 o 7 ' E, 4 o 34 ' N, Kuala Belalong FSC, Dipterocarp forest, site 16 – 2, 520 m alt., Fog 28, 18. iii. 1992 (N. Mawdsley) (1 ♂) (BMNH); MALAYSIA: Sabah, Sipitang, Mendolong, 26. xi. 1987, 27. xi. 1987, various dates from 13. iv. – 13. v. 1988, 19. ii. 1989, 19. iii. 1989 (S. Adebratt) (23 ♂, 16 ♀) (ZMLU, RAB). Biology: Recorded from four genera of Dipterocarpaceae (Anisoptera, Dipterocarpus, Dryobalanops, Shorea) and from ' Sapotaceae sp. ' (Ohno 1990 b).	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF5FFD1FF6799FA5294B88C.taxon	description	(Fig. 8)	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF5FFD1FF6799FA5294B88C.taxon	distribution	Distribution: G. diaphanus is widespread, occurring from India through Southeast Asia to Borneo and the Philippines. It has frequently been intercepted in dipterocarp timber imported to Japan (e. g. Schedl 1966, Ohno 1990 b) and Korea (Choo & Woo 1983). It is newly recorded here from northern Thailand. New Records: THAILAND, Chiang Mai, C [hiang] M [ai]. Univ., 300 m, ex EtOH trap, various dates from 10. i. 2005 – 4. iv. 2005, (W. Puranasakul) (1 ♂, 3 ♀) (CMU, RAB). Biology: The species, like its congeners, breeds primarily in trees of the family Dipterocarpaceae, and has been recorded from the following genera (Anisoptera, Balanocarpus, Dipterocarpus, Hopea, Parashorea, Pentacme, Shorea, Upuna, Vatica). There are a few records from trees of other families: Gonystylus (Thymeleaceae) (Browne 1961 a), Gluta or Melanorrhoea, Mangifera (Anacardiaceae), Durio (Bombacaceae), Palaquium (Sapotaceae), Pterocymbium (Sterculiaceae) (Ohno 1990 b), but these involve small numbers of specimens. Ohno (1990 b, as Genyocerus abdominalis) records a total of 9851 (6938 ♂, 2913 ♀) specimens bred from imported logs of seven genera of Dipterocarpaceae, but only 24 (21 ♂, 3 ♀) obtained from imported logs of six non-dipterocarp genera. This suggests that breeding was very rare, if it occurred at all, in the latter.	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF6FFD0FF679AEF5461BCCA.taxon	description	(Figs 9 – 10)	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF6FFD0FF679AEF5461BCCA.taxon	distribution	Distribution: East Malaysia (Sarawak), Indonesia (Maluku, Bacan I.), West Malaysia. It is newly recorded here from Brunei Darussalam and Sabah. It has been intercepted in Japan in timber from Borneo and Maluku. New Records: BRUNEI: 4 o 34 ' N, 115 o 7 ' E, Kuala Belalong FSC, Dipterocarp forest, 220 – 275 m. alt, fogging, malaise trap, flight intercept trap, various dates from 16. vi. 1991 – 6. viii. 1991, 8. ii. 1992, 10. iii. 1992 (N. Mawdsley) (8 ♂, 10 ♀) (BMNH); MALAYSIA: Sabah, Sipitang, Mendolong, 8. xii. 1987, various dates from 10. iv. 1988 – 13. v. 1988, 10 – 31. iii. 1989 (S. Adebratt) (20 ♂, 32 ♀) (ZMLU, RAB); Sabah, Poring Spring, lower montane, mixed dipterocarp forest,> 650 m., fogging Aporusa sp., 22. iii. 1993 (A. Floren) (1 ♂) (BZUW). Biology: Recorded from the following genera of Dipterocarpaceae: Anisoptera, Dipterocarpus, Dryobalanops, Shorea, Upuna, Vatica (Browne 1961 a, Ohno 1990 b) ..	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF7FFD7FF679D10520CBCC9.taxon	description	(Figs 11 – 14)	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF7FFD7FF679D10520CBCC9.taxon	description	Female: Generally resembling male, 3.7 – 3.8 mm long (excluding frontal hair brushes and part of abdomen extending beyond elytra), 2.9 – 3.0 times as long as wide. Frons with antennae inserted on upper part of frons close to upper margin of eyes, brushes of long, golden – yellow hairs present in young females both above epistoma and on upper part of frons (Figs 13 – 14), a large teardrop – shaped, shallow, shining, impunctate impression on each side, extending from vertex to lower quarter of frons, the impressions separated by a narrow, median carina, upper part of frons lateral to impressions flat, bearing on each side a large brush of hairs, extending dorsally well above antennal insertion and nearly meeting medially just below vertex, the majority of hairs directed forward, downcurved at tips, each brush extending ventrally in a narrow band close to inner margin of eye nearly to its lower margin, the lowest group of hairs longer and incurved towards midline; epistomal brushes divided into two parts on each side, the long hairs upcurved below those of the upper brushes, almost reaching vertex, lowest part of frons below impressions finely punctured, the punctures with much shorter erect hairs, normally concealed below epistomal brushes. Pronotum 1.05 – 1.1 times longer than wide, similar to that of male but with 9 – 12 large mycangial pores on each side of midline close to base (Fig. 13), rather than 0 – 2 as in male. Elytra wider than and about 1.6 times as long as pronotum, 1.5 times longer than wide, carina on base of elytra reduced and more rounded, striae less strongly impressed, becoming obsolete in posterior third of elytra, interstriae with sparse, semi – recumbent hairs, elytral apices lacking teeth, separately rounded, with more abundant short hairs. Female specimens examined: MALAYSIA: Sabah, Sipitang, Mendolong, 28. xi. 1987, 6. v. 1988, 13. v. 1988 (S. Adebratt) (5 ♀) (ZMLU, RAB). Distribution: The species is recorded only from East Malaysia (Sarawak) and West Malaysia. It is newly recorded here from Brunei Darussalam and Sabah. New records: BRUNEI: 4 o 34 ' N, 115 o 7 ' E, Kuala Belalong FSC, Dipterocarp forest, site 1, 260 m alt., Fog 8, 9. viii. 1991 (N. Mawdsley) (1 ♂) (BMNH); Temburong, nr. K. Belalong Field Stud. Centre, 4 o 33 ' N, 115 o 09 ' E, 150 m, 1. iii. 1992 (R. A. Beaver) (3 ♂); as previous except: 200 m, ex Shorea sp., 10. iii. 1992 (2 ♂) (all RAB); [MALAYSIA (Sabah)] North Borneo (SE), Forest camp, 19 km. N. of Kalabakan, 60 m., 13. x. 1962 (Y. Hirashima) (1 ♂); as previous except: 21. x. 1962 (1 ♂); as previous except: 19. x. 1962 (K. J. Kuncheria) (1 ♂) (all BPBM); MALAYSIA: Sabah, Sipitang, Mendolong, 28. xi. 1987, 6. v. 1988, 13. v. 1988 (S. Adebratt) (1 ♂, 5 ♀) (ZMLU, RAB). Biology: The only host record is from the dipterocarp tree Shorea leprosula (Browne 1962).	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF0FFD7FF679D175503B9B6.taxon	description	(Figs 15 – 18)	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF0FFD7FF679D175503B9B6.taxon	distribution	Distribution: The species occurs in East and West Malaysia and in the Philippines and has been imported in timber to Japan. Its occurrence in Vietnam (Schedl 1972 a) needs to be substantiated. Biology: Recorded hosts are all in the family Dipterocarpaceae (Anisoptera, Shorea, Vatica) (Browne 1961 a, 1985; Ohno et al. 1988).	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF0FFD6FF679A2A53BEBC35.taxon	description	(Figs 19 – 22)	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF0FFD6FF679A2A53BEBC35.taxon	materials_examined	Taxonomy: We have examined the holotype male, allotype female and a female paratype (BPBM) from Papua New Guinea and compared them with a series of males collected in Indonesia: Papua Barat (formerly Irian Jaya) (NKME). The available material suggests that the species occurs in two size classes. The holotype male from Papua New Guinea (BPBM) and nine males (NKME) from Indonesia: Papua Barat are 4.5 – 4.8 mm long; two further males (NKME) from a second locality in Papua Barat (about 100 km distant from the first, but in a different river basin) are 5.5 and 5.7 mm long. The smaller males have a total of 8 – 9 mycangial pores; the larger males have 14 – 17. We are unable to find any other significant differences between the two groups of males. Although the size difference suggests reproductive isolation, it would be premature to describe the larger males as a distinct species, because intermediates may yet be found (as in G. quadriporus (see below )). Roberts (1993) suggests that the male can be distinguished from G. biporus by the hair ornamentation of the lower part of the frons. However, the basic arrangement of hairs in four longitudinal rows is the same in both species. Alternative distinguishing characters are given in the keys above. Distribution: Known only from the island of New Guinea (Indonesia: Papua Barat, and Papua New Guinea). New Records: INDONESIA, Irian Jaya, Nabire area, road Nabire-Ilaga, km 62, 03 o 30 ' 936 '' S, 135 o 41 ' 945 '' E, 250 m NN, LEK, X. 1997 (M. Balke) (9 ♂) (NKME, RAB) (all smaller form); Wapoga River, E Asori, km 64, Kwadewa camp, 2 o 49 ' S, 136 o 28 ' E, 10. i. 1999 (A. Weigel) (2 ♂) (NKME, RAB) (larger form). Biology: The only recorded host is Anisoptera sp. (Dipterocarpaceae) (Roberts 1993).	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF1FFD6FF679EAA53D7B83B.taxon	distribution	Distribution: The species is distributed from Northeastern India (Assam) through Southeast Asia to Borneo and the Philippines. It has been intercepted in timber from Indonesia and the Philippines imported into Japan and Korea. Biology: It usually breeds in trees of the family Dipterocarpaceae (Anisoptera, Dipterocarpus, Dryobalanops, Pentacme, Shorea), but has also been recorded from Fagaceae (Quercus). Browne (1977) notes that the record from Quercus sp. came from a site at high altitude where dipterocarps were scarce.	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF1FFD6FF679AA156BFBA7B.taxon	distribution	Distribution: Philippines (Luzon?). A single female is recorded here from East Malaysia. New Record: MALAYSIA, Sabah, Sipitang, Mendolong, 17. ii. 1989 (S. Adebratt) (1 ♀) (RAB). Biology: Unknown.	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF2FFD5FF679FAD55BDBEF1.taxon	description	(Figs 23 – 25)	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF2FFD5FF679FAD55BDBEF1.taxon	distribution	Distribution: Papua New Guinea only. Biology: Specimens have been collected from sticky traps placed on felled trees of Anisoptera sp. and Hopea sp. (Dipterocarpaceae) (Roberts 1993).	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF2FFD5FF679B7D56EFBAE9.taxon	description	(Figs 27 – 29)	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF2FFD5FF679B7D56EFBAE9.taxon	distribution	Distribution: The species is recorded from West Malaysia, East Malaysia (Sabah, Sarawak), Sumatra, Thailand and the Philippines. It has been intercepted in Japan and South Korea in timber exported from the region. Biology: All recorded host trees are members of the family Dipterocarpaceae (Anisoptera, Dipterocarpus, Dryobalanops, Shorea).	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF3FFD4FF679FAD57F0BF9C.taxon	distribution	Distribution: The species is known only from the Philippines, from which it was intercepted in timber imported to Japan. The record by Schedl (1973) from Nepal must be regarded as dubious and is possibly the result of misidentification of G. pendleburyi, which occurs in the area. Biology: The species was originally taken from ' lauan' wood, probably a species of Shorea (Dipterocarpaceae).	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF3FFD4FF679DD7526CB8C4.taxon	description	(Figs 32 – 34)	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF3FFD4FF679DD7526CB8C4.taxon	distribution	Distribution: East Malaysia (Sarawak), West Malaysia. The species is newly recorded here from Brunei Darussalam and East Malaysia (Sabah). New Records: BRUNEI, 115 o 7 ' E, 4 o 34 ' N, Kuala Belalong FSC, Dipterocarp forest, 270 m, Aerial malaise, 16. vi. 1991 (N. Mawdsley) (1 ♀); as previous except: Ground F [light] I [ntercept] T [rap], 17. vi. 1991 (1 ♀); 220 m, Aerial FIT, 16. vii. 1991 (1 ♀; 210 m, Ground malaise, 8. ii. 1992 (1 ♂) (all BMNH). MALAYSIA, Sabah, Sipitang, Mendolong, 1. xii. 1987 (S. Adebratt) (9 ♂, 6 ♀); as previous except various dates from 29. iv. 1988 – 11. v. 1988 (10 ♂, 6 ♀) (ZMLU, RAB). Biology: The species has been recorded from species of Dipterocarpus, Dryobalanops, Hopea, Parashorea, Shorea and Vatica (all Dipterocarpaceae) (Browne 1961 a, b; Beaver & Browne 1979). The gallery system is described by Browne (1961 a), who suggests that the life cycle in West Malaysia spans 5 – 6 weeks.	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFF3FFD4FF67991A531FBA9A.taxon	distribution	Distribution: Philippines (Mindanao). Biology: Recorded from Shorea sp. (Dipterocarpaceae) (Browne 1981).	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFECFFCBFF679FAD563DBFFC.taxon	distribution	Distribution: Intercepted in Japan in timber imported from ' various shipment localities' in Borneo (Ohno 1990 b) and from Indonesia (Sumatera) (Ohno et al. 1986). Biology: Recorded from four genera of Dipterocarpaceae (Anisoptera, Dipterocarpus, Dryobalanops, Shorea), with nearly 3,000 specimens obtained from Dipterocarpus sp., and as single specimens from Dyera sp. (Apocyanaceae) and Tarrietia sp. (Sterculiaceae) (Ohno 1990 b). The latter two genera are unlikely to be normal hosts.	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFECFFCAFF679C6D54CABDDC.taxon	distribution	Distribution: India (including Andaman Is.), Myanmar, Pakistan, Thailand, Vietnam. It has been intercepted in timber imported to Japan from East Malaysia (Sabah, Sarawak) (Browne 1986, Ohno 1990 b) and Indonesia (Sumatera) (Ohno et al. 1986), but its presence in these countries should be confirmed. Biology: Recorded from the following genera of Dipterocarpaceae: Anisoptera, Dipterocarpus, Shorea, Vateria, Vatica. Other recorded hosts are: Canarium euphyllum (Burseraceae), Lannea coromandelica (= Odina wodier) (Anacardiaceae) and Quercus semicarpifolia (Fagaceae) (Browne 1977, Wood & Bright 1992). It is unlikely that the three latter species are regular hosts. The life history, gallery system and economic importance of the species are described by Beeson (1917, 1961). Of particular interest is the observa- tion by Beeson (1917) that the male secretes wax from minute pores on the posterior part of the elytra. This wax is used to form a short tube at the entrance to the gallery system.	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFEDFFCAFF679E0D520EBFCA.taxon	description	(Figs 37 – 39)	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
842B6D7DFFEDFFCAFF679E0D520EBFCA.taxon	distribution	Distribution: Philippines (Luzon). It is known only from specimens intercepted in Japan in timber imported from Luzon (Schedl 1966, Ohno et al. 1987). Biology: Recorded from Shorea sp. and ' lauan' (presumably Shorea sp.) (Dipterocarpaceae).	en	Beaver, R. A., L. - Y (2007): A review of the genus Genyocerus Motschulsky (Coleoptera: Curculionidae: Platypodinae), with new synonyms and keys to species. Zootaxa 1576 (1): 25-56, DOI: 10.11646/zootaxa.1576.1.3, URL: https://biotaxa.org/Zootaxa/article/view/zootaxa.1576.1.3
