identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
8B20878DFFD0FFCEE2BEFA0FFD6FFCFF.text	8B20878DFFD0FFCEE2BEFA0FFD6FFCFF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heligmonoides mirzai Smales 2009	<div><p>Heligmonoides mirzai, new species</p> <p>(Figs. 1–14)</p> <p>Material examined. – Holotype male and allotype female: BBM- NG-99529, from the small intestine of Melomys rufescens (Muridae: Hydromyinae), type host, Nauti Village, 7°15'S 146°13'E, Morobe Province, Papua New Guinea, coll. A. B. Mirza, 10 Oct.1970.</p> <p>Paratypes: 12 males, 12 females, SAM AHC 34783; collection data as above.</p> <p>Voucher specimens: BBM NG- 97928 1 male, 2 females from the small intestine of Melomys rufescens, Minava Village, Watut, 6°50'S 146°21'E, Morobe Province, coll. A. B. Mirza, 23 Oct.1969; AM W32620, 2 males, 3 females from small intestine of Melomys rufescens, Yapsiei area, 4°35'S 141°5'E, Sanduan Province, coll. T. Flannery, 17 Jan.1984; 1 male, 2 females, AM W32625 from the small intestine of Melomys species, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.11667&amp;materialsCitation.latitude=-5.016667" title="Search Plazi for locations around (long 141.11667/lat -5.016667)">Dokfuma Star Mountains</a> 5º01'S 141°07'E, West Sepik, Sanduan Province, coll. T. Flannery, 5 Apr.1984.</p> <p>Prevalence. – A total of 5 of 25 M. rufescens and 1 of 10 Melomys spp.</p> <p>Etymology. – The species name is in recognition of the extensive collecting done by A. B. Mirza in the 1960s and 1970s in New Guinea.</p> <p>Description. – Small coiled worms; prominent cephalic vesicle present; buccal capsule vestigial. Mouth opening triangular with rudimentary lips; labial and cephalic papillae not observed. Oesophagus claviform. Nerve ring surrounds oesophagus at about middle third; deirids and excretory pore at about same level, posterior to nerve ring. Synlophe of pointed longitudinal cuticular ridges extends from the posterior margin of the cephalic vesicle to level of spicules or vulva; 16–18 ridges in anterior, 21–24 in midbody. Axis of orientation of ridges passing from ventral right to dorsal left side, inclined about 75° from sagittal axis; 10–12 ridges dorsal side, 11–12 ridges ventral side in midbody. Ridges 1, 1’ large but not thickened basally, together with ridges 2, 2’ form a carene of type B. Ridges 3–7 decreasing in size, ridges 8–10 increasing in size, ridge 11 smaller; ridges 3’– 8’ increasing in size, ridges 9’–11’ decreasing in size. Posterior region of body with 19–20 (male) 20–21 (female) ridges reduced in size; 10–11 dorsal side, 9–10 ventral side ridges.</p> <p>Male (holotype and 9 paratypes): Length 2.2–3.3 (2.8) mm, maximum width 83–107 (90). Cephalic vesicle 36–46 (39.25) long. Oesophagus 350–550 (420) long; nerve ring 145 from cephalic end. Bursa asymmetrical, left lobe larger, rays of left lobe more robust; without dorsal median notch. Dorsal ray symmetrical; divided at about half its length, each branch dividing again at distal tip; terminal divisions, rays 9, 10 symmetrical; rays 8 arising at same level proximally to division of dorsal ray, left ray 8 slightly more robust than right. Rays 4, 5, 6 with common stem, reaching margin of bursa, rays 6 more slender than rays 4, 5. Rays 2, 3 with common stem, widely divergent distally, rays 3 longer than rays 2. Genital cone short, lightly sclerotized, ventral lobe with unpaired papillae 0, dorsal lip bifid, each lobe with single papilla 7. Spicules equal, filiform, proximal ends rounded, distal tips simple, straight, 250–320 (275) long. Gubernaculum 27 long.</p> <p>Female (allotype and 9 paratypes): Length 2.95–3.7 (3.4) mm, maximum width 99–107 (104). Cephalic vesicle 42.0 –49.5 (45) long. Oesophagus 350–400 (380) long; nerve ring 190, 220, excretory pore 250, 280 deirids 250, 280 from cephalic end. Posterior end twisted and flexed ventrally, with praepuce in mature worms; vulva opening about 100 from tail tip. Monodelphic; ovejector with sphincter, 95, longer than vestibule 40, and infundibulum, 45. Tail conical, tip with spike 39.5–55 (48.5) long. Eggs thin- shelled, ellipsoidal, in utero 51–69 (57) by 30–36 (33.8).</p> <p>Remarks. – Despite the fact that much of the material had been fixed in the hosts in situ, making it difficult to discern the morphology of the anterior end of the worms, it could be seen that Heligmonoides mirzai, new species, has most of the morphological features of the genus Heligmonoides and resembles H. bulbosus Ow Yang et al., 1983, in the number and arrangement of the ridges on the synlophe (Ow Yang et al., 1983, Fig 11A, B). Heligmonoides mirzai, new species, difers from H. bulbosus in spicule length (250–320 compared with 92–100), the proportions of the dorsal ray, the length of rays 3 and the shape of the female posterior end, with a praepuce and spiked tail tip.</p> <p>Hasegawa &amp; Syafruddin (1994a) originally described Maxomystrongylus musseri as Heligmonoides musseri, noting that it most closely resembled H. bulbosus, particularly in the features of the carene; supported by 3–4 slender ridges lacking thickened roots. Subsequently these authors erected the new genus Maxomystrongylus Hasegawa &amp; Syafruddin, 1997, and transferred H. musseri into it on the basis of the characters of the synlophe discussed above, having the division of the dorsal ray distal to the derivation of rays 8 and a vagina vera with a unilateral diverticulum (Hasegawa &amp; Syafruddin, 1994a). Further they commented that the systematic relationship between H. bulbosus and Maxomystrongylus remained unclear. Although H. mirzai, new species, has characters of the synlophe and a form of the dorsal ray consistent with Maxomystrongylus, it does not have a unilateral diverticulum of the vagina. Consequently it has been placed in the genus Heligmonoides together with the other Southeast Asian representatives of the genus, H. bulbosus and H. lanceolatus Ow Yang et al., 1983, pro tem. A revision of the heligmonellid genera from the Oriental and Australasian regions is needed to resolve the relationship between Heligmonoides and Maxomystrongylus and the systematic position of the Australasian species.</p> </div>	https://treatment.plazi.org/id/8B20878DFFD0FFCEE2BEFA0FFD6FFCFF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Smales, L. R.	Smales, L. R. (2009): Helminths Of Melomys Rufescens And Melomys Spp. (Muridae: Hydromyinae) From Papua New Guinea With The Descriptions Of A New Genus And Five New Species In The Heligmonellidae (Nematoda: Trichostrongyloidea). Raffles Bulletin of Zoology 57 (1): 5-15, DOI: 10.5281/zenodo.5341456
8B20878DFFD5FFCEE007FC8FFF31FAFF.text	8B20878DFFD5FFCEE007FC8FFF31FAFF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Melomystrongylus Smales 2009	<div><p>Melomystrongylus, new genus</p> <p>Diagnosis. – Trichostrongyloidea: Heligmonellidae: Nippostrongylinae. Synlophe well developed with pointed ridges; in midbody axis of orientation of ridges passing through ventral right and dorsal left sides inclined about 65° from sagittal axis in anterior body, lacking clear orientation in mid and hind body. Ventral ridge 5’ hypertrophied anteriorly. Bursa asymmetrical with larger left lobe. Dorsal ray divided distal to level of branching of rays 8 from dorsal trunk. Parasites of hydromyine murids.</p> <p>Type species. – Melomystrongylus sepikensis, new species.</p> </div>	https://treatment.plazi.org/id/8B20878DFFD5FFCEE007FC8FFF31FAFF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Smales, L. R.	Smales, L. R. (2009): Helminths Of Melomys Rufescens And Melomys Spp. (Muridae: Hydromyinae) From Papua New Guinea With The Descriptions Of A New Genus And Five New Species In The Heligmonellidae (Nematoda: Trichostrongyloidea). Raffles Bulletin of Zoology 57 (1): 5-15, DOI: 10.5281/zenodo.5341456
8B20878DFFD5FFCCE1C5FA8FFDB7FC1F.text	8B20878DFFD5FFCCE1C5FA8FFDB7FC1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Melomystrongylus sepikensis Smales 2009	<div><p>Melomystrongylus sepikensis, new species</p> <p>(Figs. 15–28)</p> <p>Material examined. – Holotype male and allotype female: BBM- NG-104657B, from the small intestine of Melomys rufescens (Muridae: Hydromyinae) type host, 2 km south of Mt Samoro, West Sepik, Sanduan Province, Papua New Guinea, coll. A. B. Mirza, 10 May 1975.</p> <p>Paratypes: 2 males, 5 females BBM-NG-105128B, from small intestine Melomys rufescens, 12 km northwest of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=146.08333&amp;materialsCitation.latitude=-5.95" title="Search Plazi for locations around (long 146.08333/lat -5.95)">Tep Tep</a>, 5°57'S 146°05'E, Madang Province, coll. A. B. Mirza, 2 Dec.1975.</p> <p>Voucher specimens: 4 males, 1 female BBM-NG-105155B, 5 females, 1 male, BBM-NG-105156B, from the small intestine of Melomys rufescens, 12 km northwest of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=146.08333&amp;materialsCitation.latitude=-5.95" title="Search Plazi for locations around (long 146.08333/lat -5.95)">Tep Tep</a>, 5°57'S 146°05'E, Madang Province, coll. A. B. Mirza, 2 Dec.1975; 1 female, BBM-NG-103717, Wanuma, 4°54'S 145°19'E, Madang Province, coll A. B. Mirza, 3 Feb.1974; 1 male, 2 females, AM W32617, from the small intestine of Melomys rufescens, Sideia Mission, 10°32'S 150°48'E, Milne Bay Province, coll. G. Hangay 31 Dec.1988; 3 males, 8 females, AM W32557, from the small intestine of</p> <p>Melomys sp., Munimun Village, Aguan, 9°53'S 149°23'E, coll P. German, 8 Aug.1992; 1 female, AM W32558, from the caecum of Melomys sp., Boulder Camp, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=150.26666&amp;materialsCitation.latitude=-9.35" title="Search Plazi for locations around (long 150.26666/lat -9.35)">Goodenough Island</a>, 9°21'S 150°16'E, Sanduan Province, coll. T. Ennis, 25 Aug.1987; 2 females, AM W32559 from the small intestine of Melomys sp., Boulder Camp, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=150.26666&amp;materialsCitation.latitude=-9.35" title="Search Plazi for locations around (long 150.26666/lat -9.35)">Goodenough Island</a>, 9°21'S 150°16'E, Sanduan Province, coll. T. Ennis, 25 Aug.1987; AM W32626, 5 males, from the small intestine of Melomys sp., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.11667&amp;materialsCitation.latitude=-5.016667" title="Search Plazi for locations around (long 141.11667/lat -5.016667)">Dokfuma Star Mountains</a> 5º01'S 141°07'E, Milne Bay Province, coll. T. Flannery, 5 May 1987.</p> <p>Prevalence. – A total of 5 of 25 M. rufescens and 3 of 10 Melomys spp.</p> <p>Etymology. – The species name reflects the locality in which this nematode is found.</p> <p>Description. – Relatively robust worms; prominent cephalic vesicle present; buccal capsule vestigial. Mouth opening triangular with rudimentary lips; surrounded by 4 cephalic papillae and 2 amphids; labial papillae not observed. Oesophagus claviform. Nerve ring surrounds oesophagus at about a third its length, digitiform deirids and excretory pore at about same level, posterior to nerve ring. Synlophe of continuous longitudinal pointed cuticular ridges extends from the posterior margin of the cephalic vesicle to just anterior to the bursa or vulva; 8–14 ridges in anterior, 14–16 in midbody. Axis of orientation passing from ventral right to dorsal left sides, inclined about 65° to sagittal axis in anterior body; 4–8 ridges dorsal side, 4–8 ridges ventral side. Ridges 1–6 decreasing in size; ridges 1'–4' decreasing in size, ridge 5' largest, ridges 6'–8' decreasing in size. Posterior region of body with 15 (male) 10–14 (female) ridges reduced in size with no clear axis of orientation; 7–8 (male), 6–7 (female) ridges dorsal side, 8 (male), 7–9 (female) ridges ventral side.</p> <p>Male (holotype, 2 paratypes and 7 vouchers): Length 3.2–4.3 (3.8) mm, maximum width 82.5–99 (90.5). Cephalic vesicle 20.5–42.5 (33) long, 34–39 (37) wide. Oesophagus 280–355 (322) long; excretory pore 169–221 (186), deirids 204 from cephalic end. Bursa slightly asymmetrical, left lobe larger; dorsal lobe with median dorsal notch. Dorsal ray symmetrical, divided at about half its length, each branch dividing again at distal tip; terminal divisions, rays 9, 10 symmetrical; rays 8 arising at same level proximal to division of dorsal ray, right ray 8 slightly longer than left. Rays 4, 5, 6 with common stem, reaching margin of bursa, rays 6, more slender, curving anteriorly, rays 4, 5 curving posteriorly. Rays 2, 3 with common stem, dividing into 2 widely divergent branches, curving posteriorly, reaching margin of bursa. Genital cone slightly extended, lightly sclerotized, ventral lobe with unpaired papilla 0 and dorsal lip with paired papillae 7. Spicules equal, filiform, tips pointed, 450–660 (610) long. Gubernaculum forming thin plate 38–56 (43) long.</p> <p>Female (allotype, 4 paratypes and 5 vouchers): Length 3.9–5.5 (4.6) mm, maximum width 80–114 (100). Cephalic vesicle 29.5–39.5 (30) long, 33–46 (35.5) wide. Oesophagus 310–390 (345) long; nerve ring 150, excretory pore 145–248 (216), deirids 145–248 (216) from cephalic end. Vulva with prominent lips, opens 60–115 (95) from tail tip. Body wall extends over vulva and tail as praepuce in mature specimens. Monodelphic; ovejector with infundibulum, 90–100, longer than vestibule, 60–80, sphincter 30, 50, shortest element. Tail conical, tip rounded, reflected ventrally, 29–42.5 (35) long. Eggs thin-shelled, ellipsoidal, in utero 49–66 (57) by 30–39 (34).</p> <p>Remarks. – Melomystrongylus new genus has all the characteristics of the subfamily Nippostrongylinae (see Durette-Desset, 1985). Lacking a carene and with a hypertrophied ventral ridge Melomystrongylus resembles Hasanuddinia Hasegawa &amp; Syafruddin, 1994. It can be readily distinguished from Hasanuddinia in that it has a single hypertrophied ventral ridge beginning from the excretory pore and ending in the anterior third of the body. Melomystrongylus has smaller similar sized ridges showing no clear axis of orientation in the mid and posterior body while the axis of orientation of the ridges is maintained along the whole length of the body in Hasanuddinia (see Hasegawa &amp; Syafruddin, 1994a). The gradient of ridge size in the anterior body in Melomystrongylus is from right to left (excepting ridge 5), but in Hasanuddinia it is from left to right.</p> </div>	https://treatment.plazi.org/id/8B20878DFFD5FFCCE1C5FA8FFDB7FC1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Smales, L. R.	Smales, L. R. (2009): Helminths Of Melomys Rufescens And Melomys Spp. (Muridae: Hydromyinae) From Papua New Guinea With The Descriptions Of A New Genus And Five New Species In The Heligmonellidae (Nematoda: Trichostrongyloidea). Raffles Bulletin of Zoology 57 (1): 5-15, DOI: 10.5281/zenodo.5341456
8B20878DFFD7FFCCE032FBEFFAC7F8E9.text	8B20878DFFD7FFCCE032FBEFFAC7F8E9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Odilia similis Smales 2009	<div><p>Odilia similis, new species</p> <p>(Figs. 29–38)</p> <p>Material examined. – Holotype male and allotype female: BBM- NG-53130, from the small intestine of Melomys rufescens (Muridae: Hydromyinae), type host, Kalolo, Morobe Province, Papua New Guinea, coll. O. R. &amp; J. W. Wilkes, 28 Aug.1966</p> <p>Paratypes: 1 male, 2 females, SAM AHC 34782; collecting data as above.</p> <p>Voucher specimens: 2 males, 3 females, BBM-NG-54020 &amp; 20 males, 18 females, BBM-NG- 53691, from the small intestine of Melomys rufescens, 10 km west of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=146.65&amp;materialsCitation.latitude=-7.2" title="Search Plazi for locations around (long 146.65/lat -7.2)">Bulolo</a>, 7°12'S 146°39'E, Morobe Province, coll. A. C. Zeigler, 7 Aug.1967; 3 males, 2 females, AM W31470, from the small intestine of Melomys rufescens, Munimun Village, Aguan, 9°53'S 149°23'E Sanduan Province, coll. P. German, 8 Aug.1992.</p> <p>Prevalence. – Four of 25 M. rufescens and one of 10 Melomys spp.</p> <p>Etymology. – The species name indicates that whilst having a unique set of characters there are no singular morphological features to highlight.</p> <p>Description. – Small coiled worms; prominent cephalic vesicle present; buccal capsule vestigial. Mouth opening triangular with rudimentary lips; labial and cephalic papillae not observed. Oesophagus claviform. Nerve ring surrounds oesophagus in middle third, deirids and excretory pore at about same level, posterior to nerve ring. Synlophe of longitudinal cuticular ridges extends from the posterior margin of the cephalic vesicle to just anterior to the bursa or vulva; 35–37 ridges in anterior, 35 in midbody of male. Axis of orientation from ventral right to dorsal left sides, inclined about 60° from sagittal axis. Ridges 1–16 increasing slightly in size; ridges 1’–10’ increasing in size, ridges 11’–21’ smaller. Posterior region of body with 28–30 ridges reduced in size; 20 ridges dorsal side, 8–10 ridges ventral side.</p> <p>Male (holotype and 9 paratypes): Length 2.1–3.2 (2.5) mm, maximum width 67–100 (84). Cephalic vesicle 39.5–49.5 (43) long, 23–29.5 (27) wide. Oesophagus 320–520 (390) long; nerve ring 130, excretory pore 180, deirids 180 from cephalic end. Bursa slightly asymmetrical, right lobe larger, rays of right lobe more robust; without dorsal median dorsal notch. Dorsal ray symmetrical, divided at about half its length, each branch dividing again at distal tip; terminal divisions, rays 9, 10 symmetrical; rays 8 arising at same level proximally to division of dorsal ray, right ray 8 slightly more robust than left. Rays 4, 5, 6 with common stem, reaching margin of bursa, rays 6 curving anteriorly, rays 4, 5 curving posteriorly. Rays 2, 3 with common stem, diverging distally, curving posteriorly, reaching margins of bursa. Genital cone short, lightly sclerotized, with papillae 0 and 7. Spicules equal, filiform, tips simple, sharply curved, 340-450 (390) long. Gubernaculum 37–41 (39) long.</p> <p>Female (allotype and 9 paratypes): Length 2.4–3.4 (2.8) mm, maximum width 66–87 (81). Cephalic vesicle 36–52 (43) long, 23–33 (28) wide. Oesophagus 250–520 (370) long; nerve ring 200, excretory pore 250, deirids 250 from cephalic end. Vulva opens 66–72.5 (71) from tail tip. Monodelphic; ovejector with sphincter 40, 40, and vestibule 40, 50, about same length, infundibulum 30, 40, and vagina, 25, 30, shorter. Tail blunt, conical, tip rounded, 16.5–24 (19) long. Eggs thin- shelled, ellipsoidal, in utero 53–63 (57) by 32–33 (32.8).</p> <p>Remarks. – Although the fixation of this material, in situ in the intestine of the host, was not ideal it was sufficient to allow adequate examination of the specimens. Odilia similis, new species, has every morphological character of the genus Odilia (Mawson, 1961) (see Durette-Desset, 1983). Following the key to the species of Odilia of Smales (2005b), O. similis, new species, with 35 ridges in the midbody of the male, falls between the group of species with up to 35 ridges, O. maxomyos Hasegawa et al. 1999, and O. praeputialis Gibbons &amp; Spratt, 1995, and the group with more than 35 ridges in the midbody, O. polyrhabdote (Mawson, 1961) and O. uromyos (Mawson, 1961). Odilia similis, new species, further differs from each of these species in having a spicule to body length ratio of 1: 6.5, simple pointed spicule tips and a female tail blunt, conical, neither twisted nor with a praepuce (Smales, 2005b).</p> </div>	https://treatment.plazi.org/id/8B20878DFFD7FFCCE032FBEFFAC7F8E9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Smales, L. R.	Smales, L. R. (2009): Helminths Of Melomys Rufescens And Melomys Spp. (Muridae: Hydromyinae) From Papua New Guinea With The Descriptions Of A New Genus And Five New Species In The Heligmonellidae (Nematoda: Trichostrongyloidea). Raffles Bulletin of Zoology 57 (1): 5-15, DOI: 10.5281/zenodo.5341456
8B20878DFFD9FFC2E1B0FC08FBE5FA7E.text	8B20878DFFD9FFC2E1B0FC08FBE5FA7E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paraheligmonelloides singauwaensis Smales 2009	<div><p>Paraheligmonelloides singauwaensis, new species</p> <p>(Figs. 44–53)</p> <p>Material examined. – Holotype male, allotype female: BBM-NG- 24762 from the small intestine of Melomys rufescens (Muridae: Hydromyinae), type host, Singauwa River, Lae, 6°07'S 146°59'E, Morobe Province, Papua New Guinea, coll. P. J. Shanahan, 7 Apr.1966.</p> <p>Paratypes: 3 males, BBM-NG-24762, collecting data as above.</p> <p>Prevalence. – Two of 25 M. rufescens.</p> <p>Etymology. – The species name is taken from the type locality.</p> <p>Description. – Small coiled worms; prominent cephalic vesicle present; buccal capsule vestigial. Mouth opening triangular with rudimentary lips; labial and cephalic papillae not observed. Oesophagus claviform. Nerve ring surrounds oesophagus in middle third; deirids and excretory pore at about same level, in posterior third of oesophagus length. Synlophe of pointed longitudinal cuticular ridges extends from the posterior margin of the cephalic vesicle to just anterior to bursa or vulva; 17 ridges in anterior, 20–22 in midbody. Axis of orientation of ridges passing from ventral right to dorsal left sides, inclined about 80° from sagittal axis; 8–11 ridges dorsal side, 9–13 ridges ventral side. Ridge 1’ more developed than ridge 1 in anterior; ridges 1’, 2’ more developed than ridges 1, 2 in midbody. Ridges 1–11 increasing in size; ridges 3’–5’ increasing in size, ridges 7’–11’ decreasing in size. Posterior region of body with 22–23 (male) 20–21 (female) ridges reducing in size, retaining axis of orientation.</p> <p>Male (holotype and 4 paratypes): Length 2.35–2.75 (2.6) mm, maximum width 99–100 (99). Cephalic vesicle 37.5–42.5 (40) long, 30.5–34 (32.5) wide. Oesophagus 320––350 (345) long; nerve ring 165, excretory pore 240 from cephalic end. Bursa asymmetrical, right lobe larger, rays of left lobe more robust; with small median dorsal notch. Dorsal ray symmetrical, divided at about 2 thirds its length, each branch dividing again at distal tip; terminal divisions, rays 9, 10 symmetrical; rays 8 arising at slightly different levels proximally to division of dorsal ray. Rays 4, 5, 6 with common stem, reaching margin of bursa, rays 6 more slender than rays 4, 5. Rays 2, 3 with common stem, robust, diverging distally, curving posteriorly, reaching margin of bursa. Genital cone short, lightly sclerotized; ventral lobe with unpaired papillae 0, dorsal lip bifid, each lobe with single papilla 7. Spicules equal, filiform, proximal ends rounded, tips simple pointed, 230–290 (265) long. Gubernaculum 34–35.5 (35) long.</p> <p>Female (allotype): Length 2.9 mm, maximum width 130. Cephalic vesicle 42.5 long, 34 wide. Oesophagus 390 long. Vulva opens 95 from tail tip. Posterior end reflected dorsally; body wall extended to form praepuce. Monodelphic; ovejector with sphincter, 80, longest, infundibulum, 50, vestibule, 30, smallest element. Tail conical tip with spike. Eggs thinshelled, ellipsoidal, in utero 45 by 30.</p> <p>Remarks. – Paraheligmonelloides singauwaensis, new species, is characterized by having a synlophe with ridge 1´much more developed than ridge 1, and with the relative sizes and orientation of the ridges as described for the southeast Asian members of the genus (Ow Yang et al., 1983; Hasegawa et al., 1999). Of these P. singauwaensis, new species, most closely resembles Paraheligmonelloides triangulus Ow Yang et al., 1983 in the number of ridges on the synlophe. It differs from P. triangulus in having more ridges in the midbody (20–22 ridges compared with 19–21 ridges), as well as in the length of the spicules (230–290 compared with 61–85) the form of the spicule tips and the size of the eggs (45 by 30 compared with 67 by 36) (Ow Yang et al., 1983).</p> </div>	https://treatment.plazi.org/id/8B20878DFFD9FFC2E1B0FC08FBE5FA7E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Smales, L. R.	Smales, L. R. (2009): Helminths Of Melomys Rufescens And Melomys Spp. (Muridae: Hydromyinae) From Papua New Guinea With The Descriptions Of A New Genus And Five New Species In The Heligmonellidae (Nematoda: Trichostrongyloidea). Raffles Bulletin of Zoology 57 (1): 5-15, DOI: 10.5281/zenodo.5341456
