taxonID	type	description	language	source
89238788841AFFECFF08FB709D86FD57.taxon	diagnosis	Diagnosis. This species can be distinguished from other species in the subgenus based on the overall body size, shape and length of the male AGP, shape of the medial lobes of tergum VIII (pseudoparameres), and shape of the posterior margin of the female SGP. Specifically, the body length is ≤ 6.8 mm. The AGP is generally narrow at the base, curving abruptly to the right at a nearly 90 degree angle, and is slightly expanded distally (Fig. 3 B). The pseudoparameres are short, symmetrical, subrectangular, with the posterior margin nearly straight, the posterolateral corners rounded, and with a pile of scattered fine setae (Fig. 3 C). The parameres are rounded with a row of long setae emanating from the middle of the dorsal surface (Fig. 3 D). The right ventral lobe of the endosoma is lightly sclerotized (Fig. 3 E). The posterior margin of the SGP has a deep, rounded central concavity almost twice as wide as deep (Fig. 3 F).	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
89238788841AFFECFF08FB709D86FD57.taxon	discussion	Discussion. This species was described from a series of specimens collected in western Mexico. The authors placed this species in the same subgenus (Syncollus) as A. circumcinctus and A. pygmaeus and indicated that it had a similar color pattern as that of the latter species, but was smaller when compared with both species (Polhemus & Polhemus 1981). The original description listed the length of the holotype at 5.75 mm, but it failed to provide a size range for paratypes. The length of our specimens is 5.20 – 6.80 mm. This is the smallest known of all species of Ambrysus in Mexico. The ground color is brown with the head, prothorax, and lateral corners of the scutellum light brown. The hemelytra are medium brown and patterned with light brown on the anterior 2 / 3 of the embolia and a conspicuous light brown triangular mark at the base of the posterior margin of the corium. The exposed connexiva are concolorous light brown (Fig. 3 A).	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
89238788841AFFECFF08FB709D86FD57.taxon	description	Variation. The color pattern on specimens distributed in southern and eastern Mexico is different from that of western populations. Specimens from western states (Oaxaca to Nayarit) posses the same color pattern reported in the type series. In general, specimens from southern and eastern Mexico (Chiapas, northern Oaxaca, San Luis Potosí, Veracruz) present a darker color pattern on the prothorax, with the hemelytra dark brown and a small, inconspicuous light brown triangular mark at the base of the posterior margin of the corium. Also, the exposed connexiva present an alternating brown and light brown checkered appearance. A single population from Michoacán (Mpio. Apatzingán) in the west presented a similar color pattern to that described from eastern populations. Structure of the AGP is variable; some specimens have a wider base and some have the posterior margin produced posteriorly, giving the impression of less abrupt curvature. Males from eastern Mexico generally present pseudoparameres with posterolateral corners that are more gradually angled as in A. chiapanecus n. sp. The SGP presents considerable variation in the shape and depth of the posterior margin. Some specimens from Jalisco presented an inverted V-shaped concavity, whereas others from Oaxaca presented a shallow concavity. The posterolateral corners of the SGP are narrow and pointed in the allotype, but we have observed variation from that of the allotype to wide and rounded posterolateral corners.	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
89238788841AFFECFF08FB709D86FD57.taxon	materials_examined	Type locality. Mexico: Nayarit: Mpio. San Blas, Aticama. Repository. Holotype is housed at USNM.	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
89238788841AFFECFF08FB709D86FD57.taxon	distribution	Distribution. This species has been reported from Belize and three western states in the Mexican Pacific Coast biogeographic province. Based on our collections, A. baeus has a characteristic distribution pattern that includes the tropical lowlands of the Mexican Pacific Coast (Chiapas, Colima, Jalisco, Michoacán, Nayarit, southern Oaxaca) and the Mexican Gulf (northern Oaxaca, San Luis Potosí, Veracruz) biogeographic provinces. Although we neither collected this species nor found museum specimens that were collected in the state of Guerrero, it is probably present in the coastal and the Balsas Basin regions of Guerrero. We collected A. baeus and A. circumcinctus syntopically at L- 1342, L- 1371, L- 1488, L- 1661, L- 1801, L- 1802, L- 1891, and L- 1900. Published records. Belize: Cayo (Sites & Shepard 2015). Mexico: Colima, Jalisco, Nayarit (Polhemus & Polhemus 1981).	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
89238788841AFFECFF08FB709D86FD57.taxon	materials_examined	Type material examined. HOLOTYPE ♂: Mex [ico]: Nay [arit]: [Mpio. San Blas], 5 ' Aticama, CL 727 A, VI- 7 - 1975, J. T. Polhemus / HOLOTYPE Ambrysus baeus J. T. Polhemus [&] D. A. Polhemus. PARATYPES: 5 ' Aticama, CL 727 A, VI- 7 - 1975, J. T. Polhemus / PARATYPE Ambrysus baeus J. T. Polhemus & D. A. Polhemus (4 ³, 3 ♀ CAS; 1 ♀ CNIN; 1 ³, 1 ♀ EMEC; 2 ³, 2 ♀ USNM). Material examined. MÉXICO: CHIAPAS: Mpio. Benemérito de las Américas, Flor de Cacao, tributario del Río Salinas, 16 ° 09 ' 24.4 '' N, 90 ° 29 ' 38 '' W, 140 m, 30 - Mar- 2015, DRV, Sites, Shepard, Barr, & PRH colls., L- 1891 (1 ³, 1 ♀ UMC); Mpio. Jiquipilas, Rancho Bonito, Pte. Rancho Bonito, 16 ° 22 ' 06 '' N, 93 ° 52 ' 04.2 '' W, 689 m, 03 - Jun- 2014, DRV & PRH colls., L- 1814 (1 ³ UMC); Mpio. La Concordia, El Altillo, Río El Dorado, 16 º 06 ' 27.5 '' N, 92 º 55 ' 29.3 '' W, 524 m, 18 - May- 2012, DRV & PRH colls., L- 1342 (1 ♀ UMC); [Mpio. Metapa], Metapa, Río Suchiate, 25 - III- 85, F. Arias (1 ³, 3 ♀ CNIN); same locality information but M. Vertiz (1 ♀ CNIN); same locality information but H. Velasco (3 ³, 2 ♀ CNIN); [Mpio. Tapachula], Tapachula, Río Huehuetan, 16 - III- 85, F. Arias (2 ♀ CNIN); Mpio. Villa Comaltitlán, Chapingo Dos, Pte. Vado Ancho, Río Vado Ancho, 15 ° 14 ' 51.8 '' N, 92 ° 36 ' 02.2 '' W, 57 m, 01 - Apr- 2015, DRV, Sites, Shepard, Barr, & PRH colls., L- 1902 (1 ♀ UMC); Mpio. Villa Comaltitlán, Lázaro Cárdenas, km 238 carr. Huixtla-Mapastepec, 15 º 10 ' 47.4 '' N, 92 º 32 ' 50.7 '' W, 16 m, 20 - May- 2012, DRV & PRH colls., L- 1353 (1 ♀ UMC); Mpio. Villa Comaltitlán, 1.36 km SE of Villa Comaltitlán, 15 ° 12 ' 45.8 '' N, 92 ° 33 ' 46.1 '' W, 48 m, 31 - Mar- 2015, DRV, Sites, Shepard, Barr, & PRH colls., L- 1900 (1 ³ UMC); Mpio. Villa Corzo, Pte. El Carmen, nr. Revolución Mexicana, 16 º 07 ' 51.4 '' N, 93 º 0.2 ' 00.9 '' W, 547 m, 08 - Jan- 2012, DRV coll., L- 1318 (1 ♀ UMC). COLIMA: [Mpio. Manzanillo], Pte. El Carrizo at Hwy. 98, 3 rd. km NE [SE] Camotlán, 7 Jan. 2005, C. B. Barr / 19 ° 13.00 ' N, 104 ° 14.66 ' W, elevation 1250 ft. (1 ♀ EMEC). JALISCO: Las Juntas Verano, 17 I 2005, Río Guapinole / William D. Shepard, leg. (3 ³, 1 ♀ EMEC); Mpio. Cabo Corrientes, Pte. Horcones, 20 º 27.548 ' N, 105 º 17.519 ' W, 330 m, 19 - Mar- 2014, J. A. Gómez, S. Smith, & R. Novelo colls. (11 ³, 5 ♀ UMC); Mpio. Tomatlán, Tomatlán, Río Tomatlán, 19 ° 56 ' 41.4 '' N, 105 ° 14 ' 36.1 '' W, 27 m, 25 - Jun- 2013, DRV & PRH colls., L- 1575 (8 ³, 6 ♀ UMC); Punta Hermanos, 5 I 2005, 980 ft, Río Horcones / William D. Shepard, leg. (1 ♀ EMEC); Río Cuitzmala, 2.5 km N of Emiliano Zapata, 6 Jan. 2005, C. B. Barr (4 ³, 2 ♀ EMEC); Río Guapinole, 1.3 rd. km E Las Adjuntas Verano-Las Adjuntas del Tuito, 17 Jan. 2005, C. B. Barr / 20 ° 28.67 ' N, 105 ° 16.45 ' W (1 ³ EMEC); Río Los Horcones, 1.1 rd. km W Las Juntas Verano / Las Juntas del Tuito, 5 Jan. 2005, C. B. Barr (3 ³, 2 ♀ EMEC); Río Purificación, 3.7 rd. km NW La Huerta, 8 Jan. 2005, C. B. Barr / 19 ° 29.50 ' N, 104 ° 39.63 ' W, elevation 740 ft. (1 ³, 1 ♀ EMEC); SE of Chamela, 6 I 2005, 180 ', Río Cuitzmala / William D. Shepard, leg (1 ³, EMEC); 4 km NW la Huerta, 8 I 2005, 740 ', Río Purificación / William D. Shepard, leg. (2 ♀ EMEC); 6.6 km SE Villa Purificación, 8 I 2005, Puente El Amborin / William D. Shepard, leg. (1 ♂, 2 ♀ EMEC). MICHOACÁN: Mpio. Apatzingán, California (arroyo), 21 - Feb- 2005, R. Novelo & Gómez-Anaya colls. (1 ³, 1 ♀ UMC); Mpio. Aquila, Aquila, Pte. Ramal Aquila, 18 º 35 ' 08.1 '' N, 103 º 30 ' 51.8 '' W, 159 m, 25 - Dec- 2012, DRV & PRH colls., L- 1487 (2 ♂, 4 ♀ UMC); Mpio. Morelos, La Placita de Morelos, Pte. La Placita, Río Aquila, 18 º 32 ' 19.9 '' N, 103 º 35 ' 17.9 '' W, 7 m, 26 - Dec- 2012, DRV & PRH colls., L- 1488 (1 ♂, 2 ♀ UMC). NAYARIT: Mpio. San Blas, El Palillo, Pte. Palillo, Arroyo Palillo, 21 ° 38 ' 27.1 '' N, 105 ° 08 ' 30.8 '' W, 43 m, 20 - Dec- 2013, DRV & PRH colls., L- 1661 (2 ♂ UMC). OAXACA: Mpio. Ayotzintepec, San José Mano Márques, 17 º 47 ' 26.2 '' N, 96 º 00 ' 15.3 '' W, 66 m, 24 - May- 2012, DRV & PRH colls., L- 1371 (1 ♀ UMC); Mpio. Candelaria Loxicha, Azulillo, Río Candelaria, 15 º 53 ' 29.6 '' N, 96 º 29 ' 30.8 '' W, 353 m, 26 - May- 2012, DRV & PRH colls., L- 1377 (8 ³, 2 ♀ UMC); Mpio. Candelaria Loxicha, km 207 carr. Oaxaca-San Pedro Pochutla, Candelaria Loxicha, Río San Juan, 15 º 55 ' 32.5 '' N, 96 º 29 ' 12.6 '' W, 400 m, 25 - May- 2012, DRV & PRH colls., L- 1375 (1 ³, 1 ♀ UMC); same but 02 - Apr- 2015, DRV, Sites, Shepard, Barr, & PRH colls., L- 1904 (8 ³, 7 ♀ UMC); Mpio. Candelaria Loxicha, km 206 carr. Oaxaca-San Pedro Pochutla, Candelaria Loxicha, Pte. San Juan, 15 º 55 ' 32.9 '' N, 96 º 29 ' 25 '' W, 403 m, 26 - May- 2012, DRV & PRH colls., L- 1376 (2 ♀ UMC); same but 02 - Apr- 2015, DRV, Sites, Shepard, Barr, & PRH colls., L- 1905 (1 ³ UMC); Mpio. Mártires de Tacubaya, Mártires de Tacubaya, 16 º 34 ' 41.1 '' N, 98 º 13 ' 08.7 '' W, 206 m, 29 - May- 2012, DRV & PRH colls., L- 1390 (15 ³, 12 ♀ UMC); [Mpio. Pluma Hidalgo, El Pacífico] Finca El Pacífico, km 34 carr. Pochutla-Oaxaca, 8 - III- 1995, 550 msnm, N 15 56 675, W 96 27 892, G. Ortega, E. Barrera (1 ♀ CNIN); Mpio. San Gabriel Mixtepec, km 194 carr. Puerto Escondido-Oaxaca, San Gabriel Mixtepec, Río Rana, 16 º 06 ' 06.3 '' N, 97 º 03 ' 52.9 '' W, 712 m, 27 - May- 2012, DRV & PRH colls., L- 1380 (1 ³, 1 ♀ UMC); same but 03 - Apr- 2015, DRV, Sites, Shepard, Barr, & PRH colls., L- 1906 (23 ³, 16 ♀ UMC); Mpio. San Pedro Huamelula, Morro Ayutla, Pte. Río Ayutla, Río Ayutla, 15 ° 54 ' 08.7 '' N, 95 ° 51 ' 13.3 '' W, 29 m, 04 - Jun- 2014, DRV & PRH colls., L- 1816 (1 ³, 1 ♀ UMC); km 179 carr. Sta. Maria Zacatepec-Putla, El Rosario, Pte. El Rosario, 16 º 51 ' 34.6 '' N, 97 º 58 ' 44.7 '' W, 361 m, 29 - May- 2012, DRV & PRH colls., L- 1394 (4 ³, 3 ♀ UMC); km 201 carr. Mártires de Tacubaya-Santa Maria Zacatepec, ca. 13 km NE of San Pedro Amuzgos, 16 º 42 ' 18.9 '' N, 98 º 00 ' 52.9 '' W, 276 m, 29 - May- 2012, DRV & PRH colls., L- 1391 (3 ♂, 2 ♀ UMC). SAN LUIS POTOSÍ: [Mpio. Huehuetlán], Huichihuayán, (Nacim. Río), 23 - Jun- 1990, R. Novelo col. (1 ♂ IEXA). VERACRUZ: [Mpio. Chicontepec], Chicontepec, 1 - 3 - 76, J. Bueno / Ambrysus baeus J. & D. Polhemus, det. J. Polhemus (1 ³ CNIN); Mpio. Paso de Ovejas, Paso de Ovejas, 19 ° 17 ' 04.1 '' N, 96 ° 26 ' 03.7 '' W, 40 m, 29 - May- 2014, DRV & PRH colls., L- 1802 (2 ♀ UMC); Mpio. Puente Nacional, Puente Nacional, Pte. Puente Nacional I, Río La Antigua, 19 ° 19 ' 28.5 '' N, 96 ° 28 ' 54.3 '' W, 67 m, 29 - May- 2014, DRV & PRH colls., L- 1801 (1 ♀ UMC); [Mpio. San Andrés Tuxtla, Los Tuxtlas], Salto [de] Eyipantla, 30 - I- 84, M. García / Fam: Naucoridae, Gen: Ambrysus Det. M. García (1 ♀ CNIN); same locality information but 01 - Apr- 2011, R. Novelo coll. (2 ³, 3 ♀ UMC); Mpio. San Rafael, El Faisán, 20 ° 11 ' 01.7 '' N, 97 ° 05 ' 38.5 '' W, 89 m, 12 - Jun- 2013, DRV, Sites, Shepard & PRH colls., L- 1547 (1 ³, 1 ♀ UMC); Mpio. Tlapacoyan, ca. 4 km SW Vista Hermosa, 19 º 56.891 N, 97 º 0 7.893 W, 261 m, 8 - Nov- 2009, Sites, Cervantes & Novelo colls., L- 1127 (1 ♀ UMC); [Mpio. Tlapacoyan ca. 4.5 km SW Vista Hermosa], waterfall at Rancho Nuevo, 19 º 56.308 ' N, 97 º 0 7.793 W, 270 m, 8 - Nov- 2009, Sites, Cervantes, & Novelo colls., L- 1125 (1 ³, 1 ♀ UMC); Mpio. Tlapacoyan, Ixtacuaco (arroyo), 30 - Mar- 2003, R. Novelo col. (4 ♀ UMC); [Mpio. Tlapacoyan], Ixtacuaco, 07 - Noviembre- 2009, R. Novelo col. (2 ³, 1 ♀ IEXA); [Mpio. Tlapacoyan], Ixtacuaco, 6 IV 2011, 103 m, Río Ixtacuaco, N 20 º 02 ', W 97 º 07 ' / William D. Shepard leg. (1 ³, 3 ♀ UMC); [Mpio.] Tlapacoyan, 1 km antes El Cuajilote (ruinas Filobobos), 8 - Nov- 2009, R. Novelo col. (2 ³, 2 ♀ IEXA); 10 mi E Xalapa, Dec. 26, 1958, Menke & Stange / Ira La Rivers collection bequeathed to the California Academy of Sciences - 1978 (1 ♂ CAS).	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888416FFE9FF08FD15985AFB29.taxon	description	Description. Brachypterous male. HOLOTYPE, length 7.36; maximum width 4.40. Paratypes (n = 7), length 7.12 – 7.76 (mean = 7.37); maximum width 4.16 – 4.56 (mean = 4.29). General shape elongate, parallel-sided; widest across embolia (Fig. 5 A). Dorsal coloration of hemelytra brown, with light brown along midline and posterior and lateral margins of corium; head, pronotum, and legs yellowish brown. Dorsal surface coarsely punctate. Ventral coloration of head and thorax yellowish brown, abdomen with dense golden brown pubescence. Head. Head length 1.44; maximum width 2.04. Mostly yellowish brown with medium brown posteriorly, coarsely punctate. Eyes convergent anteriorly, synthlipsis 0.96; thin band of cuticle along posterolateral margin of eye; eyes not raised above level of vertex or pronotum. Anterior margin between eyes slightly convex, extending anteriorly in front of eyes 11 % of head length; posterior margin between eyes strongly convex, extending posteriorly 39 % of head length. Labrum width 1.7 × length, evenly rounded. Labium with three visible yellowish brown segments, darkening distally, extending 0.34 beyond labrum not including extruded stylets. Antennal proportions 2: 6: 7: 7, length 0.48, extending to near lateral margin of eye, elongate hairs on segment 4 and distal half of 3. Thorax. Pronotum coarsely punctate, ground color yellowish brown; transverse sulcus marking anterior border of transverse band in posterior 1 / 4; transverse band pale yellow; ovoid light brown mark on either side of midline at anterior margin; dark submarginal band extending from eye to transverse band; lateral margins dark brown, convergent, evenly convex, explanate; posterior margin straight; anterior margin deeply concave between eyes to embrace convex posterior margin of head; posterolateral corners rounded; width 2.9 × length; length at midline 1.24; maximum width at posterolateral corners 3.60. Prothorax ventrally pruinose throughout; apices of propleura meeting at midline, closely appressed to prosternellum; propleuron yellowish throughout, medial 1 / 3 of posterior margin with elongate golden setae. Probasisternum with sharp median carina and without row of setae lateral to carina. Prosternellum covered by apices of propleura. Scutellum coarsely punctate, triangular, brown, light brown coloration along margins and midline, width 2.0 × length, width 2.52, length 1.26. Hemelytra dark brown, densely punctate, not reaching apex of tergum V, length 5.12 (chord measurement); corium with light brown narrow oblique line at middle and posterior and lateral margins. Clavus with thin yellow stripe at base and apex, length 4.3 × width, length 2.80, width 0.64; claval commissure brown, length 0.80. Embolium length 2.20, greatest width 0.54; lateral margin convex, yellow brown in anterior 2 / 3, becoming darker posteriorly. Oblique suture connecting claval and embolar sutures near bases. Hind wings extending to posterior margin of tergum III. Mesobasisternum midventral tumescence negligible, with sulcus on midline continuing through triangular mesosternellum. Metasternellum (= metaxyphus) transverse, subtriangular, with median carina, apex acute. Legs. All legs segments yellowish brown. Profemur posterior margin with row of tightly arranged setae in basal 2 / 3, row of short brown spines along middle third, spines generally single proximally becoming small combs of two to four spines distally; anterior margin with dense pad of setae without associated spines. Protibia and tarsus with occlusal inner surface flattened and with spatulate setae; tarsus immovable, one-segmented; pretarsal claw single, minute, triangular. Procoxa with cluster of stout, brown anteromedial spines. Meso- and metacoxae partially recessed into thorax. Meso- and metafemora with row of short, brown spines on anterior margin; spines restricted to basal half on mesofemur, nearly full length of metafemur. Mesotibia with ventrolateral, ventromedial, dorsolateral, and dorsomedial rows of stout reddish-brown spines; ventromedial rows intermixed with combs of two to five spines. Metatibia with ventrolateral, dorsolateral, and dorsomedial rows of stout reddish-brown spines. Meso- and metatibiae with semi-circlet of spines at apex of dorsal and mesal margins; two comb rows of stout spines near apex of ventral margin, lateral margin without spines at apex to accommodate flexed tarsus. Meso- and metatibiae and metatarsus with long, pale swimming hairs; hairs profuse on metatibia and - tarsus. Meso- and metapretarsi with paired claws slender, gently curved, with small basal tooth. Leg measurements as follows: foreleg, femur 2.08, tibia 1.72, tarsus 0.40; middle leg, femur 2.04, tibia 1.88, tarsomeres 1 – 3, 0.16, 0.34, 0.44; hind leg, femur 2.48, tibia 2.84, tarsomeres 1 – 3, 0.24, 0.46, 0.56. Abdomen. Dorsally with connexiva III – VIII exposed, laterotergites light brown (Fig. 5 A); lateral margin finely serrate, marginal row of short yellow setae, group of trichobothria near posterolateral corners. Posterolateral corners of II (visible ventrally) – IV narrowly rounded to right angled and not spinose, V – VII bluntly acute. Accessory genitalic process of tergum VI short, curved to right at approximately 60 degrees, dilated distally (Fig. 5 B). Medial lobes of tergum VIII (pseudoparameres) short, symmetrical, posterior margin straight to slightly convex, posterolateral corners roundedly angled ca. 45 degrees (Fig. 5 C). Ventrally entirely golden brown, with dense pile of fine hairs. Lateral margin with thin, glabrous band. Glabrous rounded to oval patches around spiracles on laterosternites II – VII. Phallosoma elongate, linear, constricted basally, sclerotized dorsally; right and left ventral lobes of endosoma membranous and papillose, both lightly sclerotized (Fig. 5 E). Parameres symmetrical, almost as long as wide, mesal margin straight, row of long dark setae emanating from middle of dorsal surface. Proctiger short, length 0.6 × width. Pygophore with elongate setae sparsely distributed over most of surface, with thick brush of elongate setae on posterior margin (Fig. 5 D). Brachypterous female. Paratypes (n = 8), length 7.68 – 8.24 (mean = 7.90); maximum width 4.40 – 4.64 (mean = 4.58). Similar to male in general structure and coloration except as follows: Hemelytra surpassing posterior margin of tergum VI. Mediosternite VII (subgenital plate) width 1.5 × length; length at midline 0.84; maximum width 1.30; posterior margin straight with small central concavity; posterolateral corners rounded, slightly produced caudad (Fig. 5 F). Macropterous male. Paratypes (n = 2), length 7.76 – 7.92 (mean = 7.84); maximum width 4.56. Setation, posterolateral corners of pronotum, scutellum proportions, embolium length generally same as for brachypterous male. Otherwise head with dark brown stripe at midline becoming wider posteriorly. Scutellum with light brown on anterior corners. Hemelytra solid black, surpassing apex of tergum V, length 5.92 (chord measurement). Clavus without thin, yellow stripe at apex, length 4.3 × width, length 2.60, width 0.60; claval commissure length 0.76. Hind wings extending to posterior margin of tergum V. Each laterotergite dark brown in anterior half and light brown posteriorly giving checkered appearance. Macropterous female. Unknown. Variation. Some brachypterous specimens from Guatemala and the four localities in Mexico presented a similar dark color pattern to that described for the macropterous male, although one of the dark brachypterous males also exhibited a thin, yellow stripe at the apex of the clavus as in the holotype. The posterolateral corners of the female subgenital plate can be slightly pointed.	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888416FFE9FF08FD15985AFB29.taxon	diagnosis	Diagnosis. This species presents a characteristic color pattern on the hemelytra with light brown along the middle and along the posterior and lateral margins of the corium that is not present in any other species in the subgenus. The pseudoparameres are noticeably short with the posterolateral corners broadly rounded at a 45 degree angle. The posterior margin of the female subgenital plate has a conspicuous, small, central concavity. Habitat description. The source of the Tzaconejá River is in the western mountains in central Chiapas. The river travels southeastward along the Lacantún Basin to drain into the Lacantún River, close to the border with Guatemala. At the type locality the river channel is wide (ca. 20 – 30 m) with limestone cobble and bedrock (Fig. 6). The collecting was conducted in riffles and in shallow areas with current by disturbing the substrate and holding the net downstream. This method was particularly difficult at the type locality because mineral deposition had cemented cobbles to the bedrock.	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888416FFE9FF08FD15985AFB29.taxon	distribution	Distribution. This species was collected from rivers in the highlands and western mountain regions of Chiapas state at elevations above 1000 meters, which are part of the Chiapas biogeographic province. We suspect this species also is present in streams of the western mountains of the Sierra Madre de Chiapas and the easternmost limit (Oaxaca) of the Sierra Madre del Sur biogeographic provinces. In Guatemala the species was collected in the central mountains of the Sierra de los Cochumatanes (Fig. 4).	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888416FFE9FF08FD15985AFB29.taxon	etymology	Etymology. The specific epithet refers to the southern Mexican state of Chiapas, where we collected this species, an area that harbors great biodiversity. Repositories. The holotype and some paratypes will be deposited in the Colección Nacional de Insectos (Mexico City); additional paratypes will be deposited in the Enns Entomology Museum (University of Missouri), United States National Museum of Natural History (Washington D. C.), and Washington State University (Pullman).	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888416FFE9FF08FD15985AFB29.taxon	materials_examined	Type material examined (brachypterous unless otherwise noted). HOLOTYPE ♂: MÉXICO: CHIAPAS: Mpio. Altamirano, Cuauhtemoc, Balneario Tzaconejá, Río Tzaconejá, 28 - Mar- 2015, 1172 m, L- 1886 / 16 º 41 ' 17.5 '' N, 92 º 01 ' 03.0 '' W, Reynoso, Sites, Shepard, Barr, & Reynoso-Hdez. colls. PARATYPES: same data as holotype (1 ♀ CNIN; 1 ³ macropterous, 1 ³, 2 ♀ UMC; 1 ♀ USNM); [Mpio. Bochil], 5 mi N of Bochil, CL 1090, 4 May 1964, J. T. & M. S. Polhemus (1 ♀ CNIN; 1 ³ macropterous, 1 ♀ UMC); same but / Ambrysus pygmaeus La R. det. J. T. Polhemus (1 ³ CNIN); Mpio. Ixtapa, Pte. Río Lajas, Río Lajas, 17 - May- 2012, 1020 m, L- 1340 / 16 50 ' 42.6 '' N, 92 54 ' 01.7 '' W, Reynoso-Velasco & Reynoso-Hernández colls. (1 ♀ UMC); same locality information but 28 - Mar- 2015, L- 1885 / Reynoso, Sites, Shepard, Barr, & Reynoso-Hdez. colls (1 ³ CNIN; 1 ³ UMC; 1 ³ USNM). GUATEMALA: BAJA VERAPAZ: Rio Quilila, Rt 5, old rd to Salama from CA- 14, 1355 m, N 15 º 13.913 ', W 90 º 17.662 ', 15 Sept 2009, R. S. Zack & J. Monzon collectors (1 ♂, 1 ♀ WSUC).	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888413FFF0FF08FA829E34F836.taxon	diagnosis	Diagnosis. This species can be distinguished from congeners based on the shape and length of the AGP, as well as the shape of the pseudoparameres. In general, the AGP presents the characteristics described by La Rivers (1951): moderately long, parallel-sided, noncapitate, blunt at tip, and uniformly curved to the right at approximately 45 degree angle (Fig. 7 B). The pseudoparameres are relatively long, rounded and slightly asymmetrical, with the right pseudoparamere moderately narrower toward the apex; with a pile of fine setae mostly on the mesal margins (Fig. 7 C). The parameres are symmetrical, longer than wide, with the mesal margin straight, and with a row of long setae emanating from the middle of the dorsal surface (Fig. 7 D). The left ventral lobe of the endosoma is lightly sclerotized (Fig. 7 E). The posterior margin of the SGP is highly variable, but in general it has a shallow concavity with a central notched lobe, and rounded to squared posterolateral corners (Fig. 7 F).	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888413FFF0FF08FA829E34F836.taxon	discussion	Discussion. This species originally was described from a single female specimen from Kerrville, Texas (Montandon 1910) and later assigned as the type species of the subgenus Syncollus by La Rivers (1965). Usinger (1946) described the species A. caliginosus from central Mexico, which was later considered by La Rivers (1953) to be a subspecies of A. circumcinctus based on the relatively larger size and darker coloration. Later, La Rivers (1967) described two more subspecies: Ambrysus c. extremus from Honduras and Costa Rica with the diagnostic feature of the median lobe in the center of the posterior margin of the SGP conspicuously produced beyond the base of the concavity; and A. c. concavus, described from central Mexico, with a deeper concavity in the posterior margin of the SGP and the median lobe barely visible. In contrast, the nominal subspecies had a nearly straight posterior margin of the SGP with a short median lobe. The characteristics of the AGP reported for A. circumcinctus (La Rivers 1951) were the same reported to be present in A. c. concavus and A. c. caliginosus, whereas A. c. extremus had a distally expanded AGP (La Rivers 1967). Considering that a subspecies is a geographical variety (Mayr 1982) with morphological uniformity in one or a few diagnostic characters (Wilson & Brown 1953), the four subspecies of A. circumcinctus should be identifiable and associated to a particular geographic area based on the shape of the posterior margin of the subgenital plate (La Rivers 1967). This is not the case in part because the subspecies described by La Rivers were based on few specimens from disparate localities and he was not able to assess the variation in subgenital plate shape. We have studied ca. 3,000 specimens from approximately 300 localities from the U. S. to Honduras; thus, this massive amount of material has enabled us to assess intraspecific phenotypic plasticity of this feature. What La Rivers considered a stable and consistent feature was an artifact of the small number of specimens and localities used to describe the subspecies. At various localities in the range of A. circumcinctus, populations exhibit the characteristics of more than one described subspecies. In one example of many, at site L- 1350 in Chiapas (southern Mexico) variation in subgenital plate shape includes individuals consistent with that of both A. c. extremus and A. c. circumcinctus. The latter subspecies purportedly occurs only in the United States and northern Mexico. Because we have examples of more than one subspecific morphological form occurring in various populations and occurrence of the morphological forms does not conform to the latitudinal geographic organization suggested by La Rivers (1967), we consider these subspecies to be consubspecific. As such, we propose A. c. caliginosus, A. c. concavus, and A. c. extremus to be junior synonyms of A. c. circumcinctus.	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888413FFF0FF08FA829E34F836.taxon	description	Variation. The coloration of the hemelytra reported for A. circumcinctus was uniformly brownish black with embolia predominantly light yellow, and dark only posteriorly (La Rivers 1951). The coloration of specimens collected in the southern United States was lighter when compared with specimens from central Mexico (La Rivers 1953) and present a conspicuous, light brown triangular mark at the base of the posterior margin of the corium. The coloration of the hemelytra of specimens collected throughout Mexico is solid dark brown with the triangular mark inconspicuous and light brown to reddish brown (Fig. 7 A), although some specimens collected in Jalisco, Michoacán, and Nayarit posses a conspicuous triangular mark similar to that present in specimens from the southern United States. A similar mark also is present in specimens of A. baeus distributed in western Mexico. Minor variation in the length of the AGP also was observed. The posterior margin of the SGP showed a high degree of phenotypic plasticity that included the posterior margin with a: a) shallow concavity with central notched lobe, b) shallow concavity with basally constricted central notched lobe, c) deeper concavity with inconspicuous central notched lobe, and d) posterior margin almost straight (without concavity) with inconspicuous central notched lobe.	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888413FFF0FF08FA829E34F836.taxon	materials_examined	Type locality. United States: Texas: Kerr Co., Kerrville. Repository. Holotype is housed at USNM.	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888413FFF0FF08FA829E34F836.taxon	distribution	Distribution. This species has the largest range among species in this subgenus in North America. It is distributed from the southern United States south to Costa Rica. In Mexico, it is widely distributed with representatives in most of the biogeographic provinces from the Nearctic and Neotropical regions, except the Californian and Baja Californian provinces where the species most likely does not occur (Fig. 8). It also is present in the provinces of the Mexican Transition Zone; however, at high elevations it is replaced by other closely related species (i. e., A. pygmaeus, A. totonacus n. sp., A. xico n. sp.). We found A. circumcinctus and A. baeus occupying the same habitat at various localities within the ranges of both species (L- 1342, L- 1371, L- 1488, L- 1661, L- 1801, L- 1802, L- 1891, L- 1900), and with A. totonacus n. sp. in the states of Hidalgo (L- 1522) and Veracruz (L- 1290, L- 1296, L- 1549). Published records. Belize: Cayo, Orange Walk (Sites & Reynoso-Velasco 2015, Sites & Shepard 2015). Costa Rica: San José (La Rivers 1967, 1971; Herrera 2013). Guatemala (Herrera 2013). Honduras: Atlántida, El Paraíso, Olancho (La Rivers 1967, 1971; Herrera 2013). Mexico: Chiapas, Estado de México, Nuevo León, San Luis Potosí, Sinaloa (La Rivers 1953, 1965, 1967, 1971; Davis 1986; Lee 1991; Herrera 2013). United States: Arizona, New Mexico, Texas (Montandon 1910; La Rivers 1951, 1953, 1965, 1967, 1971; Davis 1986; Herrera 2013).	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888413FFF0FF08FA829E34F836.taxon	materials_examined	Type material examined. HOLOTYPE ♀: UNITED STATES: [Texas: Kerr Co.], Kerrville, vi. 19.07 TX / F. C. Pratt Collector / From Coll. U. S. N. M. / 0 52 / Ambrysus circumcinctus Montandon Type 1909 / type No. 54270 U. S. N. M. PARATYPES: MÉXICO: MÉXICO: Tejupilco, Mex., Temescaltepec [Temascaltepec], VI 18 33 / H. E. Hinton, R. L. Usinger Collectors / PARATYPE Ambrysus caligimosus R. L. Usinger (4 ♀ EMEC); same but / Ambrysus circumcinctus caliginosus determined by Ira La Rivers ' 50 (1 ♀ EMEC); Tejupilco, Mex., Temescaltepec [Temascaltepec], VI 21 33 / H. E. Hinton, R. L. Usinger / PARATYPE Ambrysus caliginosus R. L. Usinger (1 ♂ EMEC). Material examined. MÉXICO: CHIAPAS: Mpio. Amatengo de la Frontera, Barrio Sabinalito, Río Agua Caliente, 15 ° 25 ' 0.1 '' N, 92 ° 08 ' 33.7 '' W, 942 m, 31 - Mar- 2015, DRV, Sites, Shepard, Barr, & PRH colls., L- 1898 (1 ♀ UMC); Mpio. Ángel Albino Corzo, Jaltenango, Río Jaltenango (Río Lagartero), 15 º 52 ' 33.7 '' N, 92 º 43 ' 53.4 '' W, 605 m, 06 - Jan- 2012, DRV coll., L- 1313 (24 ³, 20 ♀ UMC); same but 07 - Jan- 2012, L- 1314 (46 ³, 45 ♀ UMC); Mpio. Ángel Albino Corzo, km 30 carr. Jaltenango-Siltepec, Plan de Ayutla, Río El Plan de Ayutla, 15 º 42 ' 23.6 '' N, 92 º 34 ' 43.7 '' W, 846 m, 19 - May- 2012, DRV & PRH colls., L- 1346 (5 ³, 4 ♀ UMC); Mpio. Ángel Albino Corzo, 1 km S of Finca Prusia, Río Prusia, 15 º 43 ' 23.3 '' N, 92 º 47 ' 46.5 '' W, 1010 m, 18 - May- 2012, DRV & PRH colls., L- 1345 (1 ³ UMC); Mpio. Benemérito de las Américas, Flor de Cacao, tributario del Río Salinas, 16 ° 09 ' 24.4 '' N, 90 ° 29 ' 38 '' W, 140 m, 30 - Mar- 2015, DRV, Sites, Shepard, Barr, & PRH colls., L- 1891 (³ 1 UMC); Mpio. Frontera Comalapa, El Caracol, 8 km S of Frontera Comalapa, Río Grijalva, 15 ° 36 ' 07.0 '' N, 92 ° 09 ' 14.5 '' W, 677 m, 31 - Mar - 2015, DRV, Sites, Shepard, Barr, & PRH colls., L- 1897 (1 ³, 1 ♀ UMC); Mpio. Huixtla, Huixtla, Río Huixtla, 15 ° 08 ' 59.9 '' N, 92 ° 27 ' 04.6 '' W, 67 m, 31 - Mar- 2015, DRV, Sites, Shepard, Barr, & PRH colls., L- 1899 (1 ♂, 4 ♀ UMC); same but 01 - Apr- 2015, L- 1901 (6 ³, 3 ♀ UMC); Mpio. Ixtapa, Ixtapa, 16 º 48 ' 43.7 '' N, 92 º 54 ' 30 '' W, 1030 m, 17 - May- 2012, DRV & PRH colls., L- 1341 (1 ♀ UMC); Mpio. La Concordia, El Altillo, Río El Dorado, 16 º 06 ' 27.5 '' N, 92 º 55 ' 29.3 '' W, 524 m, 18 - May- 2012, DRV & PRH colls., L- 1342 (1 ♀ UMC); Mpio. La Concordia, Pte. Independencia, 16 º 04 ' 25.2 '' N, 92 º 49 ' 45.3 '' W, 547 m, 07 - Jan- 2012, DRV coll., L- 1317 (43 ³, 32 ♀ UMC); Mpio. Maravilla Tenejapa, Ixcán, Pte. Ixcán, Río Ixcán, 16 ° 04 ' 33.6 '' N, 91 ° 04 ' 12.8 '' W, 175 m, 30 - Mar- 2015, DRV, Sites, Shepard, Barr, & PRH colls., L- 1896 (7 ³, 9 ♀ UMC); [Mpio. Metapa], Metapa [de Domínguez], Río Suchiate, 25 - III- 85, M. Vértiz (1 ♀, CNIN); same but / Ambrysus circumcinctus Montd. R. W. Sites det. (1 ♀ CNIN); Mpio. Monte Cristo de Guerrero, El Paraíso, 15 º 39 ' 28.1 '' N, 92 º 36 ' 18.7 '' W, 864 m, 19 - May- 2012, DRV & PRH colls., L- 1347 (1 ³ UMC); Mpio. Ocosingo, Balneario Jataté, Río Jataté, 16 ° 52 ' 20.1 '' N, 92 ° 02 ' 18.9 '' W, 861 m, 28 - Mar- 2015, DRV, Sites, Shepard, Barr, & PRH colls., L- 1887 (1 ♀ UMC); Mpio. Palenque, Ángel Albino Corzo, Balneario, Río Chocolja, 17 ° 17 ' 32 '' N, 91 ° 37 ' 49.5 '' W, 119 m, 29 - Mar- 2015, DRV, Sites, Shepard, Barr, & PRH colls., L- 1888 (17 ³, 13 ♀ UMC); Mpio. Pichucalco, Pte. Rosita, 4 km S of Pichucalco and 800 m S of Río de Janeiro, 17 ° 29 ' 07.2 '' N, 93 ° 06 ' 13.5 '' W, 44 m, 27 - Mar- 2015, DRV, Sites, Shepard, Barr, & PRH colls. L- 1883 (2 ♀ UMC); Mpio. Siltepec, km 72 carr. Jaltenango-Siltepec, nr. Nuevo Galeana, 15 º 35 ' 54.9 '' N, 92 º 32 ' 28.9 '' W, 933 m, 19 - May- 2012, DRV & PRH colls., L- 1349 (36 ³, 25 ♀ UMC); Mpio. Siltepec, km 92 carr. Jaltenango- Siltepec, Campo Aéreo, Río Bravo, 15 º 35 ' 17.9 '' N, 92 º 28 ' 02.3 '' W, 1068 m, 19 - May- 2012, DRV & PRH colls., L- 1350 (32 ³, 31 ♀ UMC); Mpio. Solosuchiapa, El Beneficio, Río La Sierra, 17 ° 21 ' 01.1 '' N, 93 ° 00 ' 21.1 '' W, 303 m, 27 - Mar- 2015, DRV, Sites, Shepard, Barr, & PRH colls., L- 1884 (1 ³, 3 ♀ UMC); Mpio. Solosuchiapa, Solosuchiapa, Río La Sierra, 17 º 25 ' 40.5 '' N, 93 º 01 ' 35.7 '' W, 122 m, 17 - May- 2012, DRV & PRH colls., L- 1339 (2 ³ UMC); [Mpio. Tapachula], Tapachula, Puente Colorado, 31 - VIII- 85, H. Velasco (1 ♀ CNIN); [Mpio. Tecpatán], Chintul [El Progreso], 6 - III- ' 88, R. Barba, E. Barrera, A. Cadena (4 ³, 2 ♀ CNIN); same but / A. c. circumcinctus Montandon det. R. W. Sites (1 ³, 1 ♀ CNIN); Mpio. Villa Comaltitlán, 1.36 km SE of Villa Comaltitlán, 15 ° 12 ' 45.8 '' N, 92 ° 33 ' 46.1 '' W, 48 m, 31 - Mar- 2015, DRV, Sites, Shepard, Barr, & PRH colls., L- 1900 (8 ³, 8 ♀ UMC); same locality information but / William D. Shepard leg. (3 ³, 4 ♀ UMC). CHIHUAHUA: Mpio. Balleza, Balleza, Río de Balleza, 26 ° 55 ' 20.8 '' N, 106 ° 19 ' 50.1 '' W, 1574 m, 30 - Dec- 2013, DRV & PRH colls., L- 1703 (15 ³, 21 ♀ UMC); [Mpio. Camargo], Camargo, Río Conchos, 17 August 1952, J. D. Lattin Cal. Acad. Sci. Coll. / Ira La Rivers collection bequeathed to the California Academy of Sciences - 1978 (♂ 21, 15 ♀ CAS); [Mpio. Camargo], Camargo, CL 1005, 19 April 1964, J. T. & M. S. Polhemus (1 ³ SEMC); same but / Ambrysus circumcinctus Mont. Det. J. T. Polhemus (1 ♀ SEMC); [Mpio. Meoqui], Meoqui, Río San Pedro, 20 August 1952, J. D. Lattin Cal. Acad. Sci. Coll. / Ira La Rivers collection bequeathed to the California Academy of Sciences - 1978 (3 ♂, 3 ♀ CAS). COAHUILA: Mpio. Cuatrociénegas, La Poza Azul, Arroyo La Puerta Colorada, 26 ° 55 ' 48.3 '' N, 102 ° 07 ' 41.5 '' W, 738 m, 16 - May- 2014, DRV & PRH colls., L- 1763 (2 ³, 1 ♀ UMC); Mpio. Cuatrociénegas, San Juan de Boquillas, 26 ° 59 ' 58.0 '' N, 101 ° 53 ' 35.8 '' W, 697 m, 17 - May- 2014, DRV & PRH colls., L- 1764 (6 ³, 1 ♀ UMC); [Mpio. Jimenez], Río San Francisco, 27 km S of Cd. Acuña, April 0 1, 1974, R. R. & F. M. Miller (1 ³ UMMZ); Mpio. Sabinas, Sabinas (at El Vado), Río Sabinas, 27 ° 50 ' 19.5 '' N, 101 ° 07 ' 20.2 '' W, 340 m, 17 - May- 2014, DRV & PRH colls., L- 1765 (1 ³, 2 ♀ UMC); Mpio. Villa de Juaréz, Sección Veintidos Cero, Río Salado, 27 ° 26 ' 36.0 '' N, 100 ° 28 ' 38.9 '' W, 226 m, 18 - May- 2014, DRV & PRH colls., L- 1768 (32 ♂, 21 ♀ UMC); Mpio. Zaragoza, 25 km N of Zaragoza, Pte. Río San Rodrigo, Río San Rodrigo, 28 ° 43 ' 50.5 '' N, 100 ° 54 ' 45.4 '' W, 329 m, 17 - May- 2014, DRV & PRH colls., L- 1766 (11 ³, 24 ♀ UMC); Río Salado de Los Nadadores, at El Carino, 5 - IV- 1961, R. R. Miller et al., UMMZ Fish M 61 - 43 (1 ♀ UMMZ). DURANGO: Mpio. Guanaceví, El Potrero Viejo, Río Sextín, 25 ° 56 ' 28.7 '' N, 105 ° 49 ' 43.0 '' W, 1931 m, 02 - Jan- 2014, DRV & PRH colls., L- 1713 (3 ♀ UMC); Mpio. Nombre de Dios, km 239 carr. Vicente Guerrero-Durango, jct to El Venado, Pte. Melones, Río Mezquital (Río Durango), 23 ° 52 ' 17.3 '' N, 104 ° 16 ' 04.7 '' W, 1731 m, 29 - Jun- 2013, DRV & PRH colls., L- 1591 (3 ³, 4 ♀ UMC); same but 04 - Jan- 2014, L- 1721 (4 ³, 8 ♀ UMC); [Mpio. Peñón Blanco], Peñón Blanco (Belem), 10 - VII- 85, E. Barrera (1 ³, 1 ♀ CNIN); [Mpio. Peñón Blanco], Peñón Blanco, 10 - VII- 85, J. Bueno (7 ³, 3 ♀ CNIN); same but / Ambrysus circumcinctus Montandon det. J. T. Polhemus 19 [??] (2 ³, 1 ♀ CNIN); Mpio. San Bernardo, San Bernardo, Río Sextín (Río Nazas), 26 ° 00 ' 05.0 '' N, 105 ° 30 ' 17.6 '' W, 1607 m, 01 - Jan- 2014, DRV & PRH colls., L- 1711 (13 ³, 9 ♀ UMC); Mpio. Santa Maria de Tepehuanes, Los Corrales, 25 ° 12 ' 13.4 '' N, 105 ° 34 ' 22.6 '' W, 1726 m, 03 - Jan- 2014, DRV & PRH colls., L- 1717 (20 ³, 26 ♀ UMC); Mpio. Villa de Ocampo, Villa de Ocampo, Río Florido, 26 ° 26 ' 07.2 '' N, 105 ° 30 ' 24.3 '' W, 1719 m, 01 - Jan- 2014, DRV & PRH colls., L- 1710 (1 ♂, 1 ♀ UMC); 33 mi W Otinapa, Río Arenales, 12 July 1952, J. D. Lattin Cal. Acad. Sci. Coll. / Ira La Rivers collection bequeathed to the California Academy of Sciences - 1978 (2 ♂ CAS). GUERRERO: Mpio. Atenango del Río, Atenango del Río, Río Amacuzac, 18 º 06 ' 01.5 '' N, 99 º 06 ' 27.8 '' W, 631 m, 30 - Dec- 2012, DRV & PRH colls., L- 1499 (5 ³, 7 ♀ UMC); Mpio. Copalillo, Papalutla, Río Balsas (Río Papalutla), 18 º 01 ' 27 '' N, 98 º 54 ' 09.3 '' W, 627 m, 30 - Dec- 2012, DRV & PRH colls., L- 1500 (11 ³, 6 ♀ UMC); Mpio. Teloloapan, Oxtotitlán, Río Oxtotitlán, 18 º 11 ' 35.7 '' N, 99 º 56 ' 24.3 '' W, 1009 m, 29 - Dec- 2012, DRV & PRH colls., L- 1498 (1 ♀ UMC); [Mpio. Tepecoacuilco de Trujano], Apan, near San Agustín Oapan, at Balsas River, 15 November 2008 / 17 º 56 ' 43 '' N, 99 º 26 ' 0 9 W, 505 m, J. D. Amit & A. Domínguez (3 ³, 4 ♀ UMC); [Mpio. Tepecoacuilco de Trujano], 17 º 57 ' 14 '' N, 99 º 26 ' 22 '' W, Balsas River, S [an] Ag [ustín] Oapan, 1650 ft, 6 June 2006, # 302, coll. J. Amith (2 ♂ UMC). HIDALGO: Mpio. Atlapexco, Atlaltipa Huitzotlaco, Pte. Atlaltipa, 21 ° 02 ' 12.3 '' N, 98 ° 22 ' 24.7 '' W, 158 m, 27 - May- 2014, DRV & PRH colls., L- 1796 (1 ♀ UMC); Mpio. Huejutla de Reyes, Huitzquilititla, 21 ° 06 ' 39.9 '' N, 98 ° 27 ' 24.4 '' W, 203 m, 27 - May- 2014, DRV & PRH colls., L- 1795 (2 ♀ UMC); Mpio. Molango Escamilla, Malila, Río Malila, 20 ° 43 ' 43.8 '' N, 98 ° 43 ' 14.2 '' W, 1386 m, 05 - Jun- 2013, DRV, Sites, Shepard & PRH colls., L- 1518 (1 ♀ UMC); Mpio. San Felipe Orizatlán, Huextetitla, 21 ° 09 ' 44.4 '' N, 98 ° 33 ' 26.6 '' W, 154 m, 07 - Jun- 2013, DRV, Sites, Shepard & PRH colls., L- 1525 (1 ♀ UMC); Mpio. San Felipe Orizatlán, Orizatlán, Pte. Tuntitlán, Río Tuntitlán, 21 ° 10 ' 03.4 '' N, 98 ° 36 ' 58.4 '' W, 175 m, 07 - Jun- 2013, DRV, Sites, Shepard & PRH colls., L- 1527 (2 ³ UMC); Mpio. San Felipe Orizatlán, San Felipe Orizatlán, Río San Pedro, 21 ° 10 ' 13.6 '' N, 98 ° 35 ' 46.8 '' W, 173 m, 07 - Jun- 2013, DRV, Sites, Shepard & PRH colls., L- 1526 (18 ³, 27 ♀ UMC); Mpio. Tlanchinol, 3.4 km W of Chalchocotipa, road to Santa Maria Catzotipan (west of Hwy 105, km 3 + 400), 21 ° 02 ' 36.4 '' N, 98 º 36 ' 00.8 '' W, 528 m, 06 - Jun- 2013, DRV, Sites, Shepard & PRH colls., L- 1522 (2 ³, 5 ♀ UMC); Mpio. Xochiatipan, Nuevo Acatepec, Pte. Garces Principal, Río El Encinal, 20 ° 56 ' 23.7 '' N, 98 ° 16 ' 54.9 '' W, 201 m, 27 - May- 2014, DRV & PRH colls., L- 1797 (7 ♂, 4 ♀ UMC). JALISCO: Estanzuela, Puente La Estanzuela, W of Ameca, 13 Jan. 2005, C. B. Barr / 20 º 31.49 ' N, 104 º 20.16 ' W, elevation 4580 ft. (1 ♀ EMEC); Estanzuela, 13 I 2005, 4580 ', unnamed stream / William D. Shepard, leg. (1 ♀ EMEC); Hwy 200 @ km 84.5 6 I 2005, 100 ', Río San Nicolas / William D. Shepard, leg. (1 ♀ EMEC); Mpio. Atenguillo, Los Volcanes, Río Atenguillo, 20 ° 19 ' 16.5 '' N, 104 ° 31 ' 41.1 '' W, 1459 m, 27 - Jun- 2013, DRV & PRH colls., L- 1582 (5 ³, 5 ♀ UMC); [Mpio. La Huerta], SE of Chamela, 6 I 2005, 180 ', Río Cuitzmala / William D. Shepard, leg. (1 ♀ EMEC); Mpio. Puerto Vallarta, Las Palmas, 235 m, 20 º 49.515 ' N, 105 º 05.367 ' W, 16 March 2014, colls: J. A. Gómez, S. Smith & R. Novelo (1 ³ IEXA); Mpio. San Sebastian del Oeste, Pte. Los Arrayanes, Arroyo Los Arrayanes, 20 ° 49 ' 44.5 '' N, 105 ° 00 ' 45.1 '' W, 337 m, 26 - Jun- 2013, DRV & PRH colls., L- 1579 (1 ♂, 1 ♀ UMC); [Mpio. Villa Purificación], Río El Amborín, 6.6 rd, km SE Villa Purificación, 19 º 38.81 ' N, 104 º 33.92 ' W, 8 Jan. 2005, C. B. Barr (2 ♀ UMC; 1 ³, 1 ♀ EMEC); [Mpio. Villa Purificación], 6.6 km SE Villa Purificación, 8 I 2005, Puente El Amborin / William D. Shepard leg. (9 ³, 5 ♀ EMEC); Río de Tepospisaloya, ca. 6 km N Unión de Tula, 9.2 km W Tacotán, 10 Jan. 2005, C. B. Barr / 20 º 03.86 ' N, 104 º 20.91 ' W, elevation 1250 ft. (1 ³, 1 ♀ EMEC); Pte. San Buenaventura at Rd. 513, 8.5 rd. km W of Tonaya, [Río Ferreria], 11 Jan. 2005, coll. C. B. Barr / 19 º 47.12 ' N, 104 º 02.71 ' W, elevation 2360 ft. (1 ♀ EMEC); 2.4 km SE Purificación, 8 I 2005, 1280 ', Río Purificación / William D. Shepard leg. (1 ³ EMEC); 8.5 km W Tonaya, 11 I 2005, 2360 ', Puente San Buenavista [San Buenaventura], [Río Ferreria] / William D. Shepard leg. (5 ³, 4 ♀ EMEC); 9.2 km W Tacotán, 10 I 2005, 1250 ', Río de Tepospisaloya / William D. Shepard leg. (1 ♂, 2 ♀ EMEC). MÉXICO: Mpio. Amatepec, Las Adjuntas, Río Sn. Felipe, 05 - May- 2008, R. Novelo & K. Cuevas cols. (1 ♀ IEXA); [Mpio.] Amatepec, Sn. Felipe de Jesús (1000 m), 20 - Abril- 2004, R. Novelo col. (1 ³, 1 ♀ IEXA); [Mpio. Tejupilco], Bejucos, Río Bejucos, 30 - Jan- 2008, R. Novelo col. (1 ♀ IEXA); same but 22 - Abril- 2004, 700 m (15 Ƌ, 8 ♀ IEXA); [Mpio. Tejupilco], Bejucos, Río Bejucos, 08 - Dic- 2008, J. Curiel col. (1 ♀ IEXA); [Mpio. Tejupilco], Bejucos, 22 IV 2004, 1950 ', Río Bejucos / William D. Shepard, leg. (3 ³, 4 ♀ EMEC); Mpio. Tejupilco, Paso de Vigas, 18 º 53 ' 32.6 '' N, 100 º 13 ' 57.0 '' W, 1107 m, 19 - Dec- 2012, DRV & PRH colls., L- 1459 (3 ³, 2 ♀ UMC); San Felipe de Jesus, 20 IV 2004, 3280 ', Río San Felipe / William D. Shepard, leg. (1 ³ EMEC); Tejupilco, Mex., Temescaltepec [Temascaltepec], VI- 18 - 1933 / H. E. Hinton, R. L. Usinger (3 ♀ EMEC); same but VI- 19 - 1933 (9 ³, 3 ♀ EMEC); Tejupilco, Mex., Temescaltepec [Temascaltepec], VII- [no day] 1934 / H. E. Hinton (2 ³, 4 ♀ EMEC); Temescaltepec [Temascaltepec], VII- [no day] 1934 / H. E. Hinton (1 ♂ EMEC). MICHOACÁN: [Mpio. Aguililla], Aguililla, 03 - Abril- 2005, R. Novelo col. (2 ³, 2 ♀ IEXA); [Mpio. Apatzingán], Apatzingán, Pte. El Orejón, 12 - Marzo- 2004, R. Novelo col. (2 ³, 3 ♀ IEXA); [Mpio.] Apatzingán, California (arroyo), 21 - Feb- 2005, R. Novelo & J. A. Gómez cols., Ambrysus circumcinctus R. Novelo det. (1 ³ IEXA); Mpio. Aquila, Aquila, Pte. Ramal Aquila, 18 º 35 ' 08.1 '' N, 103 º 30 ' 51.8 '' W, 159 m, 25 - Dec- 2012, DRV & PRH colls., L- 1487 (1 ³, 3 ♀ UMC); Mpio. Carácuaro, Carácuaro, Río San Diego, 19 º 01 ' 04.8 '' N, 101 º 07 ' 33.2 '' W, 519 m, 22 - Dec- 2012, DRV & PRH colls., L- 1473 (2 ³, 1 ♀ UMC); [Mpio. Chinicuila], Villa Victoria, 3 - Junio- 2005, R. Novelo col. (1 ♀ IEXA); Mpio. Morelos, La Placita de Morelos, Pte. La Placita, Río Aquila, 18 º 32 ' 19.9 '' N, 103 º 35 ' 17.9 '' W, 7 m, 26 - Dec- 2012, DRV & PRH colls., L- 1488 (6 ³, 1 ♀ UMC); Mpio. Nocupétaro, San Antonio de la Huerta, under bridge, 19 º 02 ' 04.4 '' N, 101 º 15 ' 07.9 '' W, 624 m, 22 - Dec- 2012, DRV & PRH colls., L- 1475 (5 ³, 1 ♀ UMC); Mpio. Parácuaro, Las Yeguas, Río Tepalcatepec, 18 º 53 ' 42.1 '' N, 102 º 17 ' 00.3 '' W, 189 m, 24 - Dec- 2012, DRV & PRH colls., L- 1480 (1 ³, 6 ♀ UMC); Mpio. Tepalcatepec, Los Horcones, km 31 carr. Tepalcatepec-Coalcomán, Pte. Pinolapa, 19 º 00 ' 31.3 '' N, 103 º 01 ' 25.6 '' W, 554 m, 25 - Dec- 2012, DRV & PRH colls., L- 1484 (1 ♀ UMC); Mpio. Tepalcatepec, Río Pinolapa, 10 - Jul- 2006, R. Novelo, col. (1 ³, 2 ♀ IEXA); Mpio. Tepalcatepec, Tepalcatepec, 19 º 11 ' 47.3 '' N, 102 º 50 ' 44.9 '' W, 371 m, 24 - Dec- 2012, DRV & PRH colls., L- 1483 (1 ³ UMC); Mpio. Tiquicheo, Piedra China, Pte. Tiquicheo, Río Purengueo, 18 º 53 ' 37.8 '' N, 100 º 45 ' 49 '' W, 363 m, 22 - Dec- 2012, DRV & PRH colls., L- 1472 (6 ³, 6 ♀ UMC); Mpio. Tuzantla, Tuzantla, Río Tuzantla, 19 º 13 ' 18.1 '' N, 100 º 34 ' 05.5 '' W, 593 m, 21 - Dec- 2012, DRV & PRH colls., L- 1470 (4 ³, 6 ♀ UMC); Río Pinolapa, 30 - May- 2005, (4 ³ IEXA); S de Coalcomán, Aguililla, 11 - Julio- 2005, R. Novelo col. (4 ³, 9 ♀ IEXA). MORELOS: [Mpio. Amacuzac], Huajintlán, Río Amacuzac, 28 Mayo 1983, [no collector] (1 ♀ CNIN); same but D. Porras, L. Castrejón [colls.] (2 ³ CNIN); same but / A. c. circumcinctus Montandon det. R. W. Sites (1 ³ CNIN); same locality information but / Ambrysus circumcinctus Montandon det. J. T. Polhemus 19 [??] (1 ³, 2 ♀ CNIN); [Mpio. Jojutla], San Rafael Vicente Aranda, 15 - Mar- 1983, R. Novelo col. (3 ³, 4 ♀ IEXA); [Mpio. Jojutla], S. Rafael A., 13 - VII- 82, M. Espinosa, (10 ³, 7 ♀ CNIN); same but / A. c. circumcinctus Montandon det. R. W. Sites (3 ³, 1 ♀ CNIN); same locality information but / A. circumcinctus Mont. det. R. W. Sites 2009 (1 ♀ CNIN); same locality information but / Ambrysus circumcinctus Montandon det. J. T. Polhemus 19 [??] (1 ³ CNIN); [Mpio. Jojutla], S. Rafael A., 17 - VI- 82, M. Espinosa (15 ³, 11 ♀ CNIN); same but / A. c. circumcinctus Montandon det. R. W. Sites (1 ³, 4 ♀ CNIN); Mpio. Jojutla, Tehuixtla, Río Amacuzac, 18 º 32 ' 47.9 '' N, 99 º 16 ' 03.6 '' W, 854 m, 28 - Apr- 2011, DRV coll., L- 1309 (11 ³, 7 ♀ UMC); Mpio. Jojutla, Tehuixtla, Río Amacuzac, 18 º 33 ' 45.2 '' N, 99 º 16 ' 51.3 '' W, 853 m, 28 - Apr- 2011, DRV coll., L- 1308 (3 ³, 3 ♀ UMC); [Mpio.] Jojutla, Vicente Aranda, Río Amacuzac (900 m), 11 - Diciembre- 1982, P. Liljehult col. (1 ³, 1 ♀ IEXA); [Mpio.] Jojutla, Vicente Aranda, Río Amacuzac (900 m), 12 - Feb- 1983, V. García col. (27 ³, 40 ♀ IEXA); same locality information but C. Valencia col. (22 ³, 17 ♀ IEXA); [Mpio.] Jojutla, Vicente Aranda, Río Amacuzac, 27 - Marzo- 1983, G. Cardoso col. (12 ³, 14 ♀ IEXA); [Mpio.] Jojutla, Vicente Aranda, Río Amacuzac, 7 - Mayo- 1983, S. Ibáñez col. (7 ³, 7 ♀ IEXA); [Mpio. Jojutla], Vicente Aranda (900 msnm), 16 - Marzo- 2001, R. Novelo col. (4 ³, 7 ♀ IEXA); [Mpio. Jojutla], Vicente Aranda, Río Amacuzac, 04 - Abril- 2002, R. Novelo col. (4 ³, 10 ♀ IEXA); [Mpio. Tetecala], Tetecala, (900 m), 02 - Diciembre- 2000, R. Novelo col. (1 ♀ IEXA); [Mpio.] Tlaquiltenango, El Astillero [La Mezquitera], Río Cuautla (dentro del río c / corriente), 5 - Feb- 1983, I. Oliva col. (4 ³, 4 ♀ IEXA); [Mpio. Yautepec], Yautepec, Río Yautepec (1,203 m), 17 - Enero- 1983, I. Oliva col. (2 ♀ IEXA). NAYARIT: Las Adjuntas, Río Huaynamota, II 3 - VII- 92, S. Santiago, R. Barba (1 ♀ CNIN); Los Sabinos, Río Huaynamota, 10 - IV- 91 R. Barba, E. Barrera (1 ³ CNIN); same but 23 - 5 - 1991 (4 ³, 1 ♀ CNIN); same but / A. c. circumcinctus Montandon det. R. W. Sites (1 ♀ CNIN); Mpio. Acaponeta, El Recodo, Río Acaponeta, 22 ° 31 ' 31.9 '' N, 105 ° 20 ' 37.3 '' W, 34 m, 01 - Jul- 2013, DRV & PRH colls., L- 1601 (1 ♀ UMC); Mpio. Acaponeta, La Guásima, Río Acaponeta, 22 ° 24 ' 26.6 '' N, 105 ° 23 ' 48.9 '' W, 26 m, 02 - Jul- 2013, DRV & PRH colls., L- 1602 (20 ♂, 17 ♀ UMC); Mpio. Ahuacatlán, Ejido Tetitlán, Río Tetitlán, 21 ° 07 ' 51.3 '' N, 104 ° 37 ' 10.1 '' W, 747 m, 02 - Jul- 2013, DRV & PRH colls., L- 1606 (10 ³, 11 ♀ UMC); Mpio. Ahuacatlán, Puerta del Río, Balneario El Paraiso, Río Tetitlán, 21 ° 10 ' 59.3 '' N, 104 ° 38 ' 32.9 '' W, 859 m, 18 - Dec- 2013, DRV & PRH colls., L- 1651 (1 ³, 1 ♀ UMC); Mpio. El Nayar, Playa Golondrinas, Río Santiago, 26 / 5 / 1991, R. Barba, E. Barrera / A. c. circumcinctus Montandon det. R. W. Sites (1 ³ CNIN); Mpio. Ruíz, San Lorenzo, Río San Pedro, 21 ° 57 ' 04.1 '' N, 105 ° 04 ' 28.3 '' W, 23 m, 20 - Dec- 2013, DRV & PRH colls., L- 1662 (1 ³, 4 ♀ UMC); Mpio. San Blas, El Palillo, Pte. Palillo, Arroyo Palillo, 21 ° 38 ' 27.1 '' N, 105 ° 08 ' 30.8 '' W, 43 m, 20 - Dec- 2013, DRV & PRH colls., L- 1661 (4 ³, 4 ♀ UMC); Mpio. Tepic, Colorado de la Mora, 232 msnm, 12 - IV- 91, E. Barrera, J. León (1 ♂ CNIN). NUEVO LEÓN: Mpio. Cadereyta Jiménez, San Juan, Río San Juan, 25 ° 31 ' 55.1 '' N, 99 ° 50 ' 39.1 '' W, 266 m, 19 - May- 2014, DRV & PRH colls., L- 1773 (15 ³, 23 ♀ UMC); Mpio. Cd. Anáhuac, Cd. Anáhuac, Río Salado, 27 ° 14 ' 09.0 '' N, 100 ° 08 ' 24.1 '' W, 187 m, 18 - May- 2014, DRV & PRH colls., L- 1769 (6 ³, 11 ♀ UMC); Mpio. García, Río Pesquerías, 8 - XI- 85, H. Rivas (1 ³ CNIN); same locality information but R. Barba / A. c. circumcinctus Montandon det. R. W. Sites (1 ♀ CNIN); same locality information but S. Tufinio / A. c. circumcinctus Montandon det. R. W. Sites (1 ♀ CNIN); Mpio. General Terán, La Barreta, Río Conchos, 25 ° 04 ' 25.9 '' N, 98 ° 58 ' 55.9 '' W, 124 m, 20 - May- 2014, DRV & PRH colls., L- 1776 (8 ³, 8 ♀ UMC); Mpio. General Terán, La Unión, Río El Pilón, 25 ° 20 ' 27.2 '' N, 99 ° 34 ' 57.5 '' W, 243 m, 19 - May- 2014, DRV & PRH colls., L- 1775 (1 ♀ UMC); Mpio. Lampazos de Naranjo, El Ancón, Río Candela, 27 ° 02 ' 33.5 '' N, 100 ° 32 ' 13.5 '' W, 293 m, 18 - May- 2014, DRV & PRH colls., L- 1770 (1 ³ UMC); Mpio. Los Ramones, 1 km N of Hacienda El Carrizo, Río San Juan, 25 ° 31 ' 40.5 '' N, 99 ° 37 ' 20.9 '' W, 206 m, 19 - May- 2014, DRV & PRH colls., L- 1774 (24 ³, 23 ♀ UMC); [Mpio. Villaldama], Villa Aldama, [Villaldama], [Arroyo] La Coyota, 13 - XI- 85, R. Barba (3 ♀ CNIN); Pilon R. at unnamed road, across from General Terán (town), off St. Hwy 35 / 15 May 1995, DB 95 - 10, colls: D. E. Baumgardner & B. C. Henry (2 ³, 3 ♀ UMC); 17.5 mi. W Linares, V- 25 - 1981, J. T. Doyen, J. K. Liebherr (1 ♀ EMEC). OAXACA: Dominguillo, 7 - XI- 1988, E. Barrera, R. Barba, L. Cervantes (1 ³ CNIN); Mpio. Ayotzintepec, San José Mano Márques, 17 º 47 ' 26.2 '' N, 96 º 00 ' 15.3 '' W, 66 m, 24 - May- 2012, DRV & PRH colls., L- 1371 (1 ³, 1 ♀ UMC); Mpio. Guelatao de Juárez, carr. Tuxtepec-Oaxaca, 6 km W of Guelatao, Pte. Xia, 17 º 18 ' 18.3 '' N 96 º 31 ' 33.8 '' W, 1497 m, 24 - May- 2012, DRV & PRH colls., L- 1373 (11 ♂, 1 ♀ UMC); [Mpio. Ixtepec], Río Perros, just above Ixtepec, III- 27 - 1957, Elev. 340 ft, R. R. Miller (7 ³, 15 ♀ UMMZ); Mpio. Jalapa de Díaz, Sto. Domingo, Río Sto Domingo, 18 ° 02 ' 15.9 '' N, 96 ° 32 ' 57.2 '' W, 100 M, 25 - Mar- 2015, DRV, Sites, Shepard, Barr, & PRH colls., L- 1881 (2 ³ UMC); [Mpio. San Juan Bautista Cuicatlán], Santiago Dominguillo, Río Las Huertas, 12 - VII- 1996, R. Barba, A. Rojas / A. c. circumcinctus Montandon det. R. W. Sites (2 ♀ CNIN); Mpio. San Juan Bautista Cuicatlán, Santiago Dominguillo, 17 º 38 ' 46 '' N, 96 º 54 ' 36.7 '' W, 726 m, 01 - Jun- 2012, DRV & PRH colls., L- 1405 (4 ³, 2 ♀ UMC); Mpio. San Juan Cotzocón, Arroyo Carrizal, km 110 carr. Palomares-Tuxtepec, 17 º 26 ' 55.6 '' N, 95 º 26 ' 39.1 '' W, 94 m, 22 - May- 2012, DRV & PRH colls., L- 1365 (3 ³, 7 ♀ UMC); Mpio. San Miguel del Puerto, Copalita, Pte. Copalita, Río Copalita, 15 ° 49 ' 12.2 '' N, 96 ° 03 ' 57.6 '' W, 28 m, 04 - Jun- 2014, DRV & PRH colls., L- 1818 (3 ³, 4 ♀ UMC); Mpio. San Sebastian Tecomaxthahuaca, Tecomaxthahuaca, Laguna Encantadac [in lateral channel], 17 º 21 ' 57.5 '' N, 98 º 01 ' 13.3 '' W, 1662 m, 30 - May- 2012, DRV & PRH colls., L- 1399 (1 ³, 2 ♀ UMC); Mpio. Santiago Jocotepec, nr. San José Río Manso, 17 º 38 ' 44.3 '' N, 95 º 52 ' 02.1 '' W, 61 m, 23 - May- 2012, DRV & PRH colls., L- 1368 (5 ³, 2 ♀ UMC); Mpio. Santiago Jocotepec, San Jose Río Manso, Centro Ecoturístico Cerro Chango, 17 º 42 ' 06.6 '' N, 95 º 54 ' 01.6 '' W, 54 m, 23 - May- 2012, DRV & PRH colls., L- 1367 (12 ³, 3 ♀ UMC); Mpio. Santo Domingo Zanatepec, carr. Arriaga-Santo Domingo Tehuantepec, Colonia Río Ostula, Pte. Ostula, 16 º 29 ' 59.5 '' N, 94 º 26 ' 00.5 '' W, 40 m, 22 - May- 2012, DRV & PRH colls., L- 1360 (1 ³, 4 ♀ UMC); Mpio. Valerio Trujano, km 124 carr. Tehuacán-San Francisco Telixtlahuaca, jct to Tomellín, Pte. El Grande, Río Grande, 17 º 45 ' 25 '' N, 96 º 56 ' 59.1 '' W, 607 m, 01 - Jun- 2012, DRV & PRH colls., L- 1406 (4 ³, 3 ♀ UMC); Mpio. Valle Nacional, San Mateo Yetla, 12 - X- 1990, E. Ramírez, E. Barrera, A. Cadena (1 ♀ CNIN); km 21 desviación San Miguel Chimalapa, 210 msnm, 22 - III- 1990, E. Barrera, A. Cadena (1 ♀ CNIN); km 116 carr. Tuxtepec-Palomares, 22 - XI- 1990, A. Cadena, E. Barrera (1 ♀ CNIN). PUEBLA: [Mpio. Jopala], Patla, 23 - VII- 75, H. Brailovsky A. col., noct. (1 ³, 1 ♀ CNIN); same but / Ambrysus circumcinctus Montandon (Ira La Rivers) (1 ♀ CNIN); Mpio. Tlacuilotepec, carr. Plan de Ayala-Tlaxco, 2 km E of Papaloctipan, 20 º 25 ' 22.4 '' N, 97 º 56 ' 38.2 '' W, 310 m, 05 - Jun- 2012, DRV & PRH colls., L- 1417 (10 ³, 8 ♀ UMC); Mpio. Tlacuilotepec, Río San Marcos, km 1 carr. A. Camacho-Petlacotla, 24 - Julio- 1987, R. Novelo col. (5 ³, 4 ♀ IEXA); Mpio. Tlaxco, carr. Tlaxco-Cuaxtla, ca. 2 km S of Acalman, Río Los Cajones, 20 º 24 ' 26.8 '' N, 98 º 03 ' 51.5 '' W, 785 m, 05 - Jun- 2012, DRV & PRH colls., L- 1418 (1 ♀ UMC); Mpio. Xicotepec, carr. Huauchinango-Poza Rica, Miguel Ávila Camacho (La Ceiba), Pte. San Marcos, Río San Marcos, 20 º 23 ' 40 '' N, 97 º 52 ' 39.1 '' W, 239 m, 05 - Jun- 2012, DRV & PRH colls., L- 1416 (4 ³, 6 ♀ UMC); [Mpio. Xicotepec], Villa Ávila Camacho, [La Ceiba], Río San Marcos, 17 - Julio- 1990, R. Novelo col. (1 ♀ IEXA); same but 30 - Abril- 1991 (1 ³, 1 ♀ IEXA). SAN LUIS POTOSÍ: Mpio. Axtla de Terrazas, Cómoca, Río Tancuilín, 21 º 25 ' 03.8 '' N, 98 º 53 ' 43.3 '' W, 75 m, 22 - Apr- 2011, DRV coll., L- 1307 (20 ³, 13 ♀ UMC); same but 19 - Jul- 2011, L- 1311 (1 ♀ UMC); Mpio. Axtla de Terrazas, Cómoca, Río Tancuilín, 21 ° 25 ' 02.9 '' N, 98 ° 53 ' 42.8 '' W, 86 m, 08 - Jun- 2013, DRV, Sites, Shepard, & PRH colls., L- 1532 (1 ³, 5 ♀ UMC); Mpio. Huehuetlán, El Nacimiento de Huichihuayán (spring), 21 ° 27 ' 35.3 '' N, 98 ° 58 ' 38.8 '' W, 96 m, 08 - Jun- 2013, DRV, Sites, Shepard, & PRH colls., L- 1533 (1 ³, 1 ♀ UMC); Mpio. Matlapa, Los Manantiales, Río Tancuilín, 21 ° 20 ' 45.5 '' N, 98 ° 51 ' 45.8 '' W, 113 m, 07 - Jun- 2013, DRV, Sites, Shepard, & PRH colls., L- 1530 (24 ³, 21 ♀ UMC); Mpio. San Vicente Tancuayalab, Ejido El Álamo, Río Moctezuma, 21 ° 43 ' 10.9 '' N, 98 ° 33 ' 16.7 '' W, 32 m, 10 - Jun- 2013, DRV, Sites, Shepard, & PRH colls., L- 1540 (17 ³, 16 ♀ UMC); Mpio. Tamazunchale, Vega Larga, 21 º 15 ' 27.3 '' N, 98 º 50 ' 36.8 '' W, 161 m, 17 - Jul- 2011, DRV coll., L- 1310 (1 ♂, 1 ♀ UMC); Mpio. Tamazunchale, Tamazunchale, Río Moctezuma, 21 ° 15 ' 27.6 '' N, 98 ° 48 ' 20.6 '' W, 128 m, 07 - Jun- 2013, DRV, Sites, Shepard, & PRH colls., L- 1529 (3 ³, 2 ♀ UMC); Mpio. Tamazunchale, Zacatipan, Río Amajac, ca. 2 km SE of Tamazunchale, 21 ° 14 ' 38.1 '' N, 98 ° 46 ' 19.7 '' W, 111 m, 07 - Jun- 2013, DRV, Sites, Shepard, & PRH colls., L- 1528 (2 ³, 1 ♀ UMC); Mpio. Xilitla, ca. 1 km E of Xilitla, unnamed creek from Las Pozas, 21 ° 23 ' 30.6 '' N, 98 ° 59 ' 23.3 '' W, 534 m, 08 - Jun- 2013, DRV, Sites, Shepard, & PRH colls., L- 1531 (1 ³ UMC); [Mpio. Xilitla], Xilitla, 600 msnm, 24 - Jul- 1998, R. Novelo col. (1 ³ IEXA); Puente Palitla [Palictla], 28 - Junio- 1990, R. Novelo col. (2 ³, 1 ♀ IEXA); 34 Km antes Tamazunchale, 20 - V- 78, J. Bueno / Ambrysus circumcinctus Montandon det. J. T. Polhemus 19 [??] (1 ♂, 1 ♀ CNIN). SINALOA: Mpio. Concordia, Chupaderos, Río Pánuco, 23 ° 21 ' 49.2 '' N, 105 ° 57 ' 10.3 '' W, 244 m, 30 - Jun- 2013, DRV & PRH colls., L- 1597 (2 ³, 2 ♀ UMC); [Mpio. Concordia], Río Pánuco, 38 mi E Mazatlan, Sin. Mex. VIII- 28 - 52, J. D. Lattin / Ira La Rivers collection bequeathed to the California Academy of Sciences - 1978 (1 ³, CAS); [Mpio. Culiacán], Los Mayos, 24 July 1952, John D. Lattin, Cal. Acad. Sci. Coll. / Ira La Rivers collection bequeathed to the California Academy of Sciences - 1978 (2 ³, CAS); [Mpio. El Fuerte], El Fuerte, Río Fuerte, 26 º 24 ' N, 108 º 37 ' W, April 2002, W. Trimble (2 ³, 1 ♀ UMC); Mpio. Elota, Pte. Los Barriles, Arroyo El Japuino, 24 ° 03 ' 36.7 '' N, 106 ° 48 ' 03.6 '' W, 83 m, 22 - Dec- 2013, DRV & PRH colls., L- 1670 (27 ³, 29 ♀ UMC); Mpio. Elota, Pte. Río Elota, Río Elota, 23 ° 57 ' 20.7 '' N, 106 ° 42 ' 53.4 '' W, 46 m, 22 - Dec- 2013, DRV & PRH colls., L- 1669 (27 ³, 28 ♀ UMC); [Mpio. Mazatlán], Mazatlán, Mex., V- 25 - 26 - 1934 / H. E. Hinton (1 ♀ EMEC); same but / Ambrysus pygmaeus La Rivers PARATYPE (1 ♀ EMEC); Mpio. Mazatlán, Siqueiros, Siqueiros (Dam), Río Presidio, 23 ° 20 ' 47.0 '' N, 106 ° 14 ' 35.3 '' W, 42 m, 21 - Dec- 2013, DRV & PRH colls., L- 1664 (4 ³, 4 ♀ UMC); Mpio. Mazatlán, Villa Unión, Pte. Presidio, Río Presidio, 23 ° 11 ' 32.3 '' N, 106 ° 13 ' 26.3 '' W, 10 m, 01 - Jul- 2013, DRV & PRH colls., L- 1598 (10 ³, 10 ♀ UMC); Mpio. Mocorito, Mocorito, Río Evora, 25 ° 28 ' 55.1 '' N, 107 ° 54 ' 37.6 '' W, 75 m, 23 - Dec- 2013, DRV & PRH colls., L- 1674 (12 ³, 13 ♀ UMC); Mpio. Rosario, Matatán, 16 km W of El Rosario, Río Los Cedros, 23 ° 01 ' 54.1 '' N, 105 ° 43 ' 52.0 '' W, 55 m, 21 - Dec- 2013, DRV & PRH colls., L- 1663 (3 ³ UMC); Mpio. San Ignacio, San Ignacio, Río Piaxtla, 23 ° 56 ' 06.3 '' N, 106 ° 25 ' 41.0 '' W, 127 m, 21 - Dec- 2013, DRV & PRH colls., L- 1666 (1 ³ UMC); Mpio. Sinaloa de Leyva, Burague, Río Sinaloa, 25 ° 49 ' 25.4 '' N, 107 ° 57 ' 49.4 '' W, 140 m, 24 - Dec- 2013, DRV & PRH colls., L- 1678 (34 ♂, 23 ♀ UMC). SONORA: [Mpio. Álamos], Arroyo Cuchujaqui, 7 mi. SE Álamos, Son., Mex., VI- 19 - 63 / J. Doyen Collector (1 ♀ EMEC); Mpio. Álamos, El Naranjo, Arroyo El Taymuco, 27 ° 14 ' 43.3 '' N, 108 ° 45 ' 27.8 '' W, 462 m, 25 - Dec- 2013, DRV & PRH colls., L- 1682 (5 ³, 3 ♀ UMC); Mpio. Álamos, Taymuco (El Vado), Arroyo La Vinatería, 27 ° 14 ' 39.7 '' N, 108 ° 43 ' 17.3 '' W, 537 m, 25 - Dec- 2013, DRV & PRH colls., L- 1683 (1 ³, 2 ♀ UMC); [Mpio. Álamos], 8 mi S Álamos, 12 Sept 79 / Arroyo Cucyacho in River, C. Y. Kitayama (1 ³, 3 ♀ EMEC); Mpio. Onavas, Tepoca, Arroyo Tepoca, 28 ° 25 ' 50.3 '' N, 109 ° 15 ' 26.0 '' W, 564 m, 27 - Dec- 2013, DRV & PRH colls., L- 1691 (25 ³, 9 ♀ UMC); Mpio. Quiriego, Tepahui, 27 ° 23 ' 27.3 '' N, 109 ° 11 ' 03.6 '' W, 137 m, 26 - Dec- 2013, DRV & PRH colls., L- 1687 (1 ♀ UMC); Mpio. Rosario de Tesopaco, Nuri, Arroyo Nuri, 28 ° 06 ' 39.0 '' N, 109 ° 19 ' 15.5 '' W, 357 m, 27 - Dec- 2013, DRV & PRH colls., L- 1688 (7 ³, 15 ♀ UMC); Mpio. Rosario de Tesopaco, ca. 6 km N of Nuri, Arroyo Cajón de Amador, 28 ° 09 ' 50.0 '' N, 109 ° 19 ' 37.7 '' W, 324 m, 27 - Dec- 2013, DRV & PRH colls., L- 1689 (11 ³, 13 ♀ UMC); Mpio. Soyopa, ca. 15 km E of Barranca, Río Yaqui, 28 ° 34 ' 21.2 '' N, 109 ° 33 ' 11.6 '' W, 173 m, 27 - Dec- 2013, DRV & PRH colls., L- 1690 (1 ♀ UMC); 10 miles so [uth] Alamos, VI 13 1961 / F. D. Parker (1 ³, 2 ♀ UCDC). TABASCO: Mpio. Tlacotalpa, [Tapijulapa], Río de la Sierra, 29 / III / 1998, 17 º 28 ' 04 '' N, 92 º 46 ' 52 '' W, J. Bueno. R. Barba (2 ♂, 2 ♀ CNIN). TAMAULIPAS: Mpio. Aldama, Las Nubes, Pte. San Rafael, 23 ° 09 ' 56.4 '' N, 97 ° 57 ' 14.2 '' W, 37 m, 23 - May- 2014, DRV & PRH colls., L- 1789 (1 ³, 1 ♀ UMC); Mpio. Güemes, ca. 6 km W of Estación Santa Engracia, Río Corona, 24 ° 00 ' 31.4 '' N, 99 ° 10 ' 04.1 '' W, 216 m, 21 - May- 2014, DRV & PRH colls., L- 1782 (5 ³, 11 ♀ UMC); Mpio. Güemes, La Gloria, Río Corona, 24 ° 00 ' 30.5 '' N, 99 ° 09 ' 24.0 '' W, 214 m, 21 - May- 2014, DRV & PRH colls., L- 1781 (17 ³, 6 ♀ UMC); Mpio. Juamave [Jaumave], Los Nogales, 2 - xi- 99, R. Jones, matorral (1 ♀ UAQE); Mpio. Padilla, El Barretal, Pte. Purificación Poniente, Río Purificación, 24 ° 04 ' 43.4 '' N, 99 ° 07 ' 23.9 '' W, 194 m, 21 - May- 2014, DRV & PRH colls., L- 1779 (12 ³, 7 ♀ UMC); Mpio. Padilla, El Barretal, Río Purificación, 24 ° 04 ' 40.5 '' N, 99 ° 08 ' 31.3 '' W, 205 m, 21 - May- 2014, DRV & PRH colls., L- 1780 (9 ³, 7 ♀ UMC); Mpio. San Fernando, Las Norias, Arroyo Chorreras, 24 ° 36 ' 30.0 '' N, 98 ° 17 ' 54.6 '' W, 70 m, 21 - May- 2014, DRV & PRH colls., L- 1777 (36 ♂, 40 ♀ UMC); 15 km N of C. Victoria, 18 March 1983, coll. M. S. Davis (1 ³, UMC). VERACRUZ: La Palma, 6 / V / 81, G. Ortega L. (1 ³, CNIN); same but / A. c. circumcinctus Montandon det. R. W. Sites (1 ♀ CNIN); Las Palmas, 13 - V- 83, G. Ortega / Ambrysus hybridus det. M. G. Ortega / Ambrysus circumcinctus Montandon det. J. T. Polhemus 19 [??] (1 ³, 1 ♀ CNIN); Las Palmas, 13 - V- 83, G. Ortega / Ambrysus hybridus det. M. G. Ortega / A. c. circumcinctus Montandon det. R. W. Sites (1 ³ CNIN); same but 18 - V- 83 (1 ³ CNIN); Los Tuxtlas, La Palma, 13 - V- 83 G. Ortega L. / Ambrysus hybridus M. G. Ortega / Ambrysus circumcinctus Montandon det. J. T. Polhemus 19 [??] (1 ♀ CNIN); [Mpio. Atzalán], Tomata, Pte. Tomata, Río Alseseca, 06 - Abril- 2011, V. Amaya col. (1 ♀ IEXA); Mpio. Atzalán, Pte. Tomata, Río Alseseca, 19 º 55.335 ' N, 97 º 13.385 ' W, 500 m, 6 - Apr- 2011, Sites, Shepard, Amaya, Novelo colls., gravel riffles, L- 1296 (2 ♀ UMC): Mpio. Atzalán, Pte. Tomata, Río Alseseca, 19 ° 55 ' 15.7 '' N, 97 ° 13 ' 28.2 '' W, 514 m, 13 - Jun- 2013, DRV, Sites, Shepard, Novelo & PRH colls., L- 1549 (6 ♂, 2 ♀ UMC); [Mpio. Atzalán], W of Tlapacoyan, El Filo, Río Filobobos, 08 - Noviembre- 2009, R. Novelo col., Ambrysus circumcinctus R. Novelo det. (33 ³, 50 ♀ IEXA); Mpio. Catemaco, Arroyo a 3 km Margaritas, 11 - V- 2012, J. L. Villalobos (3 ³, 1 ♀ CNIN); [Mpio. Chicontepec], Chicontepec, 14 - IV- 75, H. Brailovsky / Ambrysus circumcinctus Montandon (Ira La Rivers) (1 ³ CNIN); Mpio. Citlaltepec, Citlaltepec, 21 ° 19 ' 47.7 '' N, 97 ° 53 ' 00.1 '' W, 194 m, 11 - Jun- 2013, DRV, Sites, Shepard, & PRH colls., L- 1544 (2 ³, 1 ♀ UMC); Mpio. Coatepec, Agua Alegre, Río Huehueyapan, 18 [19 º] 26.884 ' N, 96 58.867 ' W, 1198 m, 4 IV 2011 / William D. Shepard leg., L- 1290 (2 ³ UMC); [Mpio. Coatepec], Coatepec, Río Huehueyapan, 04 - Abr- 2011, R. Novelo Col. (1 ♀ IEXA); [Mpio. Coscomatepec], Coscomatepec, 29 - I- 84, L. Becerra (1 ♀ CNIN); [Mpio. Huatusco]: Huatusco, Río Jamapa (rápidos), 16 - Agosto- 1986, M. Calvillo Col. (1 ♀ IEXA); Mpio. Ixhuatlán de Madero, Naranjo Dulce, Río Vinazco, 20 ° 49 ' 24.9 '' N, 97 ° 55 ' 19.2 '' W, 86 m, 27 - May- 2014, DRV & PRH colls., L- 1798 (3 ♀ UMC); [Mpio. Jamapa], Jamapa, 1200 mts, 1 - V- 1954, C. Bolivar (2 ♀ CNIN); same but / A. c. circumcinctus Montandon det. R. W. Sites (1 ³ CNIN); Mpio. Misantla, Libertad, Río Kilate, 20 o 03 ' 06.2 " N, 96 o 58 ' 39.4 " W, 61 m, 14 - Jun- 2013, DRV, Sites, Shepard, Novelo, & PRH colls., L- 1552 (47 ³, 47 ♀ UMC); Mpio. Misantla, Pte. Chapachapa, Río Chapachapa, 19 o 58 ' 12.5 " N, 96 o 53 ' 07.6 " W, 121 m, 14 - Jun- 2013, DRV, Sites, Shepard, Novelo, & PRH colls., L- 1554 (1 ³, 5 ♀ UMC); Mpio. Paso de Ovejas, Paso de Ovejas, 19 ° 17 ' 04.1 '' N, 96 ° 26 ' 03.7 '' W, 40 m, 29 - May- 2014, DRV & PRH colls., L- 1802 (1 ♀ UMC); Mpio. Puente Nacional, Puente Nacional, Pte. Puente Nacional I, Río La Antigua, 19 ° 19 ' 28.5 '' N, 96 ° 28 ' 54.3 '' W, 67 m, 29 - May- 2014, DRV & PRH colls., L- 1801 (1 ³, 1 ♀ UMC); [Mpio. San Andrés Tuxtla], Est. Biol. Los Tuxtlas, 20 - I- 86, col. F. Arias & C. Mayorga (3 ³, 3 ♀ CNIN); [Mpio. San Andrés Tuxtla], Estación de Biología Los Tuxtlas, 28 - IV- 1991, G. Ortega, C. Mayorga / A. c. circumcinctus Montandon det. R. W. Sites (1 ♂, 1 ♀ CNIN); [Mpio. San Andrés Tuxtla], Los Tuxtlas, El Salto de Eyipantla, 6 - 5 - 81, M. García, (1 ♀ CNIN); same locality information but R. Novelo / Ambrysus circumcinctus Montandon det. J. T. Polhemus 19 [??] (1 ³ CNIN); same locality information but R. Arce (1 ³ CNIN); same but / Ambrysus circumcinctus Montandon det. J. T. Polhemus 19 [??] (1 ♀ CNIN); [Mpio. San Andrés Tuxtla], Los Tuxtlas, L [aguna] Escondida, 5 - XII- 75, J. Bueno / Ambrysus circumcinctus Montandon det. J. T. Polhemus 19 [??] / Ambrysus circumcinctus caliginosus Usinger det. R. Sites 2009 (1 ♀ CNIN); [Mpio. San Andrés Tuxtla], Tuxtlas, L [aguna] Escondida, 8 - XII- 75, J. Bueno (1 ♀ CNIN); same but / Ambrysus circumcinctus Montandon det. J. T. Polhemus 19 [??] (1 ³ CNIN); [Mpio. San Andrés Tuxtla], Los Tuxtlas, Río La Palma, Ver., 7 - V- 81, R. Novelo / Ambrysus circumcinctus Montandon det. J. T. Polhemus 19 [??] (1 ♀ CNIN); same locality information but 02 - Abril- 2011, R. Novelo col. (3 ³ IEXA); [Mpio. San Andrés Tuxtla], Los Tuxtlas, Río Máquinas, 14 - [VII or VIII, unclear] - 79, J. Bueno / Ambrysus circumcinctus Montandon det. J. T. Polhemus 19 [??] (2 ³ CNIN); [Mpio. San Andrés Tuxtla], Los Tuxtlas, Río Máquinas, 5 - V- 81, R. Novelo (1 ³ CNIN); same locality information but 5 - 5 - 81, M. García (2 ♀ CNIN); [Mpio. San Andrés Tuxtla], Los Tuxtlas, Río Máquinas, 11 - V- 81, Herández / Ambrysus circumcinctus Montandon det. J. T. Polhemus 19 [??] (1 ♀ CNIN); [Mpio. San Andrés Tuxtla], Los Tuxtlas, Río Máquinas, 12 - 5 - 83, G. Ortega / Ambrysus hybridus det. G. Ortega (1 ♀ CNIN); [Mpio. San Andrés Tuxtla], Los Tuxtlas, Río Máquinas, 02 - Abril- 2011, R. Novelo col. (2 ³, 2 ♀ IEXA); Mpio. Sta. Maria, E of Ixcatepec, 21 ° 17 ' 25.8 '' N, 97 ° 54 ' 56.8 '' W, 223 m, 11 - Jun- 2013, DRV, Sites, Shepard, & PRH colls., L- 1543 (4 ³, 7 ♀ UMC); Mpio. Tantoyuca, N of Platon Sánchez, El Remanzo, Río Calabazo, 21 ° 17 ' 08.2 '' N, 98 ° 22 ' 38.6 '' W, 41 m, 06 - Jun- 2013, DRV, Sites, Shepard, & PRH colls., L- 1524 (6 ♂, 6 ♀ UMC); [Mpio.] Tlapacoyan, El Encanto, Río Filobobos, 104 msnm, 8 - XI- 2009, 19 º 59 ' 01 '' N, 97 º 10 ' 08 '' W, L. Cervantes, R. Sites, R. Novelo (1 ³, 1 ♀ CNIN); [Mpio.] Tlapacoyan, El Encanto, Río Filobobos, 08 - Nov- 2009, R. Novelo col. (4 ³, 11 ♀ IEXA); [Mpio.] Tlapacoyan, Ixtacuaco, 30 - Marzo- 2003, R. Novelo col. (1 ♀, IEXA); Mpio. Tlapacoyan, Río Bobos, 19 o 55 ' 39.8 " N, 97 o 09 ' 45.5 " W, 281 m, 14 - Jun- 2013, DRV, Sites, Shepard, Novelo, & PRH colls., L- 1551 (97 ³, 121 ♀ UMC); Mpio. Tlapacoyan, Río Filobobos, El Encanto, 8 November 2009, L- 1123 / 19 º 59.022 ' N, 97 º 10.149 ' W, 178 m, Sites, Cervantes, Novelo (1 ³ UMC); Mpio. Tlapacoyan, Río Filobobos, 19 º 55.662 ' N, 97 º 09.764 ' W, 267 m, 8 Nov 2009, Sites, Cervantes, Novelo colls., L- 1124 (16 ³, 34 ♀ UMC); Mpio. Tlapacoyan, Rojo Gómez, Río Filobobos, 20 ° 02 ' 41.7 '' N, 97 ° 05 ' 38.5 '' W, 89 m, 12 - Jun- 2013, DRV, Sites, Shepard, & PRH colls., L- 1548 (11 ³, 8 ♀ UMC); [Mpio.] Tlapacoyan, Ruinas El Cuajilote (Filobobos) 08 - Nov- 2009, R. Novelo col. (5 ³, 2 ♀ IEXA); [Mpio. Tlapacoyan], Tlapacoyan, El Muro (arroyo) 900 msnm, 13 - Apr- 2004, R. Novelo col. (1 ³ IEXA); [Mpio. Tlapacoyan], Tlapacoyan, Río Tomata, 25 / V / 1985, J. Bueno, R. Barba (2 ♀ CNIN); [Mpio. Tlapacoyan], Tlapacoyan, 28 - Marzo- 1984, H. Velasco col. (1 ♀ IEXA); [Mpio. Zongolica], Zongolica, 11 - VIII- 74, J. Bueno (1 ³ CNIN); same but / Ambrysus circumcinctus Montandon (Ira La Rivers) [det.] (1 ³ CNIN); [Mpio. Zongolica], Zongolica, 11 - VIII- 74, J. Bueno / H. Brailovsky A. det. Ambrysus circumcinctus Montandon (1 ♀ CNIN); San Andrés Tuxtla, 30 - VII- 76, J. Bueno / Ambrysus circumcinctus Montandon det. J. T. Polhemus 19 [??] (1 ♂ CNIN). ZACATECAS: [Mpio. Moyahua de Estrada], Moyahua, 13 I 2005, 3610 ', Río Juchipila / William D. Shepard, leg. (8 ³, 7 ♀ EMEC); [Mpio. Moyahua de Estrada], Río Juchipila at Moyahua, 13 Jan. 2005, C. B. Barr / 21 º 16.29 ' N, 103 º 10.33 ' W, elevation ~ 3850 ft. (1 ³, 2 ♀ EMEC); [Mpio. Moyahua de Estrada], Río Juchipila, 9.1 rd. km S of Moyahua, 13 Jan. 2005, C. B. Barr / 21 º 12.63 ' N, 103 º 10.84 ' W, elevation ~ 3800 ft. (1 ³, 2 ♀ EMEC); [Mpio. Moyahua de Estrada], 9.1 km S Moyahua, 13 I 2005, 3610 ', Río Juchipila / William D. Shepard, leg. (1 ♂, 6 ♀ EMEC).	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888409FFFDFF08FF559CA4FECF.taxon	diagnosis	Diagnosis. This species can be distinguished from the others in this subgenus based on the shorter AGP that slightly curves to right at approximately 45 degrees and is slightly dilated distally (Fig. 9 B). The pseudoparameres are short, subrectangular, and asymmetrical with the left pseudoparamere slightly longer, with posterior margins straight, posterolateral corners rounded, and scattered fine setae (Fig. 9 C). The parameres are symmetrical, longer than wide, with the mesal margin straight, and a row of long setae emanating from the middle of the dorsal surface (Fig. 9 D). The left ventral lobe of the endosoma is lightly sclerotized (Fig. 9 E). The lateral margins of the SGP are narrowed subapically, giving it a characteristic bottleneck outline with an inverted V-shaped notch; the notch has two small pointed lobes (Fig. 9 F).	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888409FFFDFF08FF559CA4FECF.taxon	discussion	Discussion. This species was described based mainly on specimens collected in central Mexico. La Rivers (1951) noticed the prominent bulging eyes on the specimens of the type series and considered this feature diagnostic; however, this character is not particularly different from that exhibited by any other species in this subgenus in Mexico. La Rivers (1951) considered A. pygmaeus to be the most distinctive member of the oblongulus species complex, which he proposed but did not define. Later La Rivers and Nieser (1972) included four Central and South American species (A. bifidus, A. oblongulus, A. scolius, and A. stali) in the oblongulus species complex, but did not include A. pygmaeus as originally proposed (La Rivers 1951, 1953). The taxonomic status of the oblongulus species complex recently has been reviewed (Sites & Reynoso-Velasco 2015). Ambrysus pygmaeus was included in the subgenus Syncollus when the former division Coalescens was elevated to subgeneric rank (La Rivers 1965). The color pattern on this species is similar to A. circumcinctus with the hemelytra solid dark brown and embolia light brown anteriorly (Fig. 9 A). The triangular mark at the base of posterior margin of the corium is small and light brown to reddish brown.	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888409FFFDFF08FF559CA4FECF.taxon	description	Variation. La Rivers considered this a small species ranging from 7.0 to 7.5 mm in length, which is probably why he used pygmaeus as the specific epithet. Some specimens from southern Mexico (Oaxaca) are up to 8.4 mm in length. The small triangular mark at the base of posterior margin of corium is sometimes absent (Fig. 9 A). The two small pointed lobes in the notch on the posterior margin of the SGP can be inconspicuous.	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888409FFFDFF08FF559CA4FECF.taxon	materials_examined	Type locality. Mexico: Mexico: Mpio. Temascaltepec, Temascaltepec. Repository. Holotype is housed at CAS.	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888409FFFDFF08FF559CA4FECF.taxon	distribution	Distribution. This species is distributed through the montane areas of the Mexican Transition Zone that includes the Sierra Madre Occidental (Durango), Sierra Madre del Sur (Oaxaca), and Transmexican Volcanic Belt (Mexico) biogeographic provinces. It is also present on the border of the Transmexican Volcanic Belt and the Balsas Basin biogeographic provinces (Morelos). All specimens examined were collected at elevations above 1500 meters, except those from Oaxaca (L- 1396, L- 1397; Fig. 4), which were 711 and 767 m, respectively. Herrera (2013) cited a record from Guatemala based on the California Academy of Sciences online database; however, we did not find these specimens in the collection at CAS. Published records. Mexico: Estado de México (La Rivers, 1953, 1965, 1971).	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888409FFFDFF08FF559CA4FECF.taxon	materials_examined	Type material examined. HOLOTYPE ♂: [México: México: Mpio.] Temescaltepec [Temascaltepec], Mex. VI- 5 - 33 / H. E. Hinton, R. L. Usinger Collectors / Ambrysus pygmaeus La Rivers HOLOTYPE / Ambrysus pygmaeus La Rivers determined by Ira La Rivers 1948 / California Academy of Sciences Type No. 10058. PARATYPES: Temescaltepec [Temascaltepec], Mex. VI- 5 - 33 / H. E. Hinton, R. L. Usinger Collectors / Ambrysus pygmaeus La Rivers Paratype (2 ³, 2 ♀ EMEC); same but determined by Ira La Rivers ' 50 (1 ³ EMEC); Temescaltepec [Temascaltepec], Mex. VI- 5 - 33 / H. E. Hinton, R. L. Usinger Collectors / Ambrysus pygmaeus La Rivers Paratype / 79 [without terminal abdominal segments] (1 ³ EMEC); same but 80 [without terminal abdominal segments] (1 ♂ EMEC). MORELOS: Cuernavaca, Mex, 1934 / H. E. Hinton / Ambrysus pygmaeus La Rivers PARATYPE (3 ³ EMEC; 1 ³, 1 ♀ SEMC). Material examined. MÉXICO: DURANGO: La Michilía, Arroyo El Temazcal, 10 - Diciembre- 1987. R. Novelo col. (4 ³, 1 ♀ UMC); Res. Biósfera de Michilía, Arroyo el Temascal, 1 - VII- 1985, 2260 msnm, C. Castillo, P. Reyes (1 ³, 2 ♀ CNIN); Res. Biósfera de Michilía, Est. Piedra Herrada, 1 - VII- 1985, 2350 msnm, C. Castillo, P. Reyes (1 ♀ CNIN); same but 2 - VII- 1985, 2260 msnm (1 ♀ CNIN). MÉXICO: Temescaltepec [Temascaltepec], Mex. VI- 5 - 33 / H. E. Hinton, R. L. Usinger Collectors / Ambrysus pygmaeus La Rivers determined by Ira La Rivers 1948 (1 ³ EMEC); Real de Arriba, Temescaltepec [Temascaltepec], Mex., VI- 5 - 1933 / H. E. Hinton, R. L. Usinger Collectors / Ambrysus pygmaeus La Rivers 1953 (Ira La Rivers) (1 ♂ CNIN). MORELOS: Cuernavaca, Mex, 1934 / H. E. Hinton (2 ♀ EMEC); Cuernavaca, VI- 1934 Mex / H. E. Hinton Collector (1 ♀ EMEC). OAXACA: km 72 carr. Putla-Tlaxiaco, 16 km SW of Tlaxiaco, Pte. Los Pinos 17 º 11 ' 11.3 '' N, 97 º 45 ' 10.6 '' W, 2083 m, 31 - May- 2012, DRV & PRH colls., L- 1401 (1 ♀ UMC); Mpio. Putla Villa de Guerrero, Putla de Guerrero, Pte. La Cuchara, 17 º 01 ' 06.7 '' N, 97 º 55 ' 11.8 '' W, 711 m, 30 - May- 2012, DRV & PRH colls., L- 1396 (2 ♂, 2 ♀ UMC); Mpio. Putla Villa de Guerrero, Pluta de Guerrero, 17 º 02 ' 21.5 '' N, 97 º 54 ' 42.9 '' W, 767 m, 30 - May- 2012, DRV & PRH colls., L- 1397 (2 ³ UMC); Mpio. Tlaxiaco, km 81 carr. Putla-Tlaxiaco, San Juan del Río [Río de Ocotepec], Río Ocotepec, 17 º 10 ' 27 '' N, 97 º 46 ' 17.8 '' W, 1967 m, 31 - May- 2012, DRV & PRH colls., L- 1400 (1 ♂ UMC).	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
89238788841DFFE6FF08FC9398ECFDCB.taxon	discussion	Discussion. Stål (1862) established the genus Ambrysus for three Mexican species (A. melanopterus, A. pudicus, and A. signoreti) and indicated it differed from the genus Naucoris Geoffroy in having a deep depression on the anterior margin of the pronotum, a character later used by Montandon (1897) as the diagnostic feature for the subfamily Cryphocricinae. More recently, La Rivers (1951, 1953) provided detailed descriptions of the genus Ambrysus, which includes five subgenera (La Rivers 1952, 1965; Nieser et al. 1999). Of these, Acyttarus La Rivers and Picrops La Rivers are monotypic (A. funebris La Rivers and A. usingeri La Rivers, respectively) (La Rivers 1965); Melloiella De Carlo includes two species [A. lamprus Nieser, Pelli, & de Melo and A. truncaticollis (De Carlo)] (Nieser et al. 1999); Syncollus La Rivers comprises eight species after removal of A. partridgei, and the remaining 71 species belong to the subgenus Ambrysus Stål. La Rivers (1952) divided Ambrysus into two subgenera to accommodate A. usingeri with the subgenus Picrops. The features distinguishing this subgenus were the presence of four or more rows of small spines on the distal end of the hind tibia and the males bearing a short lateral process on tergum VI. In his revision of the fauna of Ambrysus from the United States, La Rivers (1951) recognized the divisions Coalescens and Disjunctus within the subgenus Ambrysus, based on the position of the propleura with respect to the prosternum. The former included species with the propleura closely appressed to the adjacent prosternum (prosternellum). In this case the propleura and the anterior part of the prosternum (probasisternum) are positioned at the same level dorsoventrally. The division Disjunctus included species with the propleura detached from the prosternellum; thus, the propleura are at a lower level (further ventrad) than the probasisternum (Fig. 1). In a later paper, La Rivers (1965) created the subgenus Acyttarus for the species A. funebris, which was the sole species lacking a costal cell in the hind wings. In addition, he elevated the Coalescens division to subgeneric rank as Syncollus (from Greek syn " together " and Latin collus " neck ") and the species originally placed in the Disjunctus division were assigned to the subgenus Ambrysus. Since La Rivers (1951) first used the shape of the male accessory process of tergum VI and the shape of the female mediosternite VII (subgenital plate) in his revision of the fauna of Ambrysus in the United States, these features have been widely used as the two main diagnostic features for distinguishing species of the genus. We have examined all species of Ambrysus from Mexico including most type specimens, and many thousands of museum and freshly collected specimens. The features used by La Rivers are somewhat instructive, although greater intraspecific variation exists in their expression than was previously considered. In addition to these, we discovered that the medial lobes of male tergum VIII (pseudoparameres) and characteristics of the ventral side of the aedeagus also are important diagnostic features at the species and species complex levels (see Sites & Reynoso- Velasco 2015). Each species complex is best characterized by a certain character or combination of these characters, which differs among complexes. For example, in the Ambrysus stali La Rivers complex, the left ventral lobe of the phallosoma is consistent among species, whereas the other three characters are more informative at the species level (Sites & Reynoso-Velasco 2015).	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888407FFFEFF08FEA89FE7FB8B.taxon	description	Description. Brachypterous male. HOLOTYPE, length 7.76; maximum width 4.76. Paratypes (n = 10), length 7.04 – 8.32 (mean = 7.69); maximum width 4.08 – 4.88 (mean = 4.45). General shape elongate, parallel-sided; widest across embolia (Fig. 10 A). Dorsal coloration of hemelytra dark brown; head, pronotum, and legs yellowish brown. Dorsal surface coarsely punctate. Ventral coloration of head and thorax medium brown, abdomen with dense golden brown pubescence. Head. Head length 1.52; maximum width 2.08. Mostly yellowish brown with dark brown stripe at midline becoming wider posteriorly, coarsely punctate. Eyes convergent anteriorly, synthlipsis 0.96; thin band of cuticle along posterolateral margin of eye; eyes not raised above level of vertex or pronotum. Anterior margin between eyes slightly convex, extending anteriorly in front of eyes 9 % of head length; posterior margin between eyes strongly convex, extending posteriorly 38 % of head length. Labrum width 2.1 × length, evenly rounded. Labium with three visible yellowish brown segments, darkening distally, extending 0.42 beyond labrum not including extruded stylets. Antennal proportions 2: 5: 10: 10, length 0.56, extending to near lateral margin of eye, elongate hairs on segment 4 and distal half of 3. Thorax. Pronotum coarsely punctate, ground color yellowish brown; transverse sulcus marking anterior border of transverse band in posterior 1 / 4; transverse band pale yellow with color extending anteriorly on midline as elongate, pointed marking; ovoid dark brown mark on either side of midline at anterior margin; dark, broad, submarginal band extending from eye to transverse band; lateral margins dark brown, convergent, posterior half convex, anterior half mostly straight, explanate; posterior margin straight; anterior margin deeply concave between eyes to embrace convex posterior margin of head; posterolateral corners rounded; width 3.1 × length; length at midline 1.28; maximum width at posterolateral corners 4.0. Prothorax ventrally pruinose throughout; apices of propleura meeting at midline, closely appressed to prosternellum; propleuron brown throughout, medial 2 / 3 of posterior margin with elongate golden setae. Probasisternum with sharp medial carina and without row of setae lateral to carina. Prosternellum covered by apices of propleura. Scutellum coarsely punctate, triangular, entirely brown with narrow light brown coloration along margins and midline, width 2.0 × length, width 2.68, length 1.32. Hemelytra densely punctate, dark brown, not reaching apex of tergum V, length 5.36 (chord measurement). Clavus with thin, yellow transverse stripe at base, length 4.5 × width, length 2.92, width 0.64; claval commissure dark brown, length 0.72. Embolium length 2.28, greatest width 0.54; lateral margin convex, yellow brown in anterior 2 / 3, becoming darker posteriorly. Oblique suture connecting claval and embolar sutures near bases. Hind wings extending to middle of tergum IV. Mesobasisternum midventral tumescence negligible, with sulcus on midline continuing through triangular mesosternellum. Metasternellum (= metaxyphus) transverse, subtriangular, with median carina, apex acute. Legs. All legs segments yellowish green. Profemur posterior margin with row of tightly arranged setae in basal 2 / 3, row of short brown spines along middle third, spines generally single proximally becoming small combs of two to five spines distally; anterior margin with dense pad of setae without associated spines. Protibia and tarsus with occlusal inner surface flattened and with spatulate setae; tarsus immovable, one-segmented; pretarsal claw single, minute, triangular. Procoxa with cluster of stout, brown anteromedial spines. Meso- and metacoxae partially recessed into thorax. Meso- and metafemora with row of short, brown spines on anterior margin; spines restricted to basal half on mesofemur, nearly full length of metafemur. Mesotibia with ventrolateral, ventromedial, dorsolateral, and dorsomedial rows of stout reddish-brown spines; ventromedial rows intermixed with combs of two to six spines. Metatibia with ventrolateral, dorsolateral, and dorsomedial rows of stout reddish-brown spines. Meso- and metatibiae with semi-circlet of spines at apex of dorsal and mesal margins; two comb rows of stout spines near apex of ventral margin, lateral margin without spines at apex to accommodate flexed tarsus. Meso- and metatibiae and metatarsus with long, pale swimming hairs; hairs profuse on metatibia and - tarsus. Meso- and metapretarsi with paired claws slender, gently curved, with small basal tooth. Leg measurements as follows: foreleg, femur 2.2, tibia 1.88, tarsus 0.40; middle leg, femur 2.2, tibia 1.96, tarsomeres 1 – 3, 0.24, 0.32, 0.48; hind leg, femur 2.60, tibia 2.88, tarsomeres 1 – 3, 0.20, 0.52, 0.56. Abdomen. Dorsally with connexiva III – VIII exposed, each laterotergite brown in anterior half and yellow in posterior half giving checkered appearance (Fig. 10 A); lateral margin finely serrate, marginal row of short yellow setae, group of trichobothria near posterolateral corners. Posterolateral corners of II (visible ventrally) – IV narrowly rounded to right angled and not spinose, V – VII bluntly acute. Accessory genitalic process of tergum VI short, curved to right at approximately 80 degrees, dilated distally (Fig. 10 B). Medial lobes of tergum VIII (pseudoparameres) short, wide, subrectangular, symmetrical, posterior margin straight; posterolateral corners more heavily sclerotized, nearly right-angled, slightly deflexed ventrad (Fig. 10 C). Ventrally entirely golden brown, with dense pile of fine hairs. Lateral margin with thin, glabrous band. Glabrous rounded to oval patches around spiracles on laterosternites II – VII. Phallosoma elongate, linear, constricted basally, sclerotized dorsally; right and left ventral lobes of endosoma papillose, lightly sclerotized (Fig. 10 E). Parameres symmetrical, longer than wide, mesal margin straight, narrowly rounded anteriorly, row of long dark setae emanating from middle of dorsal surface. Proctiger short, length 0.6 × width. Pygophore with elongate setae sparsely distributed over most of surface, with thick brush of elongate setae on posterior margin (Fig. 10 D). Brachypterous female. Paratypes (n = 10), length 7.84 – 9.04 (mean = 8.22); maximum width 4.40 – 5.28 (mean = 4.76). Similar to male in general structure and coloration except as follows: Hemelytra surpassing posterior margin of tergum VI. Mediosternite VII (subgenital plate) width 1.2 × length; length at midline 1.02; maximum width 1.30; posterior margin concave; two short, pointed lobes near center of concavity; posterolateral corners rounded, produced posteriorly further than medial lobes (Fig. 10 F). Macropterous male. Unknown. Macropterous female. Paratype (n = 1), length 8.48; maximum width 4.80. Coloration, setation, pronotum proportions, posterolateral corners of pronotum, scutellum proportions, embolium length generally same as for brachypterous female. Otherwise clavus and claval commissure correspondingly longer: Clavus length 6.33 × width, length 3.08, width 0.60; claval commissure length 0.80. Hind wings extending to posterior margin of tergum VI. Variation. The yellowish brown coloration reported here is green or yellowish green in live specimens or specimens that were dry-killed, pinned soon thereafter, and kept in darkness (Fig. 10 A). Specimens can present the transverse band on the pronotum with pale yellow coloration in the anterior half and brown posteriorly. The posterolateral corners of the subgenital plate can be slightly pointed.	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888407FFFEFF08FEA89FE7FB8B.taxon	diagnosis	Diagnosis. This species presents a characteristic color pattern on the prothorax with two dark, broad, submarginal bands. The flat and subrectangular shape of the male pseudoparameres is different from that of any other species in this subgenus in Mexico. Although the subgenital plate is similar to that in some females of A. circumcinctus, the pointed central lobes are distinctive for this species. Habitat description. Río Tizapam is a small stream that originates on the eastern slope of the Sierra Madre Oriental (Veracruz). The river is part of the Río La Antigua Basin that drains to the Gulf of Mexico. At the type locality the river is small with boulders and cobble (Fig. 11). Most of the collecting was conducted in areas with current and among the roots of riparian vegetation.	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888407FFFEFF08FEA89FE7FB8B.taxon	distribution	Distribution. In central Veracruz, this species is distributed in rivers of the Río La Antigua Basin, an area that is a transition zone between the Transmexican Volcanic Belt and the Sierra Madre Oriental biogeographic provinces. The northern distribution of the species in the states of Hidalgo and Puebla is on the eastern slopes of the highlands forming the Sierra Madre Oriental biogepgraphic province (Fig. 4). We found A. totonacus and A. circumcinctus syntopically in the states of Hidalgo (L- 1522) and Veracruz (L- 1290, L- 1296, L- 1549). Also, we found A. totonacus and A. xico n. sp. at the same locality in the state of Veracruz (L- 1557).	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888407FFFEFF08FEA89FE7FB8B.taxon	etymology	Etymology. The specific epithet refers to the Totonac culture of the Totonacapan region, which originally included parts of the east-central Mexican states of Hidalgo, Puebla, and Veracruz, the states where this species occurs. Repositories. The holotype and some paratypes will be deposited in the Colección Nacional de Insectos (Mexico City); additional paratypes will be deposited in the Colección Entomológica del Instituto de Ecología A. C. (Xalapa), Enns Entomology Museum (University of Missouri), and the United States National Museum of Natural History (Washington D. C.).	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888407FFFEFF08FEA89FE7FB8B.taxon	materials_examined	Type material examined (brachypterous unless otherwise noted). HOLOTYPE ♂: MÉXICO: VERACRUZ: Mpio. Xico, [Río Tizapam], 1.86 km SW of Xico, 19 ° 24 ' 26.7 '' N, 97 ° 00 ' 53.5 '' W, 1300 m, 16 June 2013, L- 1559 / Sites, Shepard, Novelo, Reynoso-Velasco & Reynoso-Hernández; PARATYPES: same data as holotype (1 ³, 2 ♀ UMC). HIDALGO: Mpio. Tlanchinol, 3.4 km W of Chalchocotipa, road to Santa Maria Catzotipan, 0 6 June 2013, L- 1522 / west of Hwy 105, km 3 + 400, 528 m, 21 ° 02 ' 36.4 '' N, 98 º 36 ' 00.8 '' W, Shepard, Sites, Reynoso-Velasco & Reynoso-Hernández (1 ♂, 1 ♀ UMC). PUEBLA: Cuetzalan, El Cuichat, 870 m, 12 - Jul- 2007, R. Novelo col. (3 ♂, 5 ♀ UMC). VERACRUZ: Mpio. Coatepec, Agua Alegre, Río Huehueyapan, 4 April 2011, L- 1290, colls. R. Sites & V. Amaya / elev. 1198 m, 18 [19 º] 26.884 ' N, 96 58.867 ' W (1 ³, 2 ♀ CNIN; 1 ³, 2 ♀ IEXA; 1 ³, 2 ♀ UMC); Coatepec, Arroyo Bola de Oro, 7 - Nov- 2009, R. Novelo col. (1 ³ CNIN; 1 ³ IEXA; 4 ³, 3 ♀ UMC; 1 ³, 1 ♀ USNM); Coatepec, Río Huehueyapan, 04 - Apr- 2011, R. Novelo col. (1 ³, 3 ♀ UMC; 1 ³, 1 ♀ USNM); Coatepec, Consolapa, Río Pixquiac (in riffles), 15 - May- 2004, R. Novelo col. (2 ³, 5 ♀ UMC); Mpio. Atzalán, Pte. Tomata, Río Alseseca, 19 ° 55 ' 15.7 '' N, 97 ° 13 ' 28.2 '' W, 514 m, 13 - Jun- 2013, L- 1549 / rock, gravel, riffles, and marginal veg., Sites, Shepard, Novelo, Reynoso-Hernández & Reynoso-Velasco, colls. (3 ♀ UMC); Mpio. Atzalán, Pte. Tomata, Río Alseseca, 6 April 2011, L- 1296 / 500 m, 19 ° 55.335 ' N, 97 ° 13.385 ' W, Sites, Shepard, Amaya & Novelo colls. (1 ♀ macropterous UMC); Mpio. Xico, Arroyo Avestruces, vegetated margin of rocky stream in cattle pasture, 16 - Jun- 2013, L- 1557 / 1401 m, 19 ° 23 ' 53.8 '' N, 97 ° 02 ' 29.9 '' W, Sites, Shepard, Novelo, Reynoso-Hernández & Reynoso- Velasco, colls., (3 ³, 1 ♀ UMC); Mpio. Xico, Arroyo Avestruces, 1395 m, 29 - May- 2014, L- 1799 / 19 ° 23 ' 53.8 '' N, 97 ° 02 ' 29.9 '' W, Reynoso-Velasco & Reynoso-Hernández (1 ♀ UMC).	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888404FFFBFF08FB699840F95A.taxon	description	Description. Brachypterous male. HOLOTYPE, length 7.92; maximum width 4.76. Paratypes (n = 2), length 7.92; maximum width 4.32 – 4.48 (mean = 4.40). General shape elongate, parallel-sided; widest across embolia (Fig. 12 A). Dorsal coloration of head, pronotum, and hemelytra dark brown; legs yellowish brown. Dorsal surface coarsely punctate and tuberculate. Ventral coloration of head and thorax medium brown, abdomen with dense golden brown pubescence. Head. Head length 1.56; maximum width 1.96. Mostly dark brown with yellowish brown stripe on midline posteriorly and on either side of midline between eyes, coarsely punctate and tuberculate. Eyes convergent anteriorly, synthlipsis 1.08; thin band of cuticle along posterolateral margin of eye; eyes not raised above level of vertex or pronotum. Anterior margin between eyes slightly convex, extending anteriorly in front of eyes 10 % of head length; posterior margin between eyes strongly convex, extending posteriorly 41 % of head length. Labrum width 1.9 × length, evenly rounded. Labium with three visible yellowish brown segments, darkening distally, extending 0.44 beyond labrum not including extruded stylets. Antennal proportions 3: 7: 10: 7, length 0.54, extending to near lateral margin of eye, elongate hairs on segment 4 and distal half of 3. Thorax. Pronotum coarsely punctate and tuberculate, ground color yellowish brown; transverse sulcus marking anterior border of transverse band in posterior 1 / 4; transverse band pale yellow with color extending anteriorly on midline as elongate, pointed marking; ovoid dark brown mark on either side of midline at anterior margin; dark, broad, submarginal band extending from eye to transverse band, reaching lateral margin; lateral margins dark brown, convergent, evenly convex, explanate; posterior margin straight; anterior margin deeply concave between eyes to embrace convex posterior margin of head; posterolateral corners rounded; width 2.8 × length; length at midline 1.36; maximum width at posterolateral corners 3.92. Prothorax ventrally pruinose throughout; apices of propleura meeting at midline, closely appressed to prosternellum; propleuron brown throughout, medial 2 / 3 of posterior margin with elongate golden setae. Probasisternum with sharp median carina, with row of short setae lateral to carina. Prosternellum covered by apices of propleura. Scutellum coarsely punctate and tuberculate, triangular, entirely brown with light brown on corners, width 2.0 × length, width 2.48, length 1.22. Hemelytra densely punctate and tuberculate, dark brown, surpassing apex of tergum V, length 5.44 (chord measurement). Clavus with thin, yellow transverse stripe at base, length 4.8 × width, length 2.88, width 0.60; claval commissure yellowish brown, length 0.98. Embolium length 2.36, greatest width 0.62; lateral margin convex, light brown in anterior 2 / 3, becoming darker posteriorly. Oblique suture connecting claval and embolar sutures near bases. Hind wings extending to anterior 1 / 4 of tergum V. Mesobasisternum midventral tumescence negligible, with sulcus on midline continuing through triangular mesosternellum. Metasternellum (= metaxyphus) transverse, subtriangular, with median carina, apex acute. Legs. All legs segments brown to yellowish brown. Profemur brown; posterior margin with row of tightly arranged setae in basal 2 / 3, row of short brown spines along middle third, spines generally single proximally becoming small combs of two to five spines distally; anterior margin with dense pad of setae without associated spines. Protibia and tarsus with occlusal inner surface flattened and with spatulate setae; tarsus immovable, onesegmented; pretarsal claw single, minute, triangular. Procoxa with cluster of stout, brown anteromedial spines. Meso- and metacoxae partially recessed into thorax. Meso- and metafemora with row of short, brown spines on anterior margin; spines restricted to basal half on mesofemur, nearly full length of metafemur. Mesotibia with ventrolateral, ventromedial, dorsolateral, and dorsomedial rows of stout reddish-brown spines; ventromedial rows intermixed with combs of two to five spines. Metatibia with ventrolateral, dorsolateral, and dorsomedial rows of stout reddish-brown spines. Meso- and metatibia with semi-circlet of spines at apex of dorsal and mesal margins; two comb rows of stout spines near apex of ventral margin, lateral margin without spines at apex to accommodate flexed tarsus. Meso- and metatibiae and metatarsi with long, pale swimming hairs; hairs profuse on metatibia and - tarsus. Meso- and metapretarsi with paired claws slender, gently curved, with small basal tooth. Leg measurements as follows: foreleg, femur 2.12, tibia 1.94, tarsus 0.40; middle leg, femur 2.2, tibia 1.96, tarsomeres 1 – 3, 0.18, 0.34, 0.44; hind leg, femur 2.56, tibia 2.84, tarsomeres 1 – 3, 0.20, 0.50, 0.56. Abdomen. Dorsally with connexiva III – VIII exposed, each laterotergite dark brown in posterior half and light brown in anterior half giving checkered appearance (Fig. 12 A); lateral margin finely serrate, marginal row of short yellow setae, group of trichobothria near posterolateral corners. Posterolateral corners of II (visible ventrally) – IV narrowly rounded to right angled and not spinose, V obtuse and not spinose, VI – VII bluntly acute. Accessory genitalic process of tergum VI short, curved to right at approximately 45 degrees, acuminate distally (Fig. 12 B). Medial lobes of tergum VIII (pseudoparameres) long, semirounded, asymmetrical, with left pseudoparamere slightly narrower (Fig. 12 C). Ventrally entirely golden brown, with dense pile of fine hairs. Lateral margin with thin, glabrous band. Glabrous rounded to oval patches around spiracles on laterosternites II – VII. Phallosoma elongate, linear, constricted basally, sclerotized dorsally; right and left ventral lobes of endosoma papillose, left ventral lobe lightly sclerotized (Fig. 12 E). Parameres symmetrical, almost as long as wide, mesal margin straight, narrowly rounded anteriorly, long setae on posterior half. Proctiger short, length 0.6 × width. Pygophore with elongate setae sparsely distributed over most of surface, with thick brush of elongate setae on posterior margin (Fig. 12 D). Brachypterous female. Paratypes (n = 3), length 8.48 – 9.28 (mean = 8.80); maximum width 4.88 – 5.20 (mean = 4.99). Similar to male in general structure and coloration except as follows: hemelytra surpassing posterior margin of tergum VI. Mediosternite VII (subgenital plate) width 1.4 × length; length at midline 0.88; maximum width 1.30; with deep medial concavity twice as wide as long; posterolateral corners narrowly rounded (Fig. 12 F). Macropterous male. Unknown. Macropterous female. Unknown. Variation. The small number of specimens (3 ♂, 3 ♀) did not allow us to assess variation of the diagnostic features.	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888404FFFBFF08FB699840F95A.taxon	diagnosis	Diagnosis. Ambrysus xico presents distinct, minute, individualized small tubercles on the head, prothorax, scutellum, and hemelytra not connected by ridges. The other species revised in this study present these tubercles interconnected by sclerotized ridges, resulting in a polygonal pattern. This species has darker coloration when compared with the other Mexican species of Syncollus. The dark brown submarginal band on the prothorax is wider and reaches the lateral margin anteriorly. The posterolateral corners of tergum V are not acutely angled as in the other the species, but are rounded and obtuse. The pseudoparameres are asymmetrical with the left pseudoparamere slightly longer and narrower. These pseudoparameres are similar to those of A. circumcinctus, although not as constrained medially. The female subgenital plate has a concavity that is exactly twice as wide as long and is similar to that of A. baeus, although the posterolateral corners of the latter species are narrower. Habitat description. The stream at the type locality (Fig. 13) is part of the Río La Antigua basin in the central area of Veracruz state. The stream originates on the eastern slopes of the Cofre de Perote volcano at the eastern limit of the Transmexican Volcanic Belt. This mountain stream is approximately 5 meters wide and has areas with strong current and riffles. Most of our collecting was conducted in those areas, and although species in this subgenus prefer habitats with strong current, A. xico was not commonly collected; thus this species might have other habitat preferences.	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888404FFFBFF08FB699840F95A.taxon	distribution	Distribution. This species was collected in Veracruz from three mountainous streams above 1400 meters elevation in the Río La Antigua Basin in the central part of the state, the transition zone between the Transmexican Volcanic Belt and the Sierra Madre Oriental biogeographic provinces. In the state of Querétaro, A. xico was collected in a small stream at 2000 meters elevation. Our extensive collecting suggests that A. xico could be endemic to streams at high elevation in the Transmexican Volcanic Belt biogeographic province (Fig. 4). We found A. xico and A. totonacus n. sp. syntopically at L- 1557 in the state of Veracruz.	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888404FFFBFF08FB699840F95A.taxon	etymology	Etymology. The specific epithet is a noun in apposition and refers to the municipality of Xico, in the central part of Veracruz, where most of the specimens were collected. Repositories. The holotype and one female paratype will be deposited in the Colección Nacional de Insectos (Mexico City); additional paratypes are deposited in the Enns Entomology Museum (University of Missouri) and the Essig Museum of Entomology Collection (Berkeley).	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
892387888404FFFBFF08FB699840F95A.taxon	materials_examined	Type material examined. HOLOTYPE ♂: MÉXICO: VERACRUZ: Mpio. San Andrés Tlalnehuayocan, 1.16 km NW of Tres Marias, 19 º 30 ' 55.6 '' N, 97 º 00 ' 27.4 '' W, 15 June 2013, L- 1556 / 1602 m, [Arroyo Piedra Blanca] rocky mountain stream, Reynoso-Velasco, Sites, Shepard, Novelo & Reynoso-Hernández, colls. PARATYPES: QUERÉTARO: Mpio. Amealco, Puerta de Alegrías, Arroyo las Zúñigas, 8 - VII- 2000, C. B. Barr / S of Galindo, 20 º 20 ' 28 '' N, 100 º 07 ' 10 '' W, elev. 2000 m (1 ♀ EMEC). VERACRUZ: Mpio. Xico, Arroyo Avestruces, vegetated margin of rocky stream in cattle pasture, 16 June 2013, L- 1557 / 1401 m, 19 º 23 ' 53.8 '' N, 97 º 02 ' 29.9 '' W, Sites, Shepard, Novelo, Reynoso-Hernández & Reynoso-Velasco, colls. (1 ³ UMC); same locality information but rootmats, 29 May 2014, L- 1799, DRV & PRH (1 ³ UMC); Mpio. Xico, Xico Viejo, 19 º 27 ' 14.10 '' N, 97 º 03 ' 31.5 '' W, 1780 m, 30 - Ago- 1997, R. Arce col. (1 ♀ CNIN; 1 ♀ UMC).	en	Reynoso-Velasco, Daniel, Sites, Robert W., Novelo-Gutiérrez, Rodolfo (2016): The Ambrysus Stål (Heteroptera: Naucoridae: Cryphocricinae) of Mexico: Revision of the subgenus Syncollus La Rivers. Zootaxa 4126 (1), DOI: 10.11646/zootaxa.4126.1.2
