taxonID	type	description	language	source
882FBC10FFE6CD5F90C777E4F34CFB2F.taxon	materials_examined	Type locality: Kyrgyzstan, Alai Mts., Tengizbai River. Type material (holotype) in MWM, examined. Distribution: Kyrgyzstan, Alai Mts. Habitat and biology. Flight period: July. Altitude: 2100 ̄ 3500 m.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE6CD5F90C7717DF34CF984.taxon	materials_examined	Type locality: Kazakhstan, Prov. Almaty, Kamenka, 76 ° 73´E; 43 ° 15´N. Type material (holotype) in MWM, examined. Distribution: Kazakhstan, Zailiiskii Ala-Tau Mts. Habitat and biology. Flight period: July. Altitude: 1300 ̄ 1400 m.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE6CD5890C77296F3E9FF2A.taxon	materials_examined	Type locality: Alexandropol (im südwestlichen Caucasus Gebiete, Armenien) [Gyumri (old name - Leninakan), Armenien]. Type material (holotype) in MHUB, examined. Synonymy:	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE6CD5890C77296F3E9FF2A.taxon	distribution	Distribution: Armenia, Azerbaijan, Turkey, С. Iran. Habitat and biology. Flight period: July ̄ August. Altitude: 1500 ̄ 2700 m.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE1CD5890C77577F3E9FD83.taxon	materials_examined	Type locality: Thianshan occ., Tsimgan [Uzbekistan, Chatkal Mts. Range, Bol’shoi Chimgan]. Type material (lectotype) in MHUB, examined. Distribution. Kazakhstan, Kirgyziya, Uzbekistan (Yakovlev, 2006 c). Habitat and biology. Flight period: July ̄ August. Altitude: 1400 ̄ 2300 m.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE1CD5890C77689F3E9FC18.taxon	materials_examined	Type locality: Jazgulem-Tale (südlicher Darvaz) [Tadzhikistan, Darvaz Mts. Range, Yazgulem village]. Type material (holotype) in ZMKU, examined. Distribution: Tadzhikistan (Darvaz Mts.). Habitat and biology. Flight period: July ̄ August. Altitude: 1700 ̄ 2340 m.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE1CD5890C77022F34CFAF1.taxon	materials_examined	Type locality: NW Pakistan, Swat prov., Gabral-Tal. Type material (holotype) in MNHW, examined. Distribution: NW Pakistan, Swat prov., SE Afghanistan. Habitat and biology. Flight period: July. Altitude: 2100 ̄ 2350 m.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE1CD5890C77144F34CF94F.taxon	distribution	Distribution: Afghanistan, Bamian prov. Habitat and biology. Flight period: July. Altitude: 2800 ̄ 2900 m.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE1CD5890C772E7F3C2F833.taxon	materials_examined	Type locality: Samarkand [Uzbekistan, Samarkand]. Type material (lectotype) in ZISP, examined. Distribution: Kyrgyzstan, Uzbekistan, Tajikistan, N. Iran, Afghanistan (Yakovlev 2006). Habitat and biology. Flight period: July ̄ September. Altitude: 700 ̄ 3500 m.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE0CD5990C774ACF399FDCB.taxon	materials_examined	Type locality: Provinz Samarkand [Uzbekistan, Samarkand]. Type material (cotypes) in MHUB, examined. Synonymy:	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE0CD5990C77651F3F5FCA0.taxon	materials_examined	Type locality: S. Persia, Elburs mont., Kendevan pass. Type material (holotype) in ZSM, examined. Distribution: Iran. Habitat and biology. Flight period: June ̄ August. Altitude: 1900 ̄ 2800 m.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE4CD5F90C77192F56AFDE0.taxon	materials_examined	Material. Holotype: ♂, Kyrgyzstan, Toktogul Reservoir (NE bank), 1000 m, 5. x. 2015, leg. A. Irtlach (ZISP).	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE4CD5F90C77192F56AFDE0.taxon	description	Description. Wingspan 27 mm. Antenna pale brown, bipectinate with short comb; length of comb processes in middle third of antenna equal to rod diameter. Thorax and abdomen covered with pale yellow hairs. Patagia reddish. Fore wing ochre brown with pattern of dark and pale elements. Brown smears on costal edge, narrow longitudinal smear in central cell, round pale yellow portions in medial and cubital area. Hind wing grey, without pattern. Marginal border of both wings brown, fringe yellowish, unicolorous. Male genitalia. Structure typical of specimen of the genus. Uncus large, wide, apex rounded. Gnathos arms thick, of medium length, gradually narrowing from base to apex. Gnathos medially belt-like, with even edges. Valva of medium width, widening on border between middle and distal thirds, apex rounded, slightly sclerotized. Semicircular crest with almost even edge on border between sclerotized and membranous parts on costal edge of valva. Transtilla process with wide base and narrowed apex. Juxta saddle-like with deep notch on dorsal surface and two long clavately widened lateral processes. Saccus semicircular, of medium size. Phallus slightly shorter than valva, thin, poorly curved in middle third. Vesica aperture in dorso-apical position, in length equal to ¼ of phallus, vesica without cornuti. - small size (wingspan of C. nasreddin is 32 mm); - dark basal part of fore wing (in C. nasreddin, the basal part of fore wing is pale); - grey hind wing (in C. nasreddin, the hind wing is brown); - clavate lateral processes of the juxta (in C. nasreddin, the processes are bar-like without expanded apices); - lanceolately tapered transtilla processes (in C. nasreddin, the processes are uncinately bent); - in C. nasreddin, the membranous part of valva is shorter.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE4CD5F90C77192F56AFDE0.taxon	etymology	Etymology. The new species is named after the collector, Alexander Irtlach from Bishkek, Kyrgyzstan.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE0CD5990C777EAF368FB62.taxon	materials_examined	Type locality: Kyrgyzstan, Toktogul reservoir (NE bank). Type material (holotype) in ZISP, examined. Distribution: Kyrgyzstan. Habitat and biology. Flight period: October. Altitude: 1000 m.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE0CD5990C7712FF192F9DE.taxon	materials_examined	Type material (holotype) in MHUB, examined. Distribution: Kyrgyzstan.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE0CD5990C7726DF38FF8B4.taxon	materials_examined	Type locality: Afghanistan, Khurd-Kabul, 30 km SO v. Kabul. Type material (holotype) in MNHW (coll. E. Vartian), examined. Distribution: Afghanistan (Paghman Mts.). Habitat and biology. Flight period: June-July. Altitude: 2200 ̄ 3000 m.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE3CD5A90C774ACF360FE77.taxon	materials_examined	Type locality: Turkey, prov. Hakkari, Yüksekova Mts., 2.5 km E of Guseldere pass. Type material (holotype) in MWM, examined. Distribution: Turkey, Hakkari prov. Habitat and biology. Flight period: August. Altitude: 2600 m.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE3CD5A90C775C5F20EFD17.taxon	materials_examined	Type locality: Sine-Sefid, Mian-Kotal und in Comèe [Iran]. Type material (holotype) in MNHS.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE3CD5A90C77729F34CFC64.taxon	distribution	Distribution: Iran, Iraq, Syria, Lebanon (Yakovlev 2006; Saldaitis et al. 2007). Habitat and biology. Flight period: July. Altitude: 2400 ̄ 2700 m.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE3CD5A90C77039F341FAC8.taxon	materials_examined	Type material (holotype) in MWM, examined. Distribution: E. Turkey. Habitat and biology. Flight period: June ̄ July. Altitude: 1000 m.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE3CD5A90C77165F2A9F9F7.taxon	materials_examined	Type locality: Askhabad [Turkmenistan, Ashkhabad]. Type material (lectotype) in ZISP, examined.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE3CD5A90C77249F3A8F8C4.taxon	distribution	Distribution: Turkmenistan. Habitat and biology. Flight period: August. Altitude: 1000 ̄ 2400 m.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE3CD5B90C77356F357FEC1.taxon	materials_examined	Type locality: Iraq, 200 km N. Baghdad, östlich von Kirkuk. Type material (holotype) in MWM, examined. Distribution: N. Iraq. Habitat and biology. Flight period: May. Altitude: 1000 ̄ 2000 m.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE2CD5B90C77554F3DBFD5F.taxon	materials_examined	Type locality: Nuchur [Turkmenistan]. Type material (lectotype) in ZISP, examined. Distribution: SE Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan, Iran, Afghanistan (Yakovlev, 2006). Habitat and biology. Flight period: end of April ̄ June. Altitude: 600 ̄ 3000 m.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE2CD5B90C776EDF34AFC34.taxon	materials_examined	Type locality: Uzbekistan, W. Tian-Shan, Zeravshan Mts., Kitab geolog. reserv. Type material (holotype) in MSW, examined. Distribution: Uzbekistan, W. Tian-Shan, Zeravshan Mts., Ghissar Mts. Habitat and biology. Flight period: September. Altitude: 2100 m.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE2CD5B90C77006F341FA92.taxon	materials_examined	Type material (holotype) in MWM, examined. Distribution. E. Turkey, Sivas prov. Habitat and biology. Flight period: June ̄ July. Altitude: 1600 m.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE2CD5B90C7719FF304F96B.taxon	materials_examined	Type locality: Uzbekistan, Ghissar Range, 60 km E. Shakhrisabz. Type material (holotype) in MWM, examined. Distribution: Uzbekistan, Ghissar Mts. Habitat and biology. Flight period: July. Altitude: 2500 m.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE2CD5B90C77331F194F80B.taxon	materials_examined	Type locality: Namangan, at Tosté on the river Padsha-ata. Type material (holotype) in ZISP. Possibly lost. Distribution: Uzbekistan.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFEDCD5490C774ACF3A6FE77.taxon	materials_examined	Type locality: Aschabad [Turkmenistan, Ashkhabad]. Type material (cotypes) in ZISP, examined. Distribution: Turkmenistan, Iran, Afghanistan (Yakovlev, 2006). Habitat and biology. Flight period: August. Altitude: 700 ̄ 2000 m.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFEDCD5490C775CFF304FCDB.taxon	materials_examined	Type locality: [Kyrgyzstan], Fergana Valley, 27 km S Osh. Type material (holotype) in MWM, examined. Distribution: Kyrgyzstan, Fergana Valley. Habitat and biology. Flight period: July. Altitude: 1200 m.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFEDCD5490C77774F387FBBF.taxon	materials_examined	Type locality: Thianshan occ., Bolshoj Tsimgan [Uzbekistan, Chatkal Mts. Range, Bol’shoi Chimgan]. Type material (syntypes) in MHUB, examined. Distribution: S. Kazakhstan, Kyrgyzstan, Uzbekistan, Tajikistan (Yakovlev, 2006). Habitat and biology. Flight period: May ̄ July. Altitude: 600 ̄ 2000 m.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE6CD5F90C776EDF40FFC52.taxon	materials_examined	Type species by monotypy: Cossulus argentatus Staudinger, 1887. Synonymy:	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFEDCD5490C77097F3D0F9F7.taxon	materials_examined	Type locality: Kulab [Tajikistan, Kulyab]. Type material (holotype) in MHUB, examined. Distribution: Kyrgyzstan, Uzbekistan, Tajikistan (Yakovlev, 2006). Synonymy:	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE8CD5190C7722AF2A9F900.taxon	materials_examined	Type locality: Askhabad, Pul-i-Hatum, Tekke [Turkmenistan, Ashkhabad]. Type material (lectotype) in ZISP, examined.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFF4CD4D90C7768CF392FC17.taxon	materials_examined	Type locality: 40 km SW v. Kabul, Afghan [instan]. Type material (holotype) in MNHW, examined. Distribution: E. Afghanistan, N. Pakistan (Yakovlev, 2006). Habitat and biology. Flight period: May ̄ June. Altitude: 1500 ̄ 3000 m.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFE8CD5190C7730BF388F80A.taxon	distribution	Distribution: Turkmenistan, N. Afghanistan, NE Iran (Yakovlev, 2006). Habitat and biology. Flight period: April ̄ June. Altitude: 300 ̄ 1500 m.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
882FBC10FFF4CD4D90C77025F145FAD0.taxon	materials_examined	Type material (syntypes) in ZISP. Possibly lost. Distribution: Iran.	en	Yakovlev, Roman V., Witt, Thomas J. (2017): World catalogue of the genus Cossulus Staudinger, 1887 (Lepidoptera, Cossidae) with description of Cossulus irtlachi sp. nov. from Kyrgyzstan. Zootaxa 4311 (1), DOI: 10.11646/zootaxa.4311.1.4
