taxonID	type	description	language	source
8F69AC09C709FF90D113FDC5CD381311.taxon	description	(Figs. 1 – 19)	en	Alarie, Yves, Michat, Mariano C., Hendrich, Lars, Watts, Chris H. S. (2018): Larval Description and Phylogenetic Placement of the Australian Endemic Genus Barretthydrus Lea, 1927 (Coleoptera: Dytiscidae: Hydroporinae: Hydroporini: Sternopriscina). The Coleopterists Bulletin 72 (4): 639-661, DOI: 10.1649/0010-065X-72.4.639, URL: http://dx.doi.org/10.1649/0010-065x-72.4.639
8F69AC09C709FF90D113FDC5CD381311.taxon	diagnosis	Diagnosis. Instar III of Barretthydrus can be distinguished from those of other genera of Australian Hydroporini that have been well studied (i. e., Antiporus and Paroster) by the following combination of characters: HL = 1.24 – 1.36 mm; HL / HW <1.40; nasale broad, subtriangular, not spatulate apically (Figs. 1, 15 – 16); parietals constricted at level of occipital suture (Figs. 15 – 16); primary seta FR 7 hair-like (Fig. 1); A 4 / A 3> 0.30; labial palpus composed of two palpomeres (Figs. 5 – 6); prementum lacking spinulae along lateral margins (Figs. 5 – 6); L 3 / HW <3.50; primary seta FE 7 present (Fig. 8); natatory setae present on dorsal margin of femora, tibiae, and tarsi (Fig. 18); U 1 / U 2 ’ <3.50; U 1 / HW <1.70.	en	Alarie, Yves, Michat, Mariano C., Hendrich, Lars, Watts, Chris H. S. (2018): Larval Description and Phylogenetic Placement of the Australian Endemic Genus Barretthydrus Lea, 1927 (Coleoptera: Dytiscidae: Hydroporinae: Hydroporini: Sternopriscina). The Coleopterists Bulletin 72 (4): 639-661, DOI: 10.1649/0010-065X-72.4.639, URL: http://dx.doi.org/10.1649/0010-065x-72.4.639
8F69AC09C709FF90D113FDC5CD381311.taxon	description	Instar I (Figs. 1 – 12) Description. Body: Subcylindrical, narrowing towards abdominal apex. Measurements and ratios that characterize the body shape are shown in Table 1. Head: Head capsule (Figs. 1 – 2) pearshaped, tapering posteriorly, not constricted at level of occipital suture; ecdysial suture welldeveloped, coronal suture short; frontoclypeus elongate, bluntly rounded, subtriangular with welldeveloped lateral branches, anteroventral margin with 12 ventral lamellae clypeales (Bertrand 1972); dorsal surface with 2 spine-like egg bursters (ruptor ovi of Bertrand 1972) at about mid-length; ocularium present, stemmata not visible ventrally and subdivided into 2 vertical series; epicranial plates meeting ventrally; tentorial pits visible medioventrally at about mid-length. Antenna (Figs. 3 – 4) elongate, slightly shorter than HW; composed of 4 antennomeres, A 2 and A 3 longest, A 1 shortest; A 3 ’ relatively elongate, shorter than A 4; A 3 lacking ventroapical spinula. Mandible (Fig. 7) prominent, falciform, distal half projecting inwards and upwards, apex sharp; mandibular channel present. Labium (Figs. 5 – 6), prementum subrectangular, about as long as broad, lacking lateral marginal spinulae; LP elongate, distinctly shorter than MP, composed of 2 palpomeres; LP 2 subfusiform, distinctly longer than LP 1. Maxilla (Figs. 13 – 14) with short, thick stipes, incompletely sclerotized ventrally; cardo fused to stipes; galea and lacinia absent; MP elongate, slightly shorter than antenna, composed of 3 palpomeres; MP 1 and MP 2 longest, MP 2 distinctly longer than MP 1. Thorax: Thoracic terga convex, pronotum slightly shorter than meso- and metanota combined, meso- and metanota subequal; protergite subrectangular to subovate, more developed than meso- and metatergites; [We were unable to determine the presence of an anterotransverse carina owing to the bad condition of the only specimen available.]; thoracic sterna membranous; spiracles absent. Legs: Long (Figs. 8 – 9), composed of 6 articles (sensu Lawrence 1991); L 1 shortest, L 3 longest; CO robust, elongate, TR divided into 2 parts by an annulus, FE, TI, and TA slender, subcylindrical, PTwith 2 long, slender, slightly curved claws; posterior claw shorter than anterior claw on L 1 and L 2, posterior claw longer than anterior claw on L 3; femora, tibiae, and tarsi lacking spinulae along ventral margin. Abdomen: Eight-segmented (Figs 10 – 11); segments I – VI sclerotized dorsally, membranous ventrally; segment VII sclerotized both dorsally and ventrally, ventral sclerite independent from dorsal one; tergites I – VII narrow, transverse, rounded laterally, lacking sagittal line; [We were unable to determine the presence of an anterotransverse carina owing to the bad condition of the only specimen available.]; segment VIII (= LAS) longest, completely sclerotized, ring-like, strongly constricted at point of insertion of urogomphus; projecting backwards into a very short, subconical siphon; spiracles absent lateroventrally on segments I – VII. Urogomphus very long, composed of 2 urogomphomeres; U 1 long, much longer than segment VIII; U 2 narrower, setiform, much shorter than U 1. Chaetotaxy: Similar to that of generalized Hydroporinae larva (Alarie and Harper 1990; Alarie et al. 1990; Alarie 1991; Alarie and Michat 2007 a) except for the following features (Figs. 1 – 12): Pores PAd and ANf absent; pore ANh distal; setae MX 4 and TR 2 absent; pore FEa articulated distally, close to seta FE 5; seta TI 7 short, spine-like; seta AB 10 spine-like; setae UR 2 and UR 3 contiguous, seta UR 4 articulated posteriorly; setae UR 5, UR 6, and UR 7 elongate; seta UR 8 inserted subapically. [Pore PAk and seta PA 13 could not be located. We are reluctant to consider them as lacking due to the condition of the only instar I specimen available.] Instar II No specimens were available for study. Instar III (Figs. 13 – 19) Description. As instar I except as follows: Body: Measurements and ratios that characterize the body shape in Tables 1 and 2. Head: Head capsule (Figs. 15 – 16) constricted at level of occipital suture; egg bursters lacking. Antenna elongate, distinctly shorter than HW. LP 2 subequal to slightly shorter than LP 1. MP subequal in length to antenna, MP 1 slightly shorter than MP 2 (Figs. 13 – 14). Thorax: Protergum lacking anterotransverse carina; both meso- and metathoracic terga with an anterotransverse carinae; sagittal line visible; mesopleural region with a spiracular opening on each side. Legs: Position and number of secondary setae in Table 3; natatory setae present on dorsal margin of femora, tibiae, and tarsi (Figs. 17 – 18). Abdomen: Segment VII completely sclerotized both dorsally and ventrally, all tergites with anterotransverse carina (Fig. 19); mesopleural region of segments I – VII with a spiracular opening on each side. Chaetotaxy: Head capsule with numerous secondary setae; lateroventral margin of PA with several secondary spine-like setae (Figs. 15 – 16); anteroventral margin of nasale with half circle of about 60 lamellae clypeales of different lengths, directed downwards; AN, MX, and LA lacking secondary setae; MN with 1 hair-like secondary seta on basoexternal margin; thoracic and abdominal sclerites I – VIII with numerous secondary setae mainly on posterior half; natatory setae present on dorsal margin of femora, tibiae, and tarsi; secondary leg setation in Tables 3 - 4 and Figs. 17 – 18; U with secondary setae (Fig. 19).	en	Alarie, Yves, Michat, Mariano C., Hendrich, Lars, Watts, Chris H. S. (2018): Larval Description and Phylogenetic Placement of the Australian Endemic Genus Barretthydrus Lea, 1927 (Coleoptera: Dytiscidae: Hydroporinae: Hydroporini: Sternopriscina). The Coleopterists Bulletin 72 (4): 639-661, DOI: 10.1649/0010-065X-72.4.639, URL: http://dx.doi.org/10.1649/0010-065x-72.4.639
8F69AC09C709FF90D113FDC5CD381311.taxon	discussion	Remarks. Larvae of Barrethydrus can readily be distinguished from those of other Australian Hydroporini described in detail (i. e., Paroster and Antiporus) by the presence of natatory setae on the dorsal margins of the femur, tibia, and tarsi and metric characters presented in Table 2, and, superficially, from the less well-studied genera by the strong, dark yellow banding on the body.	en	Alarie, Yves, Michat, Mariano C., Hendrich, Lars, Watts, Chris H. S. (2018): Larval Description and Phylogenetic Placement of the Australian Endemic Genus Barretthydrus Lea, 1927 (Coleoptera: Dytiscidae: Hydroporinae: Hydroporini: Sternopriscina). The Coleopterists Bulletin 72 (4): 639-661, DOI: 10.1649/0010-065X-72.4.639, URL: http://dx.doi.org/10.1649/0010-065x-72.4.639
8F69AC09C709FF90D113FDC5CD381311.taxon	distribution	Distribution. Endemic to Australia.	en	Alarie, Yves, Michat, Mariano C., Hendrich, Lars, Watts, Chris H. S. (2018): Larval Description and Phylogenetic Placement of the Australian Endemic Genus Barretthydrus Lea, 1927 (Coleoptera: Dytiscidae: Hydroporinae: Hydroporini: Sternopriscina). The Coleopterists Bulletin 72 (4): 639-661, DOI: 10.1649/0010-065X-72.4.639, URL: http://dx.doi.org/10.1649/0010-065x-72.4.639
8F69AC09C70DFF91D320FDDECB6D1706.taxon	description	(Figs. 1 – 14, 16 – 19)	en	Alarie, Yves, Michat, Mariano C., Hendrich, Lars, Watts, Chris H. S. (2018): Larval Description and Phylogenetic Placement of the Australian Endemic Genus Barretthydrus Lea, 1927 (Coleoptera: Dytiscidae: Hydroporinae: Hydroporini: Sternopriscina). The Coleopterists Bulletin 72 (4): 639-661, DOI: 10.1649/0010-065X-72.4.639, URL: http://dx.doi.org/10.1649/0010-065x-72.4.639
8F69AC09C70DFF91D320FDDECB6D1706.taxon	materials_examined	Source of Material. Larvae were collected in association with adults at the following localities: Australia, New South Wales, 20 km North of Nelligan, 15 - VIII. 1997, coll. C. H. S. Watts; Sardine Creek, 30 km North of Orbost V, 30 - XI- 1998, C. H. S. Watts leg.	en	Alarie, Yves, Michat, Mariano C., Hendrich, Lars, Watts, Chris H. S. (2018): Larval Description and Phylogenetic Placement of the Australian Endemic Genus Barretthydrus Lea, 1927 (Coleoptera: Dytiscidae: Hydroporinae: Hydroporini: Sternopriscina). The Coleopterists Bulletin 72 (4): 639-661, DOI: 10.1649/0010-065X-72.4.639, URL: http://dx.doi.org/10.1649/0010-065x-72.4.639
8F69AC09C70DFF91D320FDDECB6D1706.taxon	discussion	Diagnostic Combination (Instar III). The third instar of B. tibialis can easily be distinguished from that of the closely similar B. geminatus by the presence of a broad, blackish macula on the anterior portion of the frontoclypeus (Fig. 16) in addition to a relatively shorter head capsule compared to the length of abdominal segment VIII (HL / LAS <4.50 compared to> 4.60) and a larger total number of secondary setae on selected leg articles (Table 3).	en	Alarie, Yves, Michat, Mariano C., Hendrich, Lars, Watts, Chris H. S. (2018): Larval Description and Phylogenetic Placement of the Australian Endemic Genus Barretthydrus Lea, 1927 (Coleoptera: Dytiscidae: Hydroporinae: Hydroporini: Sternopriscina). The Coleopterists Bulletin 72 (4): 639-661, DOI: 10.1649/0010-065X-72.4.639, URL: http://dx.doi.org/10.1649/0010-065x-72.4.639
8F69AC09C70DFF91D320FDDECB6D1706.taxon	description	Instar I (Figs. 1 – 12) Description. Color: [Not available owing to the bad state of preservation of the only specimen available.] Body: Measurements and ratios that characterize the body shape in Table 1. Head: Head capsule as in Figs. 1 – 2, HL = 0.60 mm, HL / LAS = 4.61. Abdomen: As in Figs. 10 – 11, U 1 = 0.75 mm; U 1 / HW = 1.55. Instar II No specimen available for study. Instar III (Figs. 13 – 14, 16 – 19) Description. Color: Head capsule predominantly yellow; frontoclypeus with a broad, black, subapical macula (Fig. 16); head appendages dark yellow; thoracic tergites dark brown; abdominal tergites I, IV – VII dark brown, II and III dark yellow to pale brown and VIII pale yellow; urogomphi dark yellow to pale brown; legs dark brown proximally, pale yellow distally. Body: Measurements and ratios that characterize the body shape in Table 1. Head: Head capsule as in Fig. 16, HL = 1.24 – 1.30, HL / LAS <4.50. Abdomen: U 1 = 1.51 – 1.69 mm, U 1 / HW = 1.56 – 1.67 (Fig. 19). Chaetotaxy: ProFE with more than 52 secondary setae; mesoFE with more than 56 secondary setae; mesoTI with more than 47 secondary setae; mesoTA with more than 34 secondary setae; metaCO with more than 26 secondary setae; metaFE with more than 66 secondary setae; metaTI with more than 50 secondary setae; metaTA with more than 43 secondary setae.	en	Alarie, Yves, Michat, Mariano C., Hendrich, Lars, Watts, Chris H. S. (2018): Larval Description and Phylogenetic Placement of the Australian Endemic Genus Barretthydrus Lea, 1927 (Coleoptera: Dytiscidae: Hydroporinae: Hydroporini: Sternopriscina). The Coleopterists Bulletin 72 (4): 639-661, DOI: 10.1649/0010-065X-72.4.639, URL: http://dx.doi.org/10.1649/0010-065x-72.4.639
8F69AC09C70CFF9ED0EFF9D6C84417E3.taxon	materials_examined	Source of Material. Larvae were collected in association with adults at the following locality: Australia, New South Wales, 13 km NNW Dungog, Williams River, Tillegra Bridge, 109 m., 19. X. 2006, 32.19.078 S 151.41.250 E, L. Hendrich leg. (NSW 84).	en	Alarie, Yves, Michat, Mariano C., Hendrich, Lars, Watts, Chris H. S. (2018): Larval Description and Phylogenetic Placement of the Australian Endemic Genus Barretthydrus Lea, 1927 (Coleoptera: Dytiscidae: Hydroporinae: Hydroporini: Sternopriscina). The Coleopterists Bulletin 72 (4): 639-661, DOI: 10.1649/0010-065X-72.4.639, URL: http://dx.doi.org/10.1649/0010-065x-72.4.639
8F69AC09C70CFF9ED0EFF9D6C84417E3.taxon	diagnosis	Diagnostic Combination (Instar III). The third instar of B. geminatus can easily be distinguished from that of the closely similar B. tibialis by the absence of maculae on the dorsal surface of the head capsule (Fig. 15), in addition to a relatively longer head capsule compared to the length of abdominal segment VIII (HL / LAS> 4.60 compared to <4.50) and a lower total number of secondary setae on selected leg articles (Table 3).	en	Alarie, Yves, Michat, Mariano C., Hendrich, Lars, Watts, Chris H. S. (2018): Larval Description and Phylogenetic Placement of the Australian Endemic Genus Barretthydrus Lea, 1927 (Coleoptera: Dytiscidae: Hydroporinae: Hydroporini: Sternopriscina). The Coleopterists Bulletin 72 (4): 639-661, DOI: 10.1649/0010-065X-72.4.639, URL: http://dx.doi.org/10.1649/0010-065x-72.4.639
8F69AC09C70CFF9ED0EFF9D6C84417E3.taxon	description	Instar I No specimen available for study. Instar II No specimen available for study. Instar III (Fig. 15) Description. Color: Head capsule testaceous (Fig. 15); head appendages pale yellow; thoracic tergites dark brown; abdominal tergites I – III brown mesally, broadly yellow laterally, IV – VII dark brown, VIII pale yellow; urogomphi dark yellow; legs predominantly dark yellow except coxae brownish. Body: Measurements and ratios that characterize the body shape in Table 1. Head: Head capsule as in Fig. 15, HL = 1.27 – 1.36, HL / LAS> 4.60. Abdomen: U 1 = 1.29 – 1.45 mm, U 1 / HW = 1.37 – 1.49. Chaetotaxy: ProFE with less than 45 secondary setae; mesoFE with less than 51 secondary setae; mesoTI with less than 45 secondary setae; mesoTA with less than 30 secondary setae; metaCO with less than 18 secondary setae; metaFE with less than 53 secondary setae; metaTI with less than 46 secondary setae; metaTA with less than 38 secondary setae.	en	Alarie, Yves, Michat, Mariano C., Hendrich, Lars, Watts, Chris H. S. (2018): Larval Description and Phylogenetic Placement of the Australian Endemic Genus Barretthydrus Lea, 1927 (Coleoptera: Dytiscidae: Hydroporinae: Hydroporini: Sternopriscina). The Coleopterists Bulletin 72 (4): 639-661, DOI: 10.1649/0010-065X-72.4.639, URL: http://dx.doi.org/10.1649/0010-065x-72.4.639
