taxonID	type	description	language	source
9331B306FFB5FFCF8EA6FEB95B1AFD81.taxon	description	(Figs. 1 - 20)	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFB5FFCF8EA6FEB95B1AFD81.taxon	materials_examined	Type material. HOLOTYPE: J, YEMEN: Ar Rujum, 15 ° 29 ′ N 43 ° 41 ′ E, 16. i. - 9. iv. 2001, Malaise trap, A. van Harten leg. Slide with dissected specimen, Cat. No. 34252, Inv. No. 14041 (NMPC). PARATYPES: 4 JJ 4 ♀♀ (slides, NMPC), the same locality, collector and trap. ALLOTYPE: ♀, 16. x. 2000 - 15. i. 2001, Cat. No. 34253, Inv. No. 14042. OTHER PARATYPES: JJ, same date as holotype, Cat. No. 34254 - 34255, Inv. No. 14043 - 14044; J, 9. iv. - 5. vi. 2001, Cat. No. 34256, Inv. No. 14045; J, 16. x. 2000 - 15. i. 2001, Cat. No. 34257, Inv. No. 14046; ♀♀, same date as previous specimen, Cat. No. 34258 - 34260, Inv. No. 14047 - 14049. Figures based mainly on the holotype, partly on the paratypes.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFB5FFCF8EA6FEB95B1AFD81.taxon	description	Description. Male. Eyes separated, frontoclypeus with rectangular patch of insertions of hairs connected with dorso-ventral stripe irregularly arranged hair pits between upper apices of eyes (Fig. 1). Minimum distance between eyes 0.5 times as wide as diameter of facet (Fig. 10). Ratios of distance of both tangential points of eyes (dorsal and ventral) to minimum width of frons 9: 1, to facet diameter 18: 1. Antenna with 14 segments (Figs. 2, 11); scape almost ovoid, slightly longer than pedicel, the latter hardly globular; flagellomeres 1 - 9 pitchershaped, nodes bulbose, internodes (necks) shorter than nodes; antennomeres 12 - 14 progressively decreasing in size, almost globular, segment 12 quite separated, antennomeres 13 and 14 fused; sensory filament composed of two anterior and one posterior branch. Length ratios of maxillary palpomeres (Fig. 12) 1: 1: 1: 1.5, palpomere 4 not annulate, tapering gently to apex. Terminal lobe of labium (Fig. 3) with four digitiform protuberances and two long setae. Ratio of maximum length of cibarium to length of epipharynx 1.3: 1. Thoracic sclerites as in Fig. 13. Wing (Fig. 14) of characteristic psychodini-shape, clear, 1.3 mm (holotype) and 1.3 - 1.6 mm (paratypes) long, wing membrane bare, radial and medial forks complete; following veins or their parts strengthened: Sc (considerably), R 1, R 2, base of R 4, R 5, basal part of M 1 + 2, M 4 and begining of Cu (conspicuously); basal costal nodes distinct; Sc uninterrupted, bent; M 3, Cu and M 4 not connected basally; R 5 reaching to wing margin with round apex; medial wing angle 96 ° (BCD); wing indices: AB: AC: AD = 3: 3.5: 2.6 and BC: CD: BD = 1: 1.9: 2.2; maximum wing length equal to 2.3 times its maximum width. Maximum length of haltere equal to 3.3 times its maximum width. Ratios of lengths of femora, tibiae and first tarsomeres: P 1 = 2.6: 2.5: 1; P 2 = 2.9: 3.2: 1.1; P 3 = 3.3: 3.9: 1.2; fore claws generally bent, form depending on angle of view (Figs. 4, 15). Basal apodeme of male genitalia narrow, slightly bent proximally (in dorsal as well as in lateral aspect, Figs. 5, 16). Phallobasis bagshaped, with three phallomeres around gonopore, dorsal phallomeres broad, completely fused, distorted in lateral view, ventral phallomere needle-shaped, bent, shorter than dorsal one. Gonocoxites short, thick, protruding laterally in dorsal view; gonostylus thin, 1.5 times as long as gonocoxite (Figs. 5, 6), pointed. Epandrium (Figs. 7, 17) with one large field of insertions of hairs on each side. Basal aperture as well as sclerotized remainders of tergite and sternite 10 inside of epandrium missing. Hypandrium narrow (Fig. 5). Epiproct hardly visible, hypoproct small, with tongue-shaped lobulus in middle, hairy. Surstylus long, distinctly longer than epandrium, bulbose basally (Figs. 7, 17), S-shaped in dorsal view, C-shaped in lateral one, subapically with one rather large retinaculum. Female. Similar to male. Wing 1.2 - 1.4 mm long. Genitalia as figured (Figs. 8, 9, 18, 19, 20), subgenital plate (Fig. 18) bilobed, with shallow apical concavity, hairy; sensory digital organ developed; plate with a pointed sclerotized rib in medial line basally; complicated sclerotized forms of genital chamber without mesh-like structures (Figs. 8, 19, 20). Cercus (Fig. 9) hardly twice as long as subgenital plate, bent.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFB5FFCF8EA6FEB95B1AFD81.taxon	diagnosis	Differential diagnosis. Copropsychoda bulbosa sp. nov. is characterized by having the radial and medial forks of wings complete (Fig. 14), the ventral male phallomere rectangularly bent at the base and without a conspicuous ring, the gonocoxites bulbous basally (Fig. 5), and the subgenital plate of female with convergent sides (Fig. 18). The most closely related spe- Figs. 10 - 20. Copropsychoda bulbosa sp. nov., J (10 - 17) and ♀ (18 - 20). 10 – facets; 11 – apical antennomeres; 12 – maxilla and maxillary palpus; 13 – lateral view of thoracic sclerites; 14 – wing; 15 – lateral view of fore claw; 16 – lateral view of aedeagal complex; 17 – dorsal view of epandrium and surstyli; 18 – subgenital plate; 19 – dorsal view of genital chamber; 20 – same, lateral view. Scales = 0.1 mm (Figs. 10 - 13, 16 - 20); 1 mm (Fig. 14); 0.05 mm (Fig. 15). cies Palaearctic C. brevicornis (Tonnoir, 1940) has incomplete radial and medial forks, ventral phallomere with a fully developed basal ring which is only inconspicuously bent at the base, only slightly strengthened gonocoxites, and an almost parallel-sided female subgenital plate.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFB5FFCF8EA6FEB95B1AFD81.taxon	etymology	Etymology: Bulbosus (Latin, adjective) = bulbous – gonocoxites with conspicuous lateral protuberances. Bionomy. Unknown. The adults were collected from October to June in a Malaise trap at Ar Rujum.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFB5FFCF8EA6FEB95B1AFD81.taxon	distribution	Distribution. Yemen.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFB5FFCF8EA6FEB95B1AFD81.taxon	discussion	Comments on the generic classification and extent of the genus. VAILLANT (1971) described the genus Copropsychoda, separating it from Psychoda Latreille, 1796. He did not establish the type species but included only C. brevicornis (Tonnoir, 1940) in the new genus. It should therefore be recognized as the type species by monotypy. Four species of Copropsychoda are now known: C. brevicornis from Europe and West Siberia (JEŽEK 1992), C. miyatakei (Tokunaga, 1958) from Japan, C. dewulfi (Satchell, 1955) from the Belgian Congo (currently Democratic Republic of the Congo) and C. bulbosa sp. nov. from Yemen.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFB2FFC28EC6FD9E5AE8FE21.taxon	description	(Figs. 21 - 34)	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFB2FFC28EC6FD9E5AE8FE21.taxon	materials_examined	Type material. HOLOTYPE: J, YEMEN: Ar Rujum, 15 ° 29 ′ N 43 ° 41 ′ E, 16. x. 2000 - 15. i. 2001, Malaise trap, A. van Harten leg. Slide, dissected specimen, Cat. No. 34261, Inv. No. 14050 (NMPC). PARATYPES: 2 JJ (slides, NMPC), same date, Cat. No. 34262, Inv. No. 14051; 16.1. - 9.4.2001, Cat. No. 34263, Inv. No. 14052. Figures based mainly on the holotype, partially on paratypes.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFB2FFC28EC6FD9E5AE8FE21.taxon	description	Description. Male. Eyes not touching; frontoclypeus with central patch of insertions of hairs trilobed (Fig. 21), medial lobe thin and long, lateral ones short and rounded; areas near tentorial pits with rectangular border. Minimum distance between eyes 0.5 times as wide as diameter of facets (Fig. 26). Ratios of distance of tangential points of ends of eyes to minimum width of frons 21: 1, to facet diameter 10.5: 1. Antennae with 14 segments (Figs. 27, 28); scape cylindrical, short; pedicel almost globular; flagellomeres amphora-shaped. Apical antennomere with bulbose basis, spined, neck as well with a lateral spine, terminal part obovate with one apical seta. Sensory filaments long, thin, with four branches, paired. Length ratios of maxillary palpomeres 1: 1.1: 1.3: 1.6 (Fig. 29); palpomere 4 not annulate, connected basally with apical end of palpomere 3. Terminal lobe of labium as in Fig. 22, with four digitiform projections. Maximum length of cibarium equal to 1.3 times length of epipharynx. Thoracic sclerites as in Fig. 23, anepisternum with rounded inner apical part, pleurotergum with 2 - 3 hairs. Wing (Fig. 30) narrowly lancet-shaped, 2.0 mm (holotype) and 1.6 - 1.8 mm (paratypes) long, clear, clouded only between C and R 1; following veins or their parts strengthened: Sc, R 1, R 4 and M 1 + 2 basally, R 5, M 4 and Cu; basal costal nodes well visible; Sc uninterrupted; M 3, Cu and M 4 not connected basally; R 5 extending distally to reach wing margin in apex of wing; veins r-r, r-m and m-m not developed; medial wing angle 81 ° (BCD); wing indices: AB: AC: AD = 2.8: 3.5: 2.5; BC: CD: BD = 1: 1.6: 1.7; maximum wing length equal to 2.7 times its maximum width. Maximum length of haltere equal to 2.8 times its maximum width. Ratios of lengths of femora, tibiae and first tarsomeres: P 1 = 2.9: 2.8: 1; P 2 = 3.2: 3.7: 1.3; P 3 = 3.6: 4.4: 1.3; fore claws twice pointed and bent distad (see Fig. 31). Basal apodeme of male genitalia (Figs. 32, 33) narrowed and geniculate (dorsal view), widened from lateral aspect, rounded proximally; distal part of basal apodeme forked in two caudal arms; phallobasis with three phallomeres around gonopore; dorsal phallomeres completely fused, rounded apically from lateral view, pointed from dorsal aspect; ventral phallomere needle-shaped and bent. Paired external additional protuberances short, bilobed; inner lobuli rounded, with tufts of hairs, outer lobuli smaller and pointed. Gonocoxites (Figs. 32, 34) cylindrical with hemispherical lateral protuberances; gonostylus long, with thin pointed and bent tip, inner side covered with long hairs, outer side with one long basal seta. Epandrium (Figs. 24, 25) widely setose (one area on each side), only central rhomboidal aperture developed. Hypandrium narrow. Epiproct and hypoproct inconspicuous, hairy. Surstylus very long, bent in dorsal view, almost straight in lateral view, with one long subapical retinaculum. Female. Unknown.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFB2FFC28EC6FD9E5AE8FE21.taxon	diagnosis	Differential diagnosis. Male of Falsologima verrucosa sp. nov. is characterized by having the paired external additional protuberances near phallomeres short and bilobed, of which the inner lobule is rounded with a tuft of hairs and the outer one is smaller and pointed, large and almost equally long phallomeres, and the minimum distance between eyes equal to 0.5 times the diameter of facet. The species thus differs from F. savaiiensis (Edwards, 1928) which lacks the paired external additional protuberances of the male genitalia, has a very short rudimental ventral phallomere in contrast to fused dorsal phallomeres, and the minimum distance between eyes equal to one facet diameter.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFB2FFC28EC6FD9E5AE8FE21.taxon	etymology	Etymology. Verrucosus (Latin, adjective) = full of tubercles – in reference to the large hairy tubercles near phallomeres. Bionomy. Unknown. The adults were trapped in a Malaise trap at Ar Rujum between October and April.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFB2FFC28EC6FD9E5AE8FE21.taxon	distribution	Distribution. Yemen.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFB2FFC28EC6FD9E5AE8FE21.taxon	discussion	Comments on the generic classification and extent of the genus. Falsologima Ježek & van Harten, 1996 was erected for Psychoda auct. (nec Latreille, 1796) partim with the tropicopolitan Psychoda savaiiensis Edwards, 1928 as the type species. The genus now further includes F. guamensis (Quate, 1965) from Guam, the Philippines, New Guinea, New Britain and New Ireland; F. quadrilosa (Quate & Quate, 1967) from New Guinea; F. quadropsis (Quate et Quate, 1967) from New Guinea; F. serpentina (Quate, 1965) from the Philippines and New Guinea; and F. verrucosa sp. nov. from Yemen.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFBFFFC28D7FFE7E5BE5FA97.taxon	type_taxon	Type species. Psychana rujumensis sp. nov. by present designation.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFBFFFC28D7FFE7E5BE5FA97.taxon	diagnosis	Differential diagnosis. Antennomeres 13 - 14 fused, (apical) antennomere 15 separated, conical, meropleural suture conspicuously or at least slightly developed. This contrasts with the genera Ypsydocha Ježek, 1984 and Tinearia Schellenberg, 1803 in which antennomeres 13 - 15 are fused, antennomere 15 is not conical and the meropleural suture is missing. The mentioned characters seem to be unique for this genus, especially in the shape of the apical flagellomere.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFBFFFC28D7FFE7E5BE5FA97.taxon	etymology	Etymology. A mutation of Psychoda (gender feminine).	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFBFFFC28D7FFE7E5BE5FA97.taxon	discussion	Comments on the generic classification and extent of the genus. JEŽEK (1983, 1984) discussed in detail the position of six new genera excluded from the genus Psychoda auct. in the light of relationships among other genera. The complex taxonomical situation of the tribe Psychodini, comprising chiefly the clearly polyphyletic Psychoda s. lat. (nec Latreille, 1796) will not be fully resolved by the description of the new genus Psychana and may, in fact, become even more complicated. Sound comparative morphological studies, based not only on the last flagellomeres, are necessary for an acceptable division of the tribe. Worldwide it contains 447 species, 26 of which most probably belong to quite different tribes. Unfortunately, the clarification of their taxonomic status is hampered by the unavailability or poor condition of the type material. At any rate, future revisions of particular species groups are needed. The following species belong to the new genus: Psychana absidata (Quate & Quate, 1967) comb. nov. (from Psychoda Latreille, 1794) from New Guinea, P. moleva (Quate, 1965) comb. nov. (from Psychoda) from the Philippines, P. pellucida (Quate, 1965) comb. nov. (from Psychoda) from Malaysia, P. pinguicula (Quate & Quate, 1967) comb. nov. (from Psychoda) from New Guinea, and P. rujumensis sp. nov. from Yemen.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFBFFFC68EB0FA8858AAFE61.taxon	description	(Figs. 35 - 53)	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFBFFFC68EB0FA8858AAFE61.taxon	materials_examined	Type material. HOLOTYPE: J, YEMEN: Ar Rujum, 15 ° 29 ′ N 43 ° 41 ′ E, 16. x. 2000 - 15. i. 2001, Malaise trap, A. van Harten leg. Slide, dissected specimen, Cat. No. 34264, Inv. No. 14053 (NMPC). PARATYPES: 3 JJ 6 ♀♀ (slides, NMPC), the same locality, collector and trap. ALLO- TYPE: ♀, 16. x. 2000 - 15. i. 2001, Cat. No. 34265, Inv. No. 14054. OTHER PARATYPES: JJ:, same date, Cat. No. 34266 - 34267, Inv. No. 14055 - 14056; 16. i. - 9. iv. 2001, Cat. No. 34268, Inv. No. 14057; ♀♀, 16. x. 2000 - 15. i. 2001, Cat. No. 34269 - 34271, Inv. No. 14058 - 14060; 16. i. - 9. iv. 2001, Cat. No. 34272, Inv. No. 14061; 9. iv. - 5. vi. 2001, Cat. No. 34273, Inv. No. 14062. Figures based mainly on the holotype, partially on paratypes.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFBFFFC68EB0FA8858AAFE61.taxon	description	Description. Male. Eyes separated, C-shaped (Fig 35). Minimum distance between upper part of eyes hardly as wide as diameter of facet (Fig. 36). Eye bridge formed by four rows of facets. Frontoclypeus with large central three-lobed patch of insertions of hairs; medial stripe longest, not connected with vertex pits. Antennae 15 - segmented (Figs. 37, 38); scape very short, wide, hardly globular; flagellomeres pitcher-shaped, symmetrical; last three flagellomeres gradually reduced; antennomere 13 pyriform, antennomere 14 almost globular with very short neck, both segments fused; antennomere 15 minute, conical, entirely separated. Sensory filaments long, paired, with three branches. Length ratio of maxillary palpomeres 1: 1.1: 1: 1.6 (Fig. 45), palpomere 4 not annulate. Terminal lobe of labium with six digital protuberances (Fig. 44). Maximum length of cibarium equal to 1.3 times length of epipharynx. Thoracic sclerites as in Fig. 46, meropleural sclerite inconspicuously divided. Wing (Fig. 39) widely lancet-shaped, 1.7 mm (holotype) and 1.6 - 1.9 mm (paratypes) long, almost clear, clouded between fore margin and R 1 in small narrow area; following veins or their parts strengthened: Sc (conspicuously), R 1, R 5, M 1 + 2, M 4 and Cu; basal costal nodes well visible, Sc uninterrupted; M 3 and Cu hardly with connection to M 4; R 5 extends in apex of wing. Medial wing angle 79 ° (BCD); wing indices: AB: AC: AD = 1.4: 2.1: 1.6; BC: CD: BD = 1: 1.1: 1.4; maximum length of wing equal to 2.2 times its maximum width. Maximum length of haltere equal to 2.8 times its maximum width. Ratios of lengths of femora, tibiae and first tarsomeres: P 1 = 2.8: 2.6: 1; P 2 = 2.9: 3.2: 1.2; P 3 = 3.3: 3.8: 1.3. Fore claws as in Fig. 40. Basal apodeme (Figs. 41, 47) of male genitalia bent, narrow from dorsal view, widened from lateral one, pointed proximally; distal part of basal apodeme forked in two caudal arms. Copulatory organ with smooth surface outside, phallomeres of characteristic shape, sclerotized in some parts of lobes. Gonocoxite (Figs. 42, 47) long, cylindrical in lateral view and protruding laterally in dorsal view. Gonostylus somewhat shorter than gonocoxite, with acuminate apex, slightly arched. Epandrium (Figs. 43, 48) with two large patches of hair areas connected caudally, inside with two conspicuous spine-shaped protuberances, sclerotized remainders of tergite and sternite 10 missing. Basal aperture developed. Hypandrium (Figs. 41, 47) narrow, bare, slightly widened in middle. Epiproct and hypoproct small (Figs. 43, 48), hairy. Surstylus long, C-shaped, twice as long as epandrium, subapically with one retinaculum. Female. Wing 1.6 - 2.0 mm long. Subgenital plate (Fig. 49) of characteristic shape, hairy, with large bilobed sclerotized basis; two small caudal lobuli separated by medial cleft. Sensory digit developed. Mesh-like structures of complicated sclerotized forms in the area of genital chamber not developed (Figs. 50, 51, 52). Cercus (Fig. 53) slightly bent, more then three times as long as its basal diameter.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFBFFFC68EB0FA8858AAFE61.taxon	diagnosis	Differential diagnosis. The new species is similar to Psychana umbratica Quate, 1965, in the shape of the terminal flagellomere, the form of male gonopode as well as surstylus and the shape of female subgenital plate, but is quite different in the structures of male genitalia. Psychana rujumensis sp. nov. is diagnosed by the conspicuously developed basal apodeme and ventral phallomere, short and partially fused dorsal phallomeres, and conspicuously sclerotized sides of the base of female subgenital plate (present as largely widened patches). Psychana umbratica has a rudimentary basal apodeme of the male genitalia and ventral phallomere, very long and entirely fused dorsal paired phallomeres, and a narrowly sclerotized base of subgenital plate.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFBFFFC68EB0FA8858AAFE61.taxon	etymology	Etymology. Named after the type locality.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFBFFFC68EB0FA8858AAFE61.taxon	distribution	Distribution. Yemen. Bionomy. Unknown. The adults were trapped in a Malaise trap at Ar Rujum from October to June.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFBBFFC58EA7FDBE5B5BFEC1.taxon	description	(Figs. 54 - 69)	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFBBFFC58EA7FDBE5B5BFEC1.taxon	materials_examined	Type material. HOLOTYPE: J, YEMEN: Ar Rujum, 15 ° 29 ′ N 43 ° 41 ′ E, 9. iv. - 5. vi. 2001, Malaise trap, A. van Harten leg. Slide, dissected specimen, Cat. No. 34274, Inv. No. 14063 (NMPC). PARATYPES: 2 JJ (slides, NMPC), 16. x. 2000 - 15.1.2001, the same data, Cat. No. 34275 - 34276, Inv. No. 14064 - 14065. Figures based mainly on the holotype, partially on paratypes.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFBBFFC58EA7FDBE5B5BFEC1.taxon	description	Description. Male. Head (Fig. 54) about 1.2 times as broad as long. Vertex reduced, conspicuously low, sunken and hairy. Upper edges of eyes separated by distance of 1.2 facet diameter (Fig. 61). Frontoclypeus with patch of scars extending to upper edges of eyes on each side and in midline; medial extension stretching between eye-bridges to level of second or third row of facets. Interocular suture absent. Eye-bridges composed of four rows of facets. Ratios of distance of apices of ends of eyes to minimum width of frons 7.6: 1, to facet diameter 8.8: 1. Antennae with 15 segments (Figs. 55, 62); scape very short, somewhat widened distally, 1.3 times as long as broad; pedicel almost globular, cut distally in peripheral margin; flagellomeres amphora-shaped, distal ones with rather long necks; last three flagellomeres separated. Antennomeres 14 and 15 of almost equal size, smaller in contrast to antennomere 13, antennomeres 13 and 14 with very short necks; apical antennomere cut distally. Sensory filaments (ascoids) larger than flagellomeres (Fig. 62), paired, with three branches. Length ratios of maxillary palpomeres 1.3: 1.1: 1: 1.5, palpomere 4 not annulate (Fig. 58). Terminal lobe of labium with five digitiform projections and two long setae (Fig. 57). Maximum length of cibarium equal to 1.2 times length of epipharynx. Labrum as in Fig. 56, thoracic sclerites as in Fig. 63. Wing 1.3 mm (holotype) to 1.4 mm (paratypes) long, pointed, narrow, clear, anterior edge hardly less convex than posterior one, the latter not enlarged in cubital region (Fig. 64); following veins or their parts strengthened: Sc, R 1, base of R 4, R 5, basal part of M 1 + 2, M 4 and Cu; reduced R 2 recurved and jointed to R 1; inconspicuous and reduced R 2 + 3 connected basally to base of R 4; radial fork not developed in contrast to medial one. Basal costal nodes well visible, Sc long and bent; M 4 and Cu connected basally, M 3 free; R 5 in apex of wing; wing length equal to 2.5 times its maximum width. Maximum length of haltere equal to 3 times its maximum width. Ratios of lengths of femora, tibiae and first tarsomeres: P 1 = 2.2: 2.1: 1; P 2 = 2.3: 2.9: 1.2; P 3 = 2.5: 3.2: 1.2. Fore claws tapering and bent distally (Figs. 65, 66). Basal apodeme of male genitalia (Figs. 59, 67) compressed laterally, narrow and almost straight in dorsal view, conspicuously widened and proximally rounded in lateral view; distal part of basal apodeme forked into two caudal parts connected with aedeagal complex. Phallobasis of irregular shape with three phallomeres around gonopore; dorsal phallomeres long, thick, fused, of grooved form (lateral view), rounded on top; ventral phallomere slightly shorter, hooked and pointed, parallel to dorsal phallomeres; circular loop in basis of aedeagal complex of characteristic shape. Gonocoxite (Figs. 67, 68) very short, thick, protruded laterally. Gonostylus elongate, somewhat bent, enlarged in basal parts, longer than gonocoxite from dorsal view, setose. Epandrium (Figs. 60, 69) broad and short, with single large central opening, two large triangular patches of scars connected distally in medial line. Sclerotized remainders of tergite and sternite 10 inside of epandrium reduced to two almost triangular areas. Hypandrium narrow (Figs. 59, 67), bare. Epiproct and hypoproct in a form of hairy fold, inconspicuous (Fig. 69). Epiproct bilobed, hypoproct with one lobulus. Surstylus approximately twice as long as epandrium, slightly twisted in lateral view and inconspicuously C-shaped in dorsal view, with many scars, one long retinaculum subapically (Figs. 60, 69). Female. Unknown.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFBBFFC58EA7FDBE5B5BFEC1.taxon	diagnosis	Differential diagnosis. Psychoda obscuripennis sp. nov. differs from P. uniformata Haseman, 1907 in having the veins peculiarly ordered and the radial fork not developed (Fig. 64), R 2 and R 2 + 3 reduced, the rest of R 2 connected with R 1, and R 2 + 3 with R 4 basally; the phallobasis is bare, with the ventral phallomere conspicuously shorter than dorsal ones (Figs. 59, 67); sclerotized remainders of tergite and sternite 10 are reduced but present; and the surstyli are twice as long as the epandrium (Figs. 60, 69). In males of P. uniformata the radial fork is developed, R 2 and R 2 + 3 are fully present, R 2 lacks a connection with R 1, R 2 + 3 is not jointed with R 4; the phallobasis is provided with two tufts of long hairs, the ventral phallomere is almost as long as the dorsal one; the remnants of tergite and sternite 10 are missing; and the surstyli are as long as the epandrium.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFBBFFC58EA7FDBE5B5BFEC1.taxon	etymology	Etymology. Obscurus (Latin, adjective) = unclear, obscure; penna (Latin, substantive) = wing – in reference to the peculiar order of wing veins. Bionomy. Unknown. The adults were trapped in a Malaise trap at Ar Rujum from October to June.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFBBFFC58EA7FDBE5B5BFEC1.taxon	distribution	Distribution. Yemen.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFBBFFC58EA7FDBE5B5BFEC1.taxon	discussion	Comments on the generic classification. The diagnosis, synonymy, biology and distribution of the genus Psychoda Latreille, 1796 in a strict sense (type species: Tipula phalaenoides Linnaeus, 1758) were given by JEŽEK (1984, 1990). The taxonomical position of the genus and its relationship to other genera were discussed by JEŽEK (1983).	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFB8FFDB8E3DFEDE5965F994.taxon	description	(Figs. 70 - 84)	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFB8FFDB8E3DFEDE5965F994.taxon	description	(1967): 166; DUCKHOUSE (1973): 240; IPE et al. (1986): 130; DUCKHOUSE & LEWIS (1989): 176; WAGNER (1990): 47. Psychoda bifurcata Tokunaga, 1958: 378 (female only). Psychoda hamatifera Tokunaga, 1958: 385 (male only). Chodopsycha harrisi: JEŽEK (1984): 138; SASAKAWA (1986): 3; ANONYMUS (1990): 13 - 14.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFB8FFDB8E3DFEDE5965F994.taxon	materials_examined	Material examined. YEMEN: Ar Rujum, 15 ° 29 ′ N 43 ° 41 ′ E, Malaise trap, A. van Harten leg., 8 ♀♀, dissected, slides (NMPC): 16. i. - 9. iv. 2001, Cat. No. 34277 - 34282, Inv. No. 14066 - 14071; 9. iv. - 5.6.2001, Cat. No. 34283, Inv. No. 14072; 16. x. 2000 - 15. i. 2001, Cat. No. 34284, Inv. No. 14073. Figures based on Inv. No. 14066 - 14070.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFB8FFDB8E3DFEDE5965F994.taxon	description	Redescription. Female. Upper edges of eyes separated (Fig. 70), ocular bridge composed of four rows of facets (Fig. 71). Lower part of frontoclypeus with large, almost rectangular patch of inserted hairs, narrow medial stripe of pits ending between upper edges of eyes, and short stripe on both sides. Minimum distance between upper edges of eyes slightly larger than facet diameter. Ratios of distance of tangential points of ends of eyes to minimum width of frons 6.3: 1, to facet diameter 7.3: 1. Antennae with 16 segments (Figs. 72, 77). Scape cylindrical, twice as long as pedicel, the latter is bowl-shaped. Flagellomeres 1 - 10 amphora-shaped, last four flagellomeres fused, almost globular and almost gradually reduced, last three flagellomeres minute (apical one smallest) and in comparison to preceding segments with long neck. Antennomeres 13 and 15 spined. Sensory filaments (Fig. 77) conspicuous and well visible, with three branches. Length ratio of maxillary palpomeres 1.1: 1: 1: 1.4 (Fig. 73). All palpomeres with transversal rows of small bristles. Terminal lobe of labium with four digitiform projections and three long setae (Fig. 78). Maximum length of cibarium equal to 1.2 times length of epipharynx. Thoracic sclerites as in Fig. 79. Wing (Fig. 74) widely lancet-shaped, 2.0 - 2.4 mm long, inconspicuously clouded, conspicuously and narrowly so only between costal margin and R 1 and in short line between R 1 and R 2 + 3. Vein R 5 bordered by narrow longitudinal cloud; following veins or their parts strengthened: R 1, Sc, base of R 4, R 5, base of M 1 + 2, M 4 and Cu; basal costal nodes distinct, Sc uninterrupted, twice constricted and bent; M 3 and Cu without connection to M 4; R 5 in apex of wing; veins r-r, r-m and m-m not visible; medial wing angle (BCD) 100 °; wing indices: AB: AC: AD = 2.9: 3.4: 2.7; BC: CD: BD = 1: 1.6: 2; maximum length of wing equal to 2.4 times its maximum width. Maximum length of haltere equal to 3.5 times its maximum width. Ratios of lengths of femora, tibiae and first tarsomeres: P 1 = 2.7: 2.9: 1; P 2 = 3.3: 3.9: 1.2; P 3 = 3.5: 4.3: 1.2. Fore claws S-shaped (Figs. 75, 80). Subgenital plate of characteristic shape (Fig. 81) with three lobes: middle lobe long, Y-shaped, external lobes shorter, pointed, with small inner lobuli, hairy; sensory digit of middle lobe developed, with two setae and characteristic three-armed sclerites below, connected with genital chamber (Figs. 82, 83, 84); plate with numerous long setae distally. Cercus long, slightly bent (Fig. 76). Male. Unknown to us; shortly described and figured (male genitalia) by SATCHELL (1950). Differential diagnosis. The female of this species is similar to Psychodula minuta (Banks, 1894) by its size and the fusion of the last flagellomeres; however, its subgenital plate (Fig. 81) uniquely shaped, composed of three characteristic lobes (Figs. 82, 83, 84), and bearing numerous long setae. In P. minuta, the subgenital plate is compact, shortly tongue-shaped and without deep clefts or long setae, but with two small, apical, horizontally oriented lobuli. Bionomy. DUCKHOUSE (1966) stated that the species breeds in cow dung, chicken dung or wet garden rubbish.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFB8FFDB8E3DFEDE5965F994.taxon	distribution	Distribution. New Zealand, Australia, Papua New Guinea, Pacific Islands, Hawaii, Japan (Ryukyu Islands), the Philippines, Indonesia, Borneo, Malaysia and India. New to the fauna of the Afrotropical region; DUCKHOUSE & LEWIS (1980) did not list this species.	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
9331B306FFB8FFDB8E3DFEDE5965F994.taxon	discussion	Comments on the generic classification. The diagnosis, synonymy, biology and distribution of the genus Psychodula Ježek, 1984 (type species: Psychoda minuta Banks, 1894), separat- ed from the genus Psychoda Latreille, 1796, were discussed by JEŽEK (1984, 1990). The taxonomical position of the genus and its relationships to other genera were discussed by JEŽEK (1983).	en	Ježek, Jan, Harten, Antonius van (2005): Further new taxa and little-known species of non-biting moth flies (Diptera, Psychodidae) from Yemen. Acta Entomologica Musei Nationalis Pragae 45: 199-220, DOI: 10.5281/zenodo.5176593
