identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
95DCD953A7CA50F6BD3741C2BF00246E.text	95DCD953A7CA50F6BD3741C2BF00246E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ceratophysella granulata Stach 1949	<div><p>Ceratophysella granulata Stach, 1949</p><p>Ceratophysella granulata Stach 1949: 133</p><p>Material.</p><p>Poland ( Carpathians): ISEZ: 2 syntypes on slide (formerly in alcohol), Tatra Mts, Dziura cave, 15.VII.1909, leg. J. Stach; 29 spp. on slides, Tatra Mts, leg. J. Stach; Beskid Maly Mts , male, juv., Zagórze near Skawce, Grota Piaskowa cave, 350 m. a.s.l., XI.1951, leg. Szymczakowski; DIBEC: Tatra Mts (leg. D. Skarżyński): 74 females, 17 males, litter of dwarf mountain pine shrubs on the slopes of the Gładkie Upłaziańskie, at an altitude of 1500-1600 m a.s.l., 13.VII.2001, 14.IX.2002, 18.IX.2004, 14. VIII. 2009; 4 females, 2 males, 2 juv., Chuda Turnia, moss on rocks, 1800 m a.s.l., 19.VIII.2004; 11 females, 5 males, 2 juv., Kraków Gorge, spruce forest litter and mosses on rocks, 1050-1150 m a.s.l., 19.VIII.2004; 4 males, Mylna cave, mosses in the entrance, 1090 m a.s.l., 19.VIII.2004; female, Raptawicka cave, mosses in the entrance, 1150 m a.s.l., 19.VIII.2004; female, juv., Dziura cave, litter in the entrance, 1000 m a.s.l., 24.VIII.1991; 20 females, 15 males, litter of dwarf mountain pine shrubs on the slopes of Sucha Czuba, 1600-1700 m a.s.l., 17.IX.2004; Beskid Zywiecki Mts : 3 females, 5 males, 6 juv., Babia Góra, litter of spruce forest and dwarf mountain pine shrubs, 1300-1500 m a.s.l., 4.VI.1999, leg. A. Smolis; 4 females, 3 males, 5 juv., Pilsko, litter of spruce forest and dwarf mountain pine shrubs, 1300-1500 m a.s.l., 21.IX.2004, leg. D. Skarżyński; Pieniny Mts, 4 females, male, 2 juv., Ociemny valley, 500-600 m a.s.l., mosses on rocks, 26.V.1994, leg. R. J. Pomorski; Beskid Wyspowy Mts, 10 females, male, Zbójecka cave near Limanowa, 900 m a.s.l., bat guano, 12.VII.2007, leg. K. Piksa. Slovakia ( Western Carpathians): ISEZ: female, male, Pieniny Mts, Aksamitka cave, VII.1931, leg. Grochmalicki. PJSU: 4 females, 2 males, Veľká Fatra Mts, HornaTufna cave near Horny Harmanec village , 975 m a.s.l., entrance hall, cave sediment, 26-31.VIII.1999, leg. Ľ . Kováč, 301-99, 302-99; female, Zapadne Tatry Mts , Brestovska cave near Zuberec village, entrance hall, cave sediment, 22.V.-13.IX.2006, leg. A. Mock, 671-06; 2 males, 3 juv., Belianske Tatry Mts, Kamzíčia jaskyňa cave near Zdiar village , 2002 m a.s.l., 15 m from entrance, cave sediment, 13.IX.1991, leg. Ľ . Kováč; Low Tatras Mts: Demänovská jaskyňa slobody cave near Demaenova village , 812 m a.s.l.: female, male, rotten wood, 11.V.2000, leg. P. Ľuptáčik, 84-00; 2 females, male, Mramorové riečisko, bait, 11.V.-27.IX.2000, leg. Ľ . Kováč, 131-00, 133-00; Demänovská ľadová jaskyňa cave, 740 m a.s.l.: 2 females, entrance, talus deposit, 12.V.-28.IX.2000, Ľ . Kováč, 168-00; female, cave entrance, wood, 28.IX.2000, leg. P. Ľuptáčik, 170-00; female, Pustá jaskyňa cave, Hlinená chodba, surface of water puddle, 24.VI.2015, leg. Ľ . Kováč, 82-15; Pieniny Mts, Aksamitka cave near Haligovce village, 756 m a.s.l., leg. Ľ . Kováč: 6 females, 4 males, Blatistý dóm, rotten wood, cave sediment, 12.III.-26.V.1998, 45-98, 197-98, 198-98, 199-98, 200-98; 2 females, Dóm priekopníkov, bat guano, 26.V.1998, 201-98; 2 females, 1 juv., Dóm priekopníkov, cave sediment, 23.VIII.-7.X.1999, 363-98, 364-98, 366-98; male, soil of herbal cushion in front of cave, 26.V.1998, 202-98; Levočské vrchy Mts, Jaskyňa pod Jankovcom 2 cave near Ľubica village: 3 females, male, Hall A, bat guano, Hall II rotten wood, 5.XI.2010, leg. P. Ľuptáčik, 747-10, 758-10; male, passage B, surface of water puddle, 5.XI.2010, leg. Z. Višňovská, 755-10; Slovak Paradise: Dobšinská ľadová jaskyňa cave near Stratena village , 969 m a.s.l.: female, male, moss on rocks in front of cave, 23.VII.1997, leg. Ľ . Kováč, 126-130-97, 2 females, humus and soil in front of cave, 7.X.2004. leg. V. Šustr, 4 females, male, Psie Diery cave, cave sediment, 6.II.1997, leg. V. Košel, female, Vojenska cave , 20 m from entrance, bait, 6.XII.1998, leg. V. Košel, 22-99, 3 females, cave sediment, 28.I.-6.II.1997, leg. V. Košel, 3 females, Klastorna cave , cave sediment, 27.I.-4.II.1997, leg. V. Košel, female, 2 males, Duca cave , Dóm, bait, 4.XII.1998, leg. V. Košel, 25-99, male, Stratenska cave , 200 m from entrance, rotten wood, 9.X.1997, leg. Ľ . Kováč, 184-97; male, Muránska planina Plateau, Bobacka cave near Muranska Huta village , 30 m from entrance, cave sediment, 5.X.-9.XI.2000, Ľ . Kováč, 219-00; Čierna hora Mts: 2 females, Veľká ružínska jaskyňa cave near Mala Lodina village , 614 m a.s.l., 100 m from entrance, cave sediment, 10.VIII.-14.X.1996, leg. Ľ . Kováč, 1327-96, male, juv., Malý Ružinok Valley, Tilio-Acerion, humus and soil, rotten wood, 19.IX.2009, 23.IV.2010, leg. Ľ . Kováč, 610-09, 139-10; male, Slovak Karst, Šingliarova priepasť cave near Honce village, 680 m a.s.l, 1st Hall, rotten wood, 4.V.2008, leg. P. Ľuptáčik, 215-08 .</p><p>Remarks.</p><p>Specificaton of C. granulata morphology is provided by Skarżyński (2004a). COI sequences of this species were examined and deposited on BOLD and GenBank by Porco et al. (2012). Ceratophysella granulata, with strong tegumentary granulation and distinct fields of coarse granules, resembles C. stachi sp. nov. and four other European species of the C. armata -group: C. lawrencei, C. neomeridionalis, C. scotica, and C. silvatica . Differences between these species are presented in Table 4. The presence of true C. granulata is so far confirmed only for the Polish Carpathians (Tatra Mts, Pieniny Mts, Beskid Żywiecki Mts, Beskid Wyspowy Mts, Beskid Mały Mts) and Slovak ( Veľká Fatra Mts, Západné Tatry Mts, Belianske Tatry Mts, Low Tatras Mts, Pieniny Mts, Levočské vrchy Mts, Slovak Paradise, Muránska planina Plateau, Čierna hora Mts, Slovak Karst) (Fig. 1), where it inhabits cold and humid places: mosses in the alpine zone, litter and mosses in the dwarf mountain pine zone, deep gorges (litter and mosses), and caves (mosses, litter, and rotten wood at cave the entrance, and bat guano and cave sediments even 100 m from the entrance) in mountain forests zone. At the end of the Pleistocene, this psychro- and hygrophilous species was probably more common in the periglacial region, and due to the warming Holocene climate, its range became limited to scattered, high-mountain refuges and cold caves and other subterranean habitats at lower elevations. Based on the current distribution data, it is concluded that C. granulata is endemic to the Western Carpathians. However, to verify this thesis, additional research should be undertaken covering the rest of the Carpathians, the Alps, and other mountainous areas of central Europe.</p></div>	https://treatment.plazi.org/id/95DCD953A7CA50F6BD3741C2BF00246E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Skarzynski, Dariusz;Smolis, Adrian;Kovac, Ľubomir;Porco, David	Skarzynski, Dariusz, Smolis, Adrian, Kovac, Ľubomir, Porco, David (2021): A new European species of Ceratophysella (Collembola, Hypogastruridae) revealed by morphological data and DNA barcodes. ZooKeys 1021: 1-18, DOI: http://dx.doi.org/10.3897/zookeys.1021.63147, URL: http://dx.doi.org/10.3897/zookeys.1021.63147
3048909794415CAFAF7712BC8C024C10.text	3048909794415CAFAF7712BC8C024C10.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ceratophysella stachi Skarzynski, Smolis & Porco 2021	<div><p>Ceratophysella stachi Skarzynski, Smolis &amp; Porco sp. nov. Figures 3-4, 5-9</p><p>Ceratophysella granulata: Fjellberg, 1998: 41.</p><p>Type material.</p><p>Holotype (DIBEC): female, Poland, Carpathians, Beskid Niski Mts, litter of the Carpathian beech forest on the slopes of Ostra Góra near Tylawa village, 500 m a.s.l., 20.X.2009, leg. M. Furgoł . Paratypes (DIBEC): 2 males, same data as holotype ; male, juv., 14.V.2001, leg. A. Smolis, D. Skarżyński, other data same as holotype ; 3 females, 5 males, juv., 14.V.2002, leg. A. Smolis, D. Skarżyński, other data same as holotype .</p><p>Other material.</p><p>Denmark (AF, leg. A. Fjellberg): female, Jutland, Himmerland, Rold Skov, Fagus liter, 20.III.1994, 94.024; Funen: 6 females, 2 males, Fiskerup Skov, forest stream, 24.III.1994, 94.076; female, Syltemade Adal, Fraxinus / Ulmus / Viburnum litter, 23.III.1994, 94.070 . Germany (SMNG): Brandenburg: 6 females, 3 males, 3 juv., “Wanninchen” nature reserve, bog, 1. V.1972, leg. Hiebsch , 11911; 9 females, male, “Wanninchen” nature reserve, wet heather, 1972, leg. Hiebsch, 11911; male, 2 juv., " Bergener Moor " nature reserve, moor, Sphagnum, heather, pine forest, 1. V.1972, leg. Hiebsch , 11912. Hesse: 5 females, Vogelsberg, 1985-1995, leg. W. Böhle . Mecklenburg-West Pomerania : female, Müritz National Park, Neustrelitz, soil 0-5 cm, alder swamp woods not on acid peat, 16.X.2013, leg. U. Burkhardt , 264-F865; 4 females, 2 males, Serrahn, Hauptmannsberg near Feldberg, sandy-gravelly moraine, largely unforested since the Middle Ages, 1973, leg. Hiebsch/ ILN Greifswald, 11915; female, Serrahn, Klockenbruch, active, relatively undamaged raised bogs, moss/ Sphagnum, 16.X.2013, leg. R. Lehmitz and U. Burkardt, 261-01; male, Serrahn, Klockenbruch, active, relatively undamaged raised bogs, moss/ Sphagnum, 5.XI.2014, leg. R. Lehmitz and U. Burkardt, 401-12; female, Serrahn, “Kesselmoor” nature reserve, active, relatively undamaged raised bogs, moss/ Sphagnum, 16.X.2013, leg. R. Lehmitz, 49077-08; male, Serrahn, soil 0-5 cm, Medio-European collinear woodrush beech forest, 16.X.2013, leg. U. Burkhardt , 263-03. Saxony: female and juv., Erzgebirge, Kleiner Kranichsee, Sphagnum, 22.VII. 1971, leg. W. Dunger , 7871. Luxembourg, subadult male, Vallée d’Our: Tintesmühle, near the river, 21.VII.1991, leg. M. Ursone , L-91-101 (ISEZ). Norway (AF, leg. A. Fjellberg): Akershus: 4 females, “Östmarka” nature reserve, Tappenbergvann, old spruce cones, 18.V.1995, 95.163; female, Barum, Dalivannet, 16.XI.1997. Vestfold: male, 32/93; 2 females, Larvik, N. Holtesetra, Hvarnes, lush Alnus and Fraxinus forest, 24.XI.2007, 7.290; female and male, Larvik, Granasnekollen, Hvarnes, litter, oak/beech, 25.IX.2004, 04.086; 3 females, Brunlanes, Hummerbakken, Telemark Camp., plant debris, beach, 14.XI.1993, 93.077; female, Tjøme, Sandø, N-stranda, spongy Pinus litter behind beach, 22.IV.2009, 9.086 . Poland (Carpathians, DIBEC): female, 3 males, 2 juv., Beskid Sądecki Mts: "Las Lipowy Obrożyska” nature reserve near Muszyna, 600 m a.s.l., mosses on rocks and trees, 1.V.2004, 25.VI.2005, leg. A. Smolis, D. Skarżyński; female, Roztoka Ryterska, 600 m a.s.l., litter and mosses near stream, 3. V.2004, leg. A. Smolis, D. Skarżyński; 5 females, “Uhryń” nature reserve, 850 m a.s.l., litter of fir-beech forest, 3. V.2000, leg. A. Smolis ; female, 3 males, 2 juv., “Barnowiec” nature reserve, 850 m a.s.l., mosses on rocks in an old beech forest, 10. V.2003, leg. A. Smolis, D. Skarżyński; 4 females, 2 males, SE slopes of Jaworzyna Krynicka, litter in a beech forest, ca 800 m a.s.l, 2. V.2004, leg. A. Smolis, D. Skarżyński . Bieszczady Mts : 2 females, N slopes of Krzemieniec, 1000 m a.s.l., litter in a stream valley, 19. V.2000, leg. A. Smolis. Ukraine: male, Ivano-Frankove village, Lviv District, beech and elm forest, leaf litter and soil, leg. S. Bakaeva , 2.2.4.8 (SMNHL).</p><p>Etymology.</p><p>Dedicated to Jan Stach, the excellent specialist in Collembola .</p><p>Description.</p><p>Body length 1-2 mm. Colour (in alcohol) bluish-gray to bluish-black. Tegumental granulation strong, with fields of especially coarse granules on head (large uniform field covering whole dorsal side except antennal bases), thoracic terga II-III (two large subaxial fields and two lateral ones of medium size, Fig. 3), abdominal terga I-III (variable distribution: from four - Fig. 4 to seven fields of medium size as in C. granulata; see Skarżyński 2004a: fig. 8), abdominal tergum IV (medium axial field and two lateral large ones, Fig. 4) and abdominal terga V-VI (large uniform fields covering almost whole dorsal side, Fig. 4). 6-9 granules between macrosetae p1 on abdominal tergum V.</p><p>Arrangement of setae on head typical for the genus, spine-like setae absent. Dorsal chaetotaxy of type B (Figs 3, 4). Thoracic terga II-III with setae m6 present and one additional seta outside lateral sensillum m7 present or absent. Setae p1 on abdominal tergum IV developed as macrosetae, p2 as microsetae, setae p3 present. Differentiation of dorsal setae into micro- and macrosetae distinct. Setae long (ratio p1 microsetae and p2 macrosetae on thoracic tergum II/inner edge of claws III = 1-1.3 and 1.8-2.8, respectively), thick, curved, pointed at tips and only slightly serrate. Body sensilla (s) short (ratio sensillum p4 and m7 on thoracic tergum II/inner edge of claws III = 0.6-1 and 0.4-0.6, respectively), thin and smooth. Microsensilla (ms) on thoracic tergum II present (Fig. 3). Subcoxae I, II, III with 1, 2, 3 setae, respectively.</p><p>Antennal segment IV with simple or lobed apical vesicle, subapical organite (or), microsensillum (ms), 7 (2 lateral, 5 dorsal) cylindrical, subequal sensilla and 15-25 slightly curved blunt-tipped sensilla in ventral field (Figs 5, 6). Antennal segment III organ with two long (lateral) and two short (internal), curved sensilla (Fig. 5). Microsensillum on antennal segment III present. Eversible sac between antennal segments III and IV present. Antennal segment I with 7 setae.</p><p>Ocelli: 8 + 8. Postantennal organ 1.8-2.3 times as large as single ocellus; the former with four lobes, its anterior pair larger than posterior pair. Accessory boss large (equal to or only slightly smaller than posterior lobes of postantennal organ), often granulated (Fig. 7).</p><p>Labrum with 5, 5, 4 setae; 4 prelabrals present. Maxillary head of C. armata type (Fjellberg 1984: fig. 18). Labial palp as shown in Fjellberg (1999: Fig. 4), but with 6 proximal setae. Outer maxillary lobe with one sublobal hair.</p><p>Tibiotarsi I, II, III with 19, 19, 18 setae, respectively, clavate setae absent. Claws with inner tooth and a pair of lateral teeth. Empodial appendage with broad lamelliform base and filiform apex reaching inner tooth or slightly beyond, ratio empodial appendage/ inner edge of claws = 0.4-0.7 (Fig. 8).</p><p>Ventral tube with 4 + 4 setae. Furca well developed. Ratio dens + mucro/inner edge of claw III = 1.8-2.2, ratio dens/mucro = 1.7-2.2. Dens with uniform fine granules and 7 dorsal setae (2-4 inner setae modified) (Fig. 9). Mucro wide at tip (ratio width of apical part/length of mucro = 0.4-0.6, usually 0.5), boat-like, with large outer lamella, (Fig. 9). Retinaculum with 4 + 4 teeth.</p><p>Anal spines yellowish, slightly curved, situated on high basal papillae, 1.1-1.7 times as long as inner edge of claws III (Fig. 4).</p><p>Distribution and ecology.</p><p>The range of distribution of C. stachi sp. nov. appears to be relatively wide. It is known from Denmark ( Jutland, Funen), Germany (Brandenburg, Hesse, Mecklenburg-West Pomerania, Saxony), Luxembourg, southern Norway (Akershus, Vestfold), Poland ( Carpathians: Beskid Niski, Beskid Sadecki, Bieszczady mountains) and Ukraine (Lviv District) (Fig. 1). Probably this species is distributed much more widely in Europe, but additional research is needed to prove this thesis. Ceratophysella stachi sp. nov. lives in lowlands and in the mountains (up to ca. 1000 m a.s.l) where it inhabits litter and mosses in different types of forests, and also heathlands and bogs .</p><p>Remarks.</p><p>Ceratophysella stachi sp. nov. belongs to a small European branch of species of the C. armata -group, which have strong tegumentary granulation, with distinct fields of coarse granules: C. granulata, C. lawrencei (Gisin, 1963), C. neomeridionalis (Nosek &amp; Červek, 1970), C. scotica (Carpenter &amp; Evans, 1899), and C. silvatica . It differs from all of them in the chaetotaxy of the lateral parts of the thoracic terga II-III (setae m6 present and one additional seta outside lateral sensillum m7 present or absent vs setae m6 and additional setae absent) which is exceptional within the whole C. armata -group (both characters are found in the genus, but in other groups of species: formerly classified as Mitchellania Wray, 1953 and C. denticulata). The remaining differences between C. stachi sp. nov. and related species mentioned above are summarized in Table 4.</p><p>1 Accessory boss near post-antennal organ small (about half the size of posterior lobes of post-antennal organ), mucro narrow at tip (ratio width of apical part/length of mucro = 0.22-0.52 (mean 39). 2 Fields of especially coarse granules only on abdominal terga IV-VI. 3 Thoracic terga II-III with one additional seta outside lateral sensillum m7 present or absent.</p></div>	https://treatment.plazi.org/id/3048909794415CAFAF7712BC8C024C10	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Skarzynski, Dariusz;Smolis, Adrian;Kovac, Ľubomir;Porco, David	Skarzynski, Dariusz, Smolis, Adrian, Kovac, Ľubomir, Porco, David (2021): A new European species of Ceratophysella (Collembola, Hypogastruridae) revealed by morphological data and DNA barcodes. ZooKeys 1021: 1-18, DOI: http://dx.doi.org/10.3897/zookeys.1021.63147, URL: http://dx.doi.org/10.3897/zookeys.1021.63147
