taxonID	type	description	language	source
95DCD953A7CA50F6BD3741C2BF00246E.taxon	materials_examined	Material. Poland (Carpathians): ISEZ: 2 syntypes on slide (formerly in alcohol), Tatra Mts, Dziura cave, 15. VII. 1909, leg. J. Stach; 29 spp. on slides, Tatra Mts, leg. J. Stach; Beskid Maly Mts, male, juv., Zagorze near Skawce, Grota Piaskowa cave, 350 m. a. s. l., XI. 1951, leg. Szymczakowski; DIBEC: Tatra Mts (leg. D. Skarzynski): 74 females, 17 males, litter of dwarf mountain pine shrubs on the slopes of the Gladkie Uplazianskie, at an altitude of 1500 - 1600 m a. s. l., 13. VII. 2001, 14. IX. 2002, 18. IX. 2004, 14. VIII. 2009; 4 females, 2 males, 2 juv., Chuda Turnia, moss on rocks, 1800 m a. s. l., 19. VIII. 2004; 11 females, 5 males, 2 juv., Krakow Gorge, spruce forest litter and mosses on rocks, 1050 - 1150 m a. s. l., 19. VIII. 2004; 4 males, Mylna cave, mosses in the entrance, 1090 m a. s. l., 19. VIII. 2004; female, Raptawicka cave, mosses in the entrance, 1150 m a. s. l., 19. VIII. 2004; female, juv., Dziura cave, litter in the entrance, 1000 m a. s. l., 24. VIII. 1991; 20 females, 15 males, litter of dwarf mountain pine shrubs on the slopes of Sucha Czuba, 1600 - 1700 m a. s. l., 17. IX. 2004; Beskid Zywiecki Mts: 3 females, 5 males, 6 juv., Babia Gora, litter of spruce forest and dwarf mountain pine shrubs, 1300 - 1500 m a. s. l., 4. VI. 1999, leg. A. Smolis; 4 females, 3 males, 5 juv., Pilsko, litter of spruce forest and dwarf mountain pine shrubs, 1300 - 1500 m a. s. l., 21. IX. 2004, leg. D. Skarzynski; Pieniny Mts, 4 females, male, 2 juv., Ociemny valley, 500 - 600 m a. s. l., mosses on rocks, 26. V. 1994, leg. R. J. Pomorski; Beskid Wyspowy Mts, 10 females, male, Zbojecka cave near Limanowa, 900 m a. s. l., bat guano, 12. VII. 2007, leg. K. Piksa. Slovakia (Western Carpathians): ISEZ: female, male, Pieniny Mts, Aksamitka cave, VII. 1931, leg. Grochmalicki. PJSU: 4 females, 2 males, Veľka Fatra Mts, Horna Tufna cave near Horny Harmanec village, 975 m a. s. l., entrance hall, cave sediment, 26 - 31. VIII. 1999, leg. Ľ. Kovac, 301 - 99, 302 - 99; female, Zapadne Tatry Mts, Brestovska cave near Zuberec village, entrance hall, cave sediment, 22. V. - 13. IX. 2006, leg. A. Mock, 671 - 06; 2 males, 3 juv., Belianske Tatry Mts, Kamzicia jaskyna cave near Zdiar village, 2002 m a. s. l., 15 m from entrance, cave sediment, 13. IX. 1991, leg. Ľ. Kovac; Low Tatras Mts: Demaenovska jaskyna slobody cave near Demaenova village, 812 m a. s. l.: female, male, rotten wood, 11. V. 2000, leg. P. Ľuptacik, 84 - 00; 2 females, male, Mramorove riecisko, bait, 11. V. - 27. IX. 2000, leg. Ľ. Kovac, 131 - 00, 133 - 00; Demaenovska ľadova jaskyna cave, 740 m a. s. l.: 2 females, entrance, talus deposit, 12. V. - 28. IX. 2000, Ľ. Kovac, 168 - 00; female, cave entrance, wood, 28. IX. 2000, leg. P. Ľuptacik, 170 - 00; female, Pusta jaskyna cave, Hlinena chodba, surface of water puddle, 24. VI. 2015, leg. Ľ. Kovac, 82 - 15; Pieniny Mts, Aksamitka cave near Haligovce village, 756 m a. s. l., leg. Ľ. Kovac: 6 females, 4 males, Blatisty dom, rotten wood, cave sediment, 12. III. - 26. V. 1998, 45 - 98, 197 - 98, 198 - 98, 199 - 98, 200 - 98; 2 females, Dom priekopnikov, bat guano, 26. V. 1998, 201 - 98; 2 females, 1 juv., Dom priekopnikov, cave sediment, 23. VIII. - 7. X. 1999, 363 - 98, 364 - 98, 366 - 98; male, soil of herbal cushion in front of cave, 26. V. 1998, 202 - 98; Levocske vrchy Mts, Jaskyna pod Jankovcom 2 cave near Ľubica village: 3 females, male, Hall A, bat guano, Hall II rotten wood, 5. XI. 2010, leg. P. Ľuptacik, 747 - 10, 758 - 10; male, passage B, surface of water puddle, 5. XI. 2010, leg. Z. Visnovska, 755 - 10; Slovak Paradise: Dobsinska ľadova jaskyna cave near Stratena village, 969 m a. s. l.: female, male, moss on rocks in front of cave, 23. VII. 1997, leg. Ľ. Kovac, 126 - 130 - 97, 2 females, humus and soil in front of cave, 7. X. 2004. leg. V. Sustr, 4 females, male, Psie Diery cave, cave sediment, 6. II. 1997, leg. V. Kosel, female, Vojenska cave, 20 m from entrance, bait, 6. XII. 1998, leg. V. Kosel, 22 - 99, 3 females, cave sediment, 28. I. - 6. II. 1997, leg. V. Kosel, 3 females, Klastorna cave, cave sediment, 27. I. - 4. II. 1997, leg. V. Kosel, female, 2 males, Duca cave, Dom, bait, 4. XII. 1998, leg. V. Kosel, 25 - 99, male, Stratenska cave, 200 m from entrance, rotten wood, 9. X. 1997, leg. Ľ. Kovac, 184 - 97; male, Muranska planina Plateau, Bobacka cave near Muranska Huta village, 30 m from entrance, cave sediment, 5. X. - 9. XI. 2000, Ľ. Kovac, 219 - 00; Cierna hora Mts: 2 females, Veľka ruzinska jaskyna cave near Mala Lodina village, 614 m a. s. l., 100 m from entrance, cave sediment, 10. VIII. - 14. X. 1996, leg. Ľ. Kovac, 1327 - 96, male, juv., Maly Ruzinok Valley, Tilio-Acerion, humus and soil, rotten wood, 19. IX. 2009, 23. IV. 2010, leg. Ľ. Kovac, 610 - 09, 139 - 10; male, Slovak Karst, Singliarova priepast cave near Honce village, 680 m a. s. l, 1 st Hall, rotten wood, 4. V. 2008, leg. P. Ľuptacik, 215 - 08.	en	Skarzynski, Dariusz, Smolis, Adrian, Kovac, Ľubomir, Porco, David (2021): A new European species of Ceratophysella (Collembola, Hypogastruridae) revealed by morphological data and DNA barcodes. ZooKeys 1021: 1-18, DOI: http://dx.doi.org/10.3897/zookeys.1021.63147, URL: http://dx.doi.org/10.3897/zookeys.1021.63147
3048909794415CAFAF7712BC8C024C10.taxon	materials_examined	Other material. Denmark (AF, leg. A. Fjellberg): female, Jutland, Himmerland, Rold Skov, Fagus liter, 20. III. 1994, 94.024; Funen: 6 females, 2 males, Fiskerup Skov, forest stream, 24. III. 1994, 94.076; female, Syltemade Adal, Fraxinus / Ulmus / Viburnum litter, 23. III. 1994, 94.070. Germany (SMNG): Brandenburg: 6 females, 3 males, 3 juv., " Wanninchen " nature reserve, bog, 1. V. 1972, leg. Hiebsch, 11911; 9 females, male, " Wanninchen " nature reserve, wet heather, 1972, leg. Hiebsch, 11911; male, 2 juv., " Bergener Moor " nature reserve, moor, Sphagnum, heather, pine forest, 1. V. 1972, leg. Hiebsch, 11912. Hesse: 5 females, Vogelsberg, 1985 - 1995, leg. W. Boehle. Mecklenburg-West Pomerania: female, Mueritz National Park, Neustrelitz, soil 0 - 5 cm, alder swamp woods not on acid peat, 16. X. 2013, leg. U. Burkhardt, 264 - F 865; 4 females, 2 males, Serrahn, Hauptmannsberg near Feldberg, sandy-gravelly moraine, largely unforested since the Middle Ages, 1973, leg. Hiebsch / ILN Greifswald, 11915; female, Serrahn, Klockenbruch, active, relatively undamaged raised bogs, moss / Sphagnum, 16. X. 2013, leg. R. Lehmitz and U. Burkardt, 261 - 01; male, Serrahn, Klockenbruch, active, relatively undamaged raised bogs, moss / Sphagnum, 5. XI. 2014, leg. R. Lehmitz and U. Burkardt, 401 - 12; female, Serrahn, " Kesselmoor " nature reserve, active, relatively undamaged raised bogs, moss / Sphagnum, 16. X. 2013, leg. R. Lehmitz, 49077 - 08; male, Serrahn, soil 0 - 5 cm, Medio-European collinear woodrush beech forest, 16. X. 2013, leg. U. Burkhardt, 263 - 03. Saxony: female and juv., Erzgebirge, Kleiner Kranichsee, Sphagnum, 22. VII. 1971, leg. W. Dunger, 7871. Luxembourg, subadult male, Vallee d'Our: Tintesmuehle, near the river, 21. VII. 1991, leg. M. Ursone, L- 91 - 101 (ISEZ). Norway (AF, leg. A. Fjellberg): Akershus: 4 females, " Oestmarka " nature reserve, Tappenbergvann, old spruce cones, 18. V. 1995, 95.163; female, Barum, Dalivannet, 16. XI. 1997. Vestfold: male, 32 / 93; 2 females, Larvik, N. Holtesetra, Hvarnes, lush Alnus and Fraxinus forest, 24. XI. 2007, 7.290; female and male, Larvik, Granasnekollen, Hvarnes, litter, oak / beech, 25. IX. 2004, 04.086; 3 females, Brunlanes, Hummerbakken, Telemark Camp., plant debris, beach, 14. XI. 1993, 93.077; female, Tjome, Sando, N-stranda, spongy Pinus litter behind beach, 22. IV. 2009, 9.086. Poland (Carpathians, DIBEC): female, 3 males, 2 juv., Beskid Sadecki Mts: " Las Lipowy Obrozyska " nature reserve near Muszyna, 600 m a. s. l., mosses on rocks and trees, 1. V. 2004, 25. VI. 2005, leg. A. Smolis, D. Skarzynski; female, Roztoka Ryterska, 600 m a. s. l., litter and mosses near stream, 3. V. 2004, leg. A. Smolis, D. Skarzynski; 5 females, " Uhryn " nature reserve, 850 m a. s. l., litter of fir-beech forest, 3. V. 2000, leg. A. Smolis; female, 3 males, 2 juv., " Barnowiec " nature reserve, 850 m a. s. l., mosses on rocks in an old beech forest, 10. V. 2003, leg. A. Smolis, D. Skarzynski; 4 females, 2 males, SE slopes of Jaworzyna Krynicka, litter in a beech forest, ca 800 m a. s. l, 2. V. 2004, leg. A. Smolis, D. Skarzynski. Bieszczady Mts: 2 females, N slopes of Krzemieniec, 1000 m a. s. l., litter in a stream valley, 19. V. 2000, leg. A. Smolis. Ukraine: male, Ivano-Frankove village, Lviv District, beech and elm forest, leaf litter and soil, leg. S. Bakaeva, 2.2.4.8 (SMNHL).	en	Skarzynski, Dariusz, Smolis, Adrian, Kovac, Ľubomir, Porco, David (2021): A new European species of Ceratophysella (Collembola, Hypogastruridae) revealed by morphological data and DNA barcodes. ZooKeys 1021: 1-18, DOI: http://dx.doi.org/10.3897/zookeys.1021.63147, URL: http://dx.doi.org/10.3897/zookeys.1021.63147
3048909794415CAFAF7712BC8C024C10.taxon	etymology	Etymology. Dedicated to Jan Stach, the excellent specialist in Collembola.	en	Skarzynski, Dariusz, Smolis, Adrian, Kovac, Ľubomir, Porco, David (2021): A new European species of Ceratophysella (Collembola, Hypogastruridae) revealed by morphological data and DNA barcodes. ZooKeys 1021: 1-18, DOI: http://dx.doi.org/10.3897/zookeys.1021.63147, URL: http://dx.doi.org/10.3897/zookeys.1021.63147
3048909794415CAFAF7712BC8C024C10.taxon	description	Description. Body length 1 - 2 mm. Colour (in alcohol) bluish-gray to bluish-black. Tegumental granulation strong, with fields of especially coarse granules on head (large uniform field covering whole dorsal side except antennal bases), thoracic terga II-III (two large subaxial fields and two lateral ones of medium size, Fig. 3), abdominal terga I-III (variable distribution: from four - Fig. 4 to seven fields of medium size as in C. granulata; see Skarzynski 2004 a: fig. 8), abdominal tergum IV (medium axial field and two lateral large ones, Fig. 4) and abdominal terga V-VI (large uniform fields covering almost whole dorsal side, Fig. 4). 6 - 9 granules between macrosetae p 1 on abdominal tergum V. Arrangement of setae on head typical for the genus, spine-like setae absent. Dorsal chaetotaxy of type B (Figs 3, 4). Thoracic terga II-III with setae m 6 present and one additional seta outside lateral sensillum m 7 present or absent. Setae p 1 on abdominal tergum IV developed as macrosetae, p 2 as microsetae, setae p 3 present. Differentiation of dorsal setae into micro- and macrosetae distinct. Setae long (ratio p 1 microsetae and p 2 macrosetae on thoracic tergum II / inner edge of claws III = 1 - 1.3 and 1.8 - 2.8, respectively), thick, curved, pointed at tips and only slightly serrate. Body sensilla (s) short (ratio sensillum p 4 and m 7 on thoracic tergum II / inner edge of claws III = 0.6 - 1 and 0.4 - 0.6, respectively), thin and smooth. Microsensilla (ms) on thoracic tergum II present (Fig. 3). Subcoxae I, II, III with 1, 2, 3 setae, respectively. Antennal segment IV with simple or lobed apical vesicle, subapical organite (or), microsensillum (ms), 7 (2 lateral, 5 dorsal) cylindrical, subequal sensilla and 15 - 25 slightly curved blunt-tipped sensilla in ventral field (Figs 5, 6). Antennal segment III organ with two long (lateral) and two short (internal), curved sensilla (Fig. 5). Microsensillum on antennal segment III present. Eversible sac between antennal segments III and IV present. Antennal segment I with 7 setae. Ocelli: 8 + 8. Postantennal organ 1.8 - 2.3 times as large as single ocellus; the former with four lobes, its anterior pair larger than posterior pair. Accessory boss large (equal to or only slightly smaller than posterior lobes of postantennal organ), often granulated (Fig. 7). Labrum with 5, 5, 4 setae; 4 prelabrals present. Maxillary head of C. armata type (Fjellberg 1984: fig. 18). Labial palp as shown in Fjellberg (1999: Fig. 4), but with 6 proximal setae. Outer maxillary lobe with one sublobal hair. Tibiotarsi I, II, III with 19, 19, 18 setae, respectively, clavate setae absent. Claws with inner tooth and a pair of lateral teeth. Empodial appendage with broad lamelliform base and filiform apex reaching inner tooth or slightly beyond, ratio empodial appendage / inner edge of claws = 0.4 - 0.7 (Fig. 8). Ventral tube with 4 + 4 setae. Furca well developed. Ratio dens + mucro / inner edge of claw III = 1.8 - 2.2, ratio dens / mucro = 1.7 - 2.2. Dens with uniform fine granules and 7 dorsal setae (2 - 4 inner setae modified) (Fig. 9). Mucro wide at tip (ratio width of apical part / length of mucro = 0.4 - 0.6, usually 0.5), boat-like, with large outer lamella, (Fig. 9). Retinaculum with 4 + 4 teeth. Anal spines yellowish, slightly curved, situated on high basal papillae, 1.1 - 1.7 times as long as inner edge of claws III (Fig. 4).	en	Skarzynski, Dariusz, Smolis, Adrian, Kovac, Ľubomir, Porco, David (2021): A new European species of Ceratophysella (Collembola, Hypogastruridae) revealed by morphological data and DNA barcodes. ZooKeys 1021: 1-18, DOI: http://dx.doi.org/10.3897/zookeys.1021.63147, URL: http://dx.doi.org/10.3897/zookeys.1021.63147
3048909794415CAFAF7712BC8C024C10.taxon	distribution	Distribution and ecology. The range of distribution of C. stachi sp. nov. appears to be relatively wide. It is known from Denmark (Jutland, Funen), Germany (Brandenburg, Hesse, Mecklenburg-West Pomerania, Saxony), Luxembourg, southern Norway (Akershus, Vestfold), Poland (Carpathians: Beskid Niski, Beskid Sadecki, Bieszczady mountains) and Ukraine (Lviv District) (Fig. 1). Probably this species is distributed much more widely in Europe, but additional research is needed to prove this thesis. Ceratophysella stachi sp. nov. lives in lowlands and in the mountains (up to ca. 1000 m a. s. l) where it inhabits litter and mosses in different types of forests, and also heathlands and bogs.	en	Skarzynski, Dariusz, Smolis, Adrian, Kovac, Ľubomir, Porco, David (2021): A new European species of Ceratophysella (Collembola, Hypogastruridae) revealed by morphological data and DNA barcodes. ZooKeys 1021: 1-18, DOI: http://dx.doi.org/10.3897/zookeys.1021.63147, URL: http://dx.doi.org/10.3897/zookeys.1021.63147
