taxonID	type	description	language	source
956F571BFF95FFE933249DD6AF10FDEE.taxon	type_taxon	Type species: Neognathus insolitus Willmann, 1952, by original designation	en	Khaustov, Alexander A. (2024): A new species and a new record of Caligonellidae (Acari: Raphignathoidea) from Russia. Persian Journal of Acarology 13 (1): 1-11, DOI: 10.22073/pja.v13i1.83381, URL: https://www.mendeley.com/catalogue/b71d5021-0b82-3489-970e-a1498cb1bdc2/
956F571BFF95FFEC30E79E43A9D0FBE7.taxon	diagnosis	Diagnosis	en	Khaustov, Alexander A. (2024): A new species and a new record of Caligonellidae (Acari: Raphignathoidea) from Russia. Persian Journal of Acarology 13 (1): 1-11, DOI: 10.22073/pja.v13i1.83381, URL: https://www.mendeley.com/catalogue/b71d5021-0b82-3489-970e-a1498cb1bdc2/
956F571BFF95FFEC30E79E43A9D0FBE7.taxon	description	Female – Cupuli ia well developed, tarsi III and IV with eight setae each (vs present), genua III and IV with two setae each, tarsus IV without solenidion, tibia IV without solenidion, palpgenu with one seta, stylophore without lateral projections, peritremes with 5 – 6 pairs of segments, two pairs of pseudanal setae. Description Female (Figs. 1 ‒ 3) – Length of idiosoma 265 (240 ‒ 250), width 160 (135 ‒ 140). Idiosomal dorsum (Fig. 1 A) ‒ Ovate. Dorsal idiosomal striae thin. Dorsal setae smooth, pointed. Cupules ia, im and ip large, almost round; ia located posterolaterad c 1, im posterolaterad d, and ip anterolaterad f. Lengths of dorsal setae: vi 8 (8 – 9), ve 12 (12), sci 13 (13 – 14), sce 12 (12 – 13), c 1 12 (12), c 2 12 (11 – 12), d 14 (13 – 14), e 13 (12 – 13), f 13 (11 – 12), h 1 11 (10 – 11), h 2 13 (11 – 12). Idiosomal venter (Fig. 1 B) ‒ All ventral setae smooth and pointed. Four pairs of aggenital and two pair of genital setae; cupuli ih located anterolaterad ag 4. Anal valves with two pairs of pseudanal setae. Two longitudinal groups of tiny sclerites situated between setae 4 a and ag 1. Lengths of ventral setae: 1 a 20 (20 ‒ 22), 1 b 15 (15 ‒ 18), 1 c 16 (16 ‒ 18), 2 c 19 (19 ‒ 20), 3 a 20 (20 ‒ 23), 3 c 16 (16 ‒ 20), 4 a 19 (19 ‒ 20), 4 c 16 (14 ‒ 16), ag 1 16 (16 ‒ 18), ag 2 15 (15 ‒ 18), ag 3 13 (13 ‒ 18), ag 4 11 (11 ‒ 13), g 1 15 (15 ‒ 17), g 2 15 (15 ‒ 16), ps 1 11 (11), ps 2 11 (11 ‒ 13). Gnathosoma (Fig. 2) ‒ Stylophore width 38 (38 ‒ 40); peritremes with 5 – 6 pairs of segments; cheliceral stylets 30 (30 ‒ 31), slightly curved; cheliceral levers poorly visible, about three times shorter than cheliceral stylets. Palpal supracoxal setae (ep) short, peg-like; palpal chaetotaxy: Tr 0, Fe 1 (d), Ge 1 (d), Ti 3 (d, l’, l ”), Ta 8 (1) (ba, bp, lp, va, acmϛ, ul’ϛ, ul ” ϛ, sulϛ, ω); tibial claw well developed, slightly hooked; all palpal setae smooth, eupathidia acmϛ, ul’ϛ, ul ” ϛ, sulϛ blunt-tipped, other palpal setae pointed. Subcapitulum with delicate striae in basal part; all subcapitular setae smooth and pointed; lengths of subcapitular setae: m 16 (16 ‒ 18), n 37 (37 ‒ 38), or 1 11 (11 ‒ 12), or 2 11 (11 ‒ 12), length of palptarsal solenidion ω 4 (4). Legs (Fig. 3) ‒ Lengths of legs: I 165 (170 ‒ 180), II 125 (125 ‒ 130), III 125 (125 ‒ 130), IV 155 (155 ‒ 160). Leg I (Fig. 3 A). Leg setation: Tr 1 (v’), Fe 3 (d, l’, bv ”), Ge 6 (d, l’, l ”, v’, v ”, k), Ti 5 (2) (dϛ, l’, l ”, v’, v ”, φ, φp), Ta 14 (2) (ft’ϛ, ft ” ϛ, tc’ϛ, tc ” ϛ, p’ϛ, p ” ϛ, a’, a ”, u’, u ”, vsϛ, pv’, pl’, pl ”, ω 1, ω 2). Supracoxal seta of leg I (el) peg-like; all leg setae smooth; setae d of tibia, (ft), (tc), (p) and vs of tarsus eupathid-like, blunt-tipped; other leg setae pointed; solenidion ω 1 8 (8) digitiform, solenidion ω 2 11 (11) baculiform and curved, solenidion φ 4 (4) digitiform, solenidion φp 8 (8 ‒ 9) baculiform and curved, seta k 3 (3). Leg II (Fig. 3 B). Leg setation: Tr 1 (v’), Fe 3 (d, l’, bv ”), Ge 6 (d, l’, l ”, v’, v ”, k), Ti 5 (1) (dϛ, l’, l ”, v’, v ”, φp), Ta 9 (2) (ft’, tc’ϛ, tc ” ϛ, a’, a ”, u’, u ”, pv, vs, ω 1, ω 2). All leg setae smooth; setae d of tibia and (tc) of tarsus eupathid-like, blunt-tipped; other leg setae pointed; solenidion ω 1 7 (6 ‒ 7) digitiform, solenidion ω 2 8 (8) baculiform and curved, solenidion φp 9 (7) baculiform, seta k 3 (3). Leg III (Fig. 3 C). Leg setation: Tr 2 (l’, v’), Fe 2 (d, ev’), Ge 2 (d, v’), Ti 4 (1) (d, l ”, v’, v ”, φp), Ta 8 (1) (tc’, tc ”, a’, a ”, u’, u ”, pv, vs, ω). Solenidion ω 10 (9) digitiform, solenidion φp 9 (8) baculiform. All leg setae smooth and pointed. Seta l ” of genu absent. Leg IV (Fig. 3 D). Leg setation: Tr 1 (v’), Fe 2 (d, ev’), Ge 3 (d, v’), Ti 4 (d, l ”, v’, v ”), Ta 8 (tc’, tc ”, a’, a ”, u’, u ”, pv, vs). Solenidia ω and φp absent. All leg setae smooth and pointed. Male and immatures unknown Type material Female holotype, slide ZISP T-Cal- 05, Russia, Stavropol Krai, Beshtau mountain, 44 ° 06 ' 13.1 " N, 43 ° 01 ' 13.0 " E, 30 July 2023, in soil on the meadow, 1180 m a. s. l., coll. A. A. Khaustov. Paratypes: 2 females, same data. Type deposition The holotype and one female paratype of the new species are deposited in the acarological collection of the Zoological Institute of RAS, St. Petersburg, Russia; one paratype is deposited in the mite collection of the Tyumen State University Museum of Zoology, Tyumen, Russia. Etymology The name of the new species refers to its distribution in Beshtau mountain. Differential diagnosis The female of the new species is very similar to Neognathus sibiriensis in having large palptibial claw, absence of solenidion ω on tarsus IV, absence of lateral projections on stylophore, presence of two pairs of pseudanal setae, presence of well-developed cupuli ia, presence of three setae on femur I, presence of seta k on genu II, presence of two setae on genu III, presence of only one seta on palpgenu, and presence of seta vs on tarsi III and IV. The new species differs from N. sibiriensis by the absence of solenidion on tibia IV (solenidion on tibia IV present in N. sibiriensis), presence of two setae on genu IV (three setae in N. sibiriensis), and in having 5 – 6 pairs of segments in peritremes (four pairs in N. sibiriensis).	en	Khaustov, Alexander A. (2024): A new species and a new record of Caligonellidae (Acari: Raphignathoidea) from Russia. Persian Journal of Acarology 13 (1): 1-11, DOI: 10.22073/pja.v13i1.83381, URL: https://www.mendeley.com/catalogue/b71d5021-0b82-3489-970e-a1498cb1bdc2/
956F571BFF90FFEC30BE98B8AE12FB69.taxon	type_taxon	Type species: Molothrognathus leptostylus Summers & Schlinger, 1955, by original designation	en	Khaustov, Alexander A. (2024): A new species and a new record of Caligonellidae (Acari: Raphignathoidea) from Russia. Persian Journal of Acarology 13 (1): 1-11, DOI: 10.22073/pja.v13i1.83381, URL: https://www.mendeley.com/catalogue/b71d5021-0b82-3489-970e-a1498cb1bdc2/
956F571BFF90FFE1304A993EA903FE91.taxon	diagnosis	Diagnosis	en	Khaustov, Alexander A. (2024): A new species and a new record of Caligonellidae (Acari: Raphignathoidea) from Russia. Persian Journal of Acarology 13 (1): 1-11, DOI: 10.22073/pja.v13i1.83381, URL: https://www.mendeley.com/catalogue/b71d5021-0b82-3489-970e-a1498cb1bdc2/
956F571BFF90FFE1304A993EA903FE91.taxon	description	Female – Dorsal shield absent; idiosomal striae very thick, double; three pairs of aggenital setae; three pairs of pseudanal setae; dorsal idiosomal setae short, subequal; seta vs of tarsus I absent; seta p ” of tarsus II absent; tibia I with one solenidion; stylophore elliptical. Redescription Female (Figs. 4 – 6) – Length of idiosoma 390 ‒ 410, width 205 ‒ 210. Idiosomal dorsum (Fig. 4 A) ‒ Ovate. Dorsal idiosomal striae thick, usually double; dorsal shield absent. Two pairs of subequal ocelli located posterolaterad setae sci. Cupuli ia, im and ip large, almost round; ia located just posteromesad posterior ocellus, im anterolaterad d, and ip posterolaterad f. All dorsal idiosomal setae short, subequal, smooth and pointed. Anal valves located dorsoterminal, with three pairs of pseudanal setae; setae ps 3 located dorsally. A pair of longitudinal, oblique rows of tiny subcuticular sclerites situated laterad setae c 1 and d. Lengths of dorsal setae: vi 19 ‒ 20, ve 17 ‒ 18, sci 19 ‒ 23, sce 20 ‒ 21, c 1 18 ‒ 19, c 2 19 ‒ 20, d 17 ‒ 18, e 17, f 15 ‒ 16, h 1 15 ‒ 17, h 2 16, ps 1 8, ps 2 12, ps 3 13. Idiosomal venter (Fig. 4 B) ‒ Ventral idiosoma striated, without plates. All ventral setae smooth and pointed. Setae 3 a located on the margin of coxisternal fields III. Coxisternal fields I ‒ IV sparsely punctate. Three pairs of aggenital and one pair of genital setae; cupuli ih located anterolaterad ag 3. A pair of almost parallel rows of small subcuticular sclerites situated laterad setae 4 a. Lengths of ventral setae: 1 a 25, 1 b 22 ‒ 23, 1 c 13 ‒ 14, 2 c 13 ‒ 14, 3 a 25 ‒ 26, 3 b 14 ‒ 15, 4 a 18 ‒ 19, 4 c 13 ‒ 14, ag 1 15 ‒ 16, ag 2 12 ‒ 13, ag 3 11 ‒ 13, g 12. Gnathosoma (Fig. 5) ‒ Stylophore short and wide, elliptical in shape; longitudinally striated in basal half and punctate in distal one, its width 67 ‒ 69; peritremes tubular, typical for the genus, their anterior ends situated near anterior margin of stylophore; peritremes inside with fine striae; cheliceral levers large, bean-shaped, about 1.5 times shorter than cheliceral stylets. Palpal supracoxal setae (ep) very small and located in deep depression with terminal pore; palpal chaetotaxy: Tr 0, Fe 1 (d), Ge 1 (d), Ti 3 (d, l’, l ”), Ta 7 (1) (ba, bp, va, acmϛ, ul’ϛ, ul ” ϛ, sulϛ, ω); tibial claw large, slightly hooked; all palpal setae smooth, eupathidia acmϛ, ul’ϛ, ul ” ϛ, and sulϛ distinctly widened distally, with four subapical flat lobes; seta va usually blunt-tipped, sometimes distinctly widened distally, with four subapical flat lobes (Fig. 2 A); seta bp slightly flattened and blunt-tipped distally; seta ba pointed; solenidion ω very small, rod-shaped. Subcapitulum striated in basal half and punctate in distal one; all subcapitular setae smooth and pointed. Rostrum of subcapitulum short and wide, almost V-shaped split between setae or 2. Lengths of subcapitular setae: m 30 ‒ 32, or 1 11 ‒ 12, or 2 11 ‒ 12, length of palptarsal solenidion ω 3, length of cheliceral stylet 28. Legs (Fig. 6) ‒ Lengths of legs (excluding claws): I 235, II 180 ‒ 185, III 200 ‒ 210, IV 225 ‒ 230. Leg I (Fig. 6 A) longer than other legs. Leg setation: Tr 1 (v’), Fe 2 (d, bv ”), Ge 6 (d, l’, l ”, v’, v ”, k), Ti 5 (1) (d, l’, l ”, v’, v ”, φp), Ta 14 (1) (ft’, ft ”, tc’ϛ, tc ” ϛ, p’ϛ, p ” ϛ, a’, a ”, u’, u ”, pv’, pv ”, pl’, pl ”, ω). Supracoxal seta of leg I (el) located in deep depression with terminal pore; all leg setae smooth; seta vs absent, setae (tc) and (p) eupathid-like, blunt-tipped, with very small subapical projections, other leg setae pointed; solenidion ω 8 digitiform, solenidion φp 10 ‒ 11 baculiform; seta k 3 weakly clavate. Leg II (Fig. 6 B). Leg setation: Tr 1 (v’), Fe 2 (d, bv ”), Ge 4 (d, l’, l ”, v’), Ti 5 (d, l’, l ”, v’, v ”), Ta 9 (1) (ft’, tc’, tc ”, a’, a ”, u’, u ”, pv’, pv ”, ω). All leg setae smooth and pointed; seta p ” ϛ absent; solenidion ω 6 ‒ 7 digitiform. Leg III (Fig. 6 C). Leg setation: Tr 1 (v’), Fe 2 (d, ev’), Ge 2 (d, v’), Ti 4 (d, l ”, v’, v ”), Ta 8 (tc’, tc ”, a’, a ”, u’, u ”, pv’, pv ”). All leg setae smooth and pointed. Leg IV (Fig. 6 D). Leg setation: Tr 1 (v’), Fe 2 (d, ev’), Ge 2 (d, v’), Ti 4 (d, l ”, v’, v ”), Ta 8 (tc’, tc ”, a’, a ”, u’, u ”, pv’, pv ”). All leg setae smooth and pointed. Male (Fig. 7) – Idiosoma narrower posteriorly than in female. Length of idiosoma 285, width 185. Idiosomal dorsum (Fig. 7 A) ‒ Striation and setal lengths similar with those of female, except presence of almost smooth pygidial shield with setae h 1 and h 2. Only two pairs of pseudanal setae. Lengths of dorsal setae: vi 15, ve 13, sci 16, sce 18, c 1 16, c 2 16, d 15, e 14, f 14, h 1 11, h 2 14, ps 1 11, ps 2 12. Aedeagus weakly sclerotized, long, and slightly curved. Idiosomal venter (Fig. 7 B) ‒ Ventral idiosoma completely striated. Two pairs of aggenital setae; genital setae absent. Other characters as in female. Lengths of ventral setae: 1 a 18, 1 b 19, 1 c 13, 2 c 13, 3 a 22, 3 b 12, 4 a 16, 4 c 15, ag 1 16, ag 2 16. Gnathosoma as in female. Width of stylophore 52, lengths of setae: m 22, or 1 10, or 2 10, length of palptarsal solenidion ω 3. Legs as in female, except absence of seta v’ on trochanters I ‒ IV. Lengths of legs: I 195, II 155, III 165, IV 185. Solenidion ω I 5, φp 7, ω II 5, seta k I 3. Immatures unknown. Material examined Two females, one male, Russia, Tyumen Oblast, city of Tyumen, Gagarin Park, moss on soil, 57 ° 10 ' N, 65 ° 36 ' E, 28 March 2023, collected by A. A. Khaustov. Remarks	en	Khaustov, Alexander A. (2024): A new species and a new record of Caligonellidae (Acari: Raphignathoidea) from Russia. Persian Journal of Acarology 13 (1): 1-11, DOI: 10.22073/pja.v13i1.83381, URL: https://www.mendeley.com/catalogue/b71d5021-0b82-3489-970e-a1498cb1bdc2/
