identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
94541A33FF8EFFD7A97FEBDCFC64FCF1.text	94541A33FF8EFFD7A97FEBDCFC64FCF1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Jordanius Afifeh & Yağmur & Al-Saraireh & Amr 2024	<div><p>Jordanius gen. n.</p><p>(Figures 1, 5–6, 9–78, 363-364, 369–370) http://zoobank.org/ urn:lsid:zoobank.</p><p>org:act: BADFD5DD-EDB4-478D-A26E-445207962048</p><p>TYPE SPECIES. Scorpio granulomanus Al-Saraireh, Yağmur, Abu Afifeh &amp; Amr, 2023 (see description below) .</p><p>ETYMOLOGY. The generic name is derived from Jordan, the country to which this genus is endemic.</p><p>DIAGNOSIS. Medium-sized species. Adult males are about 47.29– 52.46 mm, and females are about 48.13–52.58 mm in length. Anterior margin of carapace bilobed, with 13–17 medium to large setae and a distinct depression. There is a protrusion in the middle of the depression. Carapace acarinate, only the posteriomedian carinae are visible. Carapace shagreened, but the triangular region between median eyes and the anterior margin is smooth in males; smooth and lustrous with small granules laterally in females. The external surface of chela manus densely covered with large, raised, rounded and somewhat pointed granules, the posterior margin of the external surface of chela manus bears very large spinoid protrudings. Internal surface of chela manus with small to moderate pointed granules. Chela manus flattened and elongated; manus length to chela depth ratio almost the same in both sexes (1.24–1.29). Chela fingers are relatively long with small pointed gradually decreased granules anteriorly. Fixed and movable fingers with 4 or 5 strong accessory denticles. Tergites I–VI matt, shagreened in males with scattered small granules; smooth and lustrous with flattened medium granules in females. Metasoma I with 10 carinae, II-IV with 8, and V with 7 carinae. Lateral inframedian carinae on segment I indistinct. Ventral submedian and ventrolateral carinae moderate to strong, with moderate rounded granules, size and numbers of granules decrease from segment I toward segment IV, nearly obsolete on segment IV. Ventromedian and ventrolateral carinae strong with spaced large and pointed granules on segment V, pointed granules gradually increase posteriorly on ventrolateral carina of segment V. Pectinal teeth number 11–13 in males, 9–13 in females. Trichobothriotaxy of type C, orthobothriotaxic (Vachon, 1974), pedipalp chela trichobothrium it placed basally along fixed finger near ib. Tibial spurs absent. Tarsi armed only with prolateral pedal spurs, tarsi of legs I to IV with 5/8–6/8– 7/9–7/10 internal and external spines arranged in series.</p><p>AFFINITIES. The new genus Jordanius gen. n. is a fossorial borrowing scorpion belonging to the family Scorpionidae Latreille, 1802; this genus is closely related to the genus Scorpio, differing primarily in the shape, elongation, and pattern of granulation of pedipalp chela. The new genus can be separated from the genus Scorpio by having remarkably elongated chela manus, and very large, dense and pointed granules on the dorsoexternal surface of the chela manus, in addition genus Scorpio has generally smooth chela fingers, whereas Jordanius gen. n. has small pointed gradually decreased granules anteriorly on chela fingers.</p><p>Figures 1 and 2 show the general difference in elongation and pattern of granulation in chela manus between Jordanius gen. n. and Scorpio .</p></div>	https://treatment.plazi.org/id/94541A33FF8EFFD7A97FEBDCFC64FCF1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Afifeh, Bassam Abu;Yağmur, Ersen Aydın;Al-Saraireh, Mohammad;Amr, Zuhair	Afifeh, Bassam Abu, Yağmur, Ersen Aydın, Al-Saraireh, Mohammad, Amr, Zuhair (2024): Revision of the genus Scorpio in Jordan, with a description of a new genus and three new species (Scorpiones: Scorpionidae). Euscorpius 391: 1-66, DOI: 10.5281/zenodo.14660305
94541A33FF8EFFD6AB6AEA42FCC6F9B1.text	94541A33FF8EFFD6AB6AEA42FCC6F9B1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Jordanius granulomanus (Al-Saraireh, Yagmur, Abu Afifeh & Amr 2023) Afifeh & Yağmur & Al-Saraireh & Amr 2024	<div><p>Jordanius granulomanus (Al-Saraireh et al., 2023), comb. n.</p><p>(Figures 1, 75–76, 364, 370)</p><p>Scorpio granulomanus: Al-Saraireh et al., 2023: 1–16, figs. 1–59.</p><p>TYPE LOCALITY AND TYPE DEPOSITORY. Jordan, Jerash Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.82247&amp;materialsCitation.latitude=32.244" title="Search Plazi for locations around (long 35.82247/lat 32.244)">Dibbeen Forest Reserve</a>, 32°14'38.40''N 35°49'20.90''E, 783 m a. s. l. , AZMM.</p><p>TYPE MATERIAL EXAMINED. Jordan: Jerash Governorate, Dibbeen Forest Reserve, 32°14'46.9''N 35°49'20.00''E, 745 m a. s.l., 2♂ (paratypes), 6 September 2013, leg. Z. Amr (RSCN); Jerash Governorate, Dibbeen Forest Reserve, 32°14'52.70''N 35°49'23.80''E, 741 m a. s. l., 1♂ 9♀ (paratypes), 22 April 2022, leg. B. Abu Afifeh &amp; M. Al-Saraireh (BAPC); Jerash Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.823277&amp;materialsCitation.latitude=32.24797" title="Search Plazi for locations around (long 35.823277/lat 32.24797)">Dibbeen Forest Reserve</a>, 32°14'38.40''N 35°49'20.90''E, 783 m a. s. l., 1♂ (holotype) (AZMM /Sco- 2022:01), 1♀ (paratype) (AZMM /Sco-2022:02), 1♂ 10♀ (paratypes), 20 May 2022, leg. B. Abu Afifeh &amp; M. Al-Saraireh (BAPC); Jerash Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.823277&amp;materialsCitation.latitude=32.24797" title="Search Plazi for locations around (long 35.823277/lat 32.24797)">Dibbeen Forest Reserve</a>, 32°14'52.70''N 35°49'23.80''E, 741 m a. s. l., 2♀ (paratypes), 26 June 2022, leg. B. Abu Afifeh &amp; M. Al-Saraireh (BAPC) .</p><p>DIAGNOSIS. Medium-sized species compared with its congeners. Adults are about 51 mm in length. Carapace is light reddish brown in males and dark reddish brown in females with reticulations in both sexes. Mesosoma basically, reddish brown in males and reddish black in females. Venter dark yellow in males and light brown in females. Pedipalp femur and patella dark yellow, chela reddish yellow in both sexes. Metasoma and legs dark yellow in males and light brownish yellow in females. Anterior margin of carapace bilobed, carapace finely granulated, the triangular region between the median eyes and the anterior margin is smooth with scattered moderate granules in males, with fine and dense granules laterally and scattered moderate granules anteriorly and posteriorly in females. Dorsoexternal carinae of pedipalp femur distinct with separate granules distally, sometimes with several rows of spaced granules, intercarinal surface smooth with numerous setae, dorsoexternal surface with numerous rounded and pointed granules of different size, internal surface with some large, pointed granules. Patella with two carinae. Dorsomedian carina distinct with swollen and some fused granules and the ventrointernal carinae are obsolete with few swollen granules, intercarinal surface is smooth with numerous setae. Chela manus densely covered with large rounded and somewhat pointed granules, the posterior margin of the external surface of chela manus bears very large spinoid protrusions. Internal surface of chela manus with small to moderate pointed granules along the dorsal margin. Chela manus flattened and elongated; manus length to chela depth ratio almost the same in both sexes with an average of 1.27. Chela fingers are relatively long with small pointed gradually decreased granules anteriorly. Fixed and movable fingers with 4 or 5 strong accessory denticles. Tergites I–VI matte, completely covered with fine granules in males; glossy and smooth, at most slightly shagreened laterally in females. Tergite VII finely granular with moderate, pointed granules posteriorly and laterally in males, but smooth anteriorly, granulate posteriorly and laterally in females. Sternites III–VI wrinkled medially, almost smooth laterally in males, smooth and shiny in females with numerous moderate setae on the posterior margins. Sternite VII granular and feebly wrinkled medially; shagreened laterally with four tuberculated carinae with lobated granules; the posterior margin bears numerous moderate setae. Metasoma: Segment I with 10 carinae, segment II–IV with 8, and segment V with 5 carinae. Lateral inframedian carinae on segment I indistinct. Ventral submedian and ventrolateral carinae moderate to strong, with moderate rounded granules, size and numbers of granules decrease from segment I toward segment IV, nearly obsolete on segment IV. Ventromedian and ventrolateral carinae strong with spaced large and pointed granules on segment V, pointed granules gradually increase posteriorly on ventrolateral carina of segment V. Pectinal teeth number 9–12 in males, 8–11 in females. Genital operculum is heart-shaped in females, ovoid in males. Trichobothriotaxy of type C, orthobothriotaxic (Vachon, 1974), pedipalp chela trichobothrium it placed basally along fixed finger near ib. Tibial spurs absent. Tarsi armed only with prolateral pedal spurs, tarsi of legs I to IV with 4/7–6/8–7/8–7/8 internal and external spines arranged in series.</p></div>	https://treatment.plazi.org/id/94541A33FF8EFFD6AB6AEA42FCC6F9B1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Afifeh, Bassam Abu;Yağmur, Ersen Aydın;Al-Saraireh, Mohammad;Amr, Zuhair	Afifeh, Bassam Abu, Yağmur, Ersen Aydın, Al-Saraireh, Mohammad, Amr, Zuhair (2024): Revision of the genus Scorpio in Jordan, with a description of a new genus and three new species (Scorpiones: Scorpionidae). Euscorpius 391: 1-66, DOI: 10.5281/zenodo.14660305
94541A33FF8FFFC3ABB7EF82FB2FFB0B.text	94541A33FF8FFFC3ABB7EF82FB2FFB0B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Jordanius maysaraensis Afifeh & Yağmur & Al-Saraireh & Amr 2024	<div><p>Jordanius maysaraensis gen. et sp. n.</p><p>(Figures 5, 9–74, 77–78, 363, 369, 374; Table 1) http: //zoobank. org/urn: lsid: zoobank. org: act: C6B0D55E -</p><p>6C8D -47D4-82C5-D2F2697A842D</p><p>TYPE LOCALITY AND TYPE DEPOSITORY. Jordan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.66228&amp;materialsCitation.latitude=32.134003" title="Search Plazi for locations around (long 35.66228/lat 32.134003)">Al Ardha</a> road, 2.7 km NW of Maysara, 32°08'02.4''N 35°39'44.2''E, Balqa Governorate, 447 m a. s. l. , AZMM.</p><p>TYPE MATERIAL EXAMINED. Jordan: Balqa Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.66228&amp;materialsCitation.latitude=32.134003" title="Search Plazi for locations around (long 35.66228/lat 32.134003)">Al Ardha</a> road, 2.7 km NW of Maysara, 32°08'02.4''N 35°39'44.2''E, 447 m a. s. l., 1♂ (holotype) (AZMM /Sco-</p><p>2022:10), 1♀ (paratype) (AZMM /Sco-2022:11), 5♀, 11 May 2022, leg. B. Abu Afifeh (BAPC); Balqa Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.662003&amp;materialsCitation.latitude=32.13392" title="Search Plazi for locations around (long 35.662003/lat 32.13392)">Al Ardha</a> road, 2.7 km NW of Maysara, 32°08'02.1''N 35°39'43.2''E, 434 m a. s. l. , 3♂, 2 subadult ♂, 2 subadult ♀, 2 juvs. ♂, 13 May 2022, leg. B. Abu Afifeh (BAPC); Balqa Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.66236&amp;materialsCitation.latitude=32.133392" title="Search Plazi for locations around (long 35.66236/lat 32.133392)">Al Ardha</a> road, 2.7 km NW of Maysara, 32°08'00.2''N 35°39'44.5''E, 438 m a. s. l. , 5♂ 4♀, 16 May 2022, leg. B. Abu Afifeh (BAPC); Balqa Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.654057&amp;materialsCitation.latitude=32.181335" title="Search Plazi for locations around (long 35.654057/lat 32.181335)">Zai</a> water pipeline road, 470 m E of Zai water pump station (PS2), 32°10'52.8''N 35°39'14.6''E, 124 m a. s. l. , 4♀, 26 Jan 2023, leg. B. Abu Afifeh (BAPC); Ajloun Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.623497&amp;materialsCitation.latitude=32.269527" title="Search Plazi for locations around (long 35.623497/lat 32.269527)">1.7 km W of Kufranjeh Dam</a>, 32°16'10.3''N 35°37'24.6''E, 122 m a. s. l. , 6♀, 7 subadult ♀, 2 subadult ♂, 28 Jan 2023, leg. B. Abu Afifeh (BAPC) .</p><p>ETYMOLOGY. The specific epithet refers to Maysara town in Balqa Governorate, Jordan.</p><p>DIAGNOSIS. Medium-sized species. Total length in adults is about 52.50 mm in both sexes. General coloration dark yellow to light yellowish brown. Carapace anterior margin distinctly bilobed. Carapace acarinate, only posteriomedian carinae visible, carapace shagreened but triangular region between median eyes and anterior margin smooth with scattered moderate and flattened granules in males; smooth and lustrous with small granules laterally in females. The external surface of chela manus densely covered with large, raised, rounded and somewhat pointed granules, the posterior margin of the external surface of chela manus bears very large spinoid protrudings. Internal surface of chela manus with small to moderate pointed granules along the dorsal margin. Chela manus flattened and elongated; manus length to chela depth ratio almost the same in both sexes. Chela fingers are relatively long with small pointed gradually decreased granules anteriorly. Fixed and movable fingers with 4 or 5 strong accessory denticles. Tergites I–VI matt, shagreened in males with scattered small granules in males; smooth and lustrous with flattened medium granules in females. Metasoma I with 10 carinae, II–IV with 8, and V with 7 carinae. Lateral inframedian carinae on segment I indistinct. Ventral submedian and ventrolateral carinae moderate to strong, with moderate rounded granules, size and numbers of granules decrease from segment I toward segment IV, nearly obsolete on segment IV. Ventromedian and ventrolateral carinae strong with spaced large and pointed granules on segment V, pointed granules gradually increase posteriorly on ventrolateral carina of segment V. Telson globular in females and relatively slender in males. Pectinal teeth number 11–13 in males, 10–12 in females. Genital operculum ovoid in males, cordate or heart-shaped in females, anterior margin slightly convex, posterior area slightly but widely elongated and a small posterior indentation exist in females. Trichobothriotaxy of type C, orthobothriotaxic (Vachon, 1974), pedipalp chela trichobothrium it placed basally along fixed finger near ib. Tibial spurs absent. Tarsi armed only with prolateral pedal spurs, tarsi of legs I to IV with 5/8–6/8–7/9–7/10 internal and external spines arranged in series.</p><p>DESCRIPTION: (♂ holotype, unless otherwise specified). Coloration (Figs. 5, 9–12). General coloration dark yellow to light yellowish brown. Prosoma: Carapace is light olive yellow with light brown triangular reticulations limited by the lateral eyes at the base and posterior median furrow in both sexes. Between median eyes is brownish black and around median eyes is light brown. Mesosoma: Basically, light olive yellow with brown pigmentation and reticulations on tergites; overlap area of segments yellow therefore yellow bands visible on posterior margins of segments I-VI. Venter dark olive yellow, poststernites yellowish. Pedipalp: femur, patella, and chela olive yellow. Coxae and sternum dark yellow, genital operculum and pectines lustrous yellow. Pedipalp: Femur and patella dark yellow, dorsal carina of patella and granules reddish brown. Chela manus dark yellow; fixed finger, and movable finger dark yellow to reddish yellow; ventrointernal surface of chela manus dark yellow; granules reddish brown or reddish black on apex; finger dental margins and movable finger condyles reddish black. Metasoma: Dark yellow to olive yellow, darker in ventral surface; granules on dorsolateral, ventrolateral, ventral and ventral submedian carinae light reddish brown. Vesicle of telson dark yellow, aculeus reddish yellow at the base and dark reddish brown at the tip. Legs: olive yellow, spinules and condyles red or reddish black. Chelicerae: Chelicera manus dark yellow with brown reticulations, fingers brown, teeth reddish black.</p><p>Carapace (Figs. 21–28). Carapace anterior margin distinctly bilobed. Carapace acarinate, only posteriomedian carinae visible, carapace shagreened but triangular region between median eyes and anterior margin smooth with scattered moderate and flattened granules in males; smooth and lustrous with small granules laterally in females. Anterior margin with 13-17 medium to large setae and distinct depression. There is a protrusion in the middle of the depression. Along anterior margin a rounded and flattened row of granules present. Median ocular tubercle distinct and located in the center of the carapace. A pair of median eyes, and three pairs of lateral eyes exist; the first two lateral eyes in equal size, and the third slightly reduced and located separately. Anteriomedian furrow moderate but posterior median and posterior lateral furrows distinct. 12 moderate setae exist on the carapace.</p><p>Chelicerae (Figs. 21–28). Cheliceral dentition characteristic for the family Scorpionidae (Vachon, 1963) . Dorsal surface of manus smooth, convex; smooth longitudinal ridges present anteriorly; prolateral and ventral surfaces densely setose; prodorsal margin bears 3 reddish macrosetae. Fixed finger dorsal and ventral surfaces densely setose, dorsal margin bears 4 teeth: distal, subdistal, median, and basal; ventral margin without teeth; movable finger dorsal surface smooth; ventral surface densely setose; dorsal margin bears 4 teeth: external distal, subdistal, median, and basal; ventral margin with internal distal teeth. The movable finger always ends in two distal teeth–one external and one internal– between which is inserted the distal tooth of the fixed finger.</p><p>Mesosoma (Figs. 13–28). Tergites I–VI acarinate. Tergites IVI matt, shagreened with scattered small granules in males; smooth and lustrous with flattened medium granules in females. Tergite VII shagreened in males, smooth in females with moderate, pointed granules posteriorly and laterally; carinae granulate and present only in posterior half of segment. Posterior margin of tergites with microsetae and a long seta on seventh tergite at end of each carina. Sternites III-VI wrinkled medially, almost smooth laterally in males, completely smooth in females. Sternite VII granular with coarse granules between lateral carinae and feebly wrinkled medially; shagreened laterally with four tuberculate carinae with lobated granules; posterior margin bears numerous moderate setae. Genital operculum ovoid in males, cordate or heart-shaped in females, anterior margin slightly convex, posterior area slightly but widely elongated and a small posterior indentation exist in females. Pectines short, dentate margin length to pectin length ratio 0.73 in male holotype. Teeth numbers 11–13 in males and 10–12 in females; three marginal and three to five median lamellae. Stigmas linear, angled 45° and conspicuous.</p><p>Metasoma and telson (Figs. 53–72). Metasoma I with 10, IIIV with 8, and V with 7 carinae. Lateral inframedian carinae on segment I indistinct. Ventral submedian and ventrolateral carinae strong on segments I–II, with moderate rounded granules, weak to moderate on III–IV, size and numbers of granules decrease from segment I toward segment IV, nearly obsolete on segment IV. Ventromedian and ventrolateral carinae strong with spaced large and pointed granules, continues in lateral of anal arch on segment V, pointed granules gradually increase posteriorly on ventrolateral carina of segment V. Dorsolateral carinae strong on segments I–IV with pointed and spaced, small granules; moderate on segment V with rounded granules. Lateral supramedian carinae moderate to strong with spaced granules on segments I–IV. Segment V with a row of granules laterally on the anterior half of segment. Intercarinal surfaces on segments I–V with fine to moderate granules in males, smooth and with scattered fine granules in females on dorsal and lateral surfaces. Segments I–V covered moderately with red setae of variable size, more dense on ventral surface. Vesicle globular, smooth dorsally, granulated ventrally and laterally; ventral aspect with numerous small, spinoid granules, vesicle with many red macrosetae even on the base of aculeus. Aculeus strongly curved and shorter than vesicle.</p><p>Pedipalp (Figs. 29–52). Pedipalp femur with three carinae; dorsointernal and ventrointernal carinae moderate with spaced rounded granules; dorsoexternal carinae of pedipalp femur distinct with separate granules distally, sometimes with several rows of spaced granules, intercarinal surface smooth with numerous setae, dorsoexternal surface with numerous rounded and pointed granules of different sizes, internal surface with some large, pointed granules. Patella with two carinae. Dorsomedian carina distinct with swollen and some fused granules and ventrointernal carinae obsolete with a few swollen granules, intercarinal surface smooth with numerous setae. The external surface of chela manus densely covered with large, raised, rounded and somewhat pointed granules, the posterior margin of the external surface of chela manus bears very large spinoid protrudings. Internal surface of chela manus with small to moderate pointed granules along dorsal margin. Ventroexternal carina strong with dark rather fused granules, ventrointernal carina smooth. Digital and external secondary carinae not visible with more distinct and darker granules only, parallel each other. Chela manus flattened and elongated. Intercarinal surface smooth with numerous distinct setae. Chela fingers are relatively long with small pointed gradually decreased granules anteriorly. Fixed and movable fingers with 4 or 5 strong accessory denticles. Trichobothriotaxy of type C; orthobothriotaxic (Vachon, 1974); femur of pedipalp with 3 (1 internal, 1 dorsal, 1 external, and 0 ventral) trichobothria; patella of pedipalp with 19 (1 internal, 2 dorsal, 13 external, and 3 ventral) trichobothria; chela of pedipalp with 26 trichobothria; of them 16 on the hand (0 internal, 2 dorsal, 10 external, and 4 ventral) trichobothria; and 10 on the fixed finger (2 internal, 4 dorsal, 4 external, and 0 ventral) trichobothria.</p><p>Legs (Figs. 73–74). Tibial spurs absent; tarsi armed with prolateral pedal spurs only, tarsi of right legs I to IV with 5/8– 6/8–7/9–7/10 internal and external spines arranged in series.</p><p>Measurements. See Table 1.</p><p>AFFINITIES. Jordanius maysaraensis gen. et sp. n. shows some morphological similarities with J. granulomanus comb. n. in the presence of large, rounded, somewhat pointed granules on the dorsal surface of chela manus; also in having a remarkably elongated chela manus. The new species can be distinguished from J. granulomanus comb. n. by:</p><p>a) having less dense and relatively smaller granulation on the external surface of chela manus (Figs. 75–78).</p><p>b) higher pectinal teeth number; 11–13 (average 12.0) in males (26 combs, n=13), 10–12 (average 10.9) in females (24 combs, n=12) in J. maysaraensis gen. et sp. n.; whereas it is 9–12 with average 10.7 in males (18 combs, n=9), 8–11 with average 9.9 in females (44 combs, n=22) in J. granulomanus comb. n.</p><p>c) general coloration of J. maysaraensis sp. n. is olive yellow (Fig. 5), whereas J. granulomanus comb. n. has generally reddish-brown coloration (Fig. 6).</p></div>	https://treatment.plazi.org/id/94541A33FF8FFFC3ABB7EF82FB2FFB0B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Afifeh, Bassam Abu;Yağmur, Ersen Aydın;Al-Saraireh, Mohammad;Amr, Zuhair	Afifeh, Bassam Abu, Yağmur, Ersen Aydın, Al-Saraireh, Mohammad, Amr, Zuhair (2024): Revision of the genus Scorpio in Jordan, with a description of a new genus and three new species (Scorpiones: Scorpionidae). Euscorpius 391: 1-66, DOI: 10.5281/zenodo.14660305
94541A33FF9AFFC3ABAEEC14FAE6FAAB.text	94541A33FF9AFFC3ABAEEC14FAE6FAAB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scorpio fuscus Linne 1758	<div><p>Scorpio fuscus (Ehrenberg in Hemprich &amp;</p><p>Ehrenberg, 1829) (Figures 3–4, 79–144, 360, 366; Table 2)</p></div>	https://treatment.plazi.org/id/94541A33FF9AFFC3ABAEEC14FAE6FAAB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Afifeh, Bassam Abu;Yağmur, Ersen Aydın;Al-Saraireh, Mohammad;Amr, Zuhair	Afifeh, Bassam Abu, Yağmur, Ersen Aydın, Al-Saraireh, Mohammad, Amr, Zuhair (2024): Revision of the genus Scorpio in Jordan, with a description of a new genus and three new species (Scorpiones: Scorpionidae). Euscorpius 391: 1-66, DOI: 10.5281/zenodo.14660305
94541A33FF91FFF4A932EF18FC5BF826.text	94541A33FF91FFF4A932EF18FC5BF826.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scorpio jordanensis Afifeh & Yağmur & Al-Saraireh & Amr 2024	<div><p>Scorpio jordanensis sp. n.</p><p>(Figures 2, 8, 145–214, 216, 218, 220, 222, 362, 368, 371; Table 3) http: //zoobank. org/urn: lsid: zoobank. org: act: 169D83E6-</p><p>E0D4-4124-8B2C -938E5F10EE48</p><p>Scorpio maurus palmatus: Amr &amp; Al-Oran, 1994: 189; Amr et al., 2004: 241, fig. 3.</p><p>Scorpio kruglovi: Amr et al., 2015: 36, fig. 2E.</p><p>TYPE LOCALITY AND TYPE DEPOSITORY. Jordan, Balqa Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.817444&amp;materialsCitation.latitude=32.09294" title="Search Plazi for locations around (long 35.817444/lat 32.09294)">Um Al-Dananeer</a>, 32°05'34.6”N 35°49'02.8”E, 773 m a. s. l. , AZMM.</p><p>TYPE MATERIAL EXAMINED. Jordan: Ma’an Governorate, near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.565887&amp;materialsCitation.latitude=30.526556" title="Search Plazi for locations around (long 35.565887/lat 30.526556)">Al-Shawback Castle</a>, 30°31'35.6''N 35°33'57.2''E, 1335 m a. s.</p><p>l., 2♂, 8 September 2013, leg. L. Prendini, Z. Amr, O. Abed, T. Al Share &amp; L. Al-Azam (RSCN); Aqaba Governorate, Al- Barrah, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.326748&amp;materialsCitation.latitude=29.546583" title="Search Plazi for locations around (long 35.326748/lat 29.546583)">Wadi Rum</a>, 29°32'47.7''N 35°19'36.3''E, 1100 m a. s. l. , 1♂, 10 September 2013, leg. Z. Amr, N. Hamidan &amp; L.Al-Azam (RSCN); Aqaba Governorate, Al-Khoshakhashe, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.431778&amp;materialsCitation.latitude=29.497807" title="Search Plazi for locations around (long 35.431778/lat 29.497807)">Wadi Rum</a>, 29°29'52.1''N 35°25'54.4''E, 1109 m a. s. l. , 1♂, 11 September 2013, leg. Z. Amr, N. Hamidan &amp; T. Al Share (RSCN); Zarqa Governorate, Abu-Al <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.049053&amp;materialsCitation.latitude=32.14703" title="Search Plazi for locations around (long 36.049053/lat 32.14703)">Zighan</a>, 32°08'49.3''N 36°02'56.6''E, 457 m a. s. l, 1♂, 6 July 2018, leg. M. Al-Saraireh and A. Gayada (BAPC); Zarqa Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.925667&amp;materialsCitation.latitude=32.158142" title="Search Plazi for locations around (long 35.925667/lat 32.158142)">Alook</a>, 32°09'29.3''N 35°55'32.4''E , 608, m a. s. l., 1♂ 1♀, 23 March 2019, leg. M. Al-Saraireh (BAPC); Amman Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.026306&amp;materialsCitation.latitude=31.820557" title="Search Plazi for locations around (long 36.026306/lat 31.820557)">Sahab</a>, 31°49'14.0''N 36°01'34.7''E, 773 m a. s. l. , 2♂, 13 August 2019, leg. M. Al-Saraireh (BAPC); Balqa Governorate, Ghor <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.612137&amp;materialsCitation.latitude=31.987028" title="Search Plazi for locations around (long 35.612137/lat 31.987028)">Ira</a>, 5 km W of Ira town, 31°59'13.3''N 35°36'43.7''E, 25 m a. s. l. , 2♂, 3 November 2019, leg. B. Abu Afifeh &amp; M. Al-Saraireh (BAPC); Amman Governorate, Ghor Sweimeh near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.589222&amp;materialsCitation.latitude=31.77036" title="Search Plazi for locations around (long 35.589222/lat 31.77036)">Dead Sea</a>, 31°46'13.3''N 35°35'21.2''E, 365 m a. s. l. , 2♂, 5 July 2020, leg. M. Al-Saraireh (BAPC); Irbid Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.655445&amp;materialsCitation.latitude=32.414974" title="Search Plazi for locations around (long 35.655445/lat 32.414974)">Kufrabeel</a>, 32°24'53.9''N 35°39'19.6''E, 370 m a. s. l. , 1♀, 26 September 2020, leg. Omar Aldhoon (BAPC); Ma’an Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.43397&amp;materialsCitation.latitude=30.386194" title="Search Plazi for locations around (long 35.43397/lat 30.386194)">Ayn Al-Baydah</a>, 5 km N of Petra, 30°23'10.3''N 35°26'02.3''E, 1075 m a. s. l. , 1 subadult ♀, 18 June 2021, leg. B. Abu Afifeh &amp; M. Al-Saraireh (BAPC); Balqa Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.817444&amp;materialsCitation.latitude=32.09294" title="Search Plazi for locations around (long 35.817444/lat 32.09294)">Um Al-Dananeer</a>, 32°05'34.6''N 35°49'02.8''E, 773 m a. s. l. , 1♂ (holotype) (AZMM /Sco- 2021:39), 4 ♂, 2 juvs. ♂, 12 August 2021, leg. B. Abu Afifeh (BAPC); Balqa Governorate, Ghor <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.61222&amp;materialsCitation.latitude=31.98275" title="Search Plazi for locations around (long 35.61222/lat 31.98275)">Ira</a>, 5 km W of Ira town, 31°58'57.9''N 35°36'44.0''E, 16 m a. s. l. , 1♀, 13 April 2022, leg. B. Abu Afifeh &amp; R. Abu Afifeh (BAPC); Balqa Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.83586&amp;materialsCitation.latitude=32.10214" title="Search Plazi for locations around (long 35.83586/lat 32.10214)">Um Al-Dananeer</a>, 32°06'07.7''N 35°50'09.1''E, 827 m a. s. l. , 1♀ (AZMM /Sco-2021:40), 21 April 2022, leg. B. Abu Afifeh; Tafilah Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.602722&amp;materialsCitation.latitude=30.692276" title="Search Plazi for locations around (long 35.602722/lat 30.692276)">Dana Biosphere Reserve</a>, 30°41'32.2''N 35°36'09.8''E, 1500 m a. s. l. , 11♂ 8♀, 25-26 May 2022, leg. B. Abu Afifeh (BAPC); Amman Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.041637&amp;materialsCitation.latitude=31.942389" title="Search Plazi for locations around (long 36.041637/lat 31.942389)">Al-Baydah</a>, 31°56'32.6''N 36°02'29.9''E, 768 m a. s. l. , 7 subadults ♂, 6 June 2022, leg. M. Al-Saraireh (BAPC); Balqa Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.823944&amp;materialsCitation.latitude=32.107525" title="Search Plazi for locations around (long 35.823944/lat 32.107525)">Um Al-Dananeer</a>, 32°06'27.1''N 35°49'26.2''E, 855 m a. s. l. , 5♂ 1♀, 28 July 2022, leg. B. Abu Afifeh (BAPC); Tafilah Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.47536&amp;materialsCitation.latitude=30.692833" title="Search Plazi for locations around (long 35.47536/lat 30.692833)">KhirbatAl-Ghuwaiba</a>, 14 km NW of Dana village, 30°41'34.2''N 35°28'31.3''E, 162 m a. s. l. , 4♀, 20 January 2023, leg. B. Abu Afifeh (BAPC); Balqa Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.62836&amp;materialsCitation.latitude=32.02133" title="Search Plazi for locations around (long 35.62836/lat 32.02133)">6.3 km E of Karameh Dam</a>, 32°01'16.8''N 35°37'42.1''E, 2 m a. s. l. , 3♀, 17 February 2023, leg. B. Abu Afifeh (BAPC); Balqa Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.662167&amp;materialsCitation.latitude=32.05639" title="Search Plazi for locations around (long 35.662167/lat 32.05639)">Um Kharrouba</a>, 32°03'23.0''N 35°39'43.8''E, 402 m a. s. l. , 1♂ 2♀, 18 March 2023, leg. B. Abu Afifeh (BAPC); Balqa Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.67814&amp;materialsCitation.latitude=32.113247" title="Search Plazi for locations around (long 35.67814/lat 32.113247)">Homrat Al Sahn</a>, 32°06'47.7''N 35°40'41.3''E, 452 m a. s. l. , 2♀, 18 March 2023, leg. B. Abu Afifeh (BAPC); Balqa Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.674168&amp;materialsCitation.latitude=32.125385" title="Search Plazi for locations around (long 35.674168/lat 32.125385)">2.5 km SW of Maysara town</a>, 32°07'31.4''N 35°40'27.0''E, 509 m a. s. l. , 1♂ 1♀, 18 March 2023, leg. B. Abu Afifeh (BAPC); Tafilah Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.602722&amp;materialsCitation.latitude=30.692276" title="Search Plazi for locations around (long 35.602722/lat 30.692276)">Dana Biosphere Reserve</a>, 30°41'32.2''N 35°36'09.8''E, 1500 m a. s. l. , 8♀, 8 July 2023, leg. B. Abu Afifeh (BAPC); Tafilah Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.597664&amp;materialsCitation.latitude=30.700724" title="Search Plazi for locations around (long 35.597664/lat 30.700724)">Ein Lahda</a> groundwater, 3.1 km NW of Dana village, 30°42'02.6''N 35°35'51.6''E, 1430 m a. s. l. , 4♂ 1♀, 8 July 2023, leg. B. Abu Afifeh (BAPC); Ma’an Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.52839&amp;materialsCitation.latitude=30.247084" title="Search Plazi for locations around (long 35.52839/lat 30.247084)">Basta</a>, 30°14'49.5''N 35°31'42.2''E, 1538 m a. s. l. , 1♂, 7 September 2023, leg. B. Abu Afifeh &amp; M. Al-Saraireh (BAPC); Aqaba Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.11086&amp;materialsCitation.latitude=29.411943" title="Search Plazi for locations around (long 35.11086/lat 29.411943)">9.5 km SW of Titin village</a>, 29°24'43.0''N 35°06'39.1''E, 1115 m a. s. l. , 14♂ 1♀, 9 May 2024, leg. B. Abu Afifeh, R. Abu Afifeh &amp; H. Bdareen (BAPC); Zarqa Governorate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.059166&amp;materialsCitation.latitude=32.134113" title="Search Plazi for locations around (long 36.059166/lat 32.134113)">As-Sukhnah</a>, 32°08'02.8''N 36°03'33.0''E, 480 m a. s. l. , 1♂, 17 May 2024, leg. B. Abu Afifeh &amp; M. Al-Saraireh (BAPC) .</p><p>ETYMOLOGY. The specific name refers to Jordan, the country in which the new species is endemic and widely distributed.</p><p>DIAGNOSIS. Medium sized species compared with its congeners. Adults are about 42.50- 53.54 mm in males and 45.70- 56.90 mm in females in length. General coloration light reddish yellow. Carapace is slightly longer than its wide; almost trapezoid in shape. Carapace anterior margin distinctly bilobed with a deep median indentation. Carapace lacks carinae, only posteriomedian carinae slightly visible. Carapace shagreened but a triangular region between median eyes and anterior margin smooth. Median ocular tubercle not distinct and located in the center of the carapace. Tergites IVII acarinate. Tergites I–VII matt and shagreened; posttergits of segments I–IV with scattered flattened granules; tergite VII moderately covered with dense coarse and pointed granules in male, while in females tergites I-VII smooth, lustrous; posttergits of segments I-IV with flattened moderate granules, tergite VII moderately covered with dense coarse and pointed granules. Pectines short; teeth numbers 10–13 in males and 9–13 in females. Metasoma I with 10, II–IV with 8, and V with 5 carinae. Metasoma I–II wider than long, segments III–V longer than wide. Ventral submedian carinae and ventrolateral carinae strong with moderate, rounded and fused granules on segments I–II, strong and obsolete granules on segments III–IV in male; strong with large lobate and spaced granules on segments I–II in females. Between ventral submedian carinae of segment I 10–20 small to medium granules exist. Dorsolateral carinae moderate and rounded with moderate spinoid granules, less distinct granules in females on segment V. Vesicle globular and smooth. Pedipalp femur with three carinae, patella with two carinae. The external surface of chela manus with very large and very flattened granules, more distinct and rounded in anterior area, posterior part with smaller granules; interior part nearly smooth in females. Posterior margin of external surface, and internal margin with large and conical granules. Digital carina not distinct but with larger and more distinct granules than other granules and not fused; external secondary carina distinct with large and fused granules; that get denser anteriorly. Chela manus flattened, slightly longer than wide. Fixed and movable fingers with 4 or 5 strong accessory denticles. Intercarinal surface with few distinct setae on manus. Trichobothriotaxy of type C; orthobothriotaxic (Vachon, 1974).</p><p>DESCRIPTION (♂ holotype, unless otherwise specified). Coloration (Figs. 8, 145–148, 211-214). General coloration light reddish yellow. Carapace: light reddish yellow, black between median eyes, and light reddish brown around median eyes, posterior area of median eyes with reddish brown reticulation; posterior margin with a yellow band in overlap area of first mesosoma segment in male; yellowish olive brown in females. Mesosoma: Tergites I-VII light reddish yellow; posterior margins of segments I-VI with yellow bands in overlap area in males; lustrous, olive brown with reticulation and yellowish olive patches in females. Coxae and sternum lustrous and reddish yellow. Genital operculum and pectines shiny yellow. Sternites III-VII reddish yellow in male; olive brown in females. Pedipalp: Femur and patella reddish yellow; carinae and granules red. Chela manus reddish yellow, granules darker, ventroexternal, ventrointernal, digital carina, external secondary carina and fingers dark yellowish red in males, manus darker, fingers lighter in females. Dental margin and condyles reddish black. Metasoma: metasomal segments I-IV uniformly light reddish yellow, segment V dark reddish yellow with reticulation in males; metasomal segments I-V uniformly reddish yellow in females. Telson dark yellow; aculeus reddish at the base and black at the tip. Legs: Tarsi and basitarsus light yellow; pretarsus, femur and tibia dark yellow. Spinules and condyles reddish brown. Chelicerae: Chelicera manus dark yellowish brown with reticulations posteriorly. Fingers brown, tips of fingers and teeth reddish.</p><p>Carapace (Figs. 157–164). Carapace is slightly longer than its wide; almost trapezoid in shape. Carapace anterior margin distinctly bilobed, with 8 medium to large setae and a deep median indentation. Carapace lacks carinae, only posteriomedian carinae slightly visible. Carapace shagreened but a triangular region between median eyes and anterior margin smooth, anterior of carapace and along anteriomedian furrow with moderately dense small granules in males; carapace smooth and lustrous with small granules laterally in females. Median ocular tubercle not distinct and located in the center of the carapace. A pair of median eyes and three pairs of lateral eyes exist; the third located separately and slightly smaller. Median eyes separated by one ocular diameter. The anteromedian furrow is narrow, shallow and more distinct in males. The posteromedian furrow is wide and deep and bifurcated as an inverted T-shaped at posterior edge and located in arrow-shaped depressions between posteriomedian carinae and posterior margin. Posterior lateral furrows wide and distinct. A few setae exist on the anterior area of carapace. Chelicerae (Figs. 149–156). Cheliceral dentition characteristic for the family Scorpionidae (Vachon, 1963); subdistal tooth and basal teeth conspicuous rather than on the movable finger. Manus lustrous, no longitudinal ridges present anteriorly.</p><p>Mesosoma (Figs. 149–156). Tergites I–VII acarinate. Tergites I–VII matt and shagreened; posttergites of segment I–IV with scattered flattened granules; tergite VII moderately covered with dense coarse and pointed granules in male; tergites I–VII smooth, lustrous; posttergits of segment I–IV with flattened moderate granules, tergite VII moderately covered with dense coarse and pointed granules in females. Sternites III–VI with scattered setae; wrinkled mideally in male; smooth and lustrous in females. Sternite VII granular between submedian carinae with four granular carinae, granules fused, granular between lateral and submedian carinae in females. Genital operculum ovoid; a small posterior indentation exists in males, while cordate in shape; anterior margin slightly convex; posterior area slightly elongated, and a small posterior indentation exist in females. Pectines short: teeth numbers 10–13 in males, and 9–13 in females. Three marginal and three to five median lamellae. Stigmas linear, angled 45° and conspicuous.</p><p>Metasoma and telson (Figs. 189–208). Metasoma I with 10, II–IV with 8, and V with 5 carinae. Metasoma I–II wider than long, III–V longer than wide. Lateral inframedian carinae and on segment I indistinct, composed by a few spaced rounded granules on anterior. Ventral submedian carinae and ventrolateral carinae strong with moderate, rounded and fused granules on metasoma I–II, strong and obsolete granules on metasoma III–IV in male; strong with large lobate and spaced granules on segments I–II in females. Dorsolateral carinae strong and crenulate with small, spaced granules on merasoma I–IV. Lateral supramedian carinae strong and obsolete on segments I–IV. Intercarinal area on segments I–IV smooth covered with scattered small sized granules at ventral and lateral surfaces, between ventral submedian carinae of segment I 10–20 small to medium granules exist; segment I–II with moderately fine granules, III–IV with scattered fine granules and without granules medially in males; segments I–IV smooth with scattered fine granules and without granules medially in females. Segment V: Ventromedian carina moderate with large, spinoid and pointed granules, bifurcated anteriorly; ventrolateral carinae strong and serrated with spaced large and conical granules, gradually increase posteriorly, continues in lateral of anal arch; dorsolateral carinae moderate and rounded with moderate spinoid granules, granules less distinct in females; segment V with a row of granules laterally at posterior half, granules less distinct in females. Lateral surfaces smooth with scattered fine granules. Ventral surface densely covered with large granules. Dorsal surface smooth covered with fine granules, without granules medially. Ventral of anal arch serrated with pointed denticles. Ventral and lateral surfaces covered with moderately variable sized setae on segments I–V. Vesicle globular and smooth; with spinoid granules ventrally, more distinct in males. Vesicle with numerous macrosetae reaching the base of aculeus. Aculeus strongly curved and shorter than vesicle.</p><p>Pedipalp (Figs. 165–188). Pedipalp femur with three carinae; dorsointernal and ventrointernal carinae moderate with spaced rounded granules; dorsoexternal carinae strong with separate large and rounded granules. Intercarinal area smooth with small sized granules dorsally and medium sized granules ventrally. Intercarinal surface with a few distinct setae. Patella with two carinae; dorsomedian carina strong with so flattened and fused granules; ventrointernal carinae moderate with rounded coarse granules. Intercarinal area of dorsal, ventral and external surfaces smooth and lustrous, external surface with moderate rounded granules; internal surface shagreened. Intercarinal surface with a few distinct setae. The external surface of chela manus with very large and very flattened granules, more distinct and rounded in anterior area, posterior part with smaller granules; interior part nearly smooth in females. Posterior margin of external surface and internal margin with large and conical granules. Internal surface of chela manus smooth with moderate pointed granules in anterior area. Ventroexternal carina strong, obsolete without granules; ventrointernal carina smooth, rounded without granules. Digital carina not distinct with granules larger and more distinct than other granules and not fused; external secondary carina distinct with large and fused granules that become denser anteriorly. Chela manus flattened, slightly longer than wide. Chela fingers are relatively short and without granules, slightly longer in females. Fixed and movable fingers with 4-5 strong accessory denticles. Intercarinal surface with few distinct setae on manus. Trichobothriotaxy of type C; orthobothriotaxic (Vachon, 1974).</p><p>Legs (Figs. 209–210). Coxa and trochanters smooth. Tarsi of right legs I to IV with 4-6/8–5-6/8-9– 6-7/8-9–6-7/9 internal and external spines arranged in series. Basitarsus of legs I with 2–3 retrolateral spines (generally with 3 spines) and on legs II with 3–4 retrolateral spines (generally with 3 spines).</p><p>Measurements. See Table 4.</p><p>AFFINITIES. S. jordanensis sp. n. shows similarities with S. palmatus regarding to several morphological characters; both are medium sized yellow species, external surface of chela manus with reduced granulation, and chela fingers are short. That led several previous authors to assign specimens of the widespread Scorpio from Jordan to S. palmatus (Wahbeh, 1976; El-Hennawy, 1988; Amr &amp; Al-Oran 1994; Amr &amp; Abu Baker, 2004). The two species can be distinguished by the following characters: (a) pedipalp chela fingers of S. jordanensis sp. n. is shorter than in S. palmatus (see table 6); (b) seventh sternite and ventral surface of segment I moderately granular in S. jordanensis sp. n. (Figs. 220, 222) whereas in S. palmatus it has a few scattered granules (Figs. 219, 221); (c) ventrolateral and ventral submedian carinae of segments I and II with spaced distinct rounded granules in S. jordanensis sp. n. (Figs. 216, 218), whereas indistinct and fused granules are present in S. palmatus (Figs. 215, 217); (d) lateral inframedian carina of segment I visible and present on posterior part in S. jordanensis sp. n. (Figs. 216, 218) whereas it is not visible and composed of 2–3 granules on posterior of segment I in S. palmatus (Figs. 215, 217).</p><p>S. jordanensis sp. n. can be distinguished from S. fuscus by several characters shown in table 6;</p><p>a) general coloration light reddish yellow in S. jordanensis sp. n. (Figs. 8, 145–148), whereas it is dark reddish brown to brownish black in S. fuscus (Figs. 3–4, 79–82);</p><p>b) S. jordanensis sp. n. is smaller in size (adults 53–56 mm in total length), than S. fuscus (adults 61–67 mm in total length); c) pectines is longer in males of S. jordanensis sp. n. than in males of S. fuscus; pectines extending well beyond coxa-trochanter articulation of leg IV in male of S. jordanensis sp. n. (Fig. 163), whereas extending approximately to the coxa-trochanter articulation of leg IV in male of S. fuscus (Fig. 97); d) external surface of pedipalp chela with isolated rounded and separated granules in male of S. jordanensis sp. n. (Figs. 171, 175), whereas male of S. fuscus has reticular fused and flattened granulation (Figs. 105, 109);</p><p>e) pectinal teeth numbers in S. jordanensis sp. n. (10–13 in males and 9–13 in females) is higher than in S. fuscus (9–10 in males and 8–11 in females);</p><p>f) metasomal segment II is always wider than long in S. jordanensis sp. n., whereas it is longer than wide or as long as wide in S. fuscus .</p></div>	https://treatment.plazi.org/id/94541A33FF91FFF4A932EF18FC5BF826	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Afifeh, Bassam Abu;Yağmur, Ersen Aydın;Al-Saraireh, Mohammad;Amr, Zuhair	Afifeh, Bassam Abu, Yağmur, Ersen Aydın, Al-Saraireh, Mohammad, Amr, Zuhair (2024): Revision of the genus Scorpio in Jordan, with a description of a new genus and three new species (Scorpiones: Scorpionidae). Euscorpius 391: 1-66, DOI: 10.5281/zenodo.14660305
94541A33FFA0FFE1A8CDED23FCB4F83E.text	94541A33FFA0FFE1A8CDED23FCB4F83E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scorpio palmatus (Ehrenberg 1828)	<div><p>Scorpio palmatus (Ehrenberg, 1828)</p><p>(Figures 215, 217, 219, 221, 223-288, 361, 367, 372; Table 4) http://zoobank.org/urn:lsid:zoobank.org:act:974207E9-</p><p>2461-49B3-BCE1-C7A4DB2A0B93</p><p>Buthus (Heterometrus) maurus palmatus: Hemprich &amp; Ehrenberg, 1828: pl. I, fig. 1.</p><p>Heterometrus palmatus: Birula, 1898: 134–135.</p><p>Scorpio palmatus: Birula, 1900: 13–14; Birula, 1908: 147– 148.</p><p>Scorpio maurus palmatus: Birula, 1909: 517–518; Birula, 1910: 166–169, pl. XI, fig. 8; pl. XII, fig. 7; Fet, 2000: 478 (complete references list until 1999).</p><p>NEOTYPE LOCALITY AND TYPE DEPOSITORY. Egypt, Alexandria, Borg El Arab, Bahig, W of Bahig village, AZMM.</p><p>MATERIAL EXAMINED. Egypt, no exact locality, “Provenance d’Egypte, 2013, Don. de Mr. J. Vanni ”, 10♂ 16♀ (IRSNB) . Egypt, Alexandria, Borg El Arab, Bahig, W of Bahig village, 1♂ (neotype), 1 September 2001 (AZMM /SCO-2001:1).</p><p>DIAGNOSIS. Medium sized species compared with its congeners. Adults are about 47.96-50.29 mm in males, and 46.52-49.63 mm in females. General coloration yellowish in males, slightly darker in females. Carapace is slightly longer than wide; almost upright trapezoid in shape. Carapace anterior margin distinctly bilobed with a deep median indentation. Carapace lacks carinae, only posteriomedian carinae slightly visible. Smooth around median eyes and anterior region of carapace, rest of carapace shagreened, laterals with small scattered granules in females. Median ocular tubercle not distinct and located in the center of the carapace. Tergites I–VII acarinate. Tergites I–VII matt and shagreened in males; smooth and, lustrous in females. Sternite VII granular between lateral carinae with four granular carinae, granules fused. Pectines short; teeth numbers 10–13 in males; and 8–12 in females (Talal et al., 2015); with three marginal and three or four median lamellae. Metasoma I with 10, II–IV with 8, and V with 5 carinae. Ventromedian carina of metasoma V moderate with large, spinoid and pointed granules; ventrolateral carinae strong and serrated with spaced large and pointed granules, gradually increase posteriorly, continues in lateral of anal arch. Vesicle globular and smooth; granulated ventrally with spinoid granules in both sexes. The external surface of chela manus with very large and flattened granules, more distinct and rounded on anterior area, posterior part nearly smooth. Internal surface of chela manus smooth with moderate pointed granules on anterior area. Chela manus flattened, slightly longer than wide. Chela fingers are relatively short without granules, slightly longer in females. Fixed and movable fingers with 4–5 strong accessory denticles. Intercarinal surface with few setae on manus, but fingers with more dense setae. Trichobothriotaxy of type C; orthobothriotaxic (Vachon, 1974).</p><p>DESCRIPTION (♂, unless otherwise specified). Coloration (Figs. 223–226). General coloration yellowish. Carapace yellow and reticulated with light reddish yellow patches in males; light reddish yellow, medially reticular. Tergites I–VII light brownish yellow and reticulated with yellow patches medially, lustrous in females. Coxae and sternum yellow. Genital operculum and pectines lustrous and light yellow. Sternites III–VII shiny light yellowish. Pedipalp femur and patella dark reddish brown with blackish reticulations. Chela manus yellow with lateral surface with very light reddish yellow reticulation and granules light reddish yellow in males, slightly darker in females. Fingers yellow in males and yellowish red in females; dental margin and condyles dark reddish in both sexes. Metasoma I–IV uniformly yellow, V light brownish yellow in males; metasoma I–III uniformly dark yellow, IV yellowish brown, segment V brownish in females. Telson yellow in males, slightly darker in females, aculeus reddish at the base and black at the tip. Legs: Tarsi, basitarsus and pretarsus yellow, femur and tibia lustrous and dark yellow. Spinules and condyles reddish brown. Chelicera manus dark yellow with reticulations anteriorly. Chelicera fingers yellow, tips of fingers and teeth reddish.</p><p>Carapace (Figs. 235–242). Carapace is slightly longer than its wide; almost upright trapezoid in shape. Carapace anterior margin distinctly bilobed with 10 medium or large setae and a deep median indentation. Carapace lacks carinae, only posteriomedian carinae slightly visible. Carapace shagreened but a triangular region between median eyes and anterior margin smooth, anterior of carapace and along anteriomedian furrow with moderately dense small granules in males, carapace smooth and lustrous with small granules laterally in females. Median ocular tubercle not distinct and located in the center of the carapace. A pair of median eyes and three pairs of lateral eyes exist; the third located separately and slightly smaller. The anteromedian furrow is narrow and shallow, and more distinct in males. The posteromedian furrow is wide and bifurcated as an inverted T-shaped at posterior edge, and located in arrow-shaped depressions between posteriomedian carinae and posterior margin. Posterior lateral furrows wide and distinct. A few setae exist on the anterior area of carapace. Chelicerae (Figs. 227–234). Cheliceral dentition characteristic for the family of Scorpionidae (Vachon, 1963); subdistal tooth and basal teeth conspicuous rather than on the movable finger. No longitudinal ridges present anteriorly.</p><p>Mesosoma (Figs. 227–234). Tergites I–VII acarinate. Tergites I–VII matt and shagreened, tergite VII with scattered medium sized granules in males, while tergites I–VII smooth, lustrous, tergite VII covered with modereately medium sized flattened granules in females. Sternites III–VI smooth and wrinkled with scattered setae in males; smooth and lustrous with scattered setae in females. Sternite VII granular between lateral carinae with four granular carinae, granules fused. Genital operculum ovoid, a small posterior indentation exists in males, while cordate; anterior margin slightly convex; posterior area slightly elongated, and a small posterior indentation exist in females. Pectines short: teeth numbers 10–12 in males and 8–11 in females; three marginal and three to four median lamellae. Stigmas linear, angled 45° and conspicuous.</p><p>Metasoma and telson (Figs. 267–286). Metasoma I slightly wider than long, II–V longer than wide. Metasoma I with 10, II–IV with 8, and V with 5 carinae. Lateral inframedian carinae on segment I indistinct, composed by a few spaced rounded granules anteriorly. Ventral submedian carinae and ventrolateral carinae strong with moderate, rounded and fused granules on segments I–II, strong and obsolete granules on segments III–IV. Dorsolateral carinae strong and crenulated with small, spaced granules on segments I–IV. Lateral supramedian carinae strong, and obsolete on segments I–IV. Intercarinal area smooth, covered with moderately medium and small sized granules in males, while smooth at ventral and lateral surfaces on segments I–IV in females, 3–4 small granules exist between ventral submedian carinae on segment I; dorsal surfaces on segments I–V shagreened in males, nearly smooth in females. Segment V: Ventromedian carina moderate with large, spinoid and pointed granules; ventrolateral carinae strong and serrated with spaced large and conical granules, gradually increase posteriorly, continues in lateral of anal arch; dorsolateral carinae moderate and rounded with moderate granules; segment V with a row of granules laterally at posterior half. Lateral surfaces shagreened in males, smooth with a few flattened granules in females. Ventral surface densely covered with large granules. Dorsal surface shagreened in males, nearly smooth in females. Ventral of anal arch serrated with pointed denticles. Ventral and lateral surfaces covered with moderately variable sized setae in segments I-V. Vesicle globular and smooth; granulated ventrally with spinoid granules in both sexes. Vesicle with numerous macrosetae reaching the base of aculeus. Aculeus strongly curved and shorter than vesicle.</p><p>Pedipalp (Figs. 243–266). Pedipalp femur with three carinae; dorsointernal and ventrointernal carinae moderate with spaced rounded granules; dorsoexternal carinae strong with separated, large and rounded granules. Intercarinal area shagreened with moderate sized granules dorsally and shagreened ventrally. Intercarinal surface with a few distinct setae. Patella with two carinae; dorsomedian carina strong with so flattened and fused granules; ventrointernal carinae obsolete without granules. Intercarinal area of dorsal, ventral and external surfaces smooth and lustrous, external surface with moderate flattened granules; internal surface shagreened. Intercarinal surface with a few distinct setae. The external surface of chela manus with very large and very flattened granules, more distinct and rounded on anterior area, posterior part nearly smooth. Posterior margin of external surface with a few large, rounded granules. Internal surface of chela manus smooth with moderate pointed granules on anterior area. Ventroexternal carina strong, obsolete without granules; ventrointernal carina smooth, rounded without granules. Digital and external secondary carinae not distinct; granules larger anteriorly; the granules seperated in males, fused in females; and become denser anteriorly. Chela manus flattened, slightly longer than wide. Chela fingers are relatively short without granules, slightly longer in females. Fixed and movable fingers with 4–5 strong accessory denticles. Intercarinal surface with few distinct setae in manus, but fingers with more dense setae. Trichobothriotaxy of type C; orthobothriotaxic (Vachon, 1974).</p><p>Legs (Fig. 287–288). Coxa and trochanters smooth. Tarsi of right legs I to IV with 4-5/7-8–4-6/7-9–5-6/8-10–5-7/8-10 internal and external spines arranged in series. Basitarsus of legs I with 2–3 retrolateral spines (generally with 2 spines) and on legs II with 3-4 retrolateral spines (generally with 3 spines). Measurements. See Table 4.</p><p>COMMENTS. Braunwalder &amp; Fet (1998: 32) noted that type specimens of Buthus (Heterometrus) palmatus are unknown. Hemprich &amp; Ehrenberg (1828) originally assigned to this taxon specimens from a very wide range they outlined as “Libyan desert (? Egypt), Egypt (Alexandria, Sinai), Lebanon (mountains), Arabia, Syria.” For the purposes of taxonomic stability, we designate here a neotype from Alexandria, Egypt .</p></div>	https://treatment.plazi.org/id/94541A33FFA0FFE1A8CDED23FCB4F83E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Afifeh, Bassam Abu;Yağmur, Ersen Aydın;Al-Saraireh, Mohammad;Amr, Zuhair	Afifeh, Bassam Abu, Yağmur, Ersen Aydın, Al-Saraireh, Mohammad, Amr, Zuhair (2024): Revision of the genus Scorpio in Jordan, with a description of a new genus and three new species (Scorpiones: Scorpionidae). Euscorpius 391: 1-66, DOI: 10.5281/zenodo.14660305
94541A33FFB8FFECABC0ED3FFCB3F824.text	94541A33FFB8FFECABC0ED3FFCB3F824.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scorpio wahbehi Afifeh & Yağmur & Al-Saraireh & Amr 2024	<div><p>Scorpio wahbehi sp. n.</p><p>(Figures 7, 289–358, 359, 365, 373; Table 5) http://zoobank.org/urn:lsid:zoobank.org:act:E7FF2A19-</p><p>6075-41D9-96DA -9F78E1D714B0</p><p>TYPE LOCALITY AND TYPE DEPOSITORY. Jordan, Zarqa Governorate, Wadi Rajel (Wadi Qarma), northeastern desert of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.954197&amp;materialsCitation.latitude=31.795416" title="Search Plazi for locations around (long 36.954197/lat 31.795416)">Jordan</a>, 14 km SE of Al Azraq, 31°47'43.5''N 36°57'15.1''E, 522 m a. s. l. , AZMM.</p><p>TYPE MATERIAL EXAMINED. Jordan, Zarqa Governorate, Wadi Rajel (Wadi Qarma), northeastern desert of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.954197&amp;materialsCitation.latitude=31.795416" title="Search Plazi for locations around (long 36.954197/lat 31.795416)">Jordan</a>, 14 km SE of Al Azraq, 31°47'43.5''N 36°57'15.1''E, 522 m a. s. l., 1♂ (holotype) (AZMM /Sco-2022:12), 1♀ (AZMM / Sco-2022:13) , 2♀, 2 May 2022, leg. B. Abu Afifeh &amp; R. Abu Afifeh; Zarqa Governorate, Wadi Rajel (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.954197&amp;materialsCitation.latitude=31.795416" title="Search Plazi for locations around (long 36.954197/lat 31.795416)">Wadi Qarma</a>), northeastern desert of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.954197&amp;materialsCitation.latitude=31.795416" title="Search Plazi for locations around (long 36.954197/lat 31.795416)">Jordan</a>, 14 km SE of Al Azraq, 31°47'43.5''N 36°57'15.1''E, 522 m a. s. l., 2♂ 3♀, 30 June 2023, leg. B. Abu Afifeh &amp; R. Abu Afifeh (BAPC); Zarqa Governorate, Wadi Rajel (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.954197&amp;materialsCitation.latitude=31.795416" title="Search Plazi for locations around (long 36.954197/lat 31.795416)">Wadi Qarma</a>), northeastern desert of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.954197&amp;materialsCitation.latitude=31.795416" title="Search Plazi for locations around (long 36.954197/lat 31.795416)">Jordan</a>, 14 km SE of Al Azraq, 31°47'43.5''N 36°57'15.1''E, 522 m a.s.l., 3♂ 5♀, 7 July 2023, leg. B. Abu Afifeh &amp; R. Abu Afifeh (BAPC) .</p><p>ETYMOLOGY. Species name is a patronym honoring Dr. Yahya Wahbeh. A Jordanian paediatrician who used to treat children from scorpion stings, he made a study of Jordanian scorpions with a primary checklist in 1976 to help healthcare system in identifying medically important species in Jordan.</p><p>DIAGNOSIS. Medium-big sized species compared with its congeners. Adults are about 61.75–64.95 mm in males, and 60.90–68.79 mm in females. General coloration reddish yellow to olive brown in male, reddish brown to reddish black in females. Carapace is slightly longer than its wide; almost trapezoid in shape. Carapace anterior margin distinctly bilobed with a deep median indentation. Carapace lacks carinae, only posteriomedian carinae slightly visible. Carapace shagreened but a triangular region between median eyes and anterior margin smooth. Median ocular tubercle not distinct and located in the center of the carapace. Tergites I–VII acarinate. Tergites I–VII matt; smooth anteriorly, shagreened posteriorly with scattered medium sized granules on tergites I–IV; tergite VII smooth anteriorly; shagreened posteriorly and covered with moderately dense coarse and pointed granules laterally in males, while tergites I–VII smooth, lustrous in females; posterior of segments III–VI with flattened moderate granules; posterior of tergite VII covered with moderately dense coarse granules laterally. Genital operculum ovoid to subhexagonal in males; anterior margin convex with a small posterior indentation exist, while circular to heart-shaped in females; anterior margin convex; posterior area slightly widely elongated; and a small posterior indentation exist. Pectines short and thin: teeth numbers 12–15 in males and 10–13 in females. Metasoma I–II with 10, III– IV with 8, and V with 5 carinae. Metasoma I slightly wider than long, II usually longer than wide, or sometimes as long as wide, III–V longer than wide. Ventral submedian carinae and ventrolateral carinae strong with moderate, rounded and fused granules on metasom I–II, strong and obsolete granules on III–IV in male; the granules more distinct on segments I–II in females. Ventromedian carina moderate with large, spinoid and pointed granules, bifurcated anteriorly on segment V. Vesicle elongated and smooth; four rows of spinoid granules exist ventrally, more distinct in males. Aculeus long and do not abruptly curved. Two parallel furrows exist on ventral of vesicle. Vesicle with numerous macrosetae reaching the base of aculeus. Pedipalp femur with three carinae, patella with two carinae. The external surface of chela manus with very large and very flattened granules, more distinct and rounded in the anterior area. Ventroexternal and ventrointernal carinae obsolete without granules. Digital carina and external secondary carinae distinct and smooth, the granules fused; and become more dense anteriorly. Chela manus flattened and slightly elongated, longer than depth. Chela fingers are relatively long without granules, longer in females. Fixed and movable fingers with 4–5 strong accessory denticles. Intercarinal surface covered densely with distinct setae on manus and fingers. Trichobothriotaxy of type C; orthobothriotaxic (Vachon, 1974).</p><p>DESCRIPTION (♂ holotype, unless otherwise specified). Coloration (Figs 7, 289–292). General coloration reddish yellow to olive brown in male, reddish brown to reddish black in females. Carapace: olive brown; posterior area of median eyes and anterior area of carapace with brownish, hardly visible reticulation; posterior margin with an olive yellow band in overlap area of first mesosoma segment in male; reddish brown; posterior area of median eyes and anterior area of carapace with hardly visible yellowish brown patches shaped reticulation in females. Tergites I–VII light olive brown; posterior margins of tergites I–VI with olive yellow bands in overlap area in male; I–VI lustrous and reddish black with reticulations medially; tergites I–VII olive brown in females. Coxae, sternum and genital operculum lustrous and dark olive yellow. Pectines light olive yellow. Sternites III–VII olive brown, poststernites of III-VI olive yellow in male; sternites III–VII yellowish olive, poststernites of IIIVI light olive yellow in females. Pedipalp femur and patella dark reddish yellow with carinae and granules light red. Chela manus dark reddish yellow to reddish brown with reddish brown reticulations, granules reddish brown; ventroexternal and ventrointernal reddish brown; digital carina and external secondary carina and fingers dark reddish brown, tip of fingers reddish yellow. Dental margin and condyles reddish black. Metasoma I–II olive brown, III–V yellowish red dorsally and laterally with reddish black reticulations, reddish black dorsally in male; metasoma I–III yellowish red dorsally and laterally, IV–V reddish black dorsally and laterally, metasoma I–II reddish brown, III–V reddish black dorsally in females. Vesicle yellowish red, aculeus dark yellow in posterior part and reddish black at the tip, granules reddish brown in male; ventroposterior area of vesicle reddish brown, granules reddish black in females. Legs with tarsi and basitarsus dark yellow; pretarsus, femur and tibia reddish yellow. Spinules and condyles reddish brown. Chelicera manus dark olive brown with reticulations posteriorly, brownish black anteriorly. Chelicera fingers brownish black, tip of fingers and teeth dark yellow, tip of teeth reddish black.</p><p>Carapace (Figs. 301–308). Carapace is slightly longer than its wide; almost trapezoid in shape. Carapace anterior margin distinctly bilobed with 8 medium to large setae and a deep median indentation. Carapace lacks carinae, only posteriomedian carinae slightly visible. Carapace shagreened but a triangular region between median eyes and anterior margin smooth, along to anteriomedian furrow with several small granules, lateral and posterior surface with scattered small granules in male; carapace smooth and lustrous with several small granules posteriorly in females. Median ocular tubercle not distinct and located in the center of the carapace. A pair of median eyes and three pairs of lateral eyes exist; the third located separately and slightly smaller. Median eyes separated by one ocular diameter. The anteromedian furrow is narrow and distinct. The posteromedian furrow is wide and deep and bifurcated as an inverted T-shaped at posterior edge and located in arrow-shaped depressions between posteriomedian carinae and posterior margin. Posterior lateral furrows wide and distinct. A few setae exist on the anterior area of carapace.</p><p>Chelicerae (Figs. 301–308). Cheliceral dentition characteristic for the family of Scorpionidae (Vachon, 1963); subdistal tooth and basal teeth conspicuous rather than on the movable finger. Manus lustrous, no longitudinal ridges present anteriorly.</p><p>Mesosoma (Figs. 293–300). Tergites I–VII acarinate, matt; smooth anteriorly; shagreened posteriorly with scattered medium sized granules on segment I–IV; tergite VII smooth anteriorly; shagreened posteriorly and covered with moderately dense coarse and pointed granules laterally in males, while tergites I–VII smooth, lustrous in females; posterior of segments III–VI with flattened moderate granules; posterior of tergite VII covered with moderately dense coarse granules laterally. Sternites III–VI with scattered setae, smooth and wrinkled in males; smooth and lustrous in females. Sternite VII with four granular carinae; intercarinal surfaces with scattered, small to medium granules anteriorly. Genital operculum ovoid to subhexagonal in males, anterior margin convex with a small posterior indentation exist, while circular to heart-shaped in females; anterior margin convex; posterior area widely elongated; and a small posterior indentation exist. Pectines short and thin: teeth numbers 12–15 in males and 10–13 in females; three marginal and seven or eight median lamellae. Stigmas linear, angled 45° and conspicuous.</p><p>Metasoma and telson (Figs. 333–352). Metasoma I–II with 10, III–IV with 8, and V with 5 carinae. Segment I slightly wider than long, II usually longer than wide, or sometimes as long as wide, segments III–V longer than wide. Lateral inframedian carinae on segment I indistinct, complete, composed of a few spaced rounded granules; on segment II composed 1–2 granules on posterior quarter. Ventral submedian carinae and ventrolateral carinae strong with moderate, rounded and fused granules on segments I–II, strong and obsolete granules on segments III–IV in male; the granules more distinct on segments I–II in females. Dorsolateral carinae strong and crenulated with small, spaced and spinoid granules on segments I–IV. Lateral supramedian carinae strong and granular on segments I–IV. Intercarinal area smooth, finely granular on lateral surfaces of segments I–IV in male, smooth in females. Dorsal surface moderately granular with fine or moderate granules, medially smooth in male, completely smooth in females on segments I–IV. Segment V: Ventromedian carina moderate with large, spinoid and pointed granules, bifurcated anteriorly; ventrolateral carinae strong and serrated with spaced large and conical granules, gradually increase posteriorly, continues in lateral of anal arch; dorsolateral carinae moderate and rounded with small spinoid granules, smooth in females; segment V with a row of granules laterally at posterior half, granules less distinct in females. Lateral surfaces smooth with scattered few fine granules in male. Ventral surface smooth with few large granules. Dorsal surface moderately granular with fine or moderate granules, medially smooth in male, completely smooth in females. Ventral of anal arch serrated with pointed denticles. Ventral and lateral surfaces covered with moderately variable sized setae in segments I-V, more dense and larger on segments IV–V. Vesicle elongated and smooth, four rows of spinoid granules exist ventrally, more distinct in males. Aculeus long and do not abruptly curved. Two parallel furrows exist on ventral of vesicle. Vesicles with numerous macrosetae reaching the base of aculeus.</p><p>Pedipalp (Figs. 309–332). Pedipalp femur with three carinae; dorsointernal and ventrointernal carinae moderate with spaced, rounded and conical granules; dorsoexternal carinae strong with separate large and rounded granules. Intercarinal area smooth with small sized granules dorsally and medium sized granules ventrally. Intercarinal surface with a few distinct setae. Patella with two carinae; dorsomedian carina strong with so flattened and fused granules; ventrointernal carinae moderate and obsolete. Intercarinal area of dorsal, ventral and external surfaces smooth and lustrous; internal surface shagreened. Intercarinal surface with a few distinct setae. The external surface of chela manus with very large and very flattened granules, more distinct and rounded in anterior area, posterior part with smaller granules; posterior internal area with less distinct or more flattened granules in male; the granules more flattened and nearly smooth in posterior internal area in females. Posterior margin of external surface and internal margin with large and conical granules in male, females lack these granules. Internal surface of chela manus smooth with moderate pointed granules in middle and anterior area in male, fewer granules in females. Ventroexternal and ventrointernal carinae obsolete without granules. Digital carina and external secondary carinae distinct and smooth, the granules fused; become denser anteriorly. Chela manus flattened and slightly elongated, longer than depth. Chela fingers are relatively long without granules, longer in females. Fixed and movable fingers with 4–5 strong accessory denticles. Intercarinal surface covered densely with distinct setae on manus and fingers. Trichobothriotaxy of type C; orthobothriotaxic (Vachon, 1974).</p><p>Legs (Figs. 353–354). Coxa and trochanters smooth. Tarsi of right legs I to IV with 4-5/7-8–5/8–6/9-10–6-7/9 internal and external spines arranged in series.</p><p>Measurements. See Table 5.</p><p>AFFINITIES. Scorpio wahbehi sp. n. can be distinguished from Scorpio kruglovi Birula, 1910 by a combination of the following characters (more details in Table 6); a) pedipalp chela hand is more elongated in S. wahbehi sp. n.; b) fixed finger of pedipalp chela is less elongated with respect to chela length in S. wahbehi sp. n.; c) external surface of pedipalp chela with flattened granules in females of S. wahbehi sp. n., whereas it is smooth in females of S. kruglovi; d) aculeus of S. kruglovi is longer and less curved than S. wahbehi sp. n., e) genital operculum is circular of S. wahbehi sp. n., whereas it is cordate in S. kruglovi .</p><p>S. wahbehi sp. n. can be distinguished from S. jordanensis sp. n. by several characters shown in Table 6: a) pedipalp chela hand is more elongated in S. wahbehi sp. n. than in S. jordanensis sp. n.; b) fixed finger of pedipalp chela is more elongated in S. wahbehi sp. n. than in S. jordanensis sp. n.; c) the average pectinal tooth numbers in S. wahbehi sp. n. is higher than S. jordanensis sp. n.; d) metasomal segment II is longer than wide or as long as wide in S. wahbehi sp. n., whereas it is always wider than long in S. jordanensis sp. n.; e) S. wahbehi sp. n. is larger in size (adults 64-68 mm), than S. jordanensis sp. n. (adults 53–56 mm); f) the basal lamellae (marginal and median) of pectines is more elongated and narrower in females of S. wahbehi sp. n. than females of S. jordanensis sp. n.; the basal median lamellae is narrowly triangular in shape in S. wahbehi sp. n. (Fig. 308), whereas it is broadly triangular in S. jordanensis sp. n. (Fig. 164); g) the external surface of pedipalp chela mostly with flattened and less distinct granules in S. wahbehi sp. n. (Figs. 316, 319), whereas it is rounded and more distinct in S. jordanensis sp. n. (Figs. 171, 175); h) general coloration is reddish yellow to olive brown in male, reddish brown to reddish black in females of S. wahbehi sp. n., whereas it is light reddish yellow in S. jordanensis sp. n.</p><p>S. wahbehi sp. n. can be distinguished from S. fuscus by several characters shown in Table 6: a) general coloration reddish yellow to olive brown in male, reddish brown to reddish black in females of S. wahbehi sp. n. (Figs. 7, 289– 292), whereas it is dark reddish brown to brownish black in S. fuscus (Figs. 3–4, 79–82); b) pedipalp chela hand is more elongated in S.wahbehi sp. n. (Figs. 309–310) than in S. fuscus (Figs. 99–100); c) the basal lamellae (marginal and median) of pectines is more elongated and narrower in females of S. wahbehi sp. n. than in females of S. fuscus; the basal median lamellae is narrowly triangular in shape in S. wahbehi sp. n. (Fig. 308), whereas it is broadly triangular in S. fuscus (Fig. 98); d) pectines is longer in S. wahbehi sp. n. than in S. fuscus; pectines extending well beyond coxa-trochanter articulation of leg IV in S. wahbehi sp. n. (Figs. 307–308), whereas extending approximately to the coxa-trochanter articulation of leg IV in</p></div>	https://treatment.plazi.org/id/94541A33FFB8FFECABC0ED3FFCB3F824	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Afifeh, Bassam Abu;Yağmur, Ersen Aydın;Al-Saraireh, Mohammad;Amr, Zuhair	Afifeh, Bassam Abu, Yağmur, Ersen Aydın, Al-Saraireh, Mohammad, Amr, Zuhair (2024): Revision of the genus Scorpio in Jordan, with a description of a new genus and three new species (Scorpiones: Scorpionidae). Euscorpius 391: 1-66, DOI: 10.5281/zenodo.14660305
94541A33FFB5FFEFAB1FEDB5FEDCF800.text	94541A33FFB5FFEFAB1FEDB5FEDCF800.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scorpio fuscus Linne 1758	<div><p>S. fuscus (Figs. 97–98); e) external surface of pedipalp chela with flattened granules in S. wahbehi sp. n., whereas S. fuscus has reticular and flattened granulation; f) the average pectinal teeth numbers in S. wahbehi sp. n. is higher than in S. fuscus .</p><p>Sexual dimorphism</p><p>In both genera Scorpio and Jordanius gen. n., males differed from females as follows: a) tergites I–VI matt, completely covered with fine granules in males, but glossy and smooth in females, Sternites III–VI wrinkled medially in males, but smooth and shining in females. b) fixed finger of pedipalp chela in adult females longer than in males; fixed finger length to chela length ratios is bigger in females than in males; in Jordanius maysaraensis gen. et sp. n. 0.36–0.39 with average 0.38 in males (n=9); 0.41–0.43 with average 0.42 in females (n=12); in S. wahbehi sp. n. 0.38–0.41 with average 0.40 in males (n=6); 0.42–0.44 with average 0.43 in females (n=10); in S. jordanensis sp. n. 0.35–0.36 with average 0.35 in males (n=9); 0.39–0.42 with average 0.40 in females (n=17). c) genital operculum generally ovoidal with minor variations in males (Figs. 359– 364), whereas it is generally heart-shaped with minor variation in females (Figs. 365–370). d) dentate margin length of pecten to pecten length is longer in males than in females; dentate margin length to pecten length ratio in Jordanius maysaraensis gen. et sp. n. 0.73–0.79 with average 0.77 in males (n=12); 0.65–0.72 with average 0.69 in females (n=12); in S. wahbehi sp. n. 0.70– 0.75 with average 0.72 in males (n=5); 0.60–0.68 with average 0.64 in females (n=10); in S. jordanensis sp. n. 0.73 in the male; 0.66 in the female. e) the average pectinal teeth numbers in males is higher than in females; in Jordanius maysaraensis gen. et sp. n. 12.1 in males (30 combs, n=15), 11.0 in females (58 combs, n=29); in S. wahbehi sp. n. 12.6 in males (12 combs, n=6), 11.0 in females (22 combs, n=11); in S. jordanensis sp. n. 11.4 in males (46 combs, n=23), 10.6 in females (36 combs, n=18). f) tergite VII finely granular with moderate, pointed granules posteriorly and laterally in males (Figs. 17, 87, 153, 297), but smooth anteriorly, granulate posteriorly and laterally in females (Figs. 18, 88, 154, 298).</p></div>	https://treatment.plazi.org/id/94541A33FFB5FFEFAB1FEDB5FEDCF800	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Afifeh, Bassam Abu;Yağmur, Ersen Aydın;Al-Saraireh, Mohammad;Amr, Zuhair	Afifeh, Bassam Abu, Yağmur, Ersen Aydın, Al-Saraireh, Mohammad, Amr, Zuhair (2024): Revision of the genus Scorpio in Jordan, with a description of a new genus and three new species (Scorpiones: Scorpionidae). Euscorpius 391: 1-66, DOI: 10.5281/zenodo.14660305
94541A33FFC9FF90A8E4E9E0FCB3F9D7.text	94541A33FFC9FF90A8E4E9E0FCB3F9D7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scorpionidae (Vachon 1963)	<div><p>Key to Family Scorpionidae in Jordan</p><p>1. External surface of chela manus densely covered with large, raised, rounded and somewhat pointed granules, the posterior margin of the external surface of chela manus bears very large spinoid protrudings, chela fingers granulated with small, pointed, gradually decreased granules anteriorly, Chela manus elongated (Manus Length/Chela Depth (1.24–1.29) (Fig. 1). ........................ .............................................. Jordanius gen. n. ……… 2</p><p>– External surface of chela manus is almost smooth, or covered with isolated, rounded, or flattened, and sometimes with reticulated fused granules, chela fingers smooth, chela manus inflated (Manus Length/Chela Depth (1.0–1.18) (Fig. 2). .. Scorpio Linnaeus, 1758 … 3</p><p>2. Coloration generally dark yellow to light yellowish brown, external surface of chela manus with moderately dense and generally small granulation. ............................ .............................. Jordanius maysaraensis gen. et sp. n.</p><p>– Coloration generally reddish brown, external surface of chela manus with dense, large and somewhat pointed granulation ............... Jordanius granulomanus, comb. n.</p><p>3. Metasomal segment II usually longer than wide, or sometimes as long as wide, generally dark coloration, adult total length&gt; 60 mm. ............................................ 4</p><p>– Metasomal segment II always wider than long, generally light coloration, adult total length &lt;55mm. ..................... ................................................ Scorpio jordanensis sp. n.</p><p>4. The basal median lamellae of pectines is narrowly triangular in shape, pectines extending well beyond coxa-trochanter articulation of leg IV, general coloration reddish yellow to olive brown in male, reddish brown to reddish black in females. ............ Scorpio wahbehi sp. n.</p><p>– The basal median lamellae of pectines are broadly triangular in shape, pectines extending approximately to the coxa-trochanter articulation of leg IV, general coloration dark reddish brown to brownish black. ........... .................................... Scorpio fuscus (Ehrenberg, 1829)</p></div>	https://treatment.plazi.org/id/94541A33FFC9FF90A8E4E9E0FCB3F9D7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Afifeh, Bassam Abu;Yağmur, Ersen Aydın;Al-Saraireh, Mohammad;Amr, Zuhair	Afifeh, Bassam Abu, Yağmur, Ersen Aydın, Al-Saraireh, Mohammad, Amr, Zuhair (2024): Revision of the genus Scorpio in Jordan, with a description of a new genus and three new species (Scorpiones: Scorpionidae). Euscorpius 391: 1-66, DOI: 10.5281/zenodo.14660305
