taxonID	type	description	language	source
78E8D3AE40CD5B96A44FFBA1906E2361.taxon	description	Figs 4 - 7, 8 - 15	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
78E8D3AE40CD5B96A44FFBA1906E2361.taxon	diagnosis	Diagnosis. Representative of subtribe Trechina due to presence of bidentate mandibles (absence of retinacle) and dorsally closed aedeagal median lobe (Jeannel 1926). Comparatively large trechines characterized by robust head, large mandibles, small but markedly protruded eyes, smooth, markedly convex tempora, cordiform pronotum, straight pronotal basal margin with large, rectangular to acute laterobasal angles, slender elytra, short metepisternum, reduced hindwings, moderately slender antenna and legs, protibia with a complete longitudinal groove on external surface, presence of a row of long adhesive hairs on apical margins of 4 th tarsomeres which are as long as the 5 th tarsomeres, and by a markedly slender median lobe of aedeagus, which possesses a small, slightly sclerotized endophallic copulatory piece. The new genus is particularly distinguished from other representatives of Trechina by unusual chaetotaxy of head capsule and elytra as follows: clypeus plurisetose, each side with 3 - 5 setae; submentum with three setae each side; anterior elytral discal seta located in the 4 th interval, adjoined to the 4 th stria.	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
78E8D3AE40CD5B96A44FFBA1906E2361.taxon	etymology	Etymology. The new genus name is given in memoriam of our dear friend and colleague, the distinguished entomologist Martin Baehr, Munich (10.03.1943 - 17.04.2019).	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
78E8D3AE40CD5B96A44FFBA1906E2361.taxon	description	Description. Head: Large and robust, without pilosity. Mandibles large, moderately slender, with bidentate dentition pattern as shown in Fig. 8. Labrum with apical margin moderately emarginated, with six setae near apical margin. Clypeus each side with three or four setae (Figs 5, 7; seldom only two setae at one of the sides). Eyes moderately small, as long as or slightly shorter than tempora, markedly convexly protruded (Figs 5, 7). Two supraorbital setae each side in normal position for Trechina. Supraorbital furrows unevenly bent in posterior half, markedly deep in front and middle portions, slightly flatter near insertion of posterior supraorbital seta. Tempora markedly convex, markedly wrinkled to the neck, smooth. Mid of head convexly elevated, with a distinct transverse depression between supraorbital area and neck (Figs 5, 7). Antennae slender, with third antennomere longest, 1 / 9 - 1 / 10 longer than first respectively fourth antennomere, and with second antennomere about 2 / 3 of length of third. Suborbital seta present. Apical tooth of mentum bifid, sensory pits of mentum present; submentum with three setae each side (Fig. 9). Prothorax: Pronotum rather small, without pilosity, moderately transverse, cordate, broadest distinctly before middle, with lateral margin markedly concave before base, and with basal margin slightly smaller than apical margin (Figs 4, 6). Disc moderately convex. Anterior margin straight or slightly concave in middle with anterior angles small but distinct, rounded, moderately protruded. Basal margin straight along internal 3 / 4, with laterobasal angles slightly shifted posteriad. Lateral margin convexly rounded in anterior 2 / 3 and concave towards laterobasal angles, latter large, rectangular or sharp at tip, sometimes slightly protruded laterally. Marginal gutter moderately broad throughout. Median longitudinal impression sharply incised, disappearing near apex, somewhat deepened before base. Anterior and posterior transverse impressions shallow and smooth. Laterobasal foveae large, internally and externally (towards lateral gutter) distinctly sloped, without punctures but with fine transversal wrinkles. Lateral and laterobasal setae present, with the former situated at or slightly anterad of maximum width of pronotum. Proepisternum glabrous and smooth. Pterothorax: Elytra without pilosity, long and very slender ovate, very slightly convex or flattened in middle of disc, in dorsal view broadest distinctly posterad middle, shoulders flatly rounded (Fig. 13), apical sinuation distinct, apex rounded with the indication of a very obtuse apical angle. Striae 1 - 8 complete, moderately deep impressed, impunctate, intervals moderately convex, parascutellar stria free, 1 / 6 - 1 / 9 of length of elytra. Recurrent preapical stria deep, long, in most specimens reaching the apex of the fifth stria. Parascutellar seta present. Anterior discal seta located in the 4 th interval, adjoined to the 4 th stria, located near the end of the anterior elytral 5 th (Fig. 13); second discal seta located at the 3 rd stria about at elytral middle (in most specimens, the posterior setiferous pore together with the 3 rd stria is switched into the 4 th interval); posterior discal seta (= subapical seta near the end of 3 rd stria) present, located about 1 / 9 of elytral length from elytral apex; subapical seta of the recurrent stria isolated, distinctly removed from this stria by distance of 2 - 3 diameters of the setiferous pore. Number and positions of the setae of the marginal umbilicate series as in Trechus s. str. Metepisternum very short, glabrous and smooth, with outer margin about as long as anterior margin. Legs: Moderately long and robust. Protibia distinctly dilated towards apex, straight, with longitudinal groove on dorsal surface complete, and with several fine setae on anterior surface near apex (Fig. 12). Two basal protarsomeres of males dilated and dentoid at the inner apical border (Fig. 10). Fourth pro-, meso- and metatarsomeres each with a row of long adhesive hairs on apical margins which are as long as the 5 th tarsomeres (Figs 10, 11). Male genitalia (Figs 14, 15): Aedeagal median lobe markedly elongated, slender tube-like, in lateral view slightly sinusoidal, with apex distinctly bent upwardly, simple, with apical lamella insignificant; basal bulb rather small with large sagittal aileron. Endophallus with a small, very slightly sclerotized copulatory piece. Parameres with 3 - 4 apical setae.	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
78E8D3AE40CD5B96A44FFBA1906E2361.taxon	distribution	Distribution. So far only known from Mt. Choke in northern Ethiopia (Fig. 1).	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
8DD625A074ED516595A21EB50BE6BB65.taxon	description	Figs 4 - 7, 8 - 15	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
8DD625A074ED516595A21EB50BE6BB65.taxon	materials_examined	Additional material. 3 males, 3 females, W-slope Mt. Choke, alt. 3370 m, 10 ° 38 ' 07 " N, 37 ° 45 ' 51 " E, 23. II. 2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek W. (CAF, CSCHM); 6 males, 7 females, W-slope Mt. Choke, alt. 3700 - 3900 m, 10 ° 42 ' 17 " N, 37 ° 50 ' 29 " E, 25. II. 2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek W. (CAF, CSCHM); 43 males, 31 females, W-slope Mt. Choke, " Shoa Kidaneberet " valley, alt. 3700 - 3800 m, 10 ° 39 ' 08 " N, 37 ° 49 ' 45 " E, 8. V. 2022, leg. J. Schmidt, Yeshitla M. (CSCHM); 10 males, 10 females, N-slope Mt. Choke, alt. 3800 - 3950 m, 10 ° 43 ' 16 " N, 37 ° 51 ' 15 " E, 26. II. 2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek W. (CSCHM); 18 males, 20 females, N-slope Mt. Choke, alt. 3750 - 3850 m, 10 ° 43 ' 51 " N, 37 ° 52 ' 15 " E, 09. V. 2022, leg. J. Schmidt, Yeshitla M. (CSCHM); 3 males, 2 females, N-slope Mt. Choke, above Gumadur, alt. 3750 - 3850 m, 10 ° 44 ' 10 " N, 37 ° 53 ' 48 " E, 05. V. 2022, leg. J. Schmidt, Yeshitla M. (CSCHM); 1 male, N-slope Mt. Choke, N of Waber, alt. 3450 - 3600 m, 10 ° 44 ' 48 " N, 37 ° 46 ' 22 " E, 07. V. 2022, leg. J. Schmidt, Yeshitla M. (CSCHM); 13 males, 5 females, Mt. Choke, eastern crater valley, alt. 3700 - 3800 m, 10 ° 42 ' 59 " N, 37 ° 54 ' 13 " E, 06. V. 2022, leg. J. Schmidt, Yeshitla M. (CSCHM).	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
8DD625A074ED516595A21EB50BE6BB65.taxon	etymology	Etymology. The specific epithet refers to the markedly separated distributional area of the taxon, which is, based on current knowledge, far away from its next relatives. It is built by the past participle of the Latin verb separare.	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
8DD625A074ED516595A21EB50BE6BB65.taxon	description	Description. See description of genus. Body length: 6.9 - 7.5 mm (O = 7.19 mm, n = 20). Proportions (n = 20): PW / HW = 1.18 - 1.25 (O = 1.21); PW / PL = 1.30 - 1.38 (O = 1.34); PW / PBW = 1.48 - 1.54 (O = 1.50); PBW / PAW = 0.92 - 1.00 (O = 0.96); EW / PW = 1.42 - 1.48 (O = 1.45); EL / EW = 1.46 - 1.56 (O = 1.51). Colour: Dark brown to blackish, moderately shiny in both sexes; palpi light brown, labrum and scapus reddish brown, basal 3 / 4 of femora light brown; antennal base in some specimens more widely brightened. Microsculpture: Same in males and females. Head with deeply engraved, rather large, almost isodiametric sculpticells on disc and supraorbital area, slightly smaller sculpticells on clypeus. Pronotum with moderately deep engraved, slightly transverse sculpticells on disc and markedly deep engraved sculpticells near base; the sculpticells are somewhat smaller than on head disc. Elytral intervals with more finely engraved sculpticells which are more transverse than on pronotum. Aedeagus. Proportion EL / AL (n = 10): 2.40 - 2.64 (O = 2.52). Median lobe in lateral view unevenly bent, dorsally with a distinct concavity before middle (Fig. 14), in dorsal view not or very slightly broadened before apex (Fig. 15).	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
8DD625A074ED516595A21EB50BE6BB65.taxon	distribution	Distribution and geographical variability. The type series was collected on the western side of the crater valley of Mt. Choke. Additional populations were collected on the north eastern side of the crater valley and on northern and western slopes of Mt. Choke. Specimens of these populations differ + / - distinctly from those of the type series and from each other by the curvature of the aedeagal median lobe, the number of lightened basal antennomeres, and the depth of the engraving of the elytral microsculpture. Slight differences were also found in the DNA sequence segments of the three investigated specimens representing three different populations (Fig. 2; Suppl. material 1). Further morphological and molecular genetic studies are needed to answer the question of whether certain populations represent separate species or subspecies.	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
23F4CEDA79AC5113BA0F78816C8FFDA3.taxon	description	Figs 17, 18, 23, 24 - 25	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
23F4CEDA79AC5113BA0F78816C8FFDA3.taxon	materials_examined	Additional material. 6 males, 9 females, Ethiopia, Amhara, Mt. Abuna Yosef, N-slope, 3800 - 3950 m, 12 ° 07 ' 52 " N, 39 ° 11 ' 39 " E, 4. III. 2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek W. (CAF, CSCHM); 6 males, 12 females, ditto, S-slope, 3850 - 3900 m, 12 ° 07 ' 29 " N, 39 ° 11 ' 21 " E, 5. III. 2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek W. (CSCHM, NHMAA); 5 males, 2 females, ditto, S-slope, 3700 - 3850 m, 12 ° 09 ' 10 " N, 39 ° 09 ' 35 " E, 5. III. 2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek W. (CSCHM).	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
23F4CEDA79AC5113BA0F78816C8FFDA3.taxon	description	Additions to the species description. Mature species with elytra including suture blackish brown (Queinnec et al. 2021: " suture distinctly brown shiny "; this information is probably based on confusion with T. habeshanicus). Body length 4.1 - 4.7 mm (Queinnec et al. 2021: "> 4.5 mm "). Length of aedeagus 0.98 - 1.05 mm. PW / PL = 1.29 - 1.37; n = 20 (Queinnec et al. 2021: PW / PL = 1.6; this value is very probably based on a measurement error). PW / TL = 1.06 - 1.15; n = 20. EL / AL = 2.15 - 2.46; n = 10.	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
23F4CEDA79AC5113BA0F78816C8FFDA3.taxon	distribution	Distribution. Endemic to the Abune Yosef Mountains of the northern Ethiopian Highlands.	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
F06932799AAE5261B94FFF356556DA17.taxon	description	Figs 21, 22, 28, 29	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
F06932799AAE5261B94FFF356556DA17.taxon	materials_examined	Additional material. 98 exx. (males, females), Ethiopia, Amhara, northern Guassa Plateau, near Guassa Comm. Lodge 3330 m, 10 ° 17 ' 17 " N, 39 ° 47 ' 54 " E, 18. V. 2022, leg. J. Schmidt, Yeshitla M. (CAF, CSCHM, NHMAA); 360 exx. (males, females), ditto, " Aste wuha " 3400 m, 10 ° 24 ' N, 39 ° 48 ' E, 19. V. 2022, leg. J. Schmidt, Yeshitla M. (CAF, CSCHM, NHMAA); 11 exx. (males, females), ditto, " Yegana Natural Forest ", river valley, 3125 m, 10 ° 26 ' 03 " N, 39 ° 47 ' 16 " E, 20. V. 2022, leg. J. Schmidt, Yeshitla M. (CSCHM).	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
F06932799AAE5261B94FFF356556DA17.taxon	description	Additions to the species description. Mature species with elytra blackish brown, and with suture and first interval reddish brown lightened in most specimens (Queinnec et al. 2021: " suture distinctly darkened "; this information is probably based on confusion with T. bipartitus). Body length 4.1 - 5.0 mm (Queinnec et al. 2021: 4.1 - 4.3 mm). Length of aedeagus 0.70 - 0.80 mm. PW / PL = 1.28 - 1.38; n = 20. PW / TL = 1.08 - 1.13; n = 20. EL / AL = 3.00 - 3.33; n = 10.	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
F06932799AAE5261B94FFF356556DA17.taxon	distribution	Distribution. Endemic to the Guassa Plateau of the northern Ethiopian Highlands.	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
712B21D4391855B8ACF80C23B31953D9.taxon	description	Figs 19, 20, 26, 27	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
712B21D4391855B8ACF80C23B31953D9.taxon	materials_examined	Additional material. 13 males, 11 females, Ethiopia, Amhara, Mt. Abuna Yosef, N-slope, 3800 - 3950 m, 12 ° 07 ' 52 " N, 39 ° 11 ' 39 " E, 4. III. 2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek W. (SCHM, NHMAA); 1 male, 1 female, ditto, S-slope, 3850 - 3900 m, 12 ° 07 ' 29 " N, 39 ° 11 ' 21 " E, 5. III. 2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek W. (CSCHM); 1 male, ditto, S-slope, 3700 - 3850 m, 12 ° 09 ' 10 " N, 39 ° 09 ' 35 " E, 5. III. 2019, leg. D. Hauth, J. Schmidt, Yeshitla M., Yitbarek W. (CSCHM).	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
712B21D4391855B8ACF80C23B31953D9.taxon	description	Additions to the species description. Body length 4.2 - 5.1 mm (Queinnec et al. 2021: 4.8 - 5.1 mm). Length of aedeagus 0.98 - 1.02 mm. PW / PL = 1.24 - 1.30; n = 20. PW / TL = 0.99 - 1.04; n = 20. EL / AL = 2.39 - 2.65; n = 10.	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
712B21D4391855B8ACF80C23B31953D9.taxon	distribution	Distribution. Endemic to the Abune Yosef mountains of the northern Ethiopian Highlands.	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
4942403632AC58E2ADAF31A673E77674.taxon	diagnosis	Diagnosis. Representative of Trechina and Trechus s. l. sensu Jeannel (1926, 1927, 1928) due to presence of bidentate mandibles (absence of retinacle), dorsally closed aedeagal median lobe, well-developed compound eyes, protibia glabrous on anterior surface, presence of two elytral dorsal setae situated in third interval, elytral intervals glabrous, 4 + 2 + 2 pattern of umbilicate setae, and two basal tarsomeres of male dilated. Externally, Trechus Abunetrechus subgen. nov. reminders a non-specialised high-altitude Trechus of moderate body size, short mandibles, antenna and legs, moderately large eyes, rounded humeri, short metepisternae and hindwings reduced to short stubs. Trechus sensu lato, Trechus Abunetrechus subgen. nov. is characterized by the combination of following character states: bisetose clypeus (Figs 18, 20, 22), smooth tempora, pronotum with fully rounded laterobasal angles and with laterobasal setae markedly protruded anteriorly (Figs 18, 20, 21), elytral striae 3 and 4 merging at level of the anterior discal seta (Fig. 23), elytral preapical seta of the third interval present and situated about at level of the elytral apical tenth, protibia with longitudinal groove on external surface; aedeagus with two moderately sclerotized portions of the endophallus arranged one behind the other in apical half of the median lobe (Figs 24 - 29).	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
4942403632AC58E2ADAF31A673E77674.taxon	etymology	Etymology. The subgenus name combines the name of the Abune Yosef Massif in northern Ethiopia, where the species of this subgenus occur, with the name of the genus Trechus.	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
4942403632AC58E2ADAF31A673E77674.taxon	description	Description. Head: Size averaged for Trechus sensu lato, without pilosity. Mandibles short, with dentition pattern as in Trechus sensu stricto. Labrum with apical margin moderately emarginated, with six setae near apical margin. Clypeus each side with one long seta (Figs 18, 20, 22; very seldom with an additional very fine seta situated interior of one of the primary setae). Eyes moderately large, convexly protruded, more than two times as long as tempora, latter moderately convex (Figs 18, 20, 22). Two supraorbital setae each side in normal position for Trechus. Supraorbital furrows moderately deep and almost evenly bent throughout. Tempora moderately convex, markedly wrinkled to the neck, smooth. Mid of head convexly elevated. Antennae short, with third antennomere slightly longer than pedicellus. Suborbital seta present. Apical tooth of mentum truncate or slightly bifid, sensory pits of mentum present but very small; submentum with 4 - 7 setae. Prothorax: Pronotum with size averaged for Trechus sensu lato, without pilosity, slightly transverse, broadest distinctly before middle, with lateral margin completely rounded towards base, and with laterobasal angles indistinct. Basal margin (between insertion points of laterobasal setae) distinctly broader than apical margin. Disc markedly convex. Anterior margin slightly or moderately concave with anterior angles shortly rounded, moderately protruded. Basal margin straight or slightly convex in middle and with outer quarters markedly shifted anteriorly towards lateral margin (Figs 18, 20, 21). Lateral margin convexly rounded throughout; laterobasal angle fully rounded or marked as a very small blunt tooth. Marginal gutter very narrow throughout. Median longitudinal impression slightly incised, disappearing near apex and base, not deepened within area of posterior transverse impression. Anterior and posterior transverse impressions very shallow, smooth or (posterior transverse impression) sometimes finely wrinkled. Laterobasal foveae rather small, moderately impressed, smooth. Lateral and laterobasal setae present, with the former situated near maximum width of pronotum. Proepisternum glabrous and smooth. Pterothorax: Elytra without pilosity, slender ovate, markedly domed towards disc, not flattened in middle of disc, in dorsal view broadest slightly posterad middle, shoulders flatly rounded, apical sinuation very slightly developed or indistinct, apex rounded or marked as an obtuse apical angle. Parascutellary stria short to moderately long, free; striae 1 - 8 almost complete, moderately deep impressed in middle of disc, less deeply towards sides, disappearing near base, crenulated, striae 3 and 4 merging at level of the anterior discal seta; intervals slightly convex. Recurrent preapical stria deep, long, connected with the apex of the fifth stria. Parascutellar seta present. Anterior discal seta situated at merging point of the 3 rd and 4 th stria, near the end of the anterior elytral 5 th (Fig. 23); second discal seta located at the 3 rd stria somewhat behind elytral middle; posterior discal seta (= subapical seta near end of 3 rd stria) present, located about 1 / 10 of elytral length from elytral apex; subapical seta of the recurrent stria isolated, removed from this stria by distance of 1 - 2 diameters of the setiferous pore. Number and positions of the setae of the marginal umbilicate series as in Trechus s. str. Metepisternum very short, glabrous and smooth, with outer margin about as long as anterior margin. Legs: Short and moderately robust. Protibia distinctly dilated towards apex, straight, glabrous, with longitudinal groove on dorsal surface complete. Two basal protarsomeres of males dilated and dentoid at the inner apical border. Chaetotaxy as in Trechus sensu stricto. Male genitalia (Figs 24 - 29): Aedeagal median lobe moderately large, in lateral view markedly curved, with apical lamella short, latter with distinct terminal capitulum; basal bulb and saggital aileron averaged. Endophallus with a moderately large, moderately sclerotized folding structure (copulatory piece) in the shape of a half-open cylinder or cone which is located in apical half of the median lobe and directed to its longitudinal axis, and with the open part of the copulatory piece facing ventrad. Apicad of this piece, an additional slightly more strongly sclerotized folding structure is developed which is shaped as a small plate (best visible in lateral view); the basal part of this piece overlaps with the apex of the more dorsal copulatory piece. Parameres with 2 - 4 apical setae.	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
4942403632AC58E2ADAF31A673E77674.taxon	distribution	Distribution. Northern Ethiopia Plateau (Fig. 1): Three species are known so far, two from Mt. Abune Yosef (T. bipartitus Raffray, T. lalibelae Queinnec & Ollivier) and one from the Guassa Plateau (T. habeshaicus Queinnec & Ollivier).	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
AD70353F6B8F5F479457FE03AE94B251.taxon	description	Figs 30 - 35	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
AD70353F6B8F5F479457FE03AE94B251.taxon	materials_examined	Additional material. For comprehensive material studied see our previous paper (Schmidt and Faille 2018). Note that in this study, T. oromiensis Magrini, Queinnec & Vigna Taglianti was erroneously treated as a distinct species. In the meantime, the following additional material was available for us: Ethiopia, Oromia, SE-slope of Mt. Chillalo, Dhaba village, alt. 3200 m, 19. II. 2020, 7.861644 ° N, 39.27711 ° E, leg. J. Schmidt, C. Wirkner, Yeshitla M. (1 female: CSCHM); ditto, Bale Mts., Web river N Dinsho, alt. 3000 m, 5. II. 2019, 07 ° 07 ' 18 " N, 39 ° 46 ' 03 " E, leg. R. Emmerich, J. Schmidt, Yeshitla M. (12 specimens: CSCHM); ditto, Bale Mts., forest remain W Dinsho, alt. 3100 m, 8. II. 2019, 07 ° 06 ' 16 " N, 39 ° 44 ' 46 " E, leg. R. Emmerich, J. Schmidt, Yeshitla M. (12 specimens: CSCHM); ditto, Bale Mts., Sebsebe Washia Forest, Salgen Valley, alt. 2720 - 2800 m, 3. II. 2019, 07 ° 02 ' 08 " N, 39 ° 36 ' 06 " E, leg. R. Emmerich, J. Schmidt, Yeshitla M. (7 specimens: CSCHM); ditto, Bale Mts., Sebsebe Washia Forest, Salgen Valley, alt. 3130 m, 4. II. 2019, 07 ° 02 ' 08 " N, 39 ° 36 ' 06 " E, leg. R. Emmerich, J. Schmidt, Yeshitla M. (30 specimens: CSCHM); ditto, Bale Mts., Angeso Valley S Goba, alt. 3050 m, 5. II. 2020, 6.932923 ° N, 39.951341 ° E, leg. J. Schmidt, C. Wirkner, Yeshitla M., Yitbarek W. (53 specimens: CSCHM); ditto, Bale Mts., Shaya Valley SW Goba, alt. 3100 - 3150 m, 6. II. 2020, 6.991843 ° N, 39.884397 ° E, leg. J. Schmidt, C. Wirkner, Yeshitla M., Yitbarek W. (15 specimens: CSCHM).	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
AD70353F6B8F5F479457FE03AE94B251.taxon	diagnosis	Diagnosis. Within the Trechus fauna of the Bale and Arsi Mountains, T. patrizii is easily recognized by absence of the posterior elytral discal seta (Schmidt and Faille 2018). Beside T. patrizii, absence of the posterior elytral discal seta is also characteristic for T. amharicus Ortuno & Novoa and T. aethiopicus Alluaud. Trechus patrizii differs from T. amharicus by two male protarsomeres dilated, by presence of an apical disc on aedeagal median lobe, and by very differently sclerotized endophallus (for comparison see Ortuno and Novoa 2011: 134, fig. 3 b, d, and Schmidt and Faille 2018: 37, figs 64 - 66). Trechus patrizii differs from T. aethiopicus by the pronotum with smaller laterobasal angles, and by the copulatory piece of the endophallus, which is long and spine-like (short and tube-like in T. aethiopicus, see Jeannel 1927: 197, figs 598, 599).	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
AD70353F6B8F5F479457FE03AE94B251.taxon	distribution	Distribution. Occurrences of T. patrizii are known from the northern slope of the Bale Mountains as well as from the northerly adjacent Arsi volcanos Chillalo, Encuolo, and Kaka (Schmidt and Faille 2018; Fig. 1).	en	Faille, Arnaud, Hofmann, Sylvia, Merene, Yeshitla, Hauth, David, Opgenoorth, Lars, Woldehawariat, Yitbarek, Schmidt, Joachim (2023): Explosive radiation versus old relicts: The complex history of Ethiopian Trechina, with description of a new genus and a new subgenus (Coleoptera, Carabidae, Trechini). Deutsche Entomologische Zeitschrift 70 (2): 311-335, DOI: http://dx.doi.org/10.3897/dez.70.107425, URL: http://dx.doi.org/10.3897/dez.70.107425
