taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
993CA639CE7BFFDBFE9FCFBF5EF452DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5370204/files/figure.png	https://doi.org/10.5281/zenodo.5370204	FIGURE 2. Worker of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species. A. Lateral habitus. B. Dorsal habitus.	FIGURE 2. Worker of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species. A. Lateral habitus. B. Dorsal habitus.	2019-10-18	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.		Zenodo	biologists	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.			
993CA639CE7BFFDBFE9FCFBF5EF452DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5370206/files/figure.png	https://doi.org/10.5281/zenodo.5370206	FIGURE 3. Worker of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species. A. Dorsal detail of head and mesosoma. B. Facial view. C. Integumental sculpture of mesosomal dorsum.	FIGURE 3. Worker of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species. A. Dorsal detail of head and mesosoma. B. Facial view. C. Integumental sculpture of mesosomal dorsum.	2019-10-18	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.		Zenodo	biologists	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.			
993CA639CE7BFFDBFE9FCFBF5EF452DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5370208/files/figure.png	https://doi.org/10.5281/zenodo.5370208	FIGURE 4. Worker of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species. A. Forewing (arrow indicates termination of radius on leading edge of wing). B. Outer surface of metatibia and metatarsus. C. Inner surface of metatibia and metatarsus.	FIGURE 4. Worker of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species. A. Forewing (arrow indicates termination of radius on leading edge of wing). B. Outer surface of metatibia and metatarsus. C. Inner surface of metatibia and metatarsus.	2019-10-18	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.		Zenodo	biologists	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.			
993CA639CE7BFFDBFE9FCFBF5EF452DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5370210/files/figure.png	https://doi.org/10.5281/zenodo.5370210	FIGURE 5. Queen of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species. A. Lateral habitus. B. Dorsal habitus.	FIGURE 5. Queen of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species. A. Lateral habitus. B. Dorsal habitus.	2019-10-18	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.		Zenodo	biologists	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.			
993CA639CE7BFFDBFE9FCFBF5EF452DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5370212/files/figure.png	https://doi.org/10.5281/zenodo.5370212	FIGURE 6. Queen of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species. A. Facial view. B. Dorsal detail of head and mesosoma. C. Outer surface of metatibia and metatarsus.	FIGURE 6. Queen of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species. A. Facial view. B. Dorsal detail of head and mesosoma. C. Outer surface of metatibia and metatarsus.	2019-10-18	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.		Zenodo	biologists	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.			
993CA639CE7BFFDBFE9FCFBF5EF452DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5370214/files/figure.png	https://doi.org/10.5281/zenodo.5370214	FIGURE 7. Drone of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species. A. Lateral habitus. B. Dorsal habitus.	FIGURE 7. Drone of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species. A. Lateral habitus. B. Dorsal habitus.	2019-10-18	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.		Zenodo	biologists	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.			
993CA639CE7BFFDBFE9FCFBF5EF452DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5370216/files/figure.png	https://doi.org/10.5281/zenodo.5370216	FIGURE 8. Drone of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species. A. Facial view. B. Outer surface of metatibia and metatarsus. C. Ventral view of metasomal apex. D. Ventral oblique aspect of metasomal apex, showing length and angulation of setae on sternal brushes.	FIGURE 8. Drone of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species. A. Facial view. B. Outer surface of metatibia and metatarsus. C. Ventral view of metasomal apex. D. Ventral oblique aspect of metasomal apex, showing length and angulation of setae on sternal brushes.	2019-10-18	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.		Zenodo	biologists	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.			
993CA639CE7BFFDBFE9FCFBF5EF452DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5370218/files/figure.png	https://doi.org/10.5281/zenodo.5370218	FIGURE 9. Male terminalia of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species. A. Sternum VI. B. Sternum VII. C. Genital capsule (left half is dorsal, right half is ventral).	FIGURE 9. Male terminalia of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species. A. Sternum VI. B. Sternum VII. C. Genital capsule (left half is dorsal, right half is ventral).	2019-10-18	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.		Zenodo	biologists	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.			
993CA639CE7BFFDBFE9FCFBF5EF452DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5370220/files/figure.png	https://doi.org/10.5281/zenodo.5370220	FIGURE 10. SEM micrograph of the damaged egg of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species. A. Egg full length with anterior end upper left. B. Anterior end showing hexagonal chorionic pattern becoming elongated toward micropyle (arrow). C. Close-up of hexagonal patterning at midbody. D. Close-up of rectangle in C. E. Micropyle (arrow) and adjoining chorion removed from egg.	FIGURE 10. SEM micrograph of the damaged egg of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species. A. Egg full length with anterior end upper left. B. Anterior end showing hexagonal chorionic pattern becoming elongated toward micropyle (arrow). C. Close-up of hexagonal patterning at midbody. D. Close-up of rectangle in C. E. Micropyle (arrow) and adjoining chorion removed from egg.	2019-10-18	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.		Zenodo	biologists	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.			
993CA639CE7BFFDBFE9FCFBF5EF452DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5370222/files/figure.png	https://doi.org/10.5281/zenodo.5370222	FIGURE 11. Diagram of entire mature larva of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species, lateral view, head toward left, showing distribution of paired segmental dorsolateral body tubercles and small size of larva.	FIGURE 11. Diagram of entire mature larva of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species, lateral view, head toward left, showing distribution of paired segmental dorsolateral body tubercles and small size of larva.	2019-10-18	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.		Zenodo	biologists	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.			
993CA639CE7BFFDBFE9FCFBF5EF452DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/6423533/files/figure.png	https://doi.org/10.5281/zenodo.6423533	FIGURE 12. SEM micrographs of mature larva of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species. A. Entire larva, oblique frontal view. B. Close-up of pro- and mesothoracic segments showing paired segmental dorsolateral tubercles (arrows) and distribution of fine spicules. C. Close-up of meso- and metathoracic segments and of first abdominal segment showing dorsolateral tubercles along left side as well as distribution of fine spicules. D. Head, frontal view, demonstrating extreme width, and showing pronounced, dome-shaped antennae. E. Same, oblique lateral view. F. Close-up of labrum and mandibles, frontal view, showing cluster of multipronged spicules at lower edge of labrum as well as sharp-pointed mandibles. G. Close-up of left side of cluster of multipronged spicules along lower edge of labrum and sharply toothed upper edge of left mandible, possibly used as comb to push provisions through prongs toward esophagus.	FIGURE 12. SEM micrographs of mature larva of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species. A. Entire larva, oblique frontal view. B. Close-up of pro- and mesothoracic segments showing paired segmental dorsolateral tubercles (arrows) and distribution of fine spicules. C. Close-up of meso- and metathoracic segments and of first abdominal segment showing dorsolateral tubercles along left side as well as distribution of fine spicules. D. Head, frontal view, demonstrating extreme width, and showing pronounced, dome-shaped antennae. E. Same, oblique lateral view. F. Close-up of labrum and mandibles, frontal view, showing cluster of multipronged spicules at lower edge of labrum as well as sharp-pointed mandibles. G. Close-up of left side of cluster of multipronged spicules along lower edge of labrum and sharply toothed upper edge of left mandible, possibly used as comb to push provisions through prongs toward esophagus.	2019-10-18	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.		Zenodo	biologists	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.			
993CA639CE7BFFDBFE9FCFBF5EF452DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5370224/files/figure.png	https://doi.org/10.5281/zenodo.5370224	FIGURE 13. Nest of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species, in La Guajira, Colombia (photographs by M.S. Engel). A. Location of nest in the side of a divi-divi tree (Libidibia coriaria (Jacq.) Schltdl.) (Fabaceae: Caesalpinioideae) (arrow shows position of nest entrance). B. Detail of nest entrance.	FIGURE 13. Nest of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species, in La Guajira, Colombia (photographs by M.S. Engel). A. Location of nest in the side of a divi-divi tree (Libidibia coriaria (Jacq.) Schltdl.) (Fabaceae: Caesalpinioideae) (arrow shows position of nest entrance). B. Detail of nest entrance.	2019-10-18	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.		Zenodo	biologists	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.			
993CA639CE7BFFDBFE9FCFBF5EF452DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5370226/files/figure.png	https://doi.org/10.5281/zenodo.5370226	FIGURE 14. Nests of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species, in La Guajira, Colombia (photographs by M.S. Engel). A. Nest in the side of an organ pipe cactus (Stenocereus griseus (Haw.) Buxb.) (Cactaceae: Cactoideae) (arrow shows position of nest entrance). B. Detail of nest entrance, from nest depicted in A. C. A single-opening entrance of a nest in Libidibia coriaria (Jacq.) Schltdl. (Fabaceae: Caesalpinioideae). D. Tomb in Riohacha cemetery, with nests embedded in sides (arrow indicates position of nest entrance). E. Detail of nest entrance, from nest depicted in D.	FIGURE 14. Nests of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species, in La Guajira, Colombia (photographs by M.S. Engel). A. Nest in the side of an organ pipe cactus (Stenocereus griseus (Haw.) Buxb.) (Cactaceae: Cactoideae) (arrow shows position of nest entrance). B. Detail of nest entrance, from nest depicted in A. C. A single-opening entrance of a nest in Libidibia coriaria (Jacq.) Schltdl. (Fabaceae: Caesalpinioideae). D. Tomb in Riohacha cemetery, with nests embedded in sides (arrow indicates position of nest entrance). E. Detail of nest entrance, from nest depicted in D.	2019-10-18	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.		Zenodo	biologists	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.			
993CA639CE7BFFDBFE9FCFBF5EF452DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5370228/files/figure.png	https://doi.org/10.5281/zenodo.5370228	FIGURE 15. Sampled nest of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species, from La Guajira, Colombia (photographs by M.S. Engel). A. Section of Libidibia coriaria (Jacq.) Schltdl. (Fabaceae: Caesalpinioideae), with side removed to expose nest within. B. Detail of brood cells in situ. C. Detail of brood cells in situ.	FIGURE 15. Sampled nest of Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species, from La Guajira, Colombia (photographs by M.S. Engel). A. Section of Libidibia coriaria (Jacq.) Schltdl. (Fabaceae: Caesalpinioideae), with side removed to expose nest within. B. Detail of brood cells in situ. C. Detail of brood cells in situ.	2019-10-18	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.		Zenodo	biologists	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.			
993CA639CE7BFFDBFE9FCFBF5EF452DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5370230/files/figure.png	https://doi.org/10.5281/zenodo.5370230	FIGURE 16. Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species, from La Guajira, Colombia. A. Diagrammatic outline of sagittal section of nest and contents (note that nest entrance was removed and would be facing outward toward the viewer), B. Photograph of medial section of brood cells. C. Photograph of honey pots and brood cells.	FIGURE 16. Trigonisca (Trigonisca) mepecheu Engel and Gonzalez, new species, from La Guajira, Colombia. A. Diagrammatic outline of sagittal section of nest and contents (note that nest entrance was removed and would be facing outward toward the viewer), B. Photograph of medial section of brood cells. C. Photograph of honey pots and brood cells.	2019-10-18	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.		Zenodo	biologists	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.			
993CA639CE7BFFDBFE9FCFBF5EF452DD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5370232/files/figure.png	https://doi.org/10.5281/zenodo.5370232	FIGURE 17. Nest of Melipona favosa (Fabricius) in the base of a large Libidibia coriaria (Jacq.) Schltdl. (Fabaceae: Caesalpinioideae), in La Guajira, Colombia (photographs by M.S. Engel). A. Worker at entrance made of mud in a radiating pattern. B. Detail of guard at nest entrance. C. Worker departing nest entrance.	FIGURE 17. Nest of Melipona favosa (Fabricius) in the base of a large Libidibia coriaria (Jacq.) Schltdl. (Fabaceae: Caesalpinioideae), in La Guajira, Colombia (photographs by M.S. Engel). A. Worker at entrance made of mud in a radiating pattern. B. Detail of guard at nest entrance. C. Worker departing nest entrance.	2019-10-18	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.		Zenodo	biologists	Engel, Michael S.;Rozen, Jerome G.;Sepúlveda-Cano, Paula A.;Smith, Corey Shepard;Thomas, Jennifer C.;Ospina-Torres, Rodulfo;Gonzalez, Victor H.			
