identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
9E6587C2FFDAFF91D0F967E1C906C2F7.text	9E6587C2FFDAFF91D0F967E1C906C2F7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thyropygus cuisinieri	<div><p>The Thyropygus cuisinieri subgroup</p><p>A subgroup of the T. allevatus group, characterized by</p><p>1. A prominent projecting lobe on the lateral surface of the anterior coxal fold (apl) (Figs. 1 A, 2A, 3A–B, 4A)</p><p>2. A slender lamellar lobe (sll) on the telopodite (not to be confused with the spatulate lobe characteristic of the T. opinatus subgroup)</p><p>3. A protruding lobe on the telopodite (lo) distal to the femoral spine</p><p>4. An additional small lobe (asl) further distal to lo on the telopodite</p><p>5. No additional projection (amp) on the anterior coxal fold (present in both opinatus and bifurcus subgroups)</p><p>6. A single femoral spine</p><p>The cuisinieri subgroup includes T. carli (Attems, 1938), T. cuisinieri Carl, 1917, T. foliaceus (Demange, 1961), new status and T. jarukchusri n. sp.</p></div>	https://treatment.plazi.org/id/9E6587C2FFDAFF91D0F967E1C906C2F7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Pimvichai, Piyatida;Enghoff, Henrik;Panha, Somsak	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak (2011): A revision of the Thyropygus allevatus group. Part 4: the T. cuisinieri subgroup (Diplopoda: Spirostreptida: Harpagophoridae). Zootaxa 2980: 37-48, DOI: 10.5281/zenodo.202125
9E6587C2FFD9FF92D0F966EDCEECC3E6.text	9E6587C2FFD9FF92D0F966EDCEECC3E6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thyropygus cuisinieri	<div><p>Key to species of the cuisinieri subgroup</p><p>1 Apical part of telopodite with a relatively broad, round transparent lamellar lobe (sll); anterior coxal fold (ac) distally with two acuminate processes, the lateral one curving mesad (Figs. 3 A, E)....... Thyropygus foliaceus (Demange, 1961), new status</p><p>1* Apical part of telopodite with a slender transparent lamellar lobe, mostly terminating in a sharp point (sll) (Figs. 1 C, 2D, 4C)..................................................................................................... 2</p><p>2 Anterior coxal fold (ac) distally with two short, broadly triangular, straight, pointed processes (Fig. 1 A).............................................................................................. Thyropygus carli Attems, 1938</p><p>2* Processes of anterior coxal fold (ac) longer, the lateral process curving mesad (Figs. 2 A, 4A)......................... 3</p><p>3 Anterior coxal fold (ac) distally with two acuminate processes: the outer a slender, sharp spine, curving mesad: the inner basally broad, longer than the outer one, pointed, pointing distad; mesal process of posterior coxal fold (pmp) in posterior view hidden behind huge, thumblike lateral process (plp); tibial spine (ti) not constricted at base (Figs. 2 A–D)........................................................................................... Thyropygus cuisinieri Carl, 1917</p><p>3* Anterior coxal fold (ac) distally with two processes: a lateral slender, sharp spine, curving mesad, and a mesal, broadly triangular, pointed process pointing distad; lateral process of posterior coxal fold (plp) much smaller, leaving mesal process of posterior coxal fold (pmp) visible in posterior view; tibial spine (ti) distinctly constricted at base (Figs. 4 A–C)........................................................................................... .. Thyropygus jarukchusri n. sp.</p></div>	https://treatment.plazi.org/id/9E6587C2FFD9FF92D0F966EDCEECC3E6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Pimvichai, Piyatida;Enghoff, Henrik;Panha, Somsak	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak (2011): A revision of the Thyropygus allevatus group. Part 4: the T. cuisinieri subgroup (Diplopoda: Spirostreptida: Harpagophoridae). Zootaxa 2980: 37-48, DOI: 10.5281/zenodo.202125
9E6587C2FFD9FF94D0F96191C980C528.text	9E6587C2FFD9FF94D0F96191C980C528.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thyropygus carli Attems 1938	<div><p>Thyropygus carli Attems, 1938</p><p>(Figs. 1 A–D)</p><p>Thyropygus cuisinieri carli Attems, 1938: 281; Hoffman 1975: 127 Thyropisthus cuisinieri carli: Attems 1942: 79; Demange 1961: 124 Thyropisthus (Duplopisthus) cuisinieri carli: Demange 1983: 563 Thyropygus carli: Enghoff 2004: 36</p><p>Material examined. SYNTYPES (2 vials) VIETNAM (S-Annam), Nhatrang, 12° 14ˏ 43˝ N, 109° 11ˏ 57˝ E. C. Dawydoff leg., (NMW 2558, NMW 2559). 2 males, 4 females VIETNAM, Nhatrang, 12° 14ˏ 43˝ N, 109° 11ˏ 57˝ E. 22 March 2000. P. Gravlund and A. R. Rasmussen leg., (ZMUC). 3 males, 1 female VIETNAM, Nhatrang, Bao Dai’s village hotel, 12° 12ˏ 39˝ N, 109° 12ˏ 59˝ E. 9–13 March 2004. J. B. Rasmussen leg., (ZMUC). 2 males VIETNAM, Cauda, near Nhatrang, 12° 15ˏ 4˝ N, 109° 11ˏ 45˝ E. October, 27 November 1959. J. Knudsen leg., (ZMUC). 1 male, 1 juvenile S. VIETNAM, Phantiet, 12° 14ˏ 31˝ N, 109° 11ˏ 14˝ E. 2002. P. Gravlund leg., (ZMUC). 1 male bought in pet shop (ZMUC).</p><p>Diagnosis. A species of the cuisinieri subgroup. Lateral process of posterior coxal fold (plp) huge, thumblike. Similar in this respect to T. cuisinieri and T. foliaceus . Differs from all other species of the T. cuisinieri subgroup by having two short, straight, broadly triangular distal processes on the anterior coxal fold (ac).</p><p>Description. Adult males with 54–61 podous rings, no apodous rings. Length 11–12 cm, width 8.0– 8.9 mm. Adult females with 57–60 podous rings, no apodous rings. Length 9–12 cm, width 7.3–9.4 mm. Color in life unknown; preserved specimens with antennae, legs pale brown; metazona dark brown; prozona whitish brown; epiproct, paraprocts and hypoproct brown.</p><p>Gonopods (Figs. 1 A–D): Anterior coxal fold (ac) (Fig. 1 A) basally slender, becoming broader towards tip, with a prominent projecting lobe (apl) on the lateral surface; distally with two short, straight, broadly triangular distal processes, the lateral one produced into a short, sharp spine, pointing distad, the mesal one broad, apically pointed, pointing distad. Posterior coxal fold (pc) (Fig. 1 B) basally with relatively high lateral paracoxites (px); mesal process (pmp) very small, short, rounded lamellar, directed distolaterad; lateral process (plp) huge, thumblike, directed distolaterad, covering pmp in posterior view. At the base of plp, a blunt ridge (br). Telopodite (Figs. 1 C–D) leaving coxite over shelf of posterior coxal fold between pmp and plp; femoral spine (fe) long, curving mesad in the horizontal plane, distinctly crenulated along inner curvature, in situ resting against plp, telopodite distally to fe with a round lobe (lo) projecting distolaterad, further distally with an additional small round lobe (asl); tibial spine (ti) very long, slender, with longitudinal crest (lc) along outer surface, recurved, its tip in situ resting close to lo; apical part with a slender, sharply pointed, transparent lamellar lobe (sll); palette (pa) simple, flattened; distally with about six to eight brownish blepharochaetae (bp).</p></div>	https://treatment.plazi.org/id/9E6587C2FFD9FF94D0F96191C980C528	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Pimvichai, Piyatida;Enghoff, Henrik;Panha, Somsak	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak (2011): A revision of the Thyropygus allevatus group. Part 4: the T. cuisinieri subgroup (Diplopoda: Spirostreptida: Harpagophoridae). Zootaxa 2980: 37-48, DOI: 10.5281/zenodo.202125
9E6587C2FFDFFF94D0F96699CEF7C0FA.text	9E6587C2FFDFFF94D0F96699CEF7C0FA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thyropygus cuisinieri Carl 1917	<div><p>Thyropygus cuisinieri Carl, 1917</p><p>(Figs. 2 A–E)</p><p>Thyropygus cuisinieri Carl, 1917: 392; Attems 1930: 157; 1936: 267; 1938: 280; Hoffman 1975: 127; Enghoff et al. 2004: 36; Enghoff 2005: 94.</p><p>Thyropisthus cuisinieri: Attems 1942: 79; Demange 1961: 122; Demange 1983: 563.</p><p>Material examined. HOLOTYPE male VIETNAM Tayninh, Conchinchine (one gonopod telopodite missing) (MHNG). 4 males, 3 females, 7 juveniles THAILAND, Koh Kut, 11° 40ˏ 47˝ N, 102° 32ˏ 56˝ E. 6 April 1959. B. Degerbøl leg., (ZMUC). 2 males VIETNAM, Phuquoc Island. Nguyen Duc Anh leg., (ZMUC). 17 males, 5 females CAMBODIA, Gulf of Thailand, SE part of Koh Tang, 10° 16.5ˏ N, 103° 8.9ˏ E. 12–14 December 2010. S. Tarasov leg., (ZMUC). 1 male, 1 juvenile, VIETNAM, Kontum, Annam, C. Dawydoff leg., (NMW).</p><p>Diagnosis. A species of the cuisinieri subgroup. Differing from all other species in the subgroup by having two distal, acuminate processes on the anterior coxal fold (ac): a lateral slender, sharp spine, curving mesad, and a mesal basally broad process, longer than the lateral one, pointed, pointing distad. Also unique in having a transverse, blunt ridge (tr) at the base of the lateral process of the posterior coxal fold (plp).</p><p>Description. Adult males with 56–66 podous rings, no apodous rings. Length 10–13 cm, width 6.3–8.8 mm. Adult females with 56–60 podous rings, no apodous rings. Length 9–11 cm, width 6.0–8.0 mm. Color in life unknown; preserved specimens with antennae, legs reddish brown; metazona dark brown; prozona brown; epiproct and margins of paraprocts reddish orange. Thai specimens with a longitudinal, brown, mid-dorsal band.</p><p>Gonopods (Figs. 2 A–E): Anterior coxal fold (ac) (Fig. 2 A) basally slender, becoming broader towards tip, with a prominent projecting lobe (apl) on the lateral surface; distally with two acuminate processes: the outer a slender, sharp spine, curving mesad; the inner basally broad, longer than the outer one, pointed, pointing distad. Posterior coxal fold (pc) (Figs. 2 B–C) basally with relatively high lateral paracoxites (px); mesal process (pmp) a short, rounded, slightly twisted lamella; lateral process (plp) thumblike, directed distolaterad. At the base of plp, a transverse, blunt ridge (tr). Some specimens with two longitudinal, blunt ridges (lr1 and lr2) basal to tr; lr1 and lr2 first parallel, but basally diverging and swollen. Telopodite (Figs. 2 D–E) leaving coxite over shelf of posterior coxal fold between pmp and plp; femoral spine (fe) very long, pointing back along telopodite shaft, very distinctly crenulated along inner curvature, in situ resting against posterior surface of ac, telopodite distally to fe with a round lobe (lo) projecting distolaterad, further distally with an additional small round lobe (asl); tibial spine (ti) very long, slender, with longitudinal crest (lc) along outer surface, recurved, its tip in situ resting close to lo; apical part with a slender, transparent lamellar lobe (sll); palette (pa) simple, flattened; distally with about seven to eight brownish blepharochaetae (bp).</p><p>Notes on material. Carl (1917) mentioned two females from the type locality which he assigned, with doubt, to T. cuisinieri . The vial containing the holotype of T. cuisinieri also contains two females, without doubt the ones referred to by Carl. They are not to be regarded as part of the type series which thus consists only of one male—the holotype. These females do not form part of the basis for our redescription of T. cuisinieri .</p><p>Notes. In some individuals, the lateral process of the posterior coxal fold (plp) is very small (Fig. 2 C), in others very distinct (Fig. 2 B). The tip of the lamellar lobe (sll) is rounded in some individuals, sharply pointed in others.</p></div>	https://treatment.plazi.org/id/9E6587C2FFDFFF94D0F96699CEF7C0FA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Pimvichai, Piyatida;Enghoff, Henrik;Panha, Somsak	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak (2011): A revision of the Thyropygus allevatus group. Part 4: the T. cuisinieri subgroup (Diplopoda: Spirostreptida: Harpagophoridae). Zootaxa 2980: 37-48, DOI: 10.5281/zenodo.202125
9E6587C2FFDEFF97D0F96298CA90C42C.text	9E6587C2FFDEFF97D0F96298CA90C42C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thyropygus foliaceus (Demange 1961) Demange 1961	<div><p>Thyropygus foliaceus (Demange, 1961), new status</p><p>(Figs. 3 A–F, 5A–B)</p><p>Thyropisthus cuisinieri (Carl) foliaceus Demange, 1961: 122</p><p>Material examined. HOLOTYPE (# 3100) male THAILAND, Ubon Ratchathani Province (Ugon in Demange, 1961). 23 March 1929. H. M. Smith leg., (USNM). 10 males, 4 females THAILAND, Roi-Et Province, Chaturaphakphiman district, Ban Nong Phue, 15° 49ˏ 34˝ N, 103° 31ˏ 11˝ E. 12 April 2009. J. Pimvichai, C. Pimvichai and P. Pimvichai leg., (CUMZ). 10 males, 10 females THAILAND, Ubon Ratchathani Province, Khong Chiam district, Phataem National Park, 15° 23ˏ 55˝ N, 105° 30ˏ 27˝ E. 25 April 2009. S. Panha, P. Pimvichai and members of Animal Systematics Research Unit leg., (CUMZ). 1 male THAILAND, Kalasin Province, Sahussakhan District, Phu-Kum-Khao, 16° 41ˏ 33˝ N, 103° 31ˏ 30˝ E. 6 June 2010. P. Pimvichai leg., (CUMZ). 1 male THAILAND, Ubon Ratchathani Province, Khemmarat district, 16° 2ˏ 14˝ N, 105° 12ˏ 28˝ E. April 2010. J. Tubtimon leg., (CUMZ).</p><p>Diagnosis. A species of the cuisinieri subgroup. Differing from all other species in the subgroup by having a relatively broad, rounded, transparent lamellar lobe (sll).</p><p>Description. Adult males with 58–64 podous rings, no apodous rings. Length 9–16 cm, width 5.8–9.7 mm. Adult females with 59–65 podous rings, no apodous rings. Length 9–14 cm, width 6.2–8.5 mm. Overall color of living animal in some populations (Fig. 5B) brownish orange, in some populations (Fig. 5A) dark brown, antennae and legs brownish orange.</p><p>Gonopods (Figs. 3 A–F): Anterior coxal fold (ac) (Fig. 3 A) basally slender, becoming broader towards tip, with a prominent projecting lobe (apl) on the lateral surface; distally with two acuminate processes: the outer a slender, sharp spine, curving mesad; the inner basally slightly broader, curving mesad. Posterior coxal fold (pc) (Fig. 3 C) basally with relatively high lateral paracoxites (px); mesal process (pmp) very small, short, rounded lamellar, directed distolaterad; lateral process (plp) large, thumblike, directed distolaterad, covering pmp in posterior view. Telopodite (Figs. 3 E–F) leaving coxite over shelf of posterior coxal fold between pmp and plp; femoral spine (fe) very long, pointing back along telopodite shaft, slightly mesad, distinctly crenulated along inner curvature, in situ resting against posterior surface of ac, telopodite distally to fe with a round lobe (lo) projecting distolaterad, further distally with an additional small round lobe (asl); tibial spine (ti) very long, slender, with longitudinal crest (lc) along outer surface, recurved, its tip in situ resting close to lo; apical part with a relatively broad, round, terminating in a slightly pointed, transparent lamellar lobe (sll); palette (pa) simple, flattened; distally with about six to eight brownish blepharochaetae (bp).</p><p>Notes. Specimens from Khong Chiam deviate in some characters; the anterior coxal fold (ac) (Fig. 3 B) is similar to that of cuisinieri . However, a transverse, blunt ridge (tr) (Fig. 3 D) at base of lateral process of posterior coxal fold (plp) is lacking, and though these specimens seem to have two longitudinal, blunt ridges (lr1 and lr2), these are not distinct as in cuisinieri . The mesal process of the posterior coxal fold (pmp) is similar to that of foliaceus, and the lamellar lobe (sll) is relatively broad (although not as broad as in foliaceus), rounded. For the time being, we propose to regards these specimens as foliaceus . In specimens from Roi-Et, the femoral spine (fe) lacks crenulation along the inner curvature.</p></div>	https://treatment.plazi.org/id/9E6587C2FFDEFF97D0F96298CA90C42C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Pimvichai, Piyatida;Enghoff, Henrik;Panha, Somsak	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak (2011): A revision of the Thyropygus allevatus group. Part 4: the T. cuisinieri subgroup (Diplopoda: Spirostreptida: Harpagophoridae). Zootaxa 2980: 37-48, DOI: 10.5281/zenodo.202125
9E6587C2FFDCFF97D0F9679DC851C080.text	9E6587C2FFDCFF97D0F9679DC851C080.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thyropygus jarukchusri	<div><p>Thyropygus jarukchusri n. sp.</p><p>(Figs. 4 A–D, 5C)</p><p>Material examined. HOLOTYPE male THAILAND, Roi-Et Province, Chaturaphakphiman district, Ban Nong Phue, 15° 49ˏ 34˝ N, 103° 31ˏ 11˝ E. 12 April 2009. J. Pimvichai, C. Pimvichai and P. Pimvichai leg., (CUMZ). – Paratypes: 21 males, 6 females, same data as holotype (CUMZ, ZMUC).</p><p>Etymology. The specific epithet is a combination of the names of PP’s parents, in recognition of their support. The name is to be treated as a noun in apposition.</p><p>Diagnosis. A species of the cuisinieri subgroup. Differing from all other species in the subgroup by smaller body size and by having the tibial spine (ti) distinctly constricted at base.</p><p>Description. Adult males with 57–59 podous rings, no apodous rings. Length ca. 6 cm, width ca. 3.9–4.4 mm. Adult females with 56–60 podous rings, no apodous rings. Length ca. 7–9 cm, width ca. 5.0– 5.2 mm. Overall color of living animal (Fig. 5 C) brownish orange, with anterior metazona dark brown.</p><p>Gonopods (Figs. 4 A–D): Anterior coxal fold (ac) (Fig. 4 A) basally almost parallel-sided, becoming broader towards tip, with a prominent projecting lobe (apl) on the lateral surface; distally with two processes: a lateral slen- der, sharp spine, curving mesad, and a mesal, broadly expanded, terminally pointed process pointing distad. Posterior coxal fold (pc) (Fig. 4 B) basally with relatively high lateral paracoxites (px); mesal process (pmp) a short, rounded, slightly twisted lamella; lateral process (plp) a small, thumblike, directed distolaterad. Telopodite (Figs. 4 C–D) leaving coxite over shelf of posterior coxal fold between pmp and plp; femoral spine (fe) long, curving mesad, without crenulated along inner curvature, in situ resting against posterior surface of ac, telopodite distally to fe with a round lobe (lo) projecting distolaterad, further distally with an additional small round lobe (asl); tibial spine (ti) very long, slender, distinctly constricted at base, with longitudinal crest (lc) along outer surface, recurved, its tip in situ resting close to lo; apical part with a slender, sharply pointed, transparent lamellar lobe (sll); palette (pa) simple, flattened; distally with about four to seven brownish blepharochaetae (bp).</p><p>Distribution (Fig. 6). Known only from the type locality, where it co-exists with T. foliaceus . Although these taxa are quite similar, there are clear difference in gonopod structure, body size (Fig. 7) and color. Furthermore, T. foliaceus and T. jarukchusri showed a divergence in COI sequences by 11.2% (74 out of 658 positions) (cf. discussion in Pimvichai et al., 2011). We regard T. jarukchusri and T. foliaceus as a sympatric species (their strict sympatry excludes the possibility of treating them as subspecies).</p></div>	https://treatment.plazi.org/id/9E6587C2FFDCFF97D0F9679DC851C080	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Pimvichai, Piyatida;Enghoff, Henrik;Panha, Somsak	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak (2011): A revision of the Thyropygus allevatus group. Part 4: the T. cuisinieri subgroup (Diplopoda: Spirostreptida: Harpagophoridae). Zootaxa 2980: 37-48, DOI: 10.5281/zenodo.202125
