taxonID	type	description	language	source
9E6587C2FFD9FF94D0F96191C980C528.taxon	materials_examined	Material examined. SYNTYPES (2 vials) VIETNAM (S-Annam), Nhatrang, 12 ° 14 ˏ 43 ˝ N, 109 ° 11 ˏ 57 ˝ E. C. Dawydoff leg., (NMW 2558, NMW 2559). 2 males, 4 females VIETNAM, Nhatrang, 12 ° 14 ˏ 43 ˝ N, 109 ° 11 ˏ 57 ˝ E. 22 March 2000. P. Gravlund and A. R. Rasmussen leg., (ZMUC). 3 males, 1 female VIETNAM, Nhatrang, Bao Dai’s village hotel, 12 ° 12 ˏ 39 ˝ N, 109 ° 12 ˏ 59 ˝ E. 9 – 13 March 2004. J. B. Rasmussen leg., (ZMUC). 2 males VIETNAM, Cauda, near Nhatrang, 12 ° 15 ˏ 4 ˝ N, 109 ° 11 ˏ 45 ˝ E. October, 27 November 1959. J. Knudsen leg., (ZMUC). 1 male, 1 juvenile S. VIETNAM, Phantiet, 12 ° 14 ˏ 31 ˝ N, 109 ° 11 ˏ 14 ˝ E. 2002. P. Gravlund leg., (ZMUC). 1 male bought in pet shop (ZMUC).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak (2011): A revision of the Thyropygus allevatus group. Part 4: the T. cuisinieri subgroup (Diplopoda: Spirostreptida: Harpagophoridae). Zootaxa 2980: 37-48, DOI: 10.5281/zenodo.202125
9E6587C2FFD9FF94D0F96191C980C528.taxon	diagnosis	Diagnosis. A species of the cuisinieri subgroup. Lateral process of posterior coxal fold (plp) huge, thumblike. Similar in this respect to T. cuisinieri and T. foliaceus. Differs from all other species of the T. cuisinieri subgroup by having two short, straight, broadly triangular distal processes on the anterior coxal fold (ac).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak (2011): A revision of the Thyropygus allevatus group. Part 4: the T. cuisinieri subgroup (Diplopoda: Spirostreptida: Harpagophoridae). Zootaxa 2980: 37-48, DOI: 10.5281/zenodo.202125
9E6587C2FFD9FF94D0F96191C980C528.taxon	description	Description. Adult males with 54 – 61 podous rings, no apodous rings. Length 11 – 12 cm, width 8.0 – 8.9 mm. Adult females with 57 – 60 podous rings, no apodous rings. Length 9 – 12 cm, width 7.3 – 9.4 mm. Color in life unknown; preserved specimens with antennae, legs pale brown; metazona dark brown; prozona whitish brown; epiproct, paraprocts and hypoproct brown. Gonopods (Figs. 1 A – D): Anterior coxal fold (ac) (Fig. 1 A) basally slender, becoming broader towards tip, with a prominent projecting lobe (apl) on the lateral surface; distally with two short, straight, broadly triangular distal processes, the lateral one produced into a short, sharp spine, pointing distad, the mesal one broad, apically pointed, pointing distad. Posterior coxal fold (pc) (Fig. 1 B) basally with relatively high lateral paracoxites (px); mesal process (pmp) very small, short, rounded lamellar, directed distolaterad; lateral process (plp) huge, thumblike, directed distolaterad, covering pmp in posterior view. At the base of plp, a blunt ridge (br). Telopodite (Figs. 1 C – D) leaving coxite over shelf of posterior coxal fold between pmp and plp; femoral spine (fe) long, curving mesad in the horizontal plane, distinctly crenulated along inner curvature, in situ resting against plp, telopodite distally to fe with a round lobe (lo) projecting distolaterad, further distally with an additional small round lobe (asl); tibial spine (ti) very long, slender, with longitudinal crest (lc) along outer surface, recurved, its tip in situ resting close to lo; apical part with a slender, sharply pointed, transparent lamellar lobe (sll); palette (pa) simple, flattened; distally with about six to eight brownish blepharochaetae (bp).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak (2011): A revision of the Thyropygus allevatus group. Part 4: the T. cuisinieri subgroup (Diplopoda: Spirostreptida: Harpagophoridae). Zootaxa 2980: 37-48, DOI: 10.5281/zenodo.202125
9E6587C2FFDFFF94D0F96699CEF7C0FA.taxon	materials_examined	Material examined. HOLOTYPE male VIETNAM Tayninh, Conchinchine (one gonopod telopodite missing) (MHNG). 4 males, 3 females, 7 juveniles THAILAND, Koh Kut, 11 ° 40 ˏ 47 ˝ N, 102 ° 32 ˏ 56 ˝ E. 6 April 1959. B. Degerbøl leg., (ZMUC). 2 males VIETNAM, Phuquoc Island. Nguyen Duc Anh leg., (ZMUC). 17 males, 5 females CAMBODIA, Gulf of Thailand, SE part of Koh Tang, 10 ° 16.5 ˏ N, 103 ° 8.9 ˏ E. 12 – 14 December 2010. S. Tarasov leg., (ZMUC). 1 male, 1 juvenile, VIETNAM, Kontum, Annam, C. Dawydoff leg., (NMW).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak (2011): A revision of the Thyropygus allevatus group. Part 4: the T. cuisinieri subgroup (Diplopoda: Spirostreptida: Harpagophoridae). Zootaxa 2980: 37-48, DOI: 10.5281/zenodo.202125
9E6587C2FFDFFF94D0F96699CEF7C0FA.taxon	diagnosis	Diagnosis. A species of the cuisinieri subgroup. Differing from all other species in the subgroup by having two distal, acuminate processes on the anterior coxal fold (ac): a lateral slender, sharp spine, curving mesad, and a mesal basally broad process, longer than the lateral one, pointed, pointing distad. Also unique in having a transverse, blunt ridge (tr) at the base of the lateral process of the posterior coxal fold (plp).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak (2011): A revision of the Thyropygus allevatus group. Part 4: the T. cuisinieri subgroup (Diplopoda: Spirostreptida: Harpagophoridae). Zootaxa 2980: 37-48, DOI: 10.5281/zenodo.202125
9E6587C2FFDFFF94D0F96699CEF7C0FA.taxon	description	Description. Adult males with 56 – 66 podous rings, no apodous rings. Length 10 – 13 cm, width 6.3 – 8.8 mm. Adult females with 56 – 60 podous rings, no apodous rings. Length 9 – 11 cm, width 6.0 – 8.0 mm. Color in life unknown; preserved specimens with antennae, legs reddish brown; metazona dark brown; prozona brown; epiproct and margins of paraprocts reddish orange. Thai specimens with a longitudinal, brown, mid-dorsal band. Gonopods (Figs. 2 A – E): Anterior coxal fold (ac) (Fig. 2 A) basally slender, becoming broader towards tip, with a prominent projecting lobe (apl) on the lateral surface; distally with two acuminate processes: the outer a slender, sharp spine, curving mesad; the inner basally broad, longer than the outer one, pointed, pointing distad. Posterior coxal fold (pc) (Figs. 2 B – C) basally with relatively high lateral paracoxites (px); mesal process (pmp) a short, rounded, slightly twisted lamella; lateral process (plp) thumblike, directed distolaterad. At the base of plp, a transverse, blunt ridge (tr). Some specimens with two longitudinal, blunt ridges (lr 1 and lr 2) basal to tr; lr 1 and lr 2 first parallel, but basally diverging and swollen. Telopodite (Figs. 2 D – E) leaving coxite over shelf of posterior coxal fold between pmp and plp; femoral spine (fe) very long, pointing back along telopodite shaft, very distinctly crenulated along inner curvature, in situ resting against posterior surface of ac, telopodite distally to fe with a round lobe (lo) projecting distolaterad, further distally with an additional small round lobe (asl); tibial spine (ti) very long, slender, with longitudinal crest (lc) along outer surface, recurved, its tip in situ resting close to lo; apical part with a slender, transparent lamellar lobe (sll); palette (pa) simple, flattened; distally with about seven to eight brownish blepharochaetae (bp).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak (2011): A revision of the Thyropygus allevatus group. Part 4: the T. cuisinieri subgroup (Diplopoda: Spirostreptida: Harpagophoridae). Zootaxa 2980: 37-48, DOI: 10.5281/zenodo.202125
9E6587C2FFDFFF94D0F96699CEF7C0FA.taxon	materials_examined	Notes on material. Carl (1917) mentioned two females from the type locality which he assigned, with doubt, to T. cuisinieri. The vial containing the holotype of T. cuisinieri also contains two females, without doubt the ones referred to by Carl. They are not to be regarded as part of the type series which thus consists only of one male — the holotype. These females do not form part of the basis for our redescription of T. cuisinieri. Notes. In some individuals, the lateral process of the posterior coxal fold (plp) is very small (Fig. 2 C), in others very distinct (Fig. 2 B). The tip of the lamellar lobe (sll) is rounded in some individuals, sharply pointed in others.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak (2011): A revision of the Thyropygus allevatus group. Part 4: the T. cuisinieri subgroup (Diplopoda: Spirostreptida: Harpagophoridae). Zootaxa 2980: 37-48, DOI: 10.5281/zenodo.202125
9E6587C2FFDEFF97D0F96298CA90C42C.taxon	materials_examined	Material examined. HOLOTYPE (# 3100) male THAILAND, Ubon Ratchathani Province (Ugon in Demange, 1961). 23 March 1929. H. M. Smith leg., (USNM). 10 males, 4 females THAILAND, Roi-Et Province, Chaturaphakphiman district, Ban Nong Phue, 15 ° 49 ˏ 34 ˝ N, 103 ° 31 ˏ 11 ˝ E. 12 April 2009. J. Pimvichai, C. Pimvichai and P. Pimvichai leg., (CUMZ). 10 males, 10 females THAILAND, Ubon Ratchathani Province, Khong Chiam district, Phataem National Park, 15 ° 23 ˏ 55 ˝ N, 105 ° 30 ˏ 27 ˝ E. 25 April 2009. S. Panha, P. Pimvichai and members of Animal Systematics Research Unit leg., (CUMZ). 1 male THAILAND, Kalasin Province, Sahussakhan District, Phu-Kum-Khao, 16 ° 41 ˏ 33 ˝ N, 103 ° 31 ˏ 30 ˝ E. 6 June 2010. P. Pimvichai leg., (CUMZ). 1 male THAILAND, Ubon Ratchathani Province, Khemmarat district, 16 ° 2 ˏ 14 ˝ N, 105 ° 12 ˏ 28 ˝ E. April 2010. J. Tubtimon leg., (CUMZ).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak (2011): A revision of the Thyropygus allevatus group. Part 4: the T. cuisinieri subgroup (Diplopoda: Spirostreptida: Harpagophoridae). Zootaxa 2980: 37-48, DOI: 10.5281/zenodo.202125
9E6587C2FFDEFF97D0F96298CA90C42C.taxon	diagnosis	Diagnosis. A species of the cuisinieri subgroup. Differing from all other species in the subgroup by having a relatively broad, rounded, transparent lamellar lobe (sll).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak (2011): A revision of the Thyropygus allevatus group. Part 4: the T. cuisinieri subgroup (Diplopoda: Spirostreptida: Harpagophoridae). Zootaxa 2980: 37-48, DOI: 10.5281/zenodo.202125
9E6587C2FFDEFF97D0F96298CA90C42C.taxon	description	Description. Adult males with 58 – 64 podous rings, no apodous rings. Length 9 – 16 cm, width 5.8 – 9.7 mm. Adult females with 59 – 65 podous rings, no apodous rings. Length 9 – 14 cm, width 6.2 – 8.5 mm. Overall color of living animal in some populations (Fig. 5 B) brownish orange, in some populations (Fig. 5 A) dark brown, antennae and legs brownish orange. Gonopods (Figs. 3 A – F): Anterior coxal fold (ac) (Fig. 3 A) basally slender, becoming broader towards tip, with a prominent projecting lobe (apl) on the lateral surface; distally with two acuminate processes: the outer a slender, sharp spine, curving mesad; the inner basally slightly broader, curving mesad. Posterior coxal fold (pc) (Fig. 3 C) basally with relatively high lateral paracoxites (px); mesal process (pmp) very small, short, rounded lamellar, directed distolaterad; lateral process (plp) large, thumblike, directed distolaterad, covering pmp in posterior view. Telopodite (Figs. 3 E – F) leaving coxite over shelf of posterior coxal fold between pmp and plp; femoral spine (fe) very long, pointing back along telopodite shaft, slightly mesad, distinctly crenulated along inner curvature, in situ resting against posterior surface of ac, telopodite distally to fe with a round lobe (lo) projecting distolaterad, further distally with an additional small round lobe (asl); tibial spine (ti) very long, slender, with longitudinal crest (lc) along outer surface, recurved, its tip in situ resting close to lo; apical part with a relatively broad, round, terminating in a slightly pointed, transparent lamellar lobe (sll); palette (pa) simple, flattened; distally with about six to eight brownish blepharochaetae (bp). Notes. Specimens from Khong Chiam deviate in some characters; the anterior coxal fold (ac) (Fig. 3 B) is similar to that of cuisinieri. However, a transverse, blunt ridge (tr) (Fig. 3 D) at base of lateral process of posterior coxal fold (plp) is lacking, and though these specimens seem to have two longitudinal, blunt ridges (lr 1 and lr 2), these are not distinct as in cuisinieri. The mesal process of the posterior coxal fold (pmp) is similar to that of foliaceus, and the lamellar lobe (sll) is relatively broad (although not as broad as in foliaceus), rounded. For the time being, we propose to regards these specimens as foliaceus. In specimens from Roi-Et, the femoral spine (fe) lacks crenulation along the inner curvature.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak (2011): A revision of the Thyropygus allevatus group. Part 4: the T. cuisinieri subgroup (Diplopoda: Spirostreptida: Harpagophoridae). Zootaxa 2980: 37-48, DOI: 10.5281/zenodo.202125
9E6587C2FFDCFF97D0F9679DC851C080.taxon	materials_examined	Material examined. HOLOTYPE male THAILAND, Roi-Et Province, Chaturaphakphiman district, Ban Nong Phue, 15 ° 49 ˏ 34 ˝ N, 103 ° 31 ˏ 11 ˝ E. 12 April 2009. J. Pimvichai, C. Pimvichai and P. Pimvichai leg., (CUMZ). – Paratypes: 21 males, 6 females, same data as holotype (CUMZ, ZMUC).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak (2011): A revision of the Thyropygus allevatus group. Part 4: the T. cuisinieri subgroup (Diplopoda: Spirostreptida: Harpagophoridae). Zootaxa 2980: 37-48, DOI: 10.5281/zenodo.202125
9E6587C2FFDCFF97D0F9679DC851C080.taxon	etymology	Etymology. The specific epithet is a combination of the names of PP’s parents, in recognition of their support. The name is to be treated as a noun in apposition.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak (2011): A revision of the Thyropygus allevatus group. Part 4: the T. cuisinieri subgroup (Diplopoda: Spirostreptida: Harpagophoridae). Zootaxa 2980: 37-48, DOI: 10.5281/zenodo.202125
9E6587C2FFDCFF97D0F9679DC851C080.taxon	diagnosis	Diagnosis. A species of the cuisinieri subgroup. Differing from all other species in the subgroup by smaller body size and by having the tibial spine (ti) distinctly constricted at base.	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak (2011): A revision of the Thyropygus allevatus group. Part 4: the T. cuisinieri subgroup (Diplopoda: Spirostreptida: Harpagophoridae). Zootaxa 2980: 37-48, DOI: 10.5281/zenodo.202125
9E6587C2FFDCFF97D0F9679DC851C080.taxon	description	Description. Adult males with 57 – 59 podous rings, no apodous rings. Length ca. 6 cm, width ca. 3.9 – 4.4 mm. Adult females with 56 – 60 podous rings, no apodous rings. Length ca. 7 – 9 cm, width ca. 5.0 – 5.2 mm. Overall color of living animal (Fig. 5 C) brownish orange, with anterior metazona dark brown. Gonopods (Figs. 4 A – D): Anterior coxal fold (ac) (Fig. 4 A) basally almost parallel-sided, becoming broader towards tip, with a prominent projecting lobe (apl) on the lateral surface; distally with two processes: a lateral slen- der, sharp spine, curving mesad, and a mesal, broadly expanded, terminally pointed process pointing distad. Posterior coxal fold (pc) (Fig. 4 B) basally with relatively high lateral paracoxites (px); mesal process (pmp) a short, rounded, slightly twisted lamella; lateral process (plp) a small, thumblike, directed distolaterad. Telopodite (Figs. 4 C – D) leaving coxite over shelf of posterior coxal fold between pmp and plp; femoral spine (fe) long, curving mesad, without crenulated along inner curvature, in situ resting against posterior surface of ac, telopodite distally to fe with a round lobe (lo) projecting distolaterad, further distally with an additional small round lobe (asl); tibial spine (ti) very long, slender, distinctly constricted at base, with longitudinal crest (lc) along outer surface, recurved, its tip in situ resting close to lo; apical part with a slender, sharply pointed, transparent lamellar lobe (sll); palette (pa) simple, flattened; distally with about four to seven brownish blepharochaetae (bp).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak (2011): A revision of the Thyropygus allevatus group. Part 4: the T. cuisinieri subgroup (Diplopoda: Spirostreptida: Harpagophoridae). Zootaxa 2980: 37-48, DOI: 10.5281/zenodo.202125
9E6587C2FFDCFF97D0F9679DC851C080.taxon	distribution	Distribution (Fig. 6). Known only from the type locality, where it co-exists with T. foliaceus. Although these taxa are quite similar, there are clear difference in gonopod structure, body size (Fig. 7) and color. Furthermore, T. foliaceus and T. jarukchusri showed a divergence in COI sequences by 11.2 % (74 out of 658 positions) (cf. discussion in Pimvichai et al., 2011). We regard T. jarukchusri and T. foliaceus as a sympatric species (their strict sympatry excludes the possibility of treating them as subspecies).	en	Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak (2011): A revision of the Thyropygus allevatus group. Part 4: the T. cuisinieri subgroup (Diplopoda: Spirostreptida: Harpagophoridae). Zootaxa 2980: 37-48, DOI: 10.5281/zenodo.202125
