taxonID	type	description	language	source
9D51B604FFFFED08FF26F8C2C2D8F9F6.taxon	description	(Figs 1 – 9)	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFFFED08FF26F8C2C2D8F9F6.taxon	materials_examined	Material examined. Holotype. Female. India, Kollangod, Palghat District, Kerala, a few kms from Sirvani, netted in grasses, 27 March 1980, Gy. Topál coll. (HNHM). Paratypes. One male, one female in MHNG, four females and four males in HNHM. Locality and date as for holotype.	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFFFED08FF26F8C2C2D8F9F6.taxon	description	Description. Female. Length of idiosoma 600 – 640 µm, width 340 – 420 µm (n = 6). Shape oval, caudally rounded. Dorsal idiosoma (Fig. 1). Dorsal shield completely separated from reduced marginal shield. Dorsal shield scutiform, 360 – 370 µm long, 250 – 260 µm wide, covered by irregular pits, bearing smooth needle-like setae (ca. 25 – 30 µm). Marginal shields divided into separate rounded platelets, each bearing one smooth and needle-like setae (ca. 29 – 32 µm). Pygidial shield divided into two large lateral quadrangular plates and a small triangular central plate. Pygidial plates covered by irregular pits, without bearing setae. A pair of smooth and needle like setae located on rounded platelets between dorsal shield and pygidial plates. Other areas of dorsal idiosoma covered by membranous cuticle. Ventral idiosoma (Fig. 2). Sternal shield weakly sclerotised, its surface mostly smooth, only anterior area has H-shaped strongly sclerotised groove. Five pairs of smooth and needle-like sternal setae (ca. 10 – 11 µm) present, St 1 situated near anterior margin of sternal shield, inserted on strongly sclerotised area. St 2 located between coxae III, St 3 and St 4 near basal edges of genital shield, St 5 situated posterior to genital shield. Inguinal shields present, covered by irregular pits, without setae. Anal shield rhomboid, its surface smooth and bearing two pairs of adanal setae (ca. 10 – 13 µm) and one postanal seta (ca. 15 µm). One pair of ventral shields present on caudal area, their surface irregularly pitted, without setae. Two pairs of smooth and needle-like setae (ca. 19 – 22 µm) without platelets situated between two inguinal shields, three pairs of long smooth setae (ca. 29 – 32 µm) placed on small platelets situated between inguinal, anal and caudal shields. Pedofossae absent. Genital shield linguliform, anteriorly rounded, its surface without ornamentation. Stigmata situated between coxae III and IV, peritremes straight, with short post-stigmatal parts (Fig. 3). Tritosternum with wide base, laciniae apically divided into three short smooth branches (Fig. 4). Gnathosoma (Fig. 4). Hypostomal setae h 1 (ca. 56 µm) and h 3 (ca. 62 µm) smooth and long, h 2 short (ca. 22 µm) and smooth, h 4 short (ca. 18 µm) and marginally pilose. Ventral surface of hypostome bearing seven rows of denticles. Palp trochanter with one pilose and one smooth seta, other setae on palp smooth. Corniculi horn-like, internal malae smooth, shorter than corniculi. Epistome marginally serrate (Fig. 4). Fixed digit of chelicera longer than movable digit. Legs. Each leg with claws on tips of tarsi, all setae on legs smooth and needle-like (Figs 5 – 8). Male. Length of idiosoma 520 – 550 µm, width 270 – 320 µm (n = 5). Shape and dorsal idiosoma as in female. Ventral idiosoma (Fig. 13). Surface of sternal shield smooth, five pairs of needle-like sternal setae (ca. 7 – 8 µm) present. St 1 placed near anterior margin of sternal shield, St 2 at level of posterior margin of coxae II, St 3 at central level of coxae III, St 4 at level of posterior margin of coxae III, St 5 near posterior margin of sternal shield. Position and shape of ventral setae and ornamentation of ventral shield as in female. Genital shield oval, without sculptural pattern, situated between coxae III and IV. Gnathosoma as in female.	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFFFED08FF26F8C2C2D8F9F6.taxon	etymology	Etymology. The new species is named for Durjodhana, the descendant of the Kuru, the legendary king, who was an ancestor of many Kuaravas in the Hindu epic, Mahabharata.	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFFFED08FF26F8C2C2D8F9F6.taxon	discussion	Notes. Because the second hypostomal setae reach the insertion of first hypostomal setae and the pygidial shield is separated from the dorsal shield, I placed the new species into the subgenus Uroseius (Apionseius) Berlese. Most species from this subgenus have an entire pygidial shield; the new species is similar to U. (A.) infirmus (Berlese) which has a divided pygidial shield. The differences between these species are shown in Table 1.	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFFBED0CFF26FF5FC696FCE3.taxon	description	(Figs 10 – 21)	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFFBED0CFF26FF5FC696FCE3.taxon	materials_examined	Material examined. Holotype. Female. India, Ngarkanda, Hugli District, West Bengal, sifted from garbage-heap in village, 0 3 January 1980, Gy. Topál coll. (HNHM). Paratypes. One female, one male deposited in MHNG, six males in HNHM. Locality and date as in holotype.	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFFBED0CFF26FF5FC696FCE3.taxon	description	Description. Female. Length of idiosoma 420 – 440 µm, width 290 – 310 µm (n = 2). Shape oval, posterior margin rounded. Dorsal idiosoma (Fig. 10). Dorsal and marginal shields fused anteriorly, all dorsal setae short (ca. 12 – 15 µm), smooth and needle-like, situated in small, rounded depressions (Fig. 11). Dorsal shield with reticulate sculptural pattern between the depressions associated with setae. Marginal shield bearing smooth and needle-like setae (ca. 12 – 14 µm) without basal depressions, with oval pits and and undulate inner margin. Ventral idiosoma (Fig. 12). Surface of sternal shield smooth. Four pairs of smooth and needle-like sternal setae present, St 1 (ca. 6 µm) inserted near anterior margin of sternal shield, St 2 (ca. 8 µm) between coxae II, St 3 (ca. 10 µm) at level of posterior margin of coxae II, St 4 (ca. 13 µm) at central level of coxae III. Ventral shield bearing numerous smooth needle-like setae associated with rounded depressions (ca. 12 – 16 µm), adanal setae and postanal seta similar in shape to ventral setae, but shorter (ad 1 ca. 7 µm, ad 2 ca. 9 µm and ps ca. 9 µm). Anal opening oval, elevated from surface of ventral shield. Metapodal lines present. Pedofossae deep, their surface smooth, with separated furrows for tarsi IV. Surface between pedofossae IV and metapodal lines with reticulate ornamenttaion. Genital shield scutiform, anteriorly pointed, its surface smooth. Stigmata situated between coxae II and III, poststigmatal part of peritremes short, pre-stigmatal area long, with three curves (Fig. 13). Tritosternum with narrow base, laciniae basally pilose, apically divided into two long and smooth lateral branches (Fig. 14). Gnathosoma (Fig. 15). Hypostomal setae h 1 placed on small protuberances, smooth and needle-like (ca. 27 µm), h 2 short (ca. 22 µm), smooth and needle-like, h 3 long (ca. 37 µm) and needle-like, with one pair of lateral spines, h 4 long (ca. 36 µm) and marginally serrate. Palp trochanter with two marginally serrate setae, other setae on palp smooth. Corniculi short and horn-like, apically with one tooth, internal malae longer than corniculi, smooth. Epistome marginally serrate. Chelicerae short and robust, movable digit as long as fixed digit, both digits bearing three teeth, internal sclerotised node present (Fig. 16). Legs. Each leg with ambulacral claws (not illustrated on leg I), all setae on legs smooth and needle-like (Figs 17 – 20). Male. Length of idiosoma 390 – 410 µm, width 250 – 280 µm (n = 7). Shape and dorsal idiosoma as in female. Ventral idiosoma (Fig. 21). Surface of sternal shield smooth, four pairs of needle-like sternal setae (ca. 8 – 10 µm) present. St 1 placed near anterior margin of sternal shield, St 2 at level of posterior margin of coxae II, St 3 at central level of coxae III, St 4 at level of posterior margin of coxae IV. Two pairs of lyriform fissures present, one pair situated near St 1, second pair near St 4. Genital shield circular, without sculptural pattern, situated between coxae IV. Gnathosoma. As in female.	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFFBED0CFF26FF5FC696FCE3.taxon	etymology	Etymology. The new species is named for Arjuna, son of Pandu and Kunti in the Hindu epic, Mahabharata. Arjuna belongs to Pandawa, the fight of Kuarawas and Pandawas is the main act of the Mahabharata.	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFFBED0CFF26FF5FC696FCE3.taxon	discussion	Notes. The new species belongs to the Trichouropoda ovalis - group (Hirschmann & WiŠniewski 1986). Within this group the common character is the presence of oval pits on the dorsal and ventral surfaces of the body. The new species can be easily separated from the other species of this group on the basis of the association of the needle-like setae with the oval pits, and on the basis of the absence of setae St 4. These character states have not been found before in this species group.	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFF9ED02FF26FC44C390FF2D.taxon	description	(Figs 22 – 32)	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFF9ED02FF26FC44C390FF2D.taxon	materials_examined	Material examined. Holotype. India, Debrapani, Darheeling District, West Bengal, extracted from litter in indigenous wood, 30 May 1980, Gy. Topál coll. (HNHM). Paratypes. One male in MHNG, one female on slide in HNHM. Locality and date same as in holotype.	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFF9ED02FF26FC44C390FF2D.taxon	description	Description. Female. Length of idiosoma 420 – 430 µm, width 360 – 370 µm (n = 2). Shape of body circular. Dorsal idiosoma (Fig. 22). Dorsal and marginal shields fused anteriorly, all dorsal setae short (ca. 7 – 9 µm), smooth and needle-like (Fig 23). Dorsal shield covered with oval pits, with very small pits between larger pits (Fig. 23). Marginal shield bearing smooth and needle-like setae (ca. 7 – 8 µm), its surface smooth. Ventral idiosoma (Fig. 24). Surface of sternal shield mostly smooth, with oval pits near its anterior margin. Five pairs of smooth, short (ca. 4 – 6 µm), needle-like sternal setae present, St 1 placed near anterior point of genital shield, St 2 at level of posterior margin of coxae II, St 3 at central level of coxae III, St 4 at level of anterior margin of coxae IV, St 5 near basal edges of genital shield. Four pairs of ventral setae and two pairs of adanal setae similar in shape and length to sternal setae, postanal seta absent. Anal opening rounded. Pedofossae deep, their surface smooth, without separate furrows for tarsi IV. Genital shield long, narrow and linguliform, anteriorly pointed, its surface covered by oval pits. Stigmata situated between coxae II and III, post-stigmatal part of peritremes absent, pre-stigmatal area long, apically hook-shaped (Fig. 25). Tritosternum with triangular base, laciniae divided into three branches (Fig. 26). Gnathosoma (Fig. 26). Hypostomal setae h 1 long (ca. 36 µm), apically pilose, h 2 short (ca. 16 µm), smooth, h 3 long (ca. 22 µm), marginally serrate, h 4 short (ca. 15 µm) and smooth. Corniculi short and horn-like, internal malae longer than corniculi, smooth. Epistome marginally pilose. Fixed digit of chelicera longer than movable digit, bearing a bulbiform sensory organ, movable digit with one central tooth, internal sclerotised node absent (Fig. 27). Legs. First leg without ambulacral claws, all setae on legs smooth and needle-like (Figs 28 – 31). Male. Length of idiosoma 440 µm, width 370 µm (n = 1). Shape and dorsal idiosoma as in female. Ventral idiosoma (Fig. 32). Five pairs of short (ca. 5 – 6 µm) needle-like sternal setae present. St 1 inserted at level of posterior margin of coxae II, St 2 at central level of coxae III, St 3 at level of anterior margin of coxae IV, St 4 near lateral margin of genital shield, St 5 near posterior margin of genital shield. Sternal shield covered by oval pits between St 1 and St 3. One pair of sensory organs can be found between St 1 and St 2. Genital shield oval, without sculptural pattern and situated between pedofossae of legs IV. Gnathosoma. As in female.	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFF9ED02FF26FC44C390FF2D.taxon	etymology	Etymology. The name of the new species refers to the region (Bengal) of India where it was collected.	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFF9ED02FF26FC44C390FF2D.taxon	discussion	Notes. Following Hirschmann's (1972 b) division of the genus Uropoda, I place the new species into the Uropoda penicillata - group. Up to now six species are known in this group (U. complicata (Berlese), U. garciai Hirschmann & Hiramatsu, U. indonensis Hiramatsu, U. micherdzinskii Hirschmann, U. penicillata Hirschmann & Zirngiebl-Nicol and U. penicillatasimilis Hirschmann). The common characters within this species group are the pilose h 1 setae, circular idiosoma, hook-shaped prestigmatal part of the peritreme, and the narrow and pointed genital shield of the female. The characteristic differences between the species of this group are presented in Table 2. Uropoda complicata (Berlese) is missing from Table 2, because most of the characters of this species are lacking from the original description and illustration (Berlese 1905). However, this species differs from the new one on the basis of the smooth surface of genital and ventral shields, which are ornamented in U. bengalica.	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFF6ED01FF26FF5FC600FBCA.taxon	description	(Figs 33 – 43)	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFF6ED01FF26FF5FC600FBCA.taxon	materials_examined	Material examined. Holotype. Female. India, Coonor, Nilgiri Hills, Tamil Nadu, Black Bridge Reserve Forest, extracted from litter, 2130 m, 14 – 17 March 1980, Gy Topál,. coll. Paratypes. One female, one male in NHMG, five females and five males in HNHM. Locality and date same as in holotype.	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFF6ED01FF26FF5FC600FBCA.taxon	description	Description. Female. Length of idiosoma 500 – 510 µm, width 410 – 420 µm (n = 5). Shape of body circular, dorsally domed. Dorsal idiosoma (Fig. 33). Dorsal and marginal shields fused anteriorly, all setae needle-like (Fig 34), with a pair of narrow branches basally, setae long (ca. 38 – 50 µm) in central area and shorter (ca. 26 – 30 µm) in lateral area of dorsal shield. Dorsal shield covered with oval pits (Fig. 34), central and caudal areas elevated. Marginal shield bearing setae similar to dorsal setae in shape, but shorter (ca. 28 – 30 µm), surface of marginal shield covered by oval pits. Ventral idiosoma (Fig. 35). Surface of sternal shield smooth. Five pairs of smooth needle-like sternal setae present, St 1 long (ca. 14 µm), placed near anterior point of genital shield, St 2 (ca. 8 µm) at level of posterior margin of coxae II, St 3 (ca. 9 µm) at level of anterior margin of coxae III, St 4 (ca. 6 µm), at level posterior margin of coxae III, St 5 (ca. 6 µm) near basal edge of genital shield. One pair of centrally located ventral setae short (ca. 7 µm) and needle-like, other ventral setae long (ca. 20 – 28 µm), divided into three or four branches; a row long smooth (ca. 40 – 45 µm) setae located between margins of ventral idiosoma and lateral parts of ventral shield. Postanal seta absent; two pairs of adanal setae present, ad 1 longer (ca. 17 µm), similar in shape to ventral setae, ad 2 short (ca. 6 µm) and needle-like. Surface of ventral shield smooth except for some irregular pits posterior to genital shield and between coxae III and IV. Anal opening oval. Pedofossae deep, their surface smooth, without separate furrows for tarsi IV. Genital shield linguliform, anteriorly pointed, its surface without sculptural pattern. Stigmata situated between coxae II and III, post-stigmatal part of peritremes absent, pre-stigmatal area long, 3 - shaped (Fig. 36). Tritosternum with narrow base, laciniae divided into four short and one long branches (Fig. 37). Gnathosoma (Fig. 38). Hypostomal setae h 1 long (ca. 26 µm) and smooth, h 2 (ca. 10 µm) and h 3 (ca. 12 µm) smooth and robust, h 4 short (ca. 7 µm), marginally serrate. Corniculi short and horn-like, internal malae longer than corniculi, marginally pilose. Epistome marginally pilose. Fixed digit of chelicera longer than movable digit, bearing a bulbiform sensory organ, internal sclerotised node absent (Fig. 39). Palp trochanter with one smooth and one marginally serrate seta, other setae on palp smooth. Legs. First leg without ambulacral claws, with a spatuliform seta on tarsi, all setae on legs smooth and needlelike (Figs 40 – 43). Male. Length of idiosoma 510 – 520 µm, width 410 – 430 µm (n = 7). Shape and dorsal idiosoma as in female. Ventral idiosoma (Fig. 44). Surface of sternal shield ornamented with oval pits, five pairs of needle-like sternal setae present. St 1 extremely long (ca. 53 µm), placed at level of posterior margins of coxae II, St 2 short (ca. 4 µm) at level of posterior margins of coxae II, St 3 short (ca. 4 µm) at central level of coxae III, St 4 (ca. 4 µm) at level of anterior margins of coxae IV, St 5 (ca. 4 µm) near lateral margin of genital shield. Two pairs of lyriform fissures situated near St 1 and near basal margin of genital shield. Genital shield oval, without sculptural pattern, bearing one pair of long (ca. 11 µm), robust eugenital setae and situated between pedofossae of legs IV. Gnathosoma. As in female.	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFF6ED01FF26FF5FC600FBCA.taxon	etymology	Etymology. I dedicate the new species to Hungarian writer Leslie L. Lawrence (László L. Lőrincz), who usually placed his stories in India.	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFF6ED01FF26FF5FC600FBCA.taxon	discussion	Notes. On the basis of the absence of a sclerotised cheliceral node and the long and needle-like first hypostomal setae, I place this species in the genus Uropoda sensu lato (Hirschmann 1972 b). The new species has a complete marginal shield, long and apically 3 - shaped peritremes, and unusual dorsal and ventral setae. It cannot be placed into any of Hirschmann’s (1972 b) species groups, because this character combination is unknown in the genus Uropoda. The shape of genital shield of the female and the apical part of peritremes are similar to the species of Uropoda penicillata - group, but the new species differs from the species of that group by the following characters: dorsal and ventral setation, the shape of the first sternal setae of the male, and the absence of male eugenital setae.	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFF4ED07FF26FBACC4B5FA48.taxon	description	(Figs 45 – 53)	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFF4ED07FF26FBACC4B5FA48.taxon	materials_examined	Material examined. Holotype. Female. India, Berijam, Palni Hills, Tamil Nandu, Nature Reserve, extracted from litter of shola, 11 April 1980, Gy. Topál coll. (HNHM). Paratypes. One female in MHNG, locality and date same as for holotype.	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFF4ED07FF26FBACC4B5FA48.taxon	description	Description. Female. Length of idiosoma 850 – 880 µm, width 610 – 630 µm (n = 2). Shape of body circular, dorsally domed. Dorsal idiosoma (Fig. 45). Dorsal and marginal shields completely separated, all dorsal setae long and needlelike, setae in anterior area shorter (ca. 75 – 83 µm) than in posterior area (ca. 235 – 265 µm). Dorsal shield without pits, caudal area strongly sclerotised and elevated, one pair of strongly sclerotised, C-shaped lines situated on margins of dorsal shield, at level of elevated area. Marginal shield reduced caudally, bearing short (ca. 55 – 60 µm) needle-like setae, except the last pair long (ca. 120 – 140 µm) and needle-like. Surface of marginal shield smooth. Caudal part of dorsal idiosoma covered by membranous cuticle, bearing three pairs of short (ca. 55 – 60 µm) needlelike setae. Ventral idiosoma (Fig. 46). Surface of sternal shield smooth. Five pairs of short (ca. 8 – 10 µm), smooth needlelike sternal setae present, St 1 placed near anterior margin of genital shield, St 2 at level of posterior margin of coxae II, St 3 at central level of coxae III, St 4 at level of posterior margin of coxae III, St 5 near basal margin of genital shield. Two pairs of ventral setae in lateral position very long (ca. 140 – 150 µm) and needle-like, other ventral setae needle-like, on central area short (ca. 20 – 22 µm) and on margins of ventral shield longer (ca. 45 – 50 µm). Postanal seta absent, adanal setae needle-like, ad 1 very short (ca. 8 µm), ad 2 as long as shortest ventral setae. Surface of ventral shield without ornamentation. Anal opening small and oval. Pedofossae reduced, metapodal line present. Genital shield linguliform, with wide process on anterior margin, its surface smooth. Stigmata situated between coxae II and III, post-stigmatal part of peritremes absent, pre-stigmatal area long, apically hooked. Tritosternum with triangular base, laciniae divided into three smooth branches (Fig. 47). Gnathosoma (Fig. 48). Hypostomal setae h 1 long (ca. 62 µm) and smooth, h 2 short (ca. 18 µm), smooth, h 3 (ca. 10 µm), h 4 short (ca. 11 µm), marginally serrate. Corniculi short and horn-like, internal malae longer than corniculi, marginally pilose. Epistome marginally pilose. Fixed digit of chelicera longer than movable digit, bearing a bulbiform sensory organ, internal sclerotised node absent (Fig. 49). Palp trochanter with one smooth and one marginally serrate seta, other setae on palp smooth. Legs. First leg without ambulacral claws, all setae on legs smooth and needle-like (Figs 50 – 53).	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFF4ED07FF26FBACC4B5FA48.taxon	etymology	Etymology. This species is named after Karna, a character of the ancient epic Mahabharata. Karna was the ally and friend of the Pandawas.	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFF4ED07FF26FBACC4B5FA48.taxon	discussion	Notes. The new species belongs to the Uropoda gibba - species group (Hirschmann 1972 b). Eight species have been described in this group, namely U. gibba Hiramatsu from Japan, U. hiramatsui Hirschmann U. hiramatsuformis Hirschmann, U. hiramatsuioides Hirschmann, U. hiramatsuisimilis Hirschmann, U. lauta Hiramatsu & Hirschmann and U. meridiana Hiramatsu & Hirschmann from New Guinea, U. matskasii Hirschmann and U. setata Kontschán from Vietnam. The species of this group are easy to recognise and easy to separate from other Uropoda species on the basis of the elevated caudal area of the dorsal shield. Unfortunately one species is described on the basis of only nymphs and larva (U. meridiana), three species are known only from males (U. hiramatsuioides, U. lauta and U. matskasii), hence I can only show the differences between those species the females of which are known. The characteristic differences between the known females are presented in Table 3.	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFF0ED1AFF26FF5FC5F8F850.taxon	description	(Figs 54 – 68)	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFF0ED1AFF26FF5FC5F8F850.taxon	materials_examined	Material examined. Holotype. Female. India, Berijam, Palni Hills, Tamil Nandu, Nature Reserve, extracted from litter of shola, 11 April 1980. Gy. Topál coll. Paratypes. One male and one female in MHNG, two females, two males, one nymph in HNHM. Locality and date same as in holotype.	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFF0ED1AFF26FF5FC5F8F850.taxon	description	Description. Female. Length of idiosoma 710 – 750 µm, width 430 – 450 µm (n = 4). Shape of body oval, caudally rounded. Dorsal idiosoma (Fig. 54). Dorsal and marginal shields fused anteriorly, all dorsal setae short (ca. 8 – 9 µm) and needle-like. Dorsal shield covered by shallow oval pits. Marginal shield anteriorly with two small, lateral horns, bearing setae similar in shape and length to dorsal setae, surface of marginal shield smooth. Caudal area of marginal shield with longer setae (ca. 19 – 20 µm) situated on small platelets (Fig. 55). Ventra l idiosoma (Fig. 56). Surface of sternal shield smooth, except anterior area, which has a reticulate pattern (Fig. 58). Five pairs of smooth and needle-like sternal setae present, St 1 short (ca. 9 µm), placed near anterior margin of genital shield, St 2 (ca. 11 µm) at level of posterior margin of coxae II, St 3 (ca. 11 µm) at central level of coxae III, St 4 (ca. 11 µm) at central level of coxae IV, St 5 (ca. 13 µm) near basal margin of genital shield. Ventral setae on anterior area of ventral shield smooth and robust (ca. 40 µm), those on caudal area of ventral shield needle-like (ca. 30 µm). Postanal seta absent, two pairs of adanal setae short and needlelike (ca. 11 – 12 µm). Surface of ventral shield covered by irregular pits, two strongly sclerotised ventral grooves situated posterior to pedofossae IV. Anal opening oval, located on a small anal mound. Pedofossae deep, their surface smooth, without separate furrows for tarsi IV. Genital shield linguliform, anteriorly with crown-like process, its surface with reticulate pattern (Fig. 58). Stigmata situated between coxae II and III, post-stigmatal part of peritremes absent, pre-stigmatal area long, apically hook-shaped, with two short lateral branches on central part (Fig. 57). Tritosternum with vase-like base, laciniae divided into three pilose branches (Fig. 59). Gnathosoma (Fig. 59). Hypostomal setae h 1 long (ca. 38 µm), smooth, h 2 (ca. 12 µm) and h 4 (ca. 11 µm) serrate, h 3 (ca. 16 µm) marginally pilose. Corniculi short and horn-like, internal malae longer than corniculi, marginally pilose. Epistome marginally pilose (Fig. 60). Fixed digit of chelicera longer than movable digit and bearing one tooth, internal sclerotised node absent (Fig. 61). Palp trochanter with two marginally serrate setae, other setae on palp smooth. Legs. Each leg with ambulacral claws and with smooth and needle-like setae (Figs 62 – 65). Male. Length of idiosoma 740 – 780 µm, width 440 – 470 µm (n = 11). Shape and dorsal idiosoma as in female. Ventral idiosoma (Fig. 66). Surface of sternal shield smooth, five pairs of short (ca. 13 – 17 µm) and needle-like sternal setae present. St 1 placed at level of anterior margin of coxae II, St 2 at level of posterior margin of coxae II, St 3 at central level of coxae III, St 4 at central level of coxae IV, St 5 at level of posterior margin of coxae IV. Genital shield oval, without sculptural pattern and situated between coxae IV. Gnathosoma. As in female. Deutonymph. Length of idiosoma 660 μm, width 440 μm (n = 1). Shape oval, posterior margin rounded. Dorsal idiosoma (Fig. 67). Dorsal shield linguliform, without ornamentation, only muscle scars visible, bearing short and needle-like setae (ca. 10 – 11 µm). Several oval platelets present on submarginal area, all of them bearing needle-like setae (ca. 12 – 13 µm). Ventra l idiosoma (Fig. 68). Sternal setae short (ca. 9 – 10 µm), St 1 placed near anterior margin of sternal shield, St 2 short, at level of posterior margin of coxae II, St 3 at central level of coxae III, St 4 at level of anterior margin of coxae IV, St 5 near posterior margin of sternal shield. Sternal shield with smooth surface. Ventrianal shield oval and ornamented by maculate sculptural pattern and bearing four pairs of needle-like setae (ca. 7 µm).	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFF0ED1AFF26FF5FC5F8F850.taxon	etymology	Etymology. I named the new species after king Pandu, father of Arjuna and four other sons in the Mahabharata.	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
9D51B604FFF0ED1AFF26FF5FC5F8F850.taxon	discussion	Notes. The new species belongs to the Discourella baloghi - species group (Hirschmann 1972 a) on the basis of the following characters: marginal shield not reduced caudally, pygidial shield and central elevation absent on dorsal shield. Including the new species, 13 species currently belong to this group. Six species of this group (D. baloghi Hirschmann & Zirngiebl-Nicol, D. baloghisimilis WiŠniewski, D. fissilis Hirschmann, D. ditricha Hirschmann, D. lindquisti Hiramatsu & Hirschmann, D. hirschmanni Hiramatsu and D. sivestrisa Hiramatsu) do not possess a crown-like apical process on the female genital shield, but the new one and six previously described species (D. bulgarica Kontschán, D. helvetica Kontschán, D. crucisimilis Hirschmann, D. aokii Hiramatsu, D. torpida Hiramatsu & Hirschmann and D. koreae Hirschmann) do have this apical process. One species is described on the basis of males (D. morikawai Hiramatsu), this species has ornamented sternal shield, but sternal shield of the male of new one is smooth. The characteristic differences among the species possessing crown-like apical process on female genital shield are presented in Table 4.	en	Kontschán, JenŐ (2012): New Uropodina mites from India (Acari: Mesostigmata). Zootaxa 3518: 25-44, DOI: 10.11646/zootaxa.3518.1.2
