identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
9C0687BBFF8DFF8DFF18CBB5A11CB13D.text	9C0687BBFF8DFF8DFF18CBB5A11CB13D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eoplatypalpus Shamshev & Vasilenko & Perkovsky 2025	<div><p>Genus Eoplatypalpus Shamshev, gen. nov.</p><p>Type species. Eoplatypalpus sachalinensis sp. nov., by present designation. Middle Eocene amber of Sakhalin Island, Russia.</p><p>Etymology. Referring to the Eocene period, with a suffix used to refer to Cretaceous genera of the subfamily (an allusion to the genus Platypalpus). The gender is masculine.</p><p>Diagnosis. Male eyes probably narrowly dichoptic, bare, ommatidia of equal size; face broader than frons; thorax somewhat elongated, with small postpronotal lobes, prosternum separated from proepisternum, scutellum with 6 setae; mid femur slender, with simple setae ventrally; wing long, narrow, of nearly equal width at basal and apical third, cells br and bm of approximately equal lengths and almost aligned apically, br narrow, bm nearly 2X as wide as br, cell cua absent (vein CuA absent but some free portion of vein CuA+CuP present as scarcely visible trace), anal lobe reduced.</p><p>Included species. Type species.</p><p>Comparison. Eoplatypalpus gen. nov. possesses bare eyes and somewhat elongated thorax with small postpronotal lobe that would suggest a comparison of this genus primarily with representatives of the tribe Tachydromiini, Symballophthalmus Becker and the Mesozoic genera. The new genus differs from all extant genera of Tachydromiini and from the Mesozoic Archiplatypalpus by the presence of six scutellar setae (versus 2–4 setae) (Kovalev 1974, 1978; Grimaldi &amp; Cumming 1999). On the other hand, Eoplatypalpus gen. nov. shares six scutellar setae with the Mesozoic Cretoplatypalpus (Taimyr amber, Russia) and with the extant Symballophthalmus (Kovalev 1978). The new genus differs from Cretoplatypalpus and Symballophthalmus primarily by the wing lacking cell cua (anal cell). However, cell cua is absent in the Mesozoic Mesoplatypalpus (Cedar Lake, Canadian amber) (Grimaldi &amp; Cumming 1999). Eoplatypalpus gen. nov. differs from Mesoplatypalpus primarily by six scutellar setae (versus eight in Mesoplatypalpus) and longer costal vein (ending at apex of vein M 1 versus ending just slightly past apex of vein R 4+ 5 in Mesoplatypalpus) (Grimaldi &amp; Cumming 1999). Finally, it should be added that the extant genus Megagrapha Melander has a combination of a separated prosternum, 6–8 scutellar setae and the wing lacking cell cua. However, this genus is placed in the tribe Drapetidini, possessing ommatrichia on the eyes and the subquadrate thorax lacking postpronotal lobes (Chillcott &amp; Teskey 1983; Sinclair &amp; Cumming 2006).</p></div>	https://treatment.plazi.org/id/9C0687BBFF8DFF8DFF18CBB5A11CB13D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shamshev, Igor V.;Vasilenko, Dmitry V.;Perkovsky, Evgeny E.	Shamshev, Igor V., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2025): A new genus of hybotid dance fly (Diptera: Hybotidae, Tachydromiinae) from the Eocene Sakhalinian amber. Zootaxa 5575 (2): 283-294, DOI: 10.11646/zootaxa.5575.2.4, URL: https://doi.org/10.11646/zootaxa.5575.2.4
9C0687BBFF8EFF8AFF18CC79A759B6B9.text	9C0687BBFF8EFF8AFF18CC79A759B6B9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eoplatypalpus sachalinensis Shamshev & Vasilenko & Perkovsky 2025	<div><p>Eoplatypalpus sachalinensis Shamshev, sp. nov.</p><p>(Figs 1–3)</p><p>Type material. HOLOTYPE, ♂, Starodubskoye, Dolinsk district, Russia, Sakhalinian amber (PIN 3387/162A; syninclusions: 1 ♂ (PIN 3387/162B) and 1 ♀ (PIN 3387/162C) paratypes; 1 ♂ of Empidoidea (PIN 3387/162D), 1 specimen of Tachydromiinae (PIN 3387/162E), and a fragment of some Diptera (PIN 3387/162F) . PARATYPES: 1 ♂, 1 ♀, Starodubskoye, Dolinsk district, Russia, Sakhalinian amber (PIN 3387/162c and PIN 3387/162d, respectively; in same amber piece with holotype) ; 1 ♀, Starodubskoye, Dolinsk district, Russia, Sakhalinian amber (PIN 3387/ 162g) .</p><p>Remarks on type series. The holotype is in an amber nodule with syninclusions (Fig. 1). The specimen is nearly complete but in a poor condition. The body is somewhat squashed, cleared and deformed; some internal cavities are probably filled with a resin and the wings are somewhat shrunken.</p><p>The paratype male PIN 3387/162B rests near the holotype (Figs 1B, 2B), complete specimen, same condition as the holotype. The paratype female PIN 3387/162C is represented by an incomplete specimen: head and anterior part of thorax missing (Fig. 1C). The paratype female PIN 3387/ 162g is included in a separate, small, narrow, reddish yellow amber nodule together with a specimen of Aranea (3387/162h). The specimen is incomplete because a part of the nodule was probably broken off: head, most part of thorax anteriorly and fore legs missing. The left wing of PIN 3387/ 162g was photographed (Fig. 3B). The wing rests nearly in one plane, only a short portion of posterior margin is slightly deformed (concave).</p><p>Diagnosis. As for genus. Medium-sized, slender species.</p><p>Description. Male (Figs 2, 3A). Since the holotype and the paratype are affected by deformation due to preservation conditions, their body lengths are not to be considered natural. Based on well preserved wing of female, body length of the species is probably about 2.5 mm.</p><p>Head. Rather small, almost rounded in profile, distinctly convex both in front and occiput; ocellar triangle prominent, bearing 2 fine, moderately long setae; vertical setae undifferentiated; occiput with some fine, short setae. Antenna with postpedicel rather elongate (natural length unclear due to deformation; also scape unnaturally elongated), tapered, dorsal margin straight, ventral margin slightly convex; stylus terminal, probably slightly longer than postpedicel, faintly pubescent. Proboscis rather weak, straight, nearly half as long as head height; palpus small, narrow, with scattered, short fine setae.</p><p>Thorax. Pruinescence or tomentum undistinguishable. Postpronotal lobe prominent, small, with fine setae. Mesoscutum entirely covered with short, fine setae, rows of acrostichals and dorsocentrals undifferentiated; row of 4 short supra-alar setae present; scutellum with 3 pairs of long, fine setae of different lengths, apical pair somewhat longer.</p><p>Legs. Long, slender, without any special raptorial modifications; covered with fine, hair-like setae. Femora of subequal widths; fore femur with somewhat longer setae ventrally than mid and hind femora; mid femur only with rows of short, fine setae ventrally. Fore tibia probably slightly thickened; mid tibia without apical projections; hind tibia slightly thickened towards apex, covered with denser and slightly longer setae subapically. Fore and mid tarsi of subequal lengths, hind tarsus somewhat longer; all tarsomeres covered with rather dense and slightly longer setulae, bearing erect setulae ventrally; claws elongate.</p><p>Wing (Fig. 3B, C). Long, narrow, of nearly equal width at basal and apical third, nearly 4X as long as width opposite apex of vein R 1; membrane hyaline, covered with distinct microtrichia. Costal vein as in Symballophthalmus, strong up to apex of M 1, very weak along lower margin; with 3–4 fine setae near base. Sc evanescent, incomplete; R 1 meeting costa at approximately midpoint of wing; Rs originating at about proximal 1/6 between humeral crossvein and apex of vein R 1, long, nearly 4.5X longer than basal portion of R 4+5; R 2+3 straight, R 4+5 and M 1 divergent at base, nearly parallel in remaining part. Cells br and bm of approximately equal lengths (along upper margin of bm) and almost aligned apically, br narrow, bm nearly 2X as wide as br. Crossvein bm-m touching or nearly so with crossvein r-m, slightly oblique. Cell cua absent (vein CuA absent but some free portion of vein CuA+CuP present as scarcely visible trace). Anal lobe scarcely prominent.</p><p>Abdomen. Tergites covered with short to moderately long, fine setae. Hypopygium moderately large, more or less elongate oval, natural rotation degree unclear due to deformation, probably rotated 45–90° (holotype hypopygium deformed and partly broken); separate sclerites unrecognisable due to deformation.</p><p>Female. Wing length 2.8 mm. Similar to male.Abdomen long, telescopic, pointed; terminal segments somewhat deformed but generally resembles Symballophthalmus . Cercus (Fig. 3D, E) long, slender, covered with setulae.</p><p>Etymology. The epithet refers to the type locality, Sakhalin Island, Russia.</p><p>Distribution. Sakhalinian amber.</p></div>	https://treatment.plazi.org/id/9C0687BBFF8EFF8AFF18CC79A759B6B9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shamshev, Igor V.;Vasilenko, Dmitry V.;Perkovsky, Evgeny E.	Shamshev, Igor V., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2025): A new genus of hybotid dance fly (Diptera: Hybotidae, Tachydromiinae) from the Eocene Sakhalinian amber. Zootaxa 5575 (2): 283-294, DOI: 10.11646/zootaxa.5575.2.4, URL: https://doi.org/10.11646/zootaxa.5575.2.4
9C0687BBFF84FF86FF18CADCA0FAB291.text	9C0687BBFF84FF86FF18CADCA0FAB291.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tachydromiini (Sinclair & Cumming 2006)	<div><p>Key to fossil genera of Tachydromiini and Cretoplatypalpus group</p><p>Remarks. We have included Symballophthalmus (tribe Symballophthalmini) in the key because fossil records of this primitive genus are highly likely. The key somewhat suffers from fragmented data in some genera, as noted.</p><p>1 Scutellum with at least 6 setae. Prosternum usually separated from proepisternum (unknown state in Mesoplatypalpus and Electrocyrtoma)...................................................................................... 2</p><p>– Scutellum with at most 4 setae. Prosternum usually fused with proepisternum (except Archiplatypalpus)................ 6</p><p>2 Wing with cell cua. Eyes holoptic or dichoptic …............................................................ 3</p><p>– Wing without cell cua. Eyes dichoptic (unknown state in male of Mesoplatypalpus)................................ 5</p><p>3 Wing cuneiform; crossvein bm-m at most slightly oblique, touching crossvein r-m (Chvála 1975: fig. 23). [Additional character: eyes holoptic.] Extant genus, no known fossils......................................... Symballophthalmus Becker</p><p>– Wing at apex of cell br as broad as at middle; crossvein bm-m strongly oblique, relatively broadly separated from crossvein r-m................................................................................................ 4</p><p>4 Wing with cell cua extending nearly to middle of cell bm (Kovalev 1978: fig. 1b; Grimaldi &amp; Cumming 1999: fig. 34). Eyes dichoptic. Scutellum with 6 setae. Mesozoic: Upper Cretaceous, Russia, Taimyr and Cedar Lake, Manitoba............................................................................................. Cretoplatypalpus Kovalev</p><p>– Wing with cell cua extending far beyond middle of cell bm (Cockerell 1917: fig. 5). Eyes probably holoptic. Scutellum with more numerous setae. Mesozoic: mid-Cretaceous, Burmese amber.......................... Electrocyrtoma Cockerell</p><p>5 Costal vein ending just slightly past apex of vein R 4+5 (Grimaldi &amp; Cumming 1999: fig. 35). Scutellum with 8 setae. Mesozoic: Upper Cretaceous, Cedar Lake, Manitoba .................................. Mesoplatypalpus Grimaldi &amp; Cumming</p><p>– Costal vein ending at apex of vein M 1. Scutellum with 6 setae. Cenozoic: middle Eocene, Sakhalinian amber............................................................................................. Eoplatypalpus gen. nov.</p><p>6 Eyes holoptic (Kovalev 1974: fig. 1). Prosternum separated from proepisternum. Mesozoic: Upper Cretaceous, Russia, Taimyr......................................................................... Archiplatypalpus Kovalev</p><p>– Eyes dichoptic. Prosternum fused with proepisternum........................................................ 7</p><p>7 Wing cell cua more or less formed, with vein CuA distinct and vein CuA+CuP present, although sometimes faint (Chvála 1975: fig. 24). Eyes with upper margins closer to level of ocellar tubercle, divergent near ocellar tubercle with distance between eyes wider than frons (Sinclair et al. 2023: fig. 9C). Mid leg usually raptorial, with femur thickened and armed with two rows of spine-like setae ventrally, tibia often ending in sharp spur. Extant genus; Cenozoic: upper Eocene, Baltic amber............................................................................................. Platypalpus Macquart</p><p>– Wing cell cua absent (vein CuA+CuP absent); vein CuA usually present (sometimes absent in recent species) (Chvála 1975: fig. 26). Eyes with upper margins extending far beyond ocellar tubercle, slightly convergent or parallel behind ocellar tubercle with distance between eyes usually slightly narrower than frons (Sinclair et al. 2023: fig. 8A). Mid leg simple, at most with some modifications in male (subbasal clusters of spine-like setae, excavations and tubercles). Extant genus; Cenozoic: uppermost Eocene, shales of Florissant Formation, Colorado, USA; upper Eocene, Baltic amber................. Tachypeza Meigen</p></div>	https://treatment.plazi.org/id/9C0687BBFF84FF86FF18CADCA0FAB291	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Shamshev, Igor V.;Vasilenko, Dmitry V.;Perkovsky, Evgeny E.	Shamshev, Igor V., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2025): A new genus of hybotid dance fly (Diptera: Hybotidae, Tachydromiinae) from the Eocene Sakhalinian amber. Zootaxa 5575 (2): 283-294, DOI: 10.11646/zootaxa.5575.2.4, URL: https://doi.org/10.11646/zootaxa.5575.2.4
