taxonID	type	description	language	source
9C0687BBFF8DFF8DFF18CBB5A11CB13D.taxon	type_taxon	Type species. Eoplatypalpus sachalinensis sp. nov., by present designation. Middle Eocene amber of Sakhalin Island, Russia.	en	Shamshev, Igor V., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2025): A new genus of hybotid dance fly (Diptera: Hybotidae, Tachydromiinae) from the Eocene Sakhalinian amber. Zootaxa 5575 (2): 283-294, DOI: 10.11646/zootaxa.5575.2.4, URL: https://doi.org/10.11646/zootaxa.5575.2.4
9C0687BBFF8DFF8DFF18CBB5A11CB13D.taxon	etymology	Etymology. Referring to the Eocene period, with a suffix used to refer to Cretaceous genera of the subfamily (an allusion to the genus Platypalpus). The gender is masculine.	en	Shamshev, Igor V., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2025): A new genus of hybotid dance fly (Diptera: Hybotidae, Tachydromiinae) from the Eocene Sakhalinian amber. Zootaxa 5575 (2): 283-294, DOI: 10.11646/zootaxa.5575.2.4, URL: https://doi.org/10.11646/zootaxa.5575.2.4
9C0687BBFF8DFF8DFF18CBB5A11CB13D.taxon	diagnosis	Diagnosis. Male eyes probably narrowly dichoptic, bare, ommatidia of equal size; face broader than frons; thorax somewhat elongated, with small postpronotal lobes, prosternum separated from proepisternum, scutellum with 6 setae; mid femur slender, with simple setae ventrally; wing long, narrow, of nearly equal width at basal and apical third, cells br and bm of approximately equal lengths and almost aligned apically, br narrow, bm nearly 2 X as wide as br, cell cua absent (vein CuA absent but some free portion of vein CuA + CuP present as scarcely visible trace), anal lobe reduced.	en	Shamshev, Igor V., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2025): A new genus of hybotid dance fly (Diptera: Hybotidae, Tachydromiinae) from the Eocene Sakhalinian amber. Zootaxa 5575 (2): 283-294, DOI: 10.11646/zootaxa.5575.2.4, URL: https://doi.org/10.11646/zootaxa.5575.2.4
9C0687BBFF8DFF8DFF18CBB5A11CB13D.taxon	description	Comparison. Eoplatypalpus gen. nov. possesses bare eyes and somewhat elongated thorax with small postpronotal lobe that would suggest a comparison of this genus primarily with representatives of the tribe Tachydromiini, Symballophthalmus Becker and the Mesozoic genera. The new genus differs from all extant genera of Tachydromiini and from the Mesozoic Archiplatypalpus by the presence of six scutellar setae (versus 2 – 4 setae) (Kovalev 1974, 1978; Grimaldi & Cumming 1999). On the other hand, Eoplatypalpus gen. nov. shares six scutellar setae with the Mesozoic Cretoplatypalpus (Taimyr amber, Russia) and with the extant Symballophthalmus (Kovalev 1978). The new genus differs from Cretoplatypalpus and Symballophthalmus primarily by the wing lacking cell cua (anal cell). However, cell cua is absent in the Mesozoic Mesoplatypalpus (Cedar Lake, Canadian amber) (Grimaldi & Cumming 1999). Eoplatypalpus gen. nov. differs from Mesoplatypalpus primarily by six scutellar setae (versus eight in Mesoplatypalpus) and longer costal vein (ending at apex of vein M 1 versus ending just slightly past apex of vein R 4 + 5 in Mesoplatypalpus) (Grimaldi & Cumming 1999). Finally, it should be added that the extant genus Megagrapha Melander has a combination of a separated prosternum, 6 – 8 scutellar setae and the wing lacking cell cua. However, this genus is placed in the tribe Drapetidini, possessing ommatrichia on the eyes and the subquadrate thorax lacking postpronotal lobes (Chillcott & Teskey 1983; Sinclair & Cumming 2006).	en	Shamshev, Igor V., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2025): A new genus of hybotid dance fly (Diptera: Hybotidae, Tachydromiinae) from the Eocene Sakhalinian amber. Zootaxa 5575 (2): 283-294, DOI: 10.11646/zootaxa.5575.2.4, URL: https://doi.org/10.11646/zootaxa.5575.2.4
9C0687BBFF8EFF8AFF18CC79A759B6B9.taxon	description	(Figs 1 – 3)	en	Shamshev, Igor V., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2025): A new genus of hybotid dance fly (Diptera: Hybotidae, Tachydromiinae) from the Eocene Sakhalinian amber. Zootaxa 5575 (2): 283-294, DOI: 10.11646/zootaxa.5575.2.4, URL: https://doi.org/10.11646/zootaxa.5575.2.4
9C0687BBFF8EFF8AFF18CC79A759B6B9.taxon	materials_examined	Type material. HOLOTYPE, ♂, Starodubskoye, Dolinsk district, Russia, Sakhalinian amber (PIN 3387 / 162 A; syninclusions: 1 ♂ (PIN 3387 / 162 B) and 1 ♀ (PIN 3387 / 162 C) paratypes; 1 ♂ of Empidoidea (PIN 3387 / 162 D), 1 specimen of Tachydromiinae (PIN 3387 / 162 E), and a fragment of some Diptera (PIN 3387 / 162 F). PARATYPES: 1 ♂, 1 ♀, Starodubskoye, Dolinsk district, Russia, Sakhalinian amber (PIN 3387 / 162 c and PIN 3387 / 162 d, respectively; in same amber piece with holotype); 1 ♀, Starodubskoye, Dolinsk district, Russia, Sakhalinian amber (PIN 3387 / 162 g).	en	Shamshev, Igor V., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2025): A new genus of hybotid dance fly (Diptera: Hybotidae, Tachydromiinae) from the Eocene Sakhalinian amber. Zootaxa 5575 (2): 283-294, DOI: 10.11646/zootaxa.5575.2.4, URL: https://doi.org/10.11646/zootaxa.5575.2.4
9C0687BBFF8EFF8AFF18CC79A759B6B9.taxon	discussion	Remarks on type series. The holotype is in an amber nodule with syninclusions (Fig. 1). The specimen is nearly complete but in a poor condition. The body is somewhat squashed, cleared and deformed; some internal cavities are probably filled with a resin and the wings are somewhat shrunken. The paratype male PIN 3387 / 162 B rests near the holotype (Figs 1 B, 2 B), complete specimen, same condition as the holotype. The paratype female PIN 3387 / 162 C is represented by an incomplete specimen: head and anterior part of thorax missing (Fig. 1 C). The paratype female PIN 3387 / 162 g is included in a separate, small, narrow, reddish yellow amber nodule together with a specimen of Aranea (3387 / 162 h). The specimen is incomplete because a part of the nodule was probably broken off: head, most part of thorax anteriorly and fore legs missing. The left wing of PIN 3387 / 162 g was photographed (Fig. 3 B). The wing rests nearly in one plane, only a short portion of posterior margin is slightly deformed (concave).	en	Shamshev, Igor V., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2025): A new genus of hybotid dance fly (Diptera: Hybotidae, Tachydromiinae) from the Eocene Sakhalinian amber. Zootaxa 5575 (2): 283-294, DOI: 10.11646/zootaxa.5575.2.4, URL: https://doi.org/10.11646/zootaxa.5575.2.4
9C0687BBFF8EFF8AFF18CC79A759B6B9.taxon	diagnosis	Diagnosis. As for genus. Medium-sized, slender species.	en	Shamshev, Igor V., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2025): A new genus of hybotid dance fly (Diptera: Hybotidae, Tachydromiinae) from the Eocene Sakhalinian amber. Zootaxa 5575 (2): 283-294, DOI: 10.11646/zootaxa.5575.2.4, URL: https://doi.org/10.11646/zootaxa.5575.2.4
9C0687BBFF8EFF8AFF18CC79A759B6B9.taxon	description	Description. Male (Figs 2, 3 A). Since the holotype and the paratype are affected by deformation due to preservation conditions, their body lengths are not to be considered natural. Based on well preserved wing of female, body length of the species is probably about 2.5 mm. Head. Rather small, almost rounded in profile, distinctly convex both in front and occiput; ocellar triangle prominent, bearing 2 fine, moderately long setae; vertical setae undifferentiated; occiput with some fine, short setae. Antenna with postpedicel rather elongate (natural length unclear due to deformation; also scape unnaturally elongated), tapered, dorsal margin straight, ventral margin slightly convex; stylus terminal, probably slightly longer than postpedicel, faintly pubescent. Proboscis rather weak, straight, nearly half as long as head height; palpus small, narrow, with scattered, short fine setae. Thorax. Pruinescence or tomentum undistinguishable. Postpronotal lobe prominent, small, with fine setae. Mesoscutum entirely covered with short, fine setae, rows of acrostichals and dorsocentrals undifferentiated; row of 4 short supra-alar setae present; scutellum with 3 pairs of long, fine setae of different lengths, apical pair somewhat longer. Legs. Long, slender, without any special raptorial modifications; covered with fine, hair-like setae. Femora of subequal widths; fore femur with somewhat longer setae ventrally than mid and hind femora; mid femur only with rows of short, fine setae ventrally. Fore tibia probably slightly thickened; mid tibia without apical projections; hind tibia slightly thickened towards apex, covered with denser and slightly longer setae subapically. Fore and mid tarsi of subequal lengths, hind tarsus somewhat longer; all tarsomeres covered with rather dense and slightly longer setulae, bearing erect setulae ventrally; claws elongate. Wing (Fig. 3 B, C). Long, narrow, of nearly equal width at basal and apical third, nearly 4 X as long as width opposite apex of vein R 1; membrane hyaline, covered with distinct microtrichia. Costal vein as in Symballophthalmus, strong up to apex of M 1, very weak along lower margin; with 3 – 4 fine setae near base. Sc evanescent, incomplete; R 1 meeting costa at approximately midpoint of wing; Rs originating at about proximal 1 / 6 between humeral crossvein and apex of vein R 1, long, nearly 4.5 X longer than basal portion of R 4 + 5; R 2 + 3 straight, R 4 + 5 and M 1 divergent at base, nearly parallel in remaining part. Cells br and bm of approximately equal lengths (along upper margin of bm) and almost aligned apically, br narrow, bm nearly 2 X as wide as br. Crossvein bm-m touching or nearly so with crossvein r-m, slightly oblique. Cell cua absent (vein CuA absent but some free portion of vein CuA + CuP present as scarcely visible trace). Anal lobe scarcely prominent. Abdomen. Tergites covered with short to moderately long, fine setae. Hypopygium moderately large, more or less elongate oval, natural rotation degree unclear due to deformation, probably rotated 45 – 90 ° (holotype hypopygium deformed and partly broken); separate sclerites unrecognisable due to deformation. Female. Wing length 2.8 mm. Similar to male. Abdomen long, telescopic, pointed; terminal segments somewhat deformed but generally resembles Symballophthalmus. Cercus (Fig. 3 D, E) long, slender, covered with setulae.	en	Shamshev, Igor V., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2025): A new genus of hybotid dance fly (Diptera: Hybotidae, Tachydromiinae) from the Eocene Sakhalinian amber. Zootaxa 5575 (2): 283-294, DOI: 10.11646/zootaxa.5575.2.4, URL: https://doi.org/10.11646/zootaxa.5575.2.4
9C0687BBFF8EFF8AFF18CC79A759B6B9.taxon	etymology	Etymology. The epithet refers to the type locality, Sakhalin Island, Russia.	en	Shamshev, Igor V., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2025): A new genus of hybotid dance fly (Diptera: Hybotidae, Tachydromiinae) from the Eocene Sakhalinian amber. Zootaxa 5575 (2): 283-294, DOI: 10.11646/zootaxa.5575.2.4, URL: https://doi.org/10.11646/zootaxa.5575.2.4
9C0687BBFF8EFF8AFF18CC79A759B6B9.taxon	distribution	Distribution. Sakhalinian amber.	en	Shamshev, Igor V., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2025): A new genus of hybotid dance fly (Diptera: Hybotidae, Tachydromiinae) from the Eocene Sakhalinian amber. Zootaxa 5575 (2): 283-294, DOI: 10.11646/zootaxa.5575.2.4, URL: https://doi.org/10.11646/zootaxa.5575.2.4
9C0687BBFF84FF86FF18CADCA0FAB291.taxon	discussion	Remarks. We have included Symballophthalmus (tribe Symballophthalmini) in the key because fossil records of this primitive genus are highly likely. The key somewhat suffers from fragmented data in some genera, as noted.	en	Shamshev, Igor V., Vasilenko, Dmitry V., Perkovsky, Evgeny E. (2025): A new genus of hybotid dance fly (Diptera: Hybotidae, Tachydromiinae) from the Eocene Sakhalinian amber. Zootaxa 5575 (2): 283-294, DOI: 10.11646/zootaxa.5575.2.4, URL: https://doi.org/10.11646/zootaxa.5575.2.4
