identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
A30D87CAFF9DFFCFF259F92BCFA3F894.text	A30D87CAFF9DFFCFF259F92BCFA3F894.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Enicospilus cederbergi Johansson 2018	<div><p>Enicospilus cederbergi sp. nov.</p><p>urn:lsid:zoobank.org:act: 9368EFAF-9601-4672-843F-A9EE71B82DB0 Figs 2 A–B, 3C–D, 4B, 5B, 7B, 10</p><p>Diagnosis</p><p>Enicospilus cederbergi sp. nov. (Fig. 2 A–B) can be distinguished from other members of the E. ramidulus species group based on the relatively stout habitus and legs, the short but numerous flagellomeres, the slightly sinuate Rs&amp;M-vein, the infuscate and thickened veins in the fore wings and the infuscate margins of the pterostigma. Specimens with an infuscate tip of the metasoma are most likely to be confused with E. ramidulus Linnaeus, 1758, while the thicker antennal segments are reminiscent of E. cerebrator Aubert, 1966 . Also similar and probably closely related to E. intermedius sp. nov. but distinguishable from that species on the stouter and less numerous flagellomeres.</p><p>Etymology</p><p>The name cederbergi refers to Björn Cederberg who is devoted to popularizing Hymenoptera in Sweden.</p><p>Material examined (n = 12 ♀♀, 5 ƋƋ)</p><p>Holotype</p><p>SWEDEN: 1 ♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=16.496&amp;materialsCitation.latitude=56.698" title="Search Plazi for locations around (long 16.496/lat 56.698)">Öland</a>, Mörbylånga, Strandskogen, 56.698° N, 16.496° E, MV-light in garden close to an oak forest and sandy meadow, 24 Jun. 2016, M. Andersson leg. (NHRS-HEVA000008145 STI: NJBC155).</p><p>Paratypes</p><p>SWEDEN: 7 ♀♀, 3 ƋƋ, Skåne, Simrishamn, Järahusen, 55.411° N, 14.195° E, MV-light trap, 8– 28 Jul. 2016, N. Ryrholm and C. Källander leg. (NHRS-HEVA000008146–NHRS-HEVA000008155; 1 Ƌ NHRS-HEVA000008155 STI:NJBC157, 1 ♀, NHRS-HEVA000008150 STI:NJBC158); 1 ♀, Skåne, Ystad, Kåseberga, 55.385° N, 14.069° E, MV-light trap in semi-open coastal meadow, 11 May– 27 Jul. 2016, N. Ryrholm and C. Källander leg. (NHRS-HEVA000008156); 2 ƋƋ, Sörmland, Tyresta, Nationalparken urskogsslingan, 59.105° N, 18.148° E, Malaise trap in old semi-open coniferous forest, 2 Jul.–21 Aug. 2003 SMTP leg. (NHRS-HEVA000008157, NHRS-HEVA000008158 STI:NJBC156); 1 Ƌ, Öland, Mörbylånga, Strandskogen, 56.698° N, 16.496° E, MV-light in garden close to an oak forest and sandy meadow, 16 Jun. 2016, B. Andersson leg. (NHRS-HEVA000008159); 1 ♀, Småland, Gnosjö, Kittlakull Store mosse, 57.172° N, 15.551° E, Malaise trap in pine bog, 12 Jul. – 21 Aug. 2003, SMPT leg. (NHRS-HEVA000008160); 1 ♀, Uppland, Häverö, Västersnäs, 60.114° N, 18.616° E, MV-light trap, 4–7 Sep. 2017, N. Ryrholm and C. Källander leg. (NHRS-HEVA000008161 STI:NJBC285); 1 ♀, Skåne, Kristianstad, Äspet (Espet) Åhus, 55.925° N, 14.305° E, Jul. 1922, K. Ander leg. (MZLU Type no. 06122:2); 1 ♀, Västergötland, Habo, Brandstorp, 58.097° N, 14.206° E, 2 Jul. 1948, T.-E. Leiler leg. (NHRS-HEVA000003806); 1 ♀, Halland, Halmstad, Örnäsudden, 56.652° N, 12.806° E, 20 Jul. 1954, B.-H. Hanson leg. (NHRS-HEVA000003807).</p><p>Description</p><p>Female</p><p>Body length 18–20 mm. Fore wing length 13–15 mm. Number of flagellomeres 58–61 (mean 59.5). Mandible strongly twisted with upper tooth two times as long as lower tooth. First flagellomere relatively stout, about 3.5–4.0 times as long as apically wide. Mid- and subapical flagellomeres about 1.5 times as long as wide (Fig. 3 C–D), slightly longer than average in E. cerebrator . Head in dorsal view always with distinct gap of about 0.2 times ocellar diameter between inner orbit of compound eye and lateral ocellus (Fig. 5B). Temples buccate, in dorsal view curved, rounded immediately behind eye, usually distinctly wider than in E. adustus (Haller, 1885) and in lateral view about 0.7 times width of compound eye. Occipital carina conspicuously curved before indicated junction with hypostomal carina. Indicated angle between occipital carina and hypostomal carina slightly acute or right angled (as in Fig. 6A). Clypeus apically truncate, moderately convex, in lateral view almost right angled, sparsely punctate, interstices shining. Mesopleuron closely punctate on a polished background, centrally becoming more rugose, intermixed with transverse striae as in typical specimens of E. adustus but interstices between punctures normally wider, about equal to diameter of punctures. Epicnemial carina ventrally complete, sinuate, often indistinct or absent dorsally well before indicated joint with propleuron. Mesoscutum with notauli weakly indicated anteriorly, entirely closely punctate. Scutellum with lateral carinae, its surface with dense punctures, punctures larger than on mesoscutum. Sides of scutellum rather parallel, apically and proximally wider than in E. ramidulus . Sclerites in fore wing (Fig. 4B) reminiscent of other species in the E. ramidulus group. Proximal sclerite entirely pigmented, approximately triangular with rounded anterior angle, central sclerite semi-circular, pigmented distally, fading to unpigmented proximally. Distal sclerite generally more prominent than in other Swedish species of the E. ramidulus group, represented by a relatively distinctly pigmented crescent along distal margin of fenestra. Fore wing veins thickened, black or dark brown. Pterostigma centrally pale brownish with margins more or less infuscate. Vein Rs&amp;M distinctly sinuate, conspicuously bent at least in lower third (Fig. 4B). Vein Rs+2 r strongly thickened, distinctly sinuate. Propodeum with anterior transverse carina strong, anterior of carina rather densely punctate, posterior entirely reticulate-rugose often with faint longitudinal striae centrally. Legs usually thicker than in other members of E. ramidulus group. Hind femur about 7–8 times as long as wide. Hind metatarsus about 10 times as long as wide. Hind claws short, more strongly curved than in E. adustus but significantly less than in E. ramidulus .</p><p>Male</p><p>Size, structure and colour as in female but generally with more flagellomeres (63–64) and striation on mesopleuron feebler in studied specimens, basically absent medio-ventrally. Parameres long, in lateral view reminiscent of E. adustus (Fig. 7B).</p><p>Based on the limited material of males and the fact that the parameres often are slightly deformed due to storage and chemical treatment, no detailed description can be made of the shape of the genitalia at this stage.</p><p>Colour</p><p>Uniformly reddish brown. Inner and outer eye margins yellowish; mandibular teeth black. Metasoma usually with infuscation from 5th tergite onwards (Fig. 2), posterior tergites usually totally infuscate, black or dark brown. Ovipositor sheath of same colour as posterior metasomal segments. In some specimens infuscation of metasomal tip is partly reduced or absent. Antennae darker in apical half, terminal segment paler.</p><p>DNA barcode</p><p>The full DNA barcode sequences of five specimens of the Swedish E. cederbergi sp. nov. specimens are available at the BOLD systems database (www.boldsystems.org, BIN; BOLD:AAI5191).</p><p>Distribution</p><p>Enicospilus cederbergi sp. nov. is so far only known from Sweden. In the BOLD database there are also specimens from Germany and Israel that seem to share the genotype, but these specimens have not been studied and might refer to E. intermedius sp. nov. In Sweden it seems to be rare but widespread and known from the southern and central parts of the country (Skåne, Småland, Öland, Uppland).</p><p>Phenology</p><p>The dates on the labels of the type series indicate that the species occurs mainly during July.</p><p>Ecology</p><p>No detailed information on the biology of E. cederbergi sp. nov. is known.The habitat of known localities ranges from coniferous to deciduous forests in coastal areas, as well as an inland pine bog and semi-open calcareous heathland.</p></div>	https://treatment.plazi.org/id/A30D87CAFF9DFFCFF259F92BCFA3F894	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johansson, Niklas	Johansson, Niklas (2018): Review of the Swedish Enicospilus (Hymenoptera; Ichneumonidae; Ophioninae) with description of three new species and an illustrated key to species. European Journal of Taxonomy 483: 1-21, DOI: 10.5852/ejt.2018.483
A30D87CAFF97FFC3F251FEB5CDBBFA12.text	A30D87CAFF97FFC3F251FEB5CDBBFA12.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Enicospilus intermedius Johansson 2018	<div><p>Enicospilus intermedius sp. nov.</p><p>urn:lsid:zoobank.org:act: 08FEDAED-6F73-4BE9-BFEA-74EBD882B0E1 Figs 8, 9 A–C, 10</p><p>Diagnosis</p><p>Enicospilus intermedius sp. nov. (Fig. 8) is superficially similar to E. myricae Broad &amp; Shaw, 2016, but it is separated from that species by the larger size, the more numerous flagellomeres and the wider head in frontal view. Also very similar to E. adustus but with face wider and head more buccate behind the eyes. The face is usually entirely testaceous without yellowish areas. The only barcoded specimen shows very little genetic differentiation from E. cederbergi sp. nov. (Fig. 10) and the two species share the same BIN. Besides the differences in colour between typical specimens, the two species are distinguishable by the shape and number of the flagellomeres.</p><p>Etymology</p><p>The species is morphologically intermediate in relation to E. adustus and E. myricae .</p><p>Material examined (n = 14 ♀♀, 3 ƋƋ)</p><p>Holotype</p><p>SWEDEN: 1 ♀, Gotland, Kräklingbo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=18.726&amp;materialsCitation.latitude=57.412" title="Search Plazi for locations around (long 18.726/lat 57.412)">Torsburgen</a>, 57.412° N, 18.726° E, MV-light in semi-open, herb rich, scots pine forest, 20 Jul. 2017, J. Törnvall leg. (NHRS-HEVA000008162 STI: NJBC273).</p><p>Paratypes</p><p>SWEDEN: 1 ♀, Uppland, Rådmansö, Bergholmen, 59.750° N, 18.953° E, MV-light trap, 29 Jul.– 18 Sep. 2017, N. Ryrholm and C. Källander leg. (NHRS-HEVA000008163); 1 ♀, Gästrikland, Gävle, Grinduga, 60.638° N, 17.294° E, MV-light trap, 9–16 Jul. 2013, N. Ryrholm and C. Källander leg. (NHRS-HEVA000008164); 1 Ƌ, 2 ♀♀, Uppland, Rådmansö, Bergholmen, 59.750° N, 18.953° E, MVlight trap, 18 Jun.–28 Jul. 2017, N. Ryrholm and C. Källander leg. (NHRS-HEVA000008165–NHRS- HEVA000008167); 1 ♀, Gotland, Visby, Roleks, 57.572° N, 18.381° E, Malaise trap in semi-open, grazed scots pine forest, 10 Apr.–06 Jun. 2005, SMTP leg. (NHRS-HEVA000008168); 1 ♀, Sörmland, Huddinge, Sofielunds återvinningsstation, 59.187° N, 18.028° E, Malaise trap in industrial area, 16 Jun.– 2 Jul. 2003, SMTP leg. (NHRS-HEVA000008169); 1 ♀, Gotland, Romakloster, Stenstugu Björke, 57.514° N, 18.428° E, MV-light, 17–24 Jul. 2017, J. Törnvall leg. (NHRS-HEVA000008170); 1 ♀, Gotland, Hamra, Suders, 56.940° N, 18.303° E, MV-light trap, 21 Apr.–25 May 2007, N. Ryrholm and C. Källander leg. (NHRS-HEVA000008171).</p><p>Description</p><p>Female</p><p>Body length 19–21 mm. Fore wing length 15–16 mm. Number of flagellomeres 62–66 (mean 64). Mandible strongly twisted with upper tooth about two times as long as lower tooth. First flagellomere slender, about 4.5–5.0 times as long as apically wide. Mid- and preapical flagellomeres about 1.7 times as long as wide, shape of flagellomeres similar to E. adustus . Head in dorsal view usually with small gap of about 0.1 times ocellar diameter between inner orbit of compound eye and lateral ocellus, distinctly narrower than in E. myricae . Ocelli larger than in E. myricae . Temples buccate, in dorsal view curved, rounded immediately behind eye, distinctly wider than in E. adustus and in lateral view about 0.7 times width of compound eye (Fig. 9C). Occipital carina conspicuously curved before indicated junction with hypostomal carina. Indicated angle between occipital carina and hypostomal carina slightly acute or right angled (as in Fig. 6A). Clypeus apically truncate, moderately convex, in lateral view almost right angled, sparsely punctate, interstices shining. Mesopleuron closely punctate on a polished background, centrally becoming more rugose, intermixed with transverse striae. Epicnemial carina ventrally complete, sinuate, often indistinct or absent dorsally well before indicated joint with propleuron. Mesoscutum with notauli weakly indicated anteriorly, entirely closely punctate. Scutellum with lateral carinae, its surface with dense punctures, punctures larger than on mesoscutum. Sclerites in fore wing identical to E. adustus (Fig. 4D) and E. myricae . Proximal sclerite entirely pigmented, approximately triangular with rounded anterior angle, central sclerite semi-circular, pigmented distally, fading to unpigmented proximally. Distal sclerite quite strong and elongate. Fore wing veins and pterostigma testaceous. Propodeum with anterior transverse carina strong, anterior of the carina rather densely punctate, posterior entirely reticulate-rugose often with faint longitudinal striae centrally. Legs of same proportions as in E. adustus .</p><p>Male</p><p>Size, structure and colour as in female. Antenna with slightly more numerous flagellomeres than in the female (65–67). Parameres long, in lateral view reminiscent of E. adustus (as in Fig. 7B). Based on the limited material of males and the fact that the parameres are often slightly deformed due to storage and chemical treatment, no detailed description can be made of the shape of the genitalia at this stage.</p><p>Colour</p><p>Uniformly testaceous. Inner and outer orbits usually only slightly paler than face (one male from Croatia with the undulation of the inner margin of compound eye and outer margin yellow); mandibular teeth black. Apical metasomal segments sometimes slightly infuscate. Ovipositor sheath usually darker than apical metasomal segments. Antennae slightly darker in apical half, but not as distinct as in typical specimens of E. myricae .</p><p>DNA barcode</p><p>The full DNA barcode sequence of one Swedish ♀ of E. intermedius sp. nov. is available at the BOLD systems database (www.boldsystems.org, BIN; BOLD:AAI5191).</p><p>Distribution</p><p>Enicospilus intermedius sp. nov. is only known from the eastern parts of Central Sweden, including Gotland (Södermanland, Uppland, Gästrikland, Gotland), and Croatia (Sibensko Kninska).</p><p>Phenology</p><p>The dates on the labels of the type series range from late May to August. The main flight period is probably during June and July. In Southern Europe the species seems to be active during April and May.</p><p>Ecology</p><p>No detailed information on the biology of E. intermedius sp. nov. is known. The habitat in Sweden seems to mainly consist of semi-open xerothermic pine forests.</p></div>	https://treatment.plazi.org/id/A30D87CAFF97FFC3F251FEB5CDBBFA12	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johansson, Niklas	Johansson, Niklas (2018): Review of the Swedish Enicospilus (Hymenoptera; Ichneumonidae; Ophioninae) with description of three new species and an illustrated key to species. European Journal of Taxonomy 483: 1-21, DOI: 10.5852/ejt.2018.483
A30D87CAFF94FFC6F26FF995CB70FA7C.text	A30D87CAFF94FFC6F26FF995CB70FA7C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Enicospilus ryrholmi Johansson 2018	<div><p>Enicospilus ryrholmi sp. nov.</p><p>urn:lsid:zoobank.org:act: 75A2F3F0-0F8E-419A-A2D9-DD55298C7534 Figs 3 G–H, 4C, 6B, 10, 11, 12A</p><p>Diagnosis</p><p>Enicospilus ryrholmi sp. nov. can be distinguished from other members of the E. ramidulus species group by the position and shape of the central sclerite; the smaller size on average; the limited number of flagellomeres; the elongate central flagellomeres; the more or less straight mesopleural part of the epicnemial carina and the less curved lower part of the occipital carina. It is most likely to be confused with small specimens of E. adustus .</p><p>Etymology</p><p>The name ryrholmi refers to the lepidopterologist Nils Ryrholm who, by sorting out and donating a large portion of Ophioninae from decades of sampling with MV-light traps, has contributed greatly to the taxonomy and knowledge of Scandinavian Enicospilus .</p><p>Material examined (n = 15 ♀♀, 2 ƋƋ)</p><p>Holotype</p><p>SWEDEN: 1 ♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=16.496&amp;materialsCitation.latitude=56.698" title="Search Plazi for locations around (long 16.496/lat 56.698)">Öland</a>, Mörbylånga, Strandskogen, 56.698° N, 16.496° E, MV-light in garden close to an oak forest and sandy meadows, 23 Jul. 2016, M. Andersson leg. (NHRS-HEVA000008172 STI: NJBC159).</p><p>Paratypes</p><p>SWEDEN: 3 ♀♀, Skåne, Klippan, Bonnarpshed, 56.087° N, 13.180° E, MV-light trap in open dry meadow, 9 Jul.–4 Aug. 2007, N. Ryrholm and C. Källander leg. (NHRS-HEVA000008173- NHRS-HEVA000008175; 1 ♀ NHRS-HEVA000008175 STI:NJBC160); 2 ♀♀, Skåne, Höganäs, Kullen, 56.293° N, 12.487° E, 19 Jul. 1975, C.H. Lindroth leg. (MZLU Type no. 06123:2-3); 1 ♀, Skåne, Höganäs, Kullen, 56.293° N, 12.487° E, 2 Aug. 1975, C.H. Lindroth leg. (MZLU Type no. 06123:4); 1 ♀, Öland, Mörbylånga, Räpplinge, 56.827° N, 16.660° E, alvar/calcareous dry meadow, 26 Jul. 1980, L.-Å. Janzon leg. (MZLU Type no. 06123:5); 2 ♀♀, Småland, Sävsjö, Södra Hägnen, 57.379° N, 14.665° E, MV-light in open dry industrial area, 16 Aug. 2017, R. Isaksson leg. (NHRS- HEVA000008176, NHRS-HEVA000008177); 1 ♀, Öland, Torslunda, Arontorp 56.637° N, 16.516° E, MV-light, 9 Jul. 2017, T. Lindberg leg. (NHRS-HEVA000008178 STI:NJBC288); 1 ♀, Gotland, Hamra, Tuvlandet, 56.966° N, 18.308° E, MV-light trap, 15 Jul.–18 Aug. 2017, N. Ryrholm and C. Källander leg. (NHRS-HEVA000008179 STI:NJBC287); 1 ♀, Öland, Borgholm, 56.880° N, 16.656° E, 19 Jul. 1964, S. Johansson leg. (MZLU Type no. 06123:7); 1 ♀, Öland, Borgholm, 56.880° N, 16.656° E, 20 Aug. 1962, S. Johansson leg. (MZLU Type no. 06123:8); 1 Ƌ, Skåne, Höganäs, Kullen, 56.293° N, 12.487° E, 28 Jun. 1975, C.H. Lindroth leg. (MZLU Type no. 06123:6); 1 ♀, Öland, Mörbylånga, Hagaberg Frö, 56.575° N, 16.448° E, 23 Jul. 1978, S. Johansson leg. (MZLU Type no. 06123:9); 1 Ƌ, Öland, Mörbylånga, Karlsro, 56.579° N, 16.421° E, 13 Jul. 1978, sweepnet in daylight, S. Johansson leg. (MZLU Type no. 06123:10).</p><p>Description</p><p>Female</p><p>Body length 16–17 mm, fore wing length 12–13 mm. Number of flagellomeres 51–56 (mean 54). Mandible strongly twisted with upper tooth distinctly longer than lower tooth, usually slightly more than two times as long as lower tooth. First flagellomere very slender, about 5 times as long as apically wide. Central- and subapical flagellomeres slender, 2.0–2.1 times as long as wide (Fig. 3 G–H). Head in dorsal view with small gap of about 0.1 times ocellar diameter between inner margin of compound eye and lateral ocellus. Temples in dorsal view narrowed behind eye as in typical specimens of E. adustus and in lateral view about 0.5 times the width of compound eye. Occipital carina only slightly curved before junction with hypostomal carina (Fig. 6B), joint sometimes indistinct or absent. Indicated angle between occipital carina and hypostomal carina acute, about 45 degrees. Clypeus apically truncate, convex in lateral view, very sparsely punctate. Mesopleuron punctate, weakly sculptured or polished, central punctures intermixed with vague transverse striae. Distance between punctures about two times diameter of punctures. Speculum without punctures, strongly polished. Epicnemial carina between pleurosternal angles and sternal part almost straight (Fig. 12A). Mesoscutum with notauli faintly indicated anteriorly, very sparsely and vaguely punctate. Sides of scutellum strongly converging posteriorly. Proximal sclerite entirely pigmented, approximately triangular with rounded anterior angle. Central sclerite varying from almost unpigmented, reminiscent in shape of E. merdarius (Gravenhorst, 1829) but narrowly pigmented distally to more distinctly sclerotized and elongate. Distance between the proximal and central sclerites often smaller than in E. adustus and E. cerebrator (Fig. 4C). Fore wing veins thin, pale brownish. Pterostigma pale. Propodeum with anterior transverse carina strong, anterior of this rather densely punctate, posterior to this entirely reticulate-rugose often with faint longitudinal striae centrally. Proportion of legs as in E. adustus . Hind femur about 10 times as long as wide. Hind metatarsus about 12 times as long as wide.</p><p>Male</p><p>Size, structure and colour as in female. Parameres in lateral view short and obtuse as in E. ramidulus (Fig. 7A).</p><p>Colour</p><p>Uniformly testaceous. Metasoma sometimes slighty infuscate posteriorly. Mandibular teeth black.</p><p>DNA barcode</p><p>The full DNA barcode sequences of four of the Swedish E. ryrholmi sp. nov. specimens are available at the BOLD systems database (www.boldsystems.org, BIN;BOLD:ADF8803).</p><p>Distribution</p><p>Enicospilus ryrholmi sp. nov. is so far only known from Sweden where it seems to be rare but widespread at least in the southern part of the country, including the Baltic islands Öland and Gotland (Skåne, Blekinge, Småland, Öland, Gotland). It can, however, be expected to occur over a wider geographical range.</p><p>Phenology</p><p>The species occurs in late summer. The documented flight period is July to August.</p><p>Ecology</p><p>No detailed information on the biology of E. ryrholmi sp. nov. is known. The habitat consists mainly of semi-open areas ranging from alvar to rocky slopes, sandy heaths, industrial landscapes and gardens. The known localities might suggest a host connected to plants depending on open sandy or rocky grounds.</p></div>	https://treatment.plazi.org/id/A30D87CAFF94FFC6F26FF995CB70FA7C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johansson, Niklas	Johansson, Niklas (2018): Review of the Swedish Enicospilus (Hymenoptera; Ichneumonidae; Ophioninae) with description of three new species and an illustrated key to species. European Journal of Taxonomy 483: 1-21, DOI: 10.5852/ejt.2018.483
A30D87CAFF8FFFDBF0BBFB49CB6FFE29.text	A30D87CAFF8FFFDBF0BBFB49CB6FFE29.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Enicospilus Stephens 1835	<div><p>Identification key to the Swedish species of Enicospilus</p><p>1. Fore wing lacking sclerites in glabrous area of discosubmarginal cell (Fig. 13C); vein Rs+2 r conspicuously curved before junction with pterostigma; large species, wing length about 20 mm .................................................................................................. E. inflexus (Ratzeburg, 1844) [For separation between E. inflexus (Ratzeburg, 1844) and E. undulatus (Gravenhorst, 1829), see Broad &amp; Shaw 2016.]</p><p>– Fore wing with at least one distinct sclerite in discosubmarginal cell (Figs 4 A–D, 13A–B); vein Rs+2 r slightly sinuate before junction with pterostigma; smaller species, wing length usually less than 16 mm ....................................................................................................................................... 2</p><p>2. Fore wing lacking any trace of central sclerite; distal sclerite very weak or absent (Fig. 13A); clypeus in lateral view flattened (Fig. 14B) ................................................. E. repentinus (Holmgren, 1860)</p><p>– Fore wing with central sclerite present, but sometimes completely translucent (Figs 4 A–D, 13B); clypeus in lateral view distinctly convex (Fig. 14A) ( E. ramidulus species group) ......................... 3</p><p>3. Fore wing with central sclerite completely translucent (Fig. 13B) ..................................................... ..................................................................................................... E. merdarius (Gravenhorst, 1829) – Fore wing with central sclerite distinctly pigmented (Fig. 4 A–D) ................................................... 4</p><p>4. Antenna with central and apical flagellomeres very slender with central and apical segments more than 2 times as long as wide (Fig. 3 G–H); number of flagellomeres 53–56; occipital carina only slightly curved before indicated junction with hypostomal carina (Fig. 6B); epicnemial carina between mesopleural angles and sternal part almost straight (Fig. 12A); central sclerite in glabrous area in fore wing usually more circular in shape, largely unpigmented; central and proximal sclerite closer to each other (Fig. 4C). Small species, fore wing length 12–13 mm ....... E. ryrholmi sp. nov.</p><p>– Antenna with central and apical flagellomeres at most 1.8 times as long as wide (Fig. 3 A–F); occipital carina distinctly curved before indicated junction with hypostomal carina, (Fig. 6A); epicnemial carina between mesopleural angles and sternal part sinuous (Fig. 12B); central sclerite in glabrous area in fore wing more semi-ovoid in shape, often elongate; central and proximal sclerites further apart (Fig. 4 A–B, D) ......................................................................................................................... 5</p><p>5. Mesosoma usually with extensive dark brown patches; central sclerite in fore wing mostly narrow, at most as long as wide (Fig. 4A); distance between central and proximal sclerites usually distinctly longer than basal side of proximal sclerite; central sclerite mostly entirely pigmented .................................................................................... E. combustus (Gravenhorst, 1829)</p><p>– Mesosoma lacking distinct dark patches, uniformly testaceous; central sclerite usually longer than wide (Fig. 4 B–C); distance between central and proximal sclerites almost equal to basal side of proximal sclerite; central sclerite mostly largely unpigmented proximally ..................................... 6</p><p>6. Antenna with central and preapical flagellomeres shorter, at most 1.5 times as long as wide (Fig. 3 A–D) ....................................................................................................................................... 7</p><p>– Antenna with central and preapical flagellomeres longer, at least 1.7 times as long as wide (Fig. 3 E–F) ........................................................................................................................................ 8</p><p>7. Antenna with 51–56 flagellomeres; temples very strongly narrowed behind eyes, head with no gap between eye and lateral ocelli (Fig. 5A); central and apical flagellomeres about 1.3 times as long as wide (Fig. 3 A–B) ..................................................................................... E. cerebrator Aubert, 1966</p><p>– Antenna with 59–62 flagellomeres; temples strongly buccate, head with distinct gap between ocelli and eye (Fig. 5B); central and apical flagellomeres about 1.5 times as long as wide (Fig. 3 C–D) ...................................................................................................... E. cederbergi sp. nov.</p><p>8. Metasoma in female abruptly black-tipped from the 5th (in females) or 6th (in males) tergite; hind tarsal claws in female conspicuously curved (Fig. 15A).................... E. ramidulus (Linnaeus, 1758)</p><p>– Metasoma never abruptly black-tipped; hind tarsal claws in female not conspicuously curved (Fig. 15B) .......................................................................................................................................... 9</p><p>9. Head in lateral view with temples narrow, at most about 0.4 times the width of compound eye (Fig. 9I); face generally with extensive yellow markings (Fig. 9 G–H); face narrow in anterior view (Fig. 9 G–H); lateral ocellus touching compound eye ................................. E. adustus (Haller, 1885)</p><p>– Head in lateral view with temples wide, about 0.7 times the width of compound eye (Fig. 9C, F); face generally more testaceous; face wider in anterior view (Fig. 9 A–B, D–E); lateral ocelli often with more or less distinct gap between lateral ocellus and compound eye .................................... 10</p><p>10. Number of flagellomeres 56–59; head in anterior view more rounded (Fig. 9 D–E); ocelli small, gap between lateral ocelli and inner margin of compound eye wide, about 0.2–0.3 times the diameter of ocellus ............................................................................................. E. myricae Broad &amp; Shaw, 2016</p><p>– Number of flagellomeres 62–67; head in anterior view distinctly transverse (Fig. 9 A–B); ocelli large, gap between lateral ocelli and inner margin of compound eye narrow, about 0.1 times the diameter of ocellus .......................................................................................... E. intermedius sp. nov</p></div>	https://treatment.plazi.org/id/A30D87CAFF8FFFDBF0BBFB49CB6FFE29	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Johansson, Niklas	Johansson, Niklas (2018): Review of the Swedish Enicospilus (Hymenoptera; Ichneumonidae; Ophioninae) with description of three new species and an illustrated key to species. European Journal of Taxonomy 483: 1-21, DOI: 10.5852/ejt.2018.483
