taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
A24087A73966042AFF7FFF51FAD2FEEC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172584/files/figure.png	https://doi.org/10.5281/zenodo.5172584	Figure 1. Habitus of Tomarus spp. a) Tomarus bidentulus (Fairmaire) dorsal view. b) Tomarus burmeisteri (Steinheil) dorsal view. c) Tomarus gyas Erichson dorsal view. d) Tomarus rosettae (Endrödi) dorsal view. e) Tomarus roigjunenti n. sp. dorsal view, holotype. f) Tomarus roigjunenti n. sp. dorsal view, allotype.	Figure 1. Habitus of Tomarus spp. a) Tomarus bidentulus (Fairmaire) dorsal view. b) Tomarus burmeisteri (Steinheil) dorsal view. c) Tomarus gyas Erichson dorsal view. d) Tomarus rosettae (Endrödi) dorsal view. e) Tomarus roigjunenti n. sp. dorsal view, holotype. f) Tomarus roigjunenti n. sp. dorsal view, allotype.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73966042AFF7FFF51FAD2FEEC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172588/files/figure.png	https://doi.org/10.5281/zenodo.5172588	Figure 2. Habitus of Tomarus spp. a) Tomarus rosettae (Endrödi) dorsal view. b) Tomarus spinipenis n. sp. dorsal view, holotype. c) Tomarus spinipenis dorsal view, allotype. d) Tomarus burmeisteri (Steinheil) dorsal view. e) Tomarus villosus (Burmeister) ventral view. f) Tomarus gyas ventral view. g) Tomarus bidentulus (Fairmaire) abdomen segments, ventral view (female). h) Tomarus bidentulus abdomen segments, ventral view (male).	Figure 2. Habitus of Tomarus spp. a) Tomarus rosettae (Endrödi) dorsal view. b) Tomarus spinipenis n. sp. dorsal view, holotype. c) Tomarus spinipenis dorsal view, allotype. d) Tomarus burmeisteri (Steinheil) dorsal view. e) Tomarus villosus (Burmeister) ventral view. f) Tomarus gyas ventral view. g) Tomarus bidentulus (Fairmaire) abdomen segments, ventral view (female). h) Tomarus bidentulus abdomen segments, ventral view (male).	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73966042AFF7FFF51FAD2FEEC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172591/files/figure.png	https://doi.org/10.5281/zenodo.5172591	Figure 3. Types of Tomarus spp. a) T. bidentulus female holotype, dorsal view. b) T. burmeisteri male holotype, dorsal view. c) T. burmeisteri male holotype, lateral view. d) T. burmeisteri male holotype parameres, caudal view. e) T. burmeisteri male holotype parameres (lateral view). f) Label for T. burmeisteri holotype. g) T. gyas male holotype, dorsal view and labels and parameres, caudal view). h) T. maimon holotype, dorsal view. i) T. rosettae male holotype, dorsal view. j) T. villosus male holotype, dorsal view. k) T. villosus male holotype, dorsal view of head. l) T. villosus male holotype, parameres, caudal view. m) Label for T. villosus holotype.	Figure 3. Types of Tomarus spp. a) T. bidentulus female holotype, dorsal view. b) T. burmeisteri male holotype, dorsal view. c) T. burmeisteri male holotype, lateral view. d) T. burmeisteri male holotype parameres, caudal view. e) T. burmeisteri male holotype parameres (lateral view). f) Label for T. burmeisteri holotype. g) T. gyas male holotype, dorsal view and labels and parameres, caudal view). h) T. maimon holotype, dorsal view. i) T. rosettae male holotype, dorsal view. j) T. villosus male holotype, dorsal view. k) T. villosus male holotype, dorsal view of head. l) T. villosus male holotype, parameres, caudal view. m) Label for T. villosus holotype.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73966042AFF7FFF51FAD2FEEC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172595/files/figure.png	https://doi.org/10.5281/zenodo.5172595	Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73966042AFF7FFF51FAD2FEEC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172599/files/figure.png	https://doi.org/10.5281/zenodo.5172599	Figure 5. Maxilla, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 5. Maxilla, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73966042AFF7FFF51FAD2FEEC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172601/files/figure.png	https://doi.org/10.5281/zenodo.5172601	Figure 6. Mandibles, ventral view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	Figure 6. Mandibles, ventral view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73966042AFF7FFF51FAD2FEEC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172605/files/figure.png	https://doi.org/10.5281/zenodo.5172605	Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73966042AFF7FFF51FAD2FEEC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172607/files/figure.png	https://doi.org/10.5281/zenodo.5172607	Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73966042AFF7FFF51FAD2FEEC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172614/files/figure.png	https://doi.org/10.5281/zenodo.5172614	Figure 10. Meso- and metatibia, dorsal view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	Figure 10. Meso- and metatibia, dorsal view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73966042AFF7FFF51FAD2FEEC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172618/files/figure.png	https://doi.org/10.5281/zenodo.5172618	Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73966042AFF7FFF51FAD2FEEC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172620/files/figure.png	https://doi.org/10.5281/zenodo.5172620	Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73966042AFF7FFF51FAD2FEEC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172622/files/figure.png	https://doi.org/10.5281/zenodo.5172622	Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73966042AFF7FFF51FAD2FEEC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172626/files/figure.png	https://doi.org/10.5281/zenodo.5172626	Figure 14. Gonocoxite, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (Gx) gonocoxite), (Gs) gonocoxal sternite.	Figure 14. Gonocoxite, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (Gx) gonocoxite), (Gs) gonocoxal sternite.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73966042AFF7FFF51FAD2FEEC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172630/files/figure.png	https://doi.org/10.5281/zenodo.5172630	Figure 15. Internal sac of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (AL) accessory lamella, (CL) copulatory lamella (CL), (Ra) raspula, (Esp) speculum, and (Fg) a complex, spike-like flagellum that varies in length and thickness.	Figure 15. Internal sac of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (AL) accessory lamella, (CL) copulatory lamella (CL), (Ra) raspula, (Esp) speculum, and (Fg) a complex, spike-like flagellum that varies in length and thickness.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73966042AFF7FFF51FAD2FEEC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172632/files/figure.png	https://doi.org/10.5281/zenodo.5172632	Figure 16. Distribution of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. maimon. d) T. gyas.	Figure 16. Distribution of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. maimon. d) T. gyas.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73965042BFF7FFE51FD7EFC6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172584/files/figure.png	https://doi.org/10.5281/zenodo.5172584	Figure 1. Habitus of Tomarus spp. a) Tomarus bidentulus (Fairmaire) dorsal view. b) Tomarus burmeisteri (Steinheil) dorsal view. c) Tomarus gyas Erichson dorsal view. d) Tomarus rosettae (Endrödi) dorsal view. e) Tomarus roigjunenti n. sp. dorsal view, holotype. f) Tomarus roigjunenti n. sp. dorsal view, allotype.	Figure 1. Habitus of Tomarus spp. a) Tomarus bidentulus (Fairmaire) dorsal view. b) Tomarus burmeisteri (Steinheil) dorsal view. c) Tomarus gyas Erichson dorsal view. d) Tomarus rosettae (Endrödi) dorsal view. e) Tomarus roigjunenti n. sp. dorsal view, holotype. f) Tomarus roigjunenti n. sp. dorsal view, allotype.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73965042BFF7FFE51FD7EFC6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172588/files/figure.png	https://doi.org/10.5281/zenodo.5172588	Figure 2. Habitus of Tomarus spp. a) Tomarus rosettae (Endrödi) dorsal view. b) Tomarus spinipenis n. sp. dorsal view, holotype. c) Tomarus spinipenis dorsal view, allotype. d) Tomarus burmeisteri (Steinheil) dorsal view. e) Tomarus villosus (Burmeister) ventral view. f) Tomarus gyas ventral view. g) Tomarus bidentulus (Fairmaire) abdomen segments, ventral view (female). h) Tomarus bidentulus abdomen segments, ventral view (male).	Figure 2. Habitus of Tomarus spp. a) Tomarus rosettae (Endrödi) dorsal view. b) Tomarus spinipenis n. sp. dorsal view, holotype. c) Tomarus spinipenis dorsal view, allotype. d) Tomarus burmeisteri (Steinheil) dorsal view. e) Tomarus villosus (Burmeister) ventral view. f) Tomarus gyas ventral view. g) Tomarus bidentulus (Fairmaire) abdomen segments, ventral view (female). h) Tomarus bidentulus abdomen segments, ventral view (male).	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73965042BFF7FFE51FD7EFC6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172591/files/figure.png	https://doi.org/10.5281/zenodo.5172591	Figure 3. Types of Tomarus spp. a) T. bidentulus female holotype, dorsal view. b) T. burmeisteri male holotype, dorsal view. c) T. burmeisteri male holotype, lateral view. d) T. burmeisteri male holotype parameres, caudal view. e) T. burmeisteri male holotype parameres (lateral view). f) Label for T. burmeisteri holotype. g) T. gyas male holotype, dorsal view and labels and parameres, caudal view). h) T. maimon holotype, dorsal view. i) T. rosettae male holotype, dorsal view. j) T. villosus male holotype, dorsal view. k) T. villosus male holotype, dorsal view of head. l) T. villosus male holotype, parameres, caudal view. m) Label for T. villosus holotype.	Figure 3. Types of Tomarus spp. a) T. bidentulus female holotype, dorsal view. b) T. burmeisteri male holotype, dorsal view. c) T. burmeisteri male holotype, lateral view. d) T. burmeisteri male holotype parameres, caudal view. e) T. burmeisteri male holotype parameres (lateral view). f) Label for T. burmeisteri holotype. g) T. gyas male holotype, dorsal view and labels and parameres, caudal view). h) T. maimon holotype, dorsal view. i) T. rosettae male holotype, dorsal view. j) T. villosus male holotype, dorsal view. k) T. villosus male holotype, dorsal view of head. l) T. villosus male holotype, parameres, caudal view. m) Label for T. villosus holotype.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73965042BFF7FFE51FD7EFC6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172595/files/figure.png	https://doi.org/10.5281/zenodo.5172595	Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73965042BFF7FFE51FD7EFC6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172599/files/figure.png	https://doi.org/10.5281/zenodo.5172599	Figure 5. Maxilla, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 5. Maxilla, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73965042BFF7FFE51FD7EFC6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172601/files/figure.png	https://doi.org/10.5281/zenodo.5172601	Figure 6. Mandibles, ventral view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	Figure 6. Mandibles, ventral view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73965042BFF7FFE51FD7EFC6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172605/files/figure.png	https://doi.org/10.5281/zenodo.5172605	Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73965042BFF7FFE51FD7EFC6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172607/files/figure.png	https://doi.org/10.5281/zenodo.5172607	Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73965042BFF7FFE51FD7EFC6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172614/files/figure.png	https://doi.org/10.5281/zenodo.5172614	Figure 10. Meso- and metatibia, dorsal view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	Figure 10. Meso- and metatibia, dorsal view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73965042BFF7FFE51FD7EFC6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172618/files/figure.png	https://doi.org/10.5281/zenodo.5172618	Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73965042BFF7FFE51FD7EFC6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172620/files/figure.png	https://doi.org/10.5281/zenodo.5172620	Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73965042BFF7FFE51FD7EFC6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172622/files/figure.png	https://doi.org/10.5281/zenodo.5172622	Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73965042BFF7FFE51FD7EFC6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172626/files/figure.png	https://doi.org/10.5281/zenodo.5172626	Figure 14. Gonocoxite, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (Gx) gonocoxite), (Gs) gonocoxal sternite.	Figure 14. Gonocoxite, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (Gx) gonocoxite), (Gs) gonocoxal sternite.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73965042BFF7FFE51FD7EFC6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172630/files/figure.png	https://doi.org/10.5281/zenodo.5172630	Figure 15. Internal sac of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (AL) accessory lamella, (CL) copulatory lamella (CL), (Ra) raspula, (Esp) speculum, and (Fg) a complex, spike-like flagellum that varies in length and thickness.	Figure 15. Internal sac of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (AL) accessory lamella, (CL) copulatory lamella (CL), (Ra) raspula, (Esp) speculum, and (Fg) a complex, spike-like flagellum that varies in length and thickness.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73965042BFF7FFE51FD7EFC6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172632/files/figure.png	https://doi.org/10.5281/zenodo.5172632	Figure 16. Distribution of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. maimon. d) T. gyas.	Figure 16. Distribution of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. maimon. d) T. gyas.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A73965042BFF7FFE51FD7EFC6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172638/files/figure.png	https://doi.org/10.5281/zenodo.5172638	Figure 18. Variation of the parameres, anterior view, of T. burmeisteri in Argentina.	Figure 18. Variation of the parameres, anterior view, of T. burmeisteri in Argentina.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739640424FF7FFBD1FCDFFA8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172584/files/figure.png	https://doi.org/10.5281/zenodo.5172584	Figure 1. Habitus of Tomarus spp. a) Tomarus bidentulus (Fairmaire) dorsal view. b) Tomarus burmeisteri (Steinheil) dorsal view. c) Tomarus gyas Erichson dorsal view. d) Tomarus rosettae (Endrödi) dorsal view. e) Tomarus roigjunenti n. sp. dorsal view, holotype. f) Tomarus roigjunenti n. sp. dorsal view, allotype.	Figure 1. Habitus of Tomarus spp. a) Tomarus bidentulus (Fairmaire) dorsal view. b) Tomarus burmeisteri (Steinheil) dorsal view. c) Tomarus gyas Erichson dorsal view. d) Tomarus rosettae (Endrödi) dorsal view. e) Tomarus roigjunenti n. sp. dorsal view, holotype. f) Tomarus roigjunenti n. sp. dorsal view, allotype.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739640424FF7FFBD1FCDFFA8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172588/files/figure.png	https://doi.org/10.5281/zenodo.5172588	Figure 2. Habitus of Tomarus spp. a) Tomarus rosettae (Endrödi) dorsal view. b) Tomarus spinipenis n. sp. dorsal view, holotype. c) Tomarus spinipenis dorsal view, allotype. d) Tomarus burmeisteri (Steinheil) dorsal view. e) Tomarus villosus (Burmeister) ventral view. f) Tomarus gyas ventral view. g) Tomarus bidentulus (Fairmaire) abdomen segments, ventral view (female). h) Tomarus bidentulus abdomen segments, ventral view (male).	Figure 2. Habitus of Tomarus spp. a) Tomarus rosettae (Endrödi) dorsal view. b) Tomarus spinipenis n. sp. dorsal view, holotype. c) Tomarus spinipenis dorsal view, allotype. d) Tomarus burmeisteri (Steinheil) dorsal view. e) Tomarus villosus (Burmeister) ventral view. f) Tomarus gyas ventral view. g) Tomarus bidentulus (Fairmaire) abdomen segments, ventral view (female). h) Tomarus bidentulus abdomen segments, ventral view (male).	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739640424FF7FFBD1FCDFFA8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172591/files/figure.png	https://doi.org/10.5281/zenodo.5172591	Figure 3. Types of Tomarus spp. a) T. bidentulus female holotype, dorsal view. b) T. burmeisteri male holotype, dorsal view. c) T. burmeisteri male holotype, lateral view. d) T. burmeisteri male holotype parameres, caudal view. e) T. burmeisteri male holotype parameres (lateral view). f) Label for T. burmeisteri holotype. g) T. gyas male holotype, dorsal view and labels and parameres, caudal view). h) T. maimon holotype, dorsal view. i) T. rosettae male holotype, dorsal view. j) T. villosus male holotype, dorsal view. k) T. villosus male holotype, dorsal view of head. l) T. villosus male holotype, parameres, caudal view. m) Label for T. villosus holotype.	Figure 3. Types of Tomarus spp. a) T. bidentulus female holotype, dorsal view. b) T. burmeisteri male holotype, dorsal view. c) T. burmeisteri male holotype, lateral view. d) T. burmeisteri male holotype parameres, caudal view. e) T. burmeisteri male holotype parameres (lateral view). f) Label for T. burmeisteri holotype. g) T. gyas male holotype, dorsal view and labels and parameres, caudal view). h) T. maimon holotype, dorsal view. i) T. rosettae male holotype, dorsal view. j) T. villosus male holotype, dorsal view. k) T. villosus male holotype, dorsal view of head. l) T. villosus male holotype, parameres, caudal view. m) Label for T. villosus holotype.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739640424FF7FFBD1FCDFFA8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172595/files/figure.png	https://doi.org/10.5281/zenodo.5172595	Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739640424FF7FFBD1FCDFFA8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172599/files/figure.png	https://doi.org/10.5281/zenodo.5172599	Figure 5. Maxilla, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 5. Maxilla, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739640424FF7FFBD1FCDFFA8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172601/files/figure.png	https://doi.org/10.5281/zenodo.5172601	Figure 6. Mandibles, ventral view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	Figure 6. Mandibles, ventral view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739640424FF7FFBD1FCDFFA8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172605/files/figure.png	https://doi.org/10.5281/zenodo.5172605	Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739640424FF7FFBD1FCDFFA8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172607/files/figure.png	https://doi.org/10.5281/zenodo.5172607	Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739640424FF7FFBD1FCDFFA8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172612/files/figure.png	https://doi.org/10.5281/zenodo.5172612	Figure 9. Protibia, dorsal view, of Tomarus spp. a) T. gyas and b) T. maimon.	Figure 9. Protibia, dorsal view, of Tomarus spp. a) T. gyas and b) T. maimon.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739640424FF7FFBD1FCDFFA8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172614/files/figure.png	https://doi.org/10.5281/zenodo.5172614	Figure 10. Meso- and metatibia, dorsal view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	Figure 10. Meso- and metatibia, dorsal view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739640424FF7FFBD1FCDFFA8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172618/files/figure.png	https://doi.org/10.5281/zenodo.5172618	Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739640424FF7FFBD1FCDFFA8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172620/files/figure.png	https://doi.org/10.5281/zenodo.5172620	Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739640424FF7FFBD1FCDFFA8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172622/files/figure.png	https://doi.org/10.5281/zenodo.5172622	Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739640424FF7FFBD1FCDFFA8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172626/files/figure.png	https://doi.org/10.5281/zenodo.5172626	Figure 14. Gonocoxite, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (Gx) gonocoxite), (Gs) gonocoxal sternite.	Figure 14. Gonocoxite, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (Gx) gonocoxite), (Gs) gonocoxal sternite.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739640424FF7FFBD1FCDFFA8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172630/files/figure.png	https://doi.org/10.5281/zenodo.5172630	Figure 15. Internal sac of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (AL) accessory lamella, (CL) copulatory lamella (CL), (Ra) raspula, (Esp) speculum, and (Fg) a complex, spike-like flagellum that varies in length and thickness.	Figure 15. Internal sac of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (AL) accessory lamella, (CL) copulatory lamella (CL), (Ra) raspula, (Esp) speculum, and (Fg) a complex, spike-like flagellum that varies in length and thickness.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739640424FF7FFBD1FCDFFA8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172632/files/figure.png	https://doi.org/10.5281/zenodo.5172632	Figure 16. Distribution of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. maimon. d) T. gyas.	Figure 16. Distribution of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. maimon. d) T. gyas.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396B0425FF7FFAF1FE4CFDCC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172591/files/figure.png	https://doi.org/10.5281/zenodo.5172591	Figure 3. Types of Tomarus spp. a) T. bidentulus female holotype, dorsal view. b) T. burmeisteri male holotype, dorsal view. c) T. burmeisteri male holotype, lateral view. d) T. burmeisteri male holotype parameres, caudal view. e) T. burmeisteri male holotype parameres (lateral view). f) Label for T. burmeisteri holotype. g) T. gyas male holotype, dorsal view and labels and parameres, caudal view). h) T. maimon holotype, dorsal view. i) T. rosettae male holotype, dorsal view. j) T. villosus male holotype, dorsal view. k) T. villosus male holotype, dorsal view of head. l) T. villosus male holotype, parameres, caudal view. m) Label for T. villosus holotype.	Figure 3. Types of Tomarus spp. a) T. bidentulus female holotype, dorsal view. b) T. burmeisteri male holotype, dorsal view. c) T. burmeisteri male holotype, lateral view. d) T. burmeisteri male holotype parameres, caudal view. e) T. burmeisteri male holotype parameres (lateral view). f) Label for T. burmeisteri holotype. g) T. gyas male holotype, dorsal view and labels and parameres, caudal view). h) T. maimon holotype, dorsal view. i) T. rosettae male holotype, dorsal view. j) T. villosus male holotype, dorsal view. k) T. villosus male holotype, dorsal view of head. l) T. villosus male holotype, parameres, caudal view. m) Label for T. villosus holotype.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396B0425FF7FFAF1FE4CFDCC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172595/files/figure.png	https://doi.org/10.5281/zenodo.5172595	Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396B0425FF7FFAF1FE4CFDCC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172599/files/figure.png	https://doi.org/10.5281/zenodo.5172599	Figure 5. Maxilla, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 5. Maxilla, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396B0425FF7FFAF1FE4CFDCC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172601/files/figure.png	https://doi.org/10.5281/zenodo.5172601	Figure 6. Mandibles, ventral view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	Figure 6. Mandibles, ventral view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396B0425FF7FFAF1FE4CFDCC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172605/files/figure.png	https://doi.org/10.5281/zenodo.5172605	Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396B0425FF7FFAF1FE4CFDCC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172607/files/figure.png	https://doi.org/10.5281/zenodo.5172607	Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396B0425FF7FFAF1FE4CFDCC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172612/files/figure.png	https://doi.org/10.5281/zenodo.5172612	Figure 9. Protibia, dorsal view, of Tomarus spp. a) T. gyas and b) T. maimon.	Figure 9. Protibia, dorsal view, of Tomarus spp. a) T. gyas and b) T. maimon.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396B0425FF7FFAF1FE4CFDCC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172614/files/figure.png	https://doi.org/10.5281/zenodo.5172614	Figure 10. Meso- and metatibia, dorsal view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	Figure 10. Meso- and metatibia, dorsal view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396B0425FF7FFAF1FE4CFDCC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172618/files/figure.png	https://doi.org/10.5281/zenodo.5172618	Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396B0425FF7FFAF1FE4CFDCC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172620/files/figure.png	https://doi.org/10.5281/zenodo.5172620	Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396B0425FF7FFAF1FE4CFDCC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172622/files/figure.png	https://doi.org/10.5281/zenodo.5172622	Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396B0425FF7FFAF1FE4CFDCC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172626/files/figure.png	https://doi.org/10.5281/zenodo.5172626	Figure 14. Gonocoxite, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (Gx) gonocoxite), (Gs) gonocoxal sternite.	Figure 14. Gonocoxite, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (Gx) gonocoxite), (Gs) gonocoxal sternite.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396B0425FF7FFAF1FE4CFDCC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172630/files/figure.png	https://doi.org/10.5281/zenodo.5172630	Figure 15. Internal sac of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (AL) accessory lamella, (CL) copulatory lamella (CL), (Ra) raspula, (Esp) speculum, and (Fg) a complex, spike-like flagellum that varies in length and thickness.	Figure 15. Internal sac of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (AL) accessory lamella, (CL) copulatory lamella (CL), (Ra) raspula, (Esp) speculum, and (Fg) a complex, spike-like flagellum that varies in length and thickness.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396B0425FF7FFAF1FE4CFDCC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172632/files/figure.png	https://doi.org/10.5281/zenodo.5172632	Figure 16. Distribution of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. maimon. d) T. gyas.	Figure 16. Distribution of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. maimon. d) T. gyas.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396B0425FF7FFAF1FE4CFDCC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172588/files/figure.png	https://doi.org/10.5281/zenodo.5172588	Figure 2. Habitus of Tomarus spp. a) Tomarus rosettae (Endrödi) dorsal view. b) Tomarus spinipenis n. sp. dorsal view, holotype. c) Tomarus spinipenis dorsal view, allotype. d) Tomarus burmeisteri (Steinheil) dorsal view. e) Tomarus villosus (Burmeister) ventral view. f) Tomarus gyas ventral view. g) Tomarus bidentulus (Fairmaire) abdomen segments, ventral view (female). h) Tomarus bidentulus abdomen segments, ventral view (male).	Figure 2. Habitus of Tomarus spp. a) Tomarus rosettae (Endrödi) dorsal view. b) Tomarus spinipenis n. sp. dorsal view, holotype. c) Tomarus spinipenis dorsal view, allotype. d) Tomarus burmeisteri (Steinheil) dorsal view. e) Tomarus villosus (Burmeister) ventral view. f) Tomarus gyas ventral view. g) Tomarus bidentulus (Fairmaire) abdomen segments, ventral view (female). h) Tomarus bidentulus abdomen segments, ventral view (male).	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396A0427FF7FFB71FA32FF4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172584/files/figure.png	https://doi.org/10.5281/zenodo.5172584	Figure 1. Habitus of Tomarus spp. a) Tomarus bidentulus (Fairmaire) dorsal view. b) Tomarus burmeisteri (Steinheil) dorsal view. c) Tomarus gyas Erichson dorsal view. d) Tomarus rosettae (Endrödi) dorsal view. e) Tomarus roigjunenti n. sp. dorsal view, holotype. f) Tomarus roigjunenti n. sp. dorsal view, allotype.	Figure 1. Habitus of Tomarus spp. a) Tomarus bidentulus (Fairmaire) dorsal view. b) Tomarus burmeisteri (Steinheil) dorsal view. c) Tomarus gyas Erichson dorsal view. d) Tomarus rosettae (Endrödi) dorsal view. e) Tomarus roigjunenti n. sp. dorsal view, holotype. f) Tomarus roigjunenti n. sp. dorsal view, allotype.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396A0427FF7FFB71FA32FF4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172595/files/figure.png	https://doi.org/10.5281/zenodo.5172595	Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396A0427FF7FFB71FA32FF4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172599/files/figure.png	https://doi.org/10.5281/zenodo.5172599	Figure 5. Maxilla, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 5. Maxilla, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396A0427FF7FFB71FA32FF4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172601/files/figure.png	https://doi.org/10.5281/zenodo.5172601	Figure 6. Mandibles, ventral view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	Figure 6. Mandibles, ventral view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396A0427FF7FFB71FA32FF4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172605/files/figure.png	https://doi.org/10.5281/zenodo.5172605	Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396A0427FF7FFB71FA32FF4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172607/files/figure.png	https://doi.org/10.5281/zenodo.5172607	Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396A0427FF7FFB71FA32FF4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172614/files/figure.png	https://doi.org/10.5281/zenodo.5172614	Figure 10. Meso- and metatibia, dorsal view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	Figure 10. Meso- and metatibia, dorsal view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396A0427FF7FFB71FA32FF4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172618/files/figure.png	https://doi.org/10.5281/zenodo.5172618	Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396A0427FF7FFB71FA32FF4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172620/files/figure.png	https://doi.org/10.5281/zenodo.5172620	Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396A0427FF7FFB71FA32FF4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172622/files/figure.png	https://doi.org/10.5281/zenodo.5172622	Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396A0427FF7FFB71FA32FF4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172626/files/figure.png	https://doi.org/10.5281/zenodo.5172626	Figure 14. Gonocoxite, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (Gx) gonocoxite), (Gs) gonocoxal sternite.	Figure 14. Gonocoxite, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (Gx) gonocoxite), (Gs) gonocoxal sternite.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396A0427FF7FFB71FA32FF4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172630/files/figure.png	https://doi.org/10.5281/zenodo.5172630	Figure 15. Internal sac of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (AL) accessory lamella, (CL) copulatory lamella (CL), (Ra) raspula, (Esp) speculum, and (Fg) a complex, spike-like flagellum that varies in length and thickness.	Figure 15. Internal sac of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (AL) accessory lamella, (CL) copulatory lamella (CL), (Ra) raspula, (Esp) speculum, and (Fg) a complex, spike-like flagellum that varies in length and thickness.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396A0427FF7FFB71FA32FF4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172634/files/figure.png	https://doi.org/10.5281/zenodo.5172634	Figure 17. Distribution of Tomarus spp. a) T. roigjunenti n. sp. b) T. rosettae. c) T. spinipenis n. sp. d) T. villosus.	Figure 17. Distribution of Tomarus spp. a) T. roigjunenti n. sp. b) T. rosettae. c) T. spinipenis n. sp. d) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396A0427FF7FFB71FA32FF4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172612/files/figure.png	https://doi.org/10.5281/zenodo.5172612	Figure 9. Protibia, dorsal view, of Tomarus spp. a) T. gyas and b) T. maimon.	Figure 9. Protibia, dorsal view, of Tomarus spp. a) T. gyas and b) T. maimon.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739680427FF7FFEB1FA06FB2C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172584/files/figure.png	https://doi.org/10.5281/zenodo.5172584	Figure 1. Habitus of Tomarus spp. a) Tomarus bidentulus (Fairmaire) dorsal view. b) Tomarus burmeisteri (Steinheil) dorsal view. c) Tomarus gyas Erichson dorsal view. d) Tomarus rosettae (Endrödi) dorsal view. e) Tomarus roigjunenti n. sp. dorsal view, holotype. f) Tomarus roigjunenti n. sp. dorsal view, allotype.	Figure 1. Habitus of Tomarus spp. a) Tomarus bidentulus (Fairmaire) dorsal view. b) Tomarus burmeisteri (Steinheil) dorsal view. c) Tomarus gyas Erichson dorsal view. d) Tomarus rosettae (Endrödi) dorsal view. e) Tomarus roigjunenti n. sp. dorsal view, holotype. f) Tomarus roigjunenti n. sp. dorsal view, allotype.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739680427FF7FFEB1FA06FB2C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172588/files/figure.png	https://doi.org/10.5281/zenodo.5172588	Figure 2. Habitus of Tomarus spp. a) Tomarus rosettae (Endrödi) dorsal view. b) Tomarus spinipenis n. sp. dorsal view, holotype. c) Tomarus spinipenis dorsal view, allotype. d) Tomarus burmeisteri (Steinheil) dorsal view. e) Tomarus villosus (Burmeister) ventral view. f) Tomarus gyas ventral view. g) Tomarus bidentulus (Fairmaire) abdomen segments, ventral view (female). h) Tomarus bidentulus abdomen segments, ventral view (male).	Figure 2. Habitus of Tomarus spp. a) Tomarus rosettae (Endrödi) dorsal view. b) Tomarus spinipenis n. sp. dorsal view, holotype. c) Tomarus spinipenis dorsal view, allotype. d) Tomarus burmeisteri (Steinheil) dorsal view. e) Tomarus villosus (Burmeister) ventral view. f) Tomarus gyas ventral view. g) Tomarus bidentulus (Fairmaire) abdomen segments, ventral view (female). h) Tomarus bidentulus abdomen segments, ventral view (male).	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739680427FF7FFEB1FA06FB2C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172591/files/figure.png	https://doi.org/10.5281/zenodo.5172591	Figure 3. Types of Tomarus spp. a) T. bidentulus female holotype, dorsal view. b) T. burmeisteri male holotype, dorsal view. c) T. burmeisteri male holotype, lateral view. d) T. burmeisteri male holotype parameres, caudal view. e) T. burmeisteri male holotype parameres (lateral view). f) Label for T. burmeisteri holotype. g) T. gyas male holotype, dorsal view and labels and parameres, caudal view). h) T. maimon holotype, dorsal view. i) T. rosettae male holotype, dorsal view. j) T. villosus male holotype, dorsal view. k) T. villosus male holotype, dorsal view of head. l) T. villosus male holotype, parameres, caudal view. m) Label for T. villosus holotype.	Figure 3. Types of Tomarus spp. a) T. bidentulus female holotype, dorsal view. b) T. burmeisteri male holotype, dorsal view. c) T. burmeisteri male holotype, lateral view. d) T. burmeisteri male holotype parameres, caudal view. e) T. burmeisteri male holotype parameres (lateral view). f) Label for T. burmeisteri holotype. g) T. gyas male holotype, dorsal view and labels and parameres, caudal view). h) T. maimon holotype, dorsal view. i) T. rosettae male holotype, dorsal view. j) T. villosus male holotype, dorsal view. k) T. villosus male holotype, dorsal view of head. l) T. villosus male holotype, parameres, caudal view. m) Label for T. villosus holotype.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739680427FF7FFEB1FA06FB2C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172595/files/figure.png	https://doi.org/10.5281/zenodo.5172595	Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739680427FF7FFEB1FA06FB2C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172599/files/figure.png	https://doi.org/10.5281/zenodo.5172599	Figure 5. Maxilla, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 5. Maxilla, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739680427FF7FFEB1FA06FB2C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172601/files/figure.png	https://doi.org/10.5281/zenodo.5172601	Figure 6. Mandibles, ventral view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	Figure 6. Mandibles, ventral view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739680427FF7FFEB1FA06FB2C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172605/files/figure.png	https://doi.org/10.5281/zenodo.5172605	Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739680427FF7FFEB1FA06FB2C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172607/files/figure.png	https://doi.org/10.5281/zenodo.5172607	Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739680427FF7FFEB1FA06FB2C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172614/files/figure.png	https://doi.org/10.5281/zenodo.5172614	Figure 10. Meso- and metatibia, dorsal view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	Figure 10. Meso- and metatibia, dorsal view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739680427FF7FFEB1FA06FB2C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172618/files/figure.png	https://doi.org/10.5281/zenodo.5172618	Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739680427FF7FFEB1FA06FB2C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172620/files/figure.png	https://doi.org/10.5281/zenodo.5172620	Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739680427FF7FFEB1FA06FB2C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172622/files/figure.png	https://doi.org/10.5281/zenodo.5172622	Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739680427FF7FFEB1FA06FB2C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172626/files/figure.png	https://doi.org/10.5281/zenodo.5172626	Figure 14. Gonocoxite, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (Gx) gonocoxite), (Gs) gonocoxal sternite.	Figure 14. Gonocoxite, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (Gx) gonocoxite), (Gs) gonocoxal sternite.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739680427FF7FFEB1FA06FB2C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172630/files/figure.png	https://doi.org/10.5281/zenodo.5172630	Figure 15. Internal sac of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (AL) accessory lamella, (CL) copulatory lamella (CL), (Ra) raspula, (Esp) speculum, and (Fg) a complex, spike-like flagellum that varies in length and thickness.	Figure 15. Internal sac of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (AL) accessory lamella, (CL) copulatory lamella (CL), (Ra) raspula, (Esp) speculum, and (Fg) a complex, spike-like flagellum that varies in length and thickness.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A739680427FF7FFEB1FA06FB2C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172634/files/figure.png	https://doi.org/10.5281/zenodo.5172634	Figure 17. Distribution of Tomarus spp. a) T. roigjunenti n. sp. b) T. rosettae. c) T. spinipenis n. sp. d) T. villosus.	Figure 17. Distribution of Tomarus spp. a) T. roigjunenti n. sp. b) T. rosettae. c) T. spinipenis n. sp. d) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396F0420FF7FFF51FBBAFD4F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172641/files/figure.png	https://doi.org/10.5281/zenodo.5172641	Figure 19. Holotype of T. rubripes (Boheman). a) Dorsal view. b) Lateral view. c) Parameres, caudal view. d) Parameres, lateral view. e) Labels. Photographs courtesy of Johannes Bergsten, NHRS. Made available by NHRS under Creative Commons Attribution 4.0 International Public License (CC-BY 4.0).	Figure 19. Holotype of T. rubripes (Boheman). a) Dorsal view. b) Lateral view. c) Parameres, caudal view. d) Parameres, lateral view. e) Labels. Photographs courtesy of Johannes Bergsten, NHRS. Made available by NHRS under Creative Commons Attribution 4.0 International Public License (CC-BY 4.0).	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396F0421FF7FFCB1FB2FFECC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172588/files/figure.png	https://doi.org/10.5281/zenodo.5172588	Figure 2. Habitus of Tomarus spp. a) Tomarus rosettae (Endrödi) dorsal view. b) Tomarus spinipenis n. sp. dorsal view, holotype. c) Tomarus spinipenis dorsal view, allotype. d) Tomarus burmeisteri (Steinheil) dorsal view. e) Tomarus villosus (Burmeister) ventral view. f) Tomarus gyas ventral view. g) Tomarus bidentulus (Fairmaire) abdomen segments, ventral view (female). h) Tomarus bidentulus abdomen segments, ventral view (male).	Figure 2. Habitus of Tomarus spp. a) Tomarus rosettae (Endrödi) dorsal view. b) Tomarus spinipenis n. sp. dorsal view, holotype. c) Tomarus spinipenis dorsal view, allotype. d) Tomarus burmeisteri (Steinheil) dorsal view. e) Tomarus villosus (Burmeister) ventral view. f) Tomarus gyas ventral view. g) Tomarus bidentulus (Fairmaire) abdomen segments, ventral view (female). h) Tomarus bidentulus abdomen segments, ventral view (male).	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396F0421FF7FFCB1FB2FFECC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172595/files/figure.png	https://doi.org/10.5281/zenodo.5172595	Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396F0421FF7FFCB1FB2FFECC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172599/files/figure.png	https://doi.org/10.5281/zenodo.5172599	Figure 5. Maxilla, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 5. Maxilla, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396F0421FF7FFCB1FB2FFECC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172601/files/figure.png	https://doi.org/10.5281/zenodo.5172601	Figure 6. Mandibles, ventral view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	Figure 6. Mandibles, ventral view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396F0421FF7FFCB1FB2FFECC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172605/files/figure.png	https://doi.org/10.5281/zenodo.5172605	Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396F0421FF7FFCB1FB2FFECC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172607/files/figure.png	https://doi.org/10.5281/zenodo.5172607	Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396F0421FF7FFCB1FB2FFECC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172614/files/figure.png	https://doi.org/10.5281/zenodo.5172614	Figure 10. Meso- and metatibia, dorsal view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	Figure 10. Meso- and metatibia, dorsal view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396F0421FF7FFCB1FB2FFECC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172618/files/figure.png	https://doi.org/10.5281/zenodo.5172618	Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396F0421FF7FFCB1FB2FFECC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172620/files/figure.png	https://doi.org/10.5281/zenodo.5172620	Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396F0421FF7FFCB1FB2FFECC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172622/files/figure.png	https://doi.org/10.5281/zenodo.5172622	Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396F0421FF7FFCB1FB2FFECC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172626/files/figure.png	https://doi.org/10.5281/zenodo.5172626	Figure 14. Gonocoxite, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (Gx) gonocoxite), (Gs) gonocoxal sternite.	Figure 14. Gonocoxite, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (Gx) gonocoxite), (Gs) gonocoxal sternite.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396F0421FF7FFCB1FB2FFECC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172630/files/figure.png	https://doi.org/10.5281/zenodo.5172630	Figure 15. Internal sac of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (AL) accessory lamella, (CL) copulatory lamella (CL), (Ra) raspula, (Esp) speculum, and (Fg) a complex, spike-like flagellum that varies in length and thickness.	Figure 15. Internal sac of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (AL) accessory lamella, (CL) copulatory lamella (CL), (Ra) raspula, (Esp) speculum, and (Fg) a complex, spike-like flagellum that varies in length and thickness.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396F0421FF7FFCB1FB2FFECC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172634/files/figure.png	https://doi.org/10.5281/zenodo.5172634	Figure 17. Distribution of Tomarus spp. a) T. roigjunenti n. sp. b) T. rosettae. c) T. spinipenis n. sp. d) T. villosus.	Figure 17. Distribution of Tomarus spp. a) T. roigjunenti n. sp. b) T. rosettae. c) T. spinipenis n. sp. d) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396E0421FF7FFC11FCDCF8EC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172588/files/figure.png	https://doi.org/10.5281/zenodo.5172588	Figure 2. Habitus of Tomarus spp. a) Tomarus rosettae (Endrödi) dorsal view. b) Tomarus spinipenis n. sp. dorsal view, holotype. c) Tomarus spinipenis dorsal view, allotype. d) Tomarus burmeisteri (Steinheil) dorsal view. e) Tomarus villosus (Burmeister) ventral view. f) Tomarus gyas ventral view. g) Tomarus bidentulus (Fairmaire) abdomen segments, ventral view (female). h) Tomarus bidentulus abdomen segments, ventral view (male).	Figure 2. Habitus of Tomarus spp. a) Tomarus rosettae (Endrödi) dorsal view. b) Tomarus spinipenis n. sp. dorsal view, holotype. c) Tomarus spinipenis dorsal view, allotype. d) Tomarus burmeisteri (Steinheil) dorsal view. e) Tomarus villosus (Burmeister) ventral view. f) Tomarus gyas ventral view. g) Tomarus bidentulus (Fairmaire) abdomen segments, ventral view (female). h) Tomarus bidentulus abdomen segments, ventral view (male).	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396E0421FF7FFC11FCDCF8EC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172591/files/figure.png	https://doi.org/10.5281/zenodo.5172591	Figure 3. Types of Tomarus spp. a) T. bidentulus female holotype, dorsal view. b) T. burmeisteri male holotype, dorsal view. c) T. burmeisteri male holotype, lateral view. d) T. burmeisteri male holotype parameres, caudal view. e) T. burmeisteri male holotype parameres (lateral view). f) Label for T. burmeisteri holotype. g) T. gyas male holotype, dorsal view and labels and parameres, caudal view). h) T. maimon holotype, dorsal view. i) T. rosettae male holotype, dorsal view. j) T. villosus male holotype, dorsal view. k) T. villosus male holotype, dorsal view of head. l) T. villosus male holotype, parameres, caudal view. m) Label for T. villosus holotype.	Figure 3. Types of Tomarus spp. a) T. bidentulus female holotype, dorsal view. b) T. burmeisteri male holotype, dorsal view. c) T. burmeisteri male holotype, lateral view. d) T. burmeisteri male holotype parameres, caudal view. e) T. burmeisteri male holotype parameres (lateral view). f) Label for T. burmeisteri holotype. g) T. gyas male holotype, dorsal view and labels and parameres, caudal view). h) T. maimon holotype, dorsal view. i) T. rosettae male holotype, dorsal view. j) T. villosus male holotype, dorsal view. k) T. villosus male holotype, dorsal view of head. l) T. villosus male holotype, parameres, caudal view. m) Label for T. villosus holotype.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396E0421FF7FFC11FCDCF8EC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172595/files/figure.png	https://doi.org/10.5281/zenodo.5172595	Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 4. Head, dorsal view of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396E0421FF7FFC11FCDCF8EC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172599/files/figure.png	https://doi.org/10.5281/zenodo.5172599	Figure 5. Maxilla, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 5. Maxilla, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396E0421FF7FFC11FCDCF8EC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172601/files/figure.png	https://doi.org/10.5281/zenodo.5172601	Figure 6. Mandibles, ventral view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	Figure 6. Mandibles, ventral view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396E0421FF7FFC11FCDCF8EC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172605/files/figure.png	https://doi.org/10.5281/zenodo.5172605	Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396E0421FF7FFC11FCDCF8EC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172607/files/figure.png	https://doi.org/10.5281/zenodo.5172607	Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396E0421FF7FFC11FCDCF8EC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172614/files/figure.png	https://doi.org/10.5281/zenodo.5172614	Figure 10. Meso- and metatibia, dorsal view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	Figure 10. Meso- and metatibia, dorsal view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396E0421FF7FFC11FCDCF8EC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172618/files/figure.png	https://doi.org/10.5281/zenodo.5172618	Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396E0421FF7FFC11FCDCF8EC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172620/files/figure.png	https://doi.org/10.5281/zenodo.5172620	Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396E0421FF7FFC11FCDCF8EC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172622/files/figure.png	https://doi.org/10.5281/zenodo.5172622	Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396E0421FF7FFC11FCDCF8EC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172626/files/figure.png	https://doi.org/10.5281/zenodo.5172626	Figure 14. Gonocoxite, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (Gx) gonocoxite), (Gs) gonocoxal sternite.	Figure 14. Gonocoxite, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (Gx) gonocoxite), (Gs) gonocoxal sternite.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396E0421FF7FFC11FCDCF8EC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172630/files/figure.png	https://doi.org/10.5281/zenodo.5172630	Figure 15. Internal sac of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (AL) accessory lamella, (CL) copulatory lamella (CL), (Ra) raspula, (Esp) speculum, and (Fg) a complex, spike-like flagellum that varies in length and thickness.	Figure 15. Internal sac of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (AL) accessory lamella, (CL) copulatory lamella (CL), (Ra) raspula, (Esp) speculum, and (Fg) a complex, spike-like flagellum that varies in length and thickness.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396E0421FF7FFC11FCDCF8EC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172634/files/figure.png	https://doi.org/10.5281/zenodo.5172634	Figure 17. Distribution of Tomarus spp. a) T. roigjunenti n. sp. b) T. rosettae. c) T. spinipenis n. sp. d) T. villosus.	Figure 17. Distribution of Tomarus spp. a) T. roigjunenti n. sp. b) T. rosettae. c) T. spinipenis n. sp. d) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396C043CFF7FFD51FF49FE2F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172588/files/figure.png	https://doi.org/10.5281/zenodo.5172588	Figure 2. Habitus of Tomarus spp. a) Tomarus rosettae (Endrödi) dorsal view. b) Tomarus spinipenis n. sp. dorsal view, holotype. c) Tomarus spinipenis dorsal view, allotype. d) Tomarus burmeisteri (Steinheil) dorsal view. e) Tomarus villosus (Burmeister) ventral view. f) Tomarus gyas ventral view. g) Tomarus bidentulus (Fairmaire) abdomen segments, ventral view (female). h) Tomarus bidentulus abdomen segments, ventral view (male).	Figure 2. Habitus of Tomarus spp. a) Tomarus rosettae (Endrödi) dorsal view. b) Tomarus spinipenis n. sp. dorsal view, holotype. c) Tomarus spinipenis dorsal view, allotype. d) Tomarus burmeisteri (Steinheil) dorsal view. e) Tomarus villosus (Burmeister) ventral view. f) Tomarus gyas ventral view. g) Tomarus bidentulus (Fairmaire) abdomen segments, ventral view (female). h) Tomarus bidentulus abdomen segments, ventral view (male).	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396C043CFF7FFD51FF49FE2F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172601/files/figure.png	https://doi.org/10.5281/zenodo.5172601	Figure 6. Mandibles, ventral view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	Figure 6. Mandibles, ventral view, of Tomarus spp. a–b) T. bidentulus. c–d) T. burmeisteri. e–f) T. gyas. g–h) T. maimon. i–j. T. roigjunenti n. sp. k–l. T. rosettae. m–n. T. spinipenis n. sp. o–p. T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396C043CFF7FFD51FF49FE2F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172599/files/figure.png	https://doi.org/10.5281/zenodo.5172599	Figure 5. Maxilla, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 5. Maxilla, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396C043CFF7FFD51FF49FE2F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172605/files/figure.png	https://doi.org/10.5281/zenodo.5172605	Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 7. Labium, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396C043CFF7FFD51FF49FE2F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172626/files/figure.png	https://doi.org/10.5281/zenodo.5172626	Figure 14. Gonocoxite, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (Gx) gonocoxite), (Gs) gonocoxal sternite.	Figure 14. Gonocoxite, ventral view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus. Components are: (Gx) gonocoxite), (Gs) gonocoxal sternite.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396C043CFF7FFD51FF49FE2F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172607/files/figure.png	https://doi.org/10.5281/zenodo.5172607	Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 8. Pronotum, dorsal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396C043CFF7FFD51FF49FE2F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172618/files/figure.png	https://doi.org/10.5281/zenodo.5172618	Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 11. Parameres, caudal view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396C043CFF7FFD51FF49FE2F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172620/files/figure.png	https://doi.org/10.5281/zenodo.5172620	Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 12. Parameres, lateral view, of: a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396C043CFF7FFD51FF49FE2F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172622/files/figure.png	https://doi.org/10.5281/zenodo.5172622	Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	Figure 13. Parameres, anterior view, of Tomarus spp. a) T. bidentulus. b) T. burmeisteri. c) T. gyas. d) T. maimon. e) T. roigjunenti n. sp. f) T. rosettae. g) T. spinipenis n. sp. h) T. villosus.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
A24087A7396C043CFF7FFD51FF49FE2F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5172612/files/figure.png	https://doi.org/10.5281/zenodo.5172612	Figure 9. Protibia, dorsal view, of Tomarus spp. a) T. gyas and b) T. maimon.	Figure 9. Protibia, dorsal view, of Tomarus spp. a) T. gyas and b) T. maimon.	2017-04-28	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa		Zenodo	biologists	Neita-Moreno, Jhon César;Alexander, Recursos Biológicos;Agustín, San;de, Villa;Leyva;Boyacá;Ratcliffe, Brett C.;Collections, Systematics Research;of, University;Museum, Nebraska State;Hall, Nebraska;of, University;Nebraska;Lincoln;Ne;Usa			
