identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
A2414922F011FF94FF5BF999FD0BFADC.text	A2414922F011FF94FF5BF999FD0BFADC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mokayathelphusa Moreno, Villalobos & Alvarez 2022	<div><p>Mokayathelphusa Moreno, Villalobos &amp; Álvarez n. gen.</p><p>urn:lsid:zoobank.org:act: A06B6E9E-4953-4B25-9DC9-3F1C592ACF4F</p><p>Diagnosis. G1 with the distal portion of principal axis twisted counterclockwise. Apical cavity oriented mesocephalically, compressed and closed cephalically by the internal angle of mesial process. Caudo-marginal projection trilobed distally, middle and distal lobes spiniform of same length, proximal lobe as elongated sheet, curved, extending through distal third of main gonopod axis. Mesial process large, subrectangular, with lateral margin ornamented with several spiniform teeth.</p><p>Type species. Mokayathelphusa angelsotoi n. sp., by present designation.</p><p>Etymology. The name of the genus is taken from the ancient tribe “Mokaya” that inhabited the Chimalapas region about 3,600 ybp. The word “Mokaya” comes from the earliest Zoque language, that means “people of the corn”. The gender of this name is feminine.</p><p>Distribution. Currently known only from the municipality of Santa María Chimalapa, State of Oaxaca, Mexico.</p><p>Remarks. The description of Mokayathelphusa n. gen. is justified by its unique male G1 morphology and clear genetic separation from related genera. The new genus can be related to other genera of the subfamily Pseudothelphusinae by the presence of the caudo-marginal projection, which ends distally in a trilobed distal plate. The phylogenetic analysis (Fig. 2) recovers it as the sister lineage to Disparithelphusa Smalley &amp; Adkison, 1984 . Morphologically the G1 of both genera present a torsion that modifies the shape and orientation of the apical cavity and have a trilobed distal plate in the caudo-marginal projection, where the distal lobes are smaller than the proximal one. They can also be differentiated by the size of the mesial process and the ornamentation of its lateral border, while in Disparithelphusa it is small, oval and without lateral ornamentation; in Mokayathelphusa n. gen., it is evidently developed and ornamented laterally.</p></div>	https://treatment.plazi.org/id/A2414922F011FF94FF5BF999FD0BFADC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Moreno-Juárez, Eric G.;Villalobos, José Luis;Álvarez, Fernando	Moreno-Juárez, Eric G., Villalobos, José Luis, Álvarez, Fernando (2022): Two new genera and one new species of freshwater crabs of the subfamily Pseudothelphusinae (Decapoda: Brachyura: Pseudothelphusidae) from southwestern Mexico. Zootaxa 5200 (1): 24-36, DOI: 10.11646/zootaxa.5200.1.2
A2414922F016FF95FF5BFA05FA83F813.text	A2414922F016FF95FF5BFA05FA83F813.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mokayathelphusa angelsotoi Moreno, Villalobos & Alvarez 2022	<div><p>Mokayathelphusa angelsotoi Moreno, Villalobos &amp; Álvarez n. sp.</p><p>(Figs. 3, 4)</p><p>urn:lsid:zoobank.org:act: 25D83587-8F68-4CF8-B0CE-4309264E971C</p><p>Type material. Holotype: CNCR 34843, ♂, CL 14.9 mm, CW 23.1 mm; tributary of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-94.70583&amp;materialsCitation.latitude=16.911388" title="Search Plazi for locations around (long -94.70583/lat 16.911388)">Río Negro</a>, near the town of Santa María Chimalapa, 16°54’41”N, 94°42’21”W; 144 m asl, Municipality of Santa María Chimalapa, Oaxaca, Mexico, coll. A. Soto, 20 July 2018.</p><p>Description. Surface of carapace flat, smooth, except finely punctate branchial region. Cardiac, mesogastric, urogastric and branchial regions, discernible. Postfrontal area swollen, curved anteriorly to reach poorly defined superior frontal margin; postfrontal lobes low, separated by narrow, shallow median groove, ending anteriorly in median notch dividing superior frontal border. Cervical groove straight, narrow, well-marked, short, not reaching anterolateral margin. Lateral margin with 30 denticles and two shallow notches, anterior notch behind external angle of orbit, posterior notch at level of cervical groove (Fig. 4a). In frontal view, front, antennules, proepistome, epistome, and merus of third maxillipeds with minute dark spots; superior frontal margin marked by acute frontal bend of carapace, inclined towards median groove, with small granules, divided by V-shaped median notch; inferior frontal margin sinuous, with blunt granules, adjacent portion to antennular cavity higher than rest; interantennular septum concealed by inferior frontal margin. Antennules and antennular fossae visible, fossae wider in middle. Antennal peduncle filling orbital hiatus.Orbital margin well defined, internal surfaces of ocular cavities with scattered dark spots and short setae; inner tooth slender, elongate, concave longitudinally, superior margin subtriangular. Opercular plate of antennal gland ovoid. Superior margin of proepistome sharp in middle, oval-shaped at lateral ends. Epistome setose, surrounding buccal cavity; laterally with rounded granules; epistomal tooth triangular, apex directed anteriorly at same level of interantennular septum, between palps of third maxillipeds when closing buccal cavity. Opening of efferent branchial channel subquadrate, superior margin curved, ratio width/length 1.25.</p><p>Third maxilliped with ischium trapezoidal, 1.33 times wider than long; merus narrower than ischium, anterior margin widely rounded with shallow, rounded notch next to insertion of palp; ratio exopod/ischium length 0.65 (Fig. 4b).</p><p>Chelipeds asymmetrical. Merus of larger cheliped with row of blunt tubercles along internal margin, carpus with strong triangular spine on internal margin. Chela with internal surface smooth, swollen; fingers gaping, curved inwards distally, cutting edges with triangular teeth (Fig. 4b). Dactylus narrow, dorsal surface with longitudinal rows of small pits, cutting edge with two proximal triangular teeth, remaining edge with alternate small and big teeth, ending in curved acute point. Propodus cutting edge with two small proximal teeth, six medial triangular and acute ones, seventh smallest, final point acute, incurved.</p><p>G1 slender, with torsion throughout its length, distal half curved mesially making apical cavity visible in mesocephalic position. In mesial view (Fig. 3a), principal axis twisted counterclockwise, distal crest of caudal surface (Cdc) cephalically oriented, projected mesially; distal portion of caudo-marginal projection directed cephalad, with distal lobe (Dl) as a strong spine with flat distal surface, delimited by well marked external crest (Ecdl); medial lobe (Ml) not visible; proximal lobe (Pl) partially evident in proximal half, as curved sheet, with subacute end; distal crest of apical cavity ornamented with acute, hooked spinules. In caudal view (Fig. 3b), principal axis with distal half twisted, mesially inclined; medial portion with curved, sharp caudal edge (Ce), lateral surface with sharp rib-like border (Le); mesial process (Mp) as laminate rectangular plate, lateral margin raised, produced into irregular crest (Lbmp), armed with acute teeth; distal crest of lateral surface (Ldc) widely rounded. In lateral view (Fig. 3c), distal half of gonopod straight, surface smooth, Ldc rounded; Lbmp thin, armed with nine irregular teeth, distally forming right angle with superior border; lobes of caudo-marginal projection partially visible, Dl and Ml conical, acute, Pl triangular. In cephalic view (Fig. 3d), distal half inclined mesially; lobes of caudo-marginal projection evident, Dl and Ml conical, acute, latero-cephalically directed, distal one moderately stouter; Pl as curved sheet, distal end slender, rounded; mesial process as thin plate with toothed lateral border (Lbmp); distal portion forming right angle with superior border (Dbmp); apical cavity U-shaped, mesial distal crest (Mdc) with row of acute, hooked spinules, field of apical spines adjacent to internal lateral surface with several chitinized setae, aperture of spermatic channel on caudal position, distal crest of lateral surface projecting distally. In distal view (Fig. 3e), apical portion evidently curved cephalad, apical cavity (Ac) barely visible; Ldc smooth laterally, mesially armed with short spinules; distal lobe of caudo-marginal projection conical, slender, acute; mesial process partially noticeable. Apical cavity only visible distally in meso-cephalic position (Fig. 3f), U-shaped, aperture on cephalic surface, closed by internal angle of mesial process (Imp); field of spines (Sf) adjacent to lateral surface, with 10 long, conical and slender corneous setae, and 20 shorter conical tubercles scattered on caudal portion of cavity; aperture of spermatic pore (Sp) in caudal position, with spinules on caudal internal surface; distal crest of mesial surface (Mdc) armed with slender spinules. Dl and Ml of caudo-marginal projection, almost totally visible, conical, subacute, fused at base, directed cephalad.</p><p>Distribution. Known only from the type locality.</p><p>Type locality. Stream tributary of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-94.70583&amp;materialsCitation.latitude=16.911388" title="Search Plazi for locations around (long -94.70583/lat 16.911388)">Río Negro</a> (16°54’41”N, 94°42’21”W; 144 m asl), Municipality of Santa María Chimalapa, Oaxaca, Mexico .</p><p>Etymology. This species is named after Ángel Fernando Soto Pozos, enthusiastic herpetologist interested in the biodiversity of the Chimalapas region, and collector of the type specimen.</p><p>Remarks. The G1 morphology of Mokayathelphusa angelsotoi sp. nov., is unique among the members of the subfamily Pseudothelphusinae . The new species is genetically related to Disparithelphusa pecki Smalley &amp; Adkison, 1984 . Both species present a trilobed caudo-marginal projection, the two short distal lobes and the elongated proximal one. However, the shape, arrangement and orientation of these lobes are completely different. Other apical elements such as the mesial process, and the form and ornamentation of apical cavity, also show differences.</p><p>Geographically, Diparithelphusa pecki occurs in northern Oaxaca, in the Sierra de Juárez near the town of Tuxtepec, whereas Mokayathelphusa angelsotoi n. sp. occurs in the Chimalapas region about 190 km to the southeast (Fig. 1). Genetically, both species are clearly different as seen in their phylogenetic placement (Fig. 2).</p></div>	https://treatment.plazi.org/id/A2414922F016FF95FF5BFA05FA83F813	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Moreno-Juárez, Eric G.;Villalobos, José Luis;Álvarez, Fernando	Moreno-Juárez, Eric G., Villalobos, José Luis, Álvarez, Fernando (2022): Two new genera and one new species of freshwater crabs of the subfamily Pseudothelphusinae (Decapoda: Brachyura: Pseudothelphusidae) from southwestern Mexico. Zootaxa 5200 (1): 24-36, DOI: 10.11646/zootaxa.5200.1.2
A2414922F015FF98FF5BFF1CFBF6FE31.text	A2414922F015FF98FF5BFF1CFBF6FE31.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Alvarezius Moreno-Juárez & Villalobos & Álvarez 2022	<div><p>Alvarezius Moreno &amp; Villalobos n. gen.</p><p>(Fig. 5)</p><p>urn:lsid:zoobank.org:act: DDF1CE7E-C440-4DD6-A41C-CEC2960509EF</p><p>Diagnosis. G1 with conical setiferous prominence on latero-caudal portion of distal border of apical cavity. Caudomarginal projection unilobed, which extends proximally reaching proximal half of gonopod principal axis, distocephalic border serrate. Mesial process subrectangular, proximal inferior angle ending in acute triangle proximally oriented.</p><p>Type species. Pseudothelpusa zongolicae Alvarez, Villalobos &amp; Moreno, 2012 .</p><p>Species included. Alvarezius zongolicae (Alvarez, Villalobos &amp; Moreno, 2012) n. comb.</p><p>Etymology. The genus name is dedicated to Dr. Fernando Álvarez, eminent Mexican carcinologist and curator of the CNCR. The gender of this name is masculine.</p><p>Distribution. Only known from Zongolica region, Veracruz, Mexico.</p><p>Remarks. When Pseudothelpusa zongolicae was first described, it was included in this genus because of the similarities in the G1 regarding the caudo-marginal projection and the mesial process as in other species of this genus, as explained by Alvarez et al. (2012). In the phylogenetic analysis presented in this paper P. zongolicae was recovered as the sister lineage to Tehuana, with mean genetic distance of 4.0% with respect to all the species of the genus, and 8.0–9.0%with respect to to species of Pseudothelphusa . Combining the genetic evidence with a new morphological description, we place P. zongolicae in the new genus Alvarezius n. gen.</p></div>	https://treatment.plazi.org/id/A2414922F015FF98FF5BFF1CFBF6FE31	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Moreno-Juárez, Eric G.;Villalobos, José Luis;Álvarez, Fernando	Moreno-Juárez, Eric G., Villalobos, José Luis, Álvarez, Fernando (2022): Two new genera and one new species of freshwater crabs of the subfamily Pseudothelphusinae (Decapoda: Brachyura: Pseudothelphusidae) from southwestern Mexico. Zootaxa 5200 (1): 24-36, DOI: 10.11646/zootaxa.5200.1.2
A2414922F01AFF99FF5BFDA8FD45FD2D.text	A2414922F01AFF99FF5BFDA8FD45FD2D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Alvarezius zongolicae (Alvarez, Villalobos & Moreno 2012) Moreno-Juárez & Villalobos & Álvarez 2022	<div><p>Alvarezius zongolicae (Alvarez, Villalobos &amp; Moreno, 2012) n. comb.</p><p>Pseudothelphusa zongolicae Alvarez, Villalobos &amp; Moreno, 2012, figs. 2, 3. — Villalobos et al., 2019: 157 (Table 1). — Álvarez et al., 2020: 10 (Supplementary material, Table 1).</p><p>Type material. Holotype: CNCR 26641; 1 ♂, CL 18.1 mm, CW 30.0 mm; spring at Choapa (18°43’32”N, 96°57’07”W, 1053 m asl), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-96.95194&amp;materialsCitation.latitude=18.725555" title="Search Plazi for locations around (long -96.95194/lat 18.725555)">Municipality of Zongolica</a>, Veracruz, Mexico; colls. E. Moreno, O. Trejo; 19 April 2011 . Paratype: CNCR 26642; 1 ♂, CL 17.7 mm, CW 28.7 mm; same locality as holotype; 12 June 2011 .</p><p>Other material examined. CNCR26643, 4♂, 1♀,CL 15.8–22.8mm,CW 25.5–37.5 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-96.94945&amp;materialsCitation.latitude=18.646944" title="Search Plazi for locations around (long -96.94945/lat 18.646944)">Macuilca</a> (18°38’49”N, 96°56’58”W, 528 m asl), Municipality of Zongolica, Veracruz, Mexico, coll. E. Moreno &amp; O. Trejo, 11 June 2011 ; CNCR 29494, 5 ♂, CL 8.9–19.9 mm CW 13.6–32.7 mm, spring of water, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-96.9475&amp;materialsCitation.latitude=18.561388" title="Search Plazi for locations around (long -96.9475/lat 18.561388)">Tepeplampa</a> (18°33’41”N, 96°56’51”W, 891 m asl), Municipality of Zongolica, Veracruz, Mexico, coll. E. Moreno &amp; S. Rodríguez, 12 December 2012 ; CNCR 29495, 6 ♂, 1 ♀, CL 11.2–23.3 mm, CW 12.5–40.6 mm, stream near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-96.941666&amp;materialsCitation.latitude=18.552776" title="Search Plazi for locations around (long -96.941666/lat 18.552776)">Tepetlampa</a> (18°33’10”N, 96°56’30”W, 110 m asl), Municipality of Zongolica, Veracruz, Mexico; colls. E. Moreno &amp; S. Rodríguez; 11 December 2012 ; CNCR 35458, 2 ♂, 2 ♀, CL 18.1–21 mm, CW 22.4–36.2 mm, same locality as holotype, coll. E. Moreno, A. Soto &amp; R. Rodríguez, 10 October 2019 ; CNCR 36478, 4 ♂, 4 ♀, CL 7.8–9.8 mm, CW 10.6–15.6 mm, spring inside cave, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-96.900116&amp;materialsCitation.latitude=18.636667" title="Search Plazi for locations around (long -96.900116/lat 18.636667)">Cintalapa</a> (18°38’12”N, 96°54’0.4”W, 1050 m asl), Municipality of Zongolica, Veracruz, Mexico; colls. E. Moreno &amp; L. Gómez; 3 April 2022 . CNCR 36479; 2 ♂, spring in <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-96.94056&amp;materialsCitation.latitude=18.605444" title="Search Plazi for locations around (long -96.94056/lat 18.605444)">Acahualco</a>, Cuahutilica (18°36’19.6”N, 96°56’26”W, 1169 m asl), Municipality of Zongolica, Veracruz, Mexico; colls. E. Moreno &amp; L. Gómez, 3 April 2022 ; CNCR 36480, 1 ♂, CL 23.2 mm, CW 39.9 mm, waterfall on the road, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-96.93931&amp;materialsCitation.latitude=18.621166" title="Search Plazi for locations around (long -96.93931/lat 18.621166)">Coapa-Pinopa</a> (18°37’16.2”N, 96°56’21.5”W, 1063 m asl), Municipality of Zongolica, Veracruz, Mexico, coll. E. Moreno &amp; L. Gómez, 3 April 2022 ; CNCR 36481, 3 ♂, CL 20.7–24.2 mm, CW 34.9–41.4 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-96.93584&amp;materialsCitation.latitude=18.604445" title="Search Plazi for locations around (long -96.93584/lat 18.604445)">Acahualco Cave</a>, Cuahutilica (18°36’16”N, 96°56’09”W, 1,111 m asl), Municipality of Zongolica, Veracruz, Mexico, coll. E. Moreno &amp; L. Gómez, 3 April 2022 .</p><p>Description. Carapace transversely oval, dorsal surface slightly convex, smooth, finely punctate, regions faintly marked, except for moderately swollen mesobranchial region (Fig. 5a). Superior frontal border present, incomplete towards middle portion, formed by incomplete row of granules, in frontal view inclined towards median groove (Fig. 5b). Inferior frontal border complete, thick, slightly rounded in dorsal view, bilobed in frontal view, extending laterally to form superior margin of orbits. Median groove deep, narrow, arising above inferior frontal border, extending posteriorly beyond postfrontal lobes. Postfrontal lobes low, well defined. Cervical grooves wide, shallow, straight towards margin of carapace, ill- defined, slightly arching towards central portion of carapace, not reaching anterolateral margin. Walking legs typical of genus. Chelipeds asymmetrical. Major chela right, palm somewhat swollen, movable finger half-length of chela, fingers not gaping; both fingers with irregular arrangement of teeth of varying sizes, both fingers curved inwards (Fig. 5c). Third maxilliped with merus showing complete rounded external margin, ischium rectangular, ratio ischium/exopod 0.85 (Fig. 5d).</p><p>G1 moderately slender, distal half of principal axis showing mesial torsion of caudo-marginal projection. In mesial view (Fig. 5e), marginal suture evident, arching distally, disappearing under caudo-marginal projection; lobe of caudo-marginal projection (Cmp) axe-shaped, large, about half length of gonopod, cephalic border smooth; caudo-marginal distal crest straight, spinulated at cephalic curvature. In cephalic view (Fig. 5f), mesial process (Mp) oriented laterally, slightly inclined proximally, tapering laterally, ending in two acute tips, superior margin closing apex cavity, extending towards internal portion of apex cavity forming central crest; caudo-marginal projection with single large, axe-shaped lobe, extending proximally slightly beyond half-length of gonopod, cephalic margin armed with small, acute teeth; crest of caudo-marginal projection becoming less thick cephalically; conical prominence (Cp) on caudal angle of lateral crest bearing apical setae. In lateral view (Fig. 5g), mesial process with two tips oriented laterally; caudo-marginal projection (Cmp) appearing as large tongue-shaped lobe; conical prominence (Cp) emerging from lateral crest. In caudal view (Fig. 5h), caudal surface showing torsion of gonopod; mesial process (Mp) subtriangular; conical prominence (Cp) of lateral crest evident; lobe of caudo-marginal projection (Cmp) elongate, slender, extending parallel to main axis of gonopod; marginal suture straight, visible along proximal two thirds of gonopod. In apical view (Fig. 5i), apex cavity approximately rectangular, field of spines (Sf) on lateral side of cavity, extending to conical prominence (Cp); central crest of apex cavity sharp, slightly undulated; distal crest of lateral surface straight; mesial process (Mp) subtriangular, distal margin rounded; lobe of caudo-marginal projection (Cmp) as undulated sheet, cephalic margin with minute spines; opening of sperm channel caudal.</p><p>Distribution. Mexico, Veracruz, in the surroundings of the Municipality of Zongolica.</p><p>Type locality. Spring at Choapa (18°43’32”N, 96°57’07”W; 1,053 m asl), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-96.95194&amp;materialsCitation.latitude=18.725555" title="Search Plazi for locations around (long -96.95194/lat 18.725555)">Municipality of Zongolica</a>, Veracruz, Mexico (CNCR 26641) .</p><p>Remarks. The term “conical projection” of the latero-caudal portion of the distal border of the apical cavity from the original description by Álvarez et al. (2012) is now changed to “conical lobe”, since the former suggests the fusion of two plates in a single apical process (Hobbs 1942, 1945; Smalley 1964); however, in this case the structure develops directly from the lateral surface.</p></div>	https://treatment.plazi.org/id/A2414922F01AFF99FF5BFDA8FD45FD2D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Moreno-Juárez, Eric G.;Villalobos, José Luis;Álvarez, Fernando	Moreno-Juárez, Eric G., Villalobos, José Luis, Álvarez, Fernando (2022): Two new genera and one new species of freshwater crabs of the subfamily Pseudothelphusinae (Decapoda: Brachyura: Pseudothelphusidae) from southwestern Mexico. Zootaxa 5200 (1): 24-36, DOI: 10.11646/zootaxa.5200.1.2
