taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
A12C6E25AC103B702CBED4E5FE0FA85C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804200/files/figure.png	https://doi.org/10.5281/zenodo.804200	FIGURE 8. Type specimens of S. purpurascens: lectotype m (A) and paralectotype f (B); S. planum: lectotype m (C) and paralectotype f (D); S. macrophallicum: holotype m (E) and allotype f (F); S. minimum lectotype m (G); S. affine lecototype m (H) (Scale bars = 1 cm).	FIGURE 8. Type specimens of S. purpurascens: lectotype m (A) and paralectotype f (B); S. planum: lectotype m (C) and paralectotype f (D); S. macrophallicum: holotype m (E) and allotype f (F); S. minimum lectotype m (G); S. affine lecototype m (H) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC103B702CBED4E5FE0FA85C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804194/files/figure.png	https://doi.org/10.5281/zenodo.804194	FIGURE 5. Male genitalia of Sphenarium: epiphallus in dorsal view (A); ectophallus in dorsal (B), lateral (C) and posterior (D) view; and endophallus in lateral (E) and dorsal (F) view. A, appendix of epiphallus; AC, apodemal plate of cingulum; AP, anterior projection of epiphallus; AS, aedeagal sclerites; ASA, apical spine of aedeagus; AV, aedeagal valve; B, bridge of epiphallus; BAS, base of aedeagal sclerites; BC, basal thickening of cingulum; BE, basal emargination of cingulum; CM, central membrane; CV, cingulum valve; DI, dorsal inflection of endophallic apodeme; EA, endophallic apodeme; ISR, inflection of supraramus; L, lophus of epiphallus; LBR, lateral borders of ramus of cingulum; LP, lateral plate of epiphallus; PZ, pseudoarch of ectophallus; RC, ramus of cingulum; S, sheath of ectophallus; SS, spermatophore sac; SZ, suprazygomal plate of cingulum; VC, ventral cleft of cingulum; VI, ventral inflection of endophallic apodeme; VMA, ventral margins of aedeagal valves; VP, ventral process of cingulum; VTC, ventral transverse thickening of cingulum; Z, zygoma of cingulum; I, length of bridge of epiphallus; II, length of lateral plate of epiphallus; III, interspace between apodemal plates of cingulum; IV, width of base of aedeagal sclerites; V, length of dorsal inflections of endophallus; VI, length of aedeagal sclerites and valves together (from the tip of aedeagus to the base of aedeagal sclerites).	FIGURE 5. Male genitalia of Sphenarium: epiphallus in dorsal view (A); ectophallus in dorsal (B), lateral (C) and posterior (D) view; and endophallus in lateral (E) and dorsal (F) view. A, appendix of epiphallus; AC, apodemal plate of cingulum; AP, anterior projection of epiphallus; AS, aedeagal sclerites; ASA, apical spine of aedeagus; AV, aedeagal valve; B, bridge of epiphallus; BAS, base of aedeagal sclerites; BC, basal thickening of cingulum; BE, basal emargination of cingulum; CM, central membrane; CV, cingulum valve; DI, dorsal inflection of endophallic apodeme; EA, endophallic apodeme; ISR, inflection of supraramus; L, lophus of epiphallus; LBR, lateral borders of ramus of cingulum; LP, lateral plate of epiphallus; PZ, pseudoarch of ectophallus; RC, ramus of cingulum; S, sheath of ectophallus; SS, spermatophore sac; SZ, suprazygomal plate of cingulum; VC, ventral cleft of cingulum; VI, ventral inflection of endophallic apodeme; VMA, ventral margins of aedeagal valves; VP, ventral process of cingulum; VTC, ventral transverse thickening of cingulum; Z, zygoma of cingulum; I, length of bridge of epiphallus; II, length of lateral plate of epiphallus; III, interspace between apodemal plates of cingulum; IV, width of base of aedeagal sclerites; V, length of dorsal inflections of endophallus; VI, length of aedeagal sclerites and valves together (from the tip of aedeagus to the base of aedeagal sclerites).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC103B702CBED4E5FE0FA85C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804204/files/figure.png	https://doi.org/10.5281/zenodo.804204	FIGURE 10. Male genital structures of S. purpurascens: morphotype 1 (A – C), morphotype 2 (D – F), morphotype 3 (G – I); and S. zapotecum sp. n. holotype (J – l). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	FIGURE 10. Male genital structures of S. purpurascens: morphotype 1 (A – C), morphotype 2 (D – F), morphotype 3 (G – I); and S. zapotecum sp. n. holotype (J – l). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC103B702CBED4E5FE0FA85C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804202/files/figure.png	https://doi.org/10.5281/zenodo.804202	FIGURE 9. External morphology of S. purpurascens: morphotype 1 m (A) and f (B), morphotype 2 m (C) and f (D), and morphotype 3 m (E) and f (F); S. zapotecum sp. n. holotype m (G) and paratype f # 23 (H); S. tarascum sp. n. holotype m (I) and paratype f # 2 (J,); S. planum m (K) and f (L); S. macrophallicum paratype m # 230 (M) and f (N); S. minimum m (O) and f (P); S. infernalis sp. n. holotype m (Q) and paratype f # 2 (R); S. rugosum morphotype 1 m (S) and f (T) (Scale bars = 1 cm).	FIGURE 9. External morphology of S. purpurascens: morphotype 1 m (A) and f (B), morphotype 2 m (C) and f (D), and morphotype 3 m (E) and f (F); S. zapotecum sp. n. holotype m (G) and paratype f # 23 (H); S. tarascum sp. n. holotype m (I) and paratype f # 2 (J,); S. planum m (K) and f (L); S. macrophallicum paratype m # 230 (M) and f (N); S. minimum m (O) and f (P); S. infernalis sp. n. holotype m (Q) and paratype f # 2 (R); S. rugosum morphotype 1 m (S) and f (T) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC103B702CBED4E5FE0FA85C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804198/files/figure.png	https://doi.org/10.5281/zenodo.804198	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC1B3B762CBED2FBFDA3A910.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804202/files/figure.png	https://doi.org/10.5281/zenodo.804202	FIGURE 9. External morphology of S. purpurascens: morphotype 1 m (A) and f (B), morphotype 2 m (C) and f (D), and morphotype 3 m (E) and f (F); S. zapotecum sp. n. holotype m (G) and paratype f # 23 (H); S. tarascum sp. n. holotype m (I) and paratype f # 2 (J,); S. planum m (K) and f (L); S. macrophallicum paratype m # 230 (M) and f (N); S. minimum m (O) and f (P); S. infernalis sp. n. holotype m (Q) and paratype f # 2 (R); S. rugosum morphotype 1 m (S) and f (T) (Scale bars = 1 cm).	FIGURE 9. External morphology of S. purpurascens: morphotype 1 m (A) and f (B), morphotype 2 m (C) and f (D), and morphotype 3 m (E) and f (F); S. zapotecum sp. n. holotype m (G) and paratype f # 23 (H); S. tarascum sp. n. holotype m (I) and paratype f # 2 (J,); S. planum m (K) and f (L); S. macrophallicum paratype m # 230 (M) and f (N); S. minimum m (O) and f (P); S. infernalis sp. n. holotype m (Q) and paratype f # 2 (R); S. rugosum morphotype 1 m (S) and f (T) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC1B3B762CBED2FBFDA3A910.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804204/files/figure.png	https://doi.org/10.5281/zenodo.804204	FIGURE 10. Male genital structures of S. purpurascens: morphotype 1 (A – C), morphotype 2 (D – F), morphotype 3 (G – I); and S. zapotecum sp. n. holotype (J – l). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	FIGURE 10. Male genital structures of S. purpurascens: morphotype 1 (A – C), morphotype 2 (D – F), morphotype 3 (G – I); and S. zapotecum sp. n. holotype (J – l). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC1B3B762CBED2FBFDA3A910.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804198/files/figure.png	https://doi.org/10.5281/zenodo.804198	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC1B3B762CBED2FBFDA3A910.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804206/files/figure.png	https://doi.org/10.5281/zenodo.804206	FIGURE 11. Bayesian phylogeny of Sphenarium species based on CO 1 sequences of Pedraza-Lara et al. (2015) and this study. The analysis was conducted using the substitution model estimated previously and under the same conditions above specified for the concatenated phylogenetic analysis in methods section. Terminals of different colours represent the 17 recognized species in this study. Black dots behind the nodes indicate PP values greater than 95 %. Black bottom bars are equal to 0.04 substitutions per site.	FIGURE 11. Bayesian phylogeny of Sphenarium species based on CO 1 sequences of Pedraza-Lara et al. (2015) and this study. The analysis was conducted using the substitution model estimated previously and under the same conditions above specified for the concatenated phylogenetic analysis in methods section. Terminals of different colours represent the 17 recognized species in this study. Black dots behind the nodes indicate PP values greater than 95 %. Black bottom bars are equal to 0.04 substitutions per site.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC193B772CBED403FE3AA95F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804202/files/figure.png	https://doi.org/10.5281/zenodo.804202	FIGURE 9. External morphology of S. purpurascens: morphotype 1 m (A) and f (B), morphotype 2 m (C) and f (D), and morphotype 3 m (E) and f (F); S. zapotecum sp. n. holotype m (G) and paratype f # 23 (H); S. tarascum sp. n. holotype m (I) and paratype f # 2 (J,); S. planum m (K) and f (L); S. macrophallicum paratype m # 230 (M) and f (N); S. minimum m (O) and f (P); S. infernalis sp. n. holotype m (Q) and paratype f # 2 (R); S. rugosum morphotype 1 m (S) and f (T) (Scale bars = 1 cm).	FIGURE 9. External morphology of S. purpurascens: morphotype 1 m (A) and f (B), morphotype 2 m (C) and f (D), and morphotype 3 m (E) and f (F); S. zapotecum sp. n. holotype m (G) and paratype f # 23 (H); S. tarascum sp. n. holotype m (I) and paratype f # 2 (J,); S. planum m (K) and f (L); S. macrophallicum paratype m # 230 (M) and f (N); S. minimum m (O) and f (P); S. infernalis sp. n. holotype m (Q) and paratype f # 2 (R); S. rugosum morphotype 1 m (S) and f (T) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC193B772CBED403FE3AA95F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804210/files/figure.png	https://doi.org/10.5281/zenodo.804210	FIGURE 12. Male genital structures of S. tarascum sp. n. holotype (A – C); S. planum (D – F); S. macrophallicum paratype # 234 (G – I); and S. minimum (J – L). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	FIGURE 12. Male genital structures of S. tarascum sp. n. holotype (A – C); S. planum (D – F); S. macrophallicum paratype # 234 (G – I); and S. minimum (J – L). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC193B772CBED403FE3AA95F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804198/files/figure.png	https://doi.org/10.5281/zenodo.804198	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC183B482CBED4CAFE1BA80B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804200/files/figure.png	https://doi.org/10.5281/zenodo.804200	FIGURE 8. Type specimens of S. purpurascens: lectotype m (A) and paralectotype f (B); S. planum: lectotype m (C) and paralectotype f (D); S. macrophallicum: holotype m (E) and allotype f (F); S. minimum lectotype m (G); S. affine lecototype m (H) (Scale bars = 1 cm).	FIGURE 8. Type specimens of S. purpurascens: lectotype m (A) and paralectotype f (B); S. planum: lectotype m (C) and paralectotype f (D); S. macrophallicum: holotype m (E) and allotype f (F); S. minimum lectotype m (G); S. affine lecototype m (H) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC183B482CBED4CAFE1BA80B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804210/files/figure.png	https://doi.org/10.5281/zenodo.804210	FIGURE 12. Male genital structures of S. tarascum sp. n. holotype (A – C); S. planum (D – F); S. macrophallicum paratype # 234 (G – I); and S. minimum (J – L). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	FIGURE 12. Male genital structures of S. tarascum sp. n. holotype (A – C); S. planum (D – F); S. macrophallicum paratype # 234 (G – I); and S. minimum (J – L). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC183B482CBED4CAFE1BA80B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804202/files/figure.png	https://doi.org/10.5281/zenodo.804202	FIGURE 9. External morphology of S. purpurascens: morphotype 1 m (A) and f (B), morphotype 2 m (C) and f (D), and morphotype 3 m (E) and f (F); S. zapotecum sp. n. holotype m (G) and paratype f # 23 (H); S. tarascum sp. n. holotype m (I) and paratype f # 2 (J,); S. planum m (K) and f (L); S. macrophallicum paratype m # 230 (M) and f (N); S. minimum m (O) and f (P); S. infernalis sp. n. holotype m (Q) and paratype f # 2 (R); S. rugosum morphotype 1 m (S) and f (T) (Scale bars = 1 cm).	FIGURE 9. External morphology of S. purpurascens: morphotype 1 m (A) and f (B), morphotype 2 m (C) and f (D), and morphotype 3 m (E) and f (F); S. zapotecum sp. n. holotype m (G) and paratype f # 23 (H); S. tarascum sp. n. holotype m (I) and paratype f # 2 (J,); S. planum m (K) and f (L); S. macrophallicum paratype m # 230 (M) and f (N); S. minimum m (O) and f (P); S. infernalis sp. n. holotype m (Q) and paratype f # 2 (R); S. rugosum morphotype 1 m (S) and f (T) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC183B482CBED4CAFE1BA80B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804198/files/figure.png	https://doi.org/10.5281/zenodo.804198	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC183B482CBED4CAFE1BA80B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804186/files/figure.png	https://doi.org/10.5281/zenodo.804186	FIGURE 2. Bayesian phylogeny of Sphenarium based on a concatenated analysis of 2527 nucleotide positions from five loci and 145 terminals (129 ingroup and 16 outgroup terminals). Higher-level phylogenetic relationships are shown on left-top box and ingroup relationships are magnified outside. Voucher and locality identifier numbers of the analysed specimens are indicated in bold characters in terminals names, except for those cases in which genetic information was retrieved from the GenBank. Different branch colours highlight the three mayor clades within the genus (brown, Clade 1; red, Clade 2; and purple, Clade 3). Different colours of the Sphenarium terminals represent the 17 taxa identified during the morphologic analysis. Black circles behind the nodes indicate PP values ± 95 %. For some important nodes we also showed the PP values in bold numbers. Black bottom bars in all cases are equal to 0.03 substitutions per site.	FIGURE 2. Bayesian phylogeny of Sphenarium based on a concatenated analysis of 2527 nucleotide positions from five loci and 145 terminals (129 ingroup and 16 outgroup terminals). Higher-level phylogenetic relationships are shown on left-top box and ingroup relationships are magnified outside. Voucher and locality identifier numbers of the analysed specimens are indicated in bold characters in terminals names, except for those cases in which genetic information was retrieved from the GenBank. Different branch colours highlight the three mayor clades within the genus (brown, Clade 1; red, Clade 2; and purple, Clade 3). Different colours of the Sphenarium terminals represent the 17 taxa identified during the morphologic analysis. Black circles behind the nodes indicate PP values ± 95 %. For some important nodes we also showed the PP values in bold numbers. Black bottom bars in all cases are equal to 0.03 substitutions per site.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC273B492CBED536FB1BA85E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804200/files/figure.png	https://doi.org/10.5281/zenodo.804200	FIGURE 8. Type specimens of S. purpurascens: lectotype m (A) and paralectotype f (B); S. planum: lectotype m (C) and paralectotype f (D); S. macrophallicum: holotype m (E) and allotype f (F); S. minimum lectotype m (G); S. affine lecototype m (H) (Scale bars = 1 cm).	FIGURE 8. Type specimens of S. purpurascens: lectotype m (A) and paralectotype f (B); S. planum: lectotype m (C) and paralectotype f (D); S. macrophallicum: holotype m (E) and allotype f (F); S. minimum lectotype m (G); S. affine lecototype m (H) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC273B492CBED536FB1BA85E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804210/files/figure.png	https://doi.org/10.5281/zenodo.804210	FIGURE 12. Male genital structures of S. tarascum sp. n. holotype (A – C); S. planum (D – F); S. macrophallicum paratype # 234 (G – I); and S. minimum (J – L). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	FIGURE 12. Male genital structures of S. tarascum sp. n. holotype (A – C); S. planum (D – F); S. macrophallicum paratype # 234 (G – I); and S. minimum (J – L). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC273B492CBED536FB1BA85E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804198/files/figure.png	https://doi.org/10.5281/zenodo.804198	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC243B4C2CBED2FBFB4EAD14.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804200/files/figure.png	https://doi.org/10.5281/zenodo.804200	FIGURE 8. Type specimens of S. purpurascens: lectotype m (A) and paralectotype f (B); S. planum: lectotype m (C) and paralectotype f (D); S. macrophallicum: holotype m (E) and allotype f (F); S. minimum lectotype m (G); S. affine lecototype m (H) (Scale bars = 1 cm).	FIGURE 8. Type specimens of S. purpurascens: lectotype m (A) and paralectotype f (B); S. planum: lectotype m (C) and paralectotype f (D); S. macrophallicum: holotype m (E) and allotype f (F); S. minimum lectotype m (G); S. affine lecototype m (H) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC243B4C2CBED2FBFB4EAD14.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804210/files/figure.png	https://doi.org/10.5281/zenodo.804210	FIGURE 12. Male genital structures of S. tarascum sp. n. holotype (A – C); S. planum (D – F); S. macrophallicum paratype # 234 (G – I); and S. minimum (J – L). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	FIGURE 12. Male genital structures of S. tarascum sp. n. holotype (A – C); S. planum (D – F); S. macrophallicum paratype # 234 (G – I); and S. minimum (J – L). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC243B4C2CBED2FBFB4EAD14.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804198/files/figure.png	https://doi.org/10.5281/zenodo.804198	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC243B4C2CBED2FBFB4EAD14.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804186/files/figure.png	https://doi.org/10.5281/zenodo.804186	FIGURE 2. Bayesian phylogeny of Sphenarium based on a concatenated analysis of 2527 nucleotide positions from five loci and 145 terminals (129 ingroup and 16 outgroup terminals). Higher-level phylogenetic relationships are shown on left-top box and ingroup relationships are magnified outside. Voucher and locality identifier numbers of the analysed specimens are indicated in bold characters in terminals names, except for those cases in which genetic information was retrieved from the GenBank. Different branch colours highlight the three mayor clades within the genus (brown, Clade 1; red, Clade 2; and purple, Clade 3). Different colours of the Sphenarium terminals represent the 17 taxa identified during the morphologic analysis. Black circles behind the nodes indicate PP values ± 95 %. For some important nodes we also showed the PP values in bold numbers. Black bottom bars in all cases are equal to 0.03 substitutions per site.	FIGURE 2. Bayesian phylogeny of Sphenarium based on a concatenated analysis of 2527 nucleotide positions from five loci and 145 terminals (129 ingroup and 16 outgroup terminals). Higher-level phylogenetic relationships are shown on left-top box and ingroup relationships are magnified outside. Voucher and locality identifier numbers of the analysed specimens are indicated in bold characters in terminals names, except for those cases in which genetic information was retrieved from the GenBank. Different branch colours highlight the three mayor clades within the genus (brown, Clade 1; red, Clade 2; and purple, Clade 3). Different colours of the Sphenarium terminals represent the 17 taxa identified during the morphologic analysis. Black circles behind the nodes indicate PP values ± 95 %. For some important nodes we also showed the PP values in bold numbers. Black bottom bars in all cases are equal to 0.03 substitutions per site.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC233B4E2CBED000FE70AB2B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804202/files/figure.png	https://doi.org/10.5281/zenodo.804202	FIGURE 9. External morphology of S. purpurascens: morphotype 1 m (A) and f (B), morphotype 2 m (C) and f (D), and morphotype 3 m (E) and f (F); S. zapotecum sp. n. holotype m (G) and paratype f # 23 (H); S. tarascum sp. n. holotype m (I) and paratype f # 2 (J,); S. planum m (K) and f (L); S. macrophallicum paratype m # 230 (M) and f (N); S. minimum m (O) and f (P); S. infernalis sp. n. holotype m (Q) and paratype f # 2 (R); S. rugosum morphotype 1 m (S) and f (T) (Scale bars = 1 cm).	FIGURE 9. External morphology of S. purpurascens: morphotype 1 m (A) and f (B), morphotype 2 m (C) and f (D), and morphotype 3 m (E) and f (F); S. zapotecum sp. n. holotype m (G) and paratype f # 23 (H); S. tarascum sp. n. holotype m (I) and paratype f # 2 (J,); S. planum m (K) and f (L); S. macrophallicum paratype m # 230 (M) and f (N); S. minimum m (O) and f (P); S. infernalis sp. n. holotype m (Q) and paratype f # 2 (R); S. rugosum morphotype 1 m (S) and f (T) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC233B4E2CBED000FE70AB2B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804212/files/figure.png	https://doi.org/10.5281/zenodo.804212	FIGURE 13. Male genital structures of S. infernalis sp. n. holotype (A – C); S. rugosum: morphotype 1 (D – F), morphotype 2 (G – I) and morphotype 3 (J – l). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	FIGURE 13. Male genital structures of S. infernalis sp. n. holotype (A – C); S. rugosum: morphotype 1 (D – F), morphotype 2 (G – I) and morphotype 3 (J – l). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC233B4E2CBED000FE70AB2B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804198/files/figure.png	https://doi.org/10.5281/zenodo.804198	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC233B4E2CBED000FE70AB2B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804186/files/figure.png	https://doi.org/10.5281/zenodo.804186	FIGURE 2. Bayesian phylogeny of Sphenarium based on a concatenated analysis of 2527 nucleotide positions from five loci and 145 terminals (129 ingroup and 16 outgroup terminals). Higher-level phylogenetic relationships are shown on left-top box and ingroup relationships are magnified outside. Voucher and locality identifier numbers of the analysed specimens are indicated in bold characters in terminals names, except for those cases in which genetic information was retrieved from the GenBank. Different branch colours highlight the three mayor clades within the genus (brown, Clade 1; red, Clade 2; and purple, Clade 3). Different colours of the Sphenarium terminals represent the 17 taxa identified during the morphologic analysis. Black circles behind the nodes indicate PP values ± 95 %. For some important nodes we also showed the PP values in bold numbers. Black bottom bars in all cases are equal to 0.03 substitutions per site.	FIGURE 2. Bayesian phylogeny of Sphenarium based on a concatenated analysis of 2527 nucleotide positions from five loci and 145 terminals (129 ingroup and 16 outgroup terminals). Higher-level phylogenetic relationships are shown on left-top box and ingroup relationships are magnified outside. Voucher and locality identifier numbers of the analysed specimens are indicated in bold characters in terminals names, except for those cases in which genetic information was retrieved from the GenBank. Different branch colours highlight the three mayor clades within the genus (brown, Clade 1; red, Clade 2; and purple, Clade 3). Different colours of the Sphenarium terminals represent the 17 taxa identified during the morphologic analysis. Black circles behind the nodes indicate PP values ± 95 %. For some important nodes we also showed the PP values in bold numbers. Black bottom bars in all cases are equal to 0.03 substitutions per site.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC213B402CBED603FEA0AFDF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804214/files/figure.png	https://doi.org/10.5281/zenodo.804214	FIGURE 14. Type specimens of S. rugosum: lectotype m (A) and paralectotype f (B); S. barrettii: lectotype m (C); S. borrei: lectotype m (D) and paralectotype f (E); S. bruneri lectotype m (F) and paralectotype f (G); S. variabile: holotype m (H) (Scale bars = 1 cm).	FIGURE 14. Type specimens of S. rugosum: lectotype m (A) and paralectotype f (B); S. barrettii: lectotype m (C); S. borrei: lectotype m (D) and paralectotype f (E); S. bruneri lectotype m (F) and paralectotype f (G); S. variabile: holotype m (H) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC213B402CBED603FEA0AFDF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804212/files/figure.png	https://doi.org/10.5281/zenodo.804212	FIGURE 13. Male genital structures of S. infernalis sp. n. holotype (A – C); S. rugosum: morphotype 1 (D – F), morphotype 2 (G – I) and morphotype 3 (J – l). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	FIGURE 13. Male genital structures of S. infernalis sp. n. holotype (A – C); S. rugosum: morphotype 1 (D – F), morphotype 2 (G – I) and morphotype 3 (J – l). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC213B402CBED603FEA0AFDF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804202/files/figure.png	https://doi.org/10.5281/zenodo.804202	FIGURE 9. External morphology of S. purpurascens: morphotype 1 m (A) and f (B), morphotype 2 m (C) and f (D), and morphotype 3 m (E) and f (F); S. zapotecum sp. n. holotype m (G) and paratype f # 23 (H); S. tarascum sp. n. holotype m (I) and paratype f # 2 (J,); S. planum m (K) and f (L); S. macrophallicum paratype m # 230 (M) and f (N); S. minimum m (O) and f (P); S. infernalis sp. n. holotype m (Q) and paratype f # 2 (R); S. rugosum morphotype 1 m (S) and f (T) (Scale bars = 1 cm).	FIGURE 9. External morphology of S. purpurascens: morphotype 1 m (A) and f (B), morphotype 2 m (C) and f (D), and morphotype 3 m (E) and f (F); S. zapotecum sp. n. holotype m (G) and paratype f # 23 (H); S. tarascum sp. n. holotype m (I) and paratype f # 2 (J,); S. planum m (K) and f (L); S. macrophallicum paratype m # 230 (M) and f (N); S. minimum m (O) and f (P); S. infernalis sp. n. holotype m (Q) and paratype f # 2 (R); S. rugosum morphotype 1 m (S) and f (T) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC213B402CBED603FEA0AFDF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804216/files/figure.png	https://doi.org/10.5281/zenodo.804216	FIGURE 15. External morphology of S. rugosum: morphotype 2 m (A) and f (B), morphotype 3 m (C) and f (D); S. crypticum sp. n. holotype m (E) and paratype f # 45 (F); S. borrei m (G) and f (H); S. variabile: morphotype 1 paratype m # 134 (I) and f (J), and morphotype 2 m (K) and f (L); S. mexicanum: morphotype 1 m (M) and f (N), and morphotype 2 m (O) and f (P); S. histrio: morphotype 1 m (Q and S) and f (R and T) (Scale bars = 1 cm).	FIGURE 15. External morphology of S. rugosum: morphotype 2 m (A) and f (B), morphotype 3 m (C) and f (D); S. crypticum sp. n. holotype m (E) and paratype f # 45 (F); S. borrei m (G) and f (H); S. variabile: morphotype 1 paratype m # 134 (I) and f (J), and morphotype 2 m (K) and f (L); S. mexicanum: morphotype 1 m (M) and f (N), and morphotype 2 m (O) and f (P); S. histrio: morphotype 1 m (Q and S) and f (R and T) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC213B402CBED603FEA0AFDF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804198/files/figure.png	https://doi.org/10.5281/zenodo.804198	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC213B402CBED603FEA0AFDF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804206/files/figure.png	https://doi.org/10.5281/zenodo.804206	FIGURE 11. Bayesian phylogeny of Sphenarium species based on CO 1 sequences of Pedraza-Lara et al. (2015) and this study. The analysis was conducted using the substitution model estimated previously and under the same conditions above specified for the concatenated phylogenetic analysis in methods section. Terminals of different colours represent the 17 recognized species in this study. Black dots behind the nodes indicate PP values greater than 95 %. Black bottom bars are equal to 0.04 substitutions per site.	FIGURE 11. Bayesian phylogeny of Sphenarium species based on CO 1 sequences of Pedraza-Lara et al. (2015) and this study. The analysis was conducted using the substitution model estimated previously and under the same conditions above specified for the concatenated phylogenetic analysis in methods section. Terminals of different colours represent the 17 recognized species in this study. Black dots behind the nodes indicate PP values greater than 95 %. Black bottom bars are equal to 0.04 substitutions per site.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC2F3B442CBED377FAD1AD14.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804216/files/figure.png	https://doi.org/10.5281/zenodo.804216	FIGURE 15. External morphology of S. rugosum: morphotype 2 m (A) and f (B), morphotype 3 m (C) and f (D); S. crypticum sp. n. holotype m (E) and paratype f # 45 (F); S. borrei m (G) and f (H); S. variabile: morphotype 1 paratype m # 134 (I) and f (J), and morphotype 2 m (K) and f (L); S. mexicanum: morphotype 1 m (M) and f (N), and morphotype 2 m (O) and f (P); S. histrio: morphotype 1 m (Q and S) and f (R and T) (Scale bars = 1 cm).	FIGURE 15. External morphology of S. rugosum: morphotype 2 m (A) and f (B), morphotype 3 m (C) and f (D); S. crypticum sp. n. holotype m (E) and paratype f # 45 (F); S. borrei m (G) and f (H); S. variabile: morphotype 1 paratype m # 134 (I) and f (J), and morphotype 2 m (K) and f (L); S. mexicanum: morphotype 1 m (M) and f (N), and morphotype 2 m (O) and f (P); S. histrio: morphotype 1 m (Q and S) and f (R and T) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC2F3B442CBED377FAD1AD14.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804218/files/figure.png	https://doi.org/10.5281/zenodo.804218	FIGURE 16. Male genital structures of S. crypticum sp. n. holotype (A – C); S. borrei (D – F); S. variabile: morphotype 1 paratype # 234 (G – I) and morphotype 2 (J – l). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	FIGURE 16. Male genital structures of S. crypticum sp. n. holotype (A – C); S. borrei (D – F); S. variabile: morphotype 1 paratype # 234 (G – I) and morphotype 2 (J – l). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC2F3B442CBED377FAD1AD14.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804198/files/figure.png	https://doi.org/10.5281/zenodo.804198	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC2F3B442CBED377FAD1AD14.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804186/files/figure.png	https://doi.org/10.5281/zenodo.804186	FIGURE 2. Bayesian phylogeny of Sphenarium based on a concatenated analysis of 2527 nucleotide positions from five loci and 145 terminals (129 ingroup and 16 outgroup terminals). Higher-level phylogenetic relationships are shown on left-top box and ingroup relationships are magnified outside. Voucher and locality identifier numbers of the analysed specimens are indicated in bold characters in terminals names, except for those cases in which genetic information was retrieved from the GenBank. Different branch colours highlight the three mayor clades within the genus (brown, Clade 1; red, Clade 2; and purple, Clade 3). Different colours of the Sphenarium terminals represent the 17 taxa identified during the morphologic analysis. Black circles behind the nodes indicate PP values ± 95 %. For some important nodes we also showed the PP values in bold numbers. Black bottom bars in all cases are equal to 0.03 substitutions per site.	FIGURE 2. Bayesian phylogeny of Sphenarium based on a concatenated analysis of 2527 nucleotide positions from five loci and 145 terminals (129 ingroup and 16 outgroup terminals). Higher-level phylogenetic relationships are shown on left-top box and ingroup relationships are magnified outside. Voucher and locality identifier numbers of the analysed specimens are indicated in bold characters in terminals names, except for those cases in which genetic information was retrieved from the GenBank. Different branch colours highlight the three mayor clades within the genus (brown, Clade 1; red, Clade 2; and purple, Clade 3). Different colours of the Sphenarium terminals represent the 17 taxa identified during the morphologic analysis. Black circles behind the nodes indicate PP values ± 95 %. For some important nodes we also showed the PP values in bold numbers. Black bottom bars in all cases are equal to 0.03 substitutions per site.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC2F3B442CBED377FAD1AD14.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804206/files/figure.png	https://doi.org/10.5281/zenodo.804206	FIGURE 11. Bayesian phylogeny of Sphenarium species based on CO 1 sequences of Pedraza-Lara et al. (2015) and this study. The analysis was conducted using the substitution model estimated previously and under the same conditions above specified for the concatenated phylogenetic analysis in methods section. Terminals of different colours represent the 17 recognized species in this study. Black dots behind the nodes indicate PP values greater than 95 %. Black bottom bars are equal to 0.04 substitutions per site.	FIGURE 11. Bayesian phylogeny of Sphenarium species based on CO 1 sequences of Pedraza-Lara et al. (2015) and this study. The analysis was conducted using the substitution model estimated previously and under the same conditions above specified for the concatenated phylogenetic analysis in methods section. Terminals of different colours represent the 17 recognized species in this study. Black dots behind the nodes indicate PP values greater than 95 %. Black bottom bars are equal to 0.04 substitutions per site.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC2B3B452CBED000FF5FACE1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804214/files/figure.png	https://doi.org/10.5281/zenodo.804214	FIGURE 14. Type specimens of S. rugosum: lectotype m (A) and paralectotype f (B); S. barrettii: lectotype m (C); S. borrei: lectotype m (D) and paralectotype f (E); S. bruneri lectotype m (F) and paralectotype f (G); S. variabile: holotype m (H) (Scale bars = 1 cm).	FIGURE 14. Type specimens of S. rugosum: lectotype m (A) and paralectotype f (B); S. barrettii: lectotype m (C); S. borrei: lectotype m (D) and paralectotype f (E); S. bruneri lectotype m (F) and paralectotype f (G); S. variabile: holotype m (H) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC2B3B452CBED000FF5FACE1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804218/files/figure.png	https://doi.org/10.5281/zenodo.804218	FIGURE 16. Male genital structures of S. crypticum sp. n. holotype (A – C); S. borrei (D – F); S. variabile: morphotype 1 paratype # 234 (G – I) and morphotype 2 (J – l). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	FIGURE 16. Male genital structures of S. crypticum sp. n. holotype (A – C); S. borrei (D – F); S. variabile: morphotype 1 paratype # 234 (G – I) and morphotype 2 (J – l). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC2B3B452CBED000FF5FACE1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804216/files/figure.png	https://doi.org/10.5281/zenodo.804216	FIGURE 15. External morphology of S. rugosum: morphotype 2 m (A) and f (B), morphotype 3 m (C) and f (D); S. crypticum sp. n. holotype m (E) and paratype f # 45 (F); S. borrei m (G) and f (H); S. variabile: morphotype 1 paratype m # 134 (I) and f (J), and morphotype 2 m (K) and f (L); S. mexicanum: morphotype 1 m (M) and f (N), and morphotype 2 m (O) and f (P); S. histrio: morphotype 1 m (Q and S) and f (R and T) (Scale bars = 1 cm).	FIGURE 15. External morphology of S. rugosum: morphotype 2 m (A) and f (B), morphotype 3 m (C) and f (D); S. crypticum sp. n. holotype m (E) and paratype f # 45 (F); S. borrei m (G) and f (H); S. variabile: morphotype 1 paratype m # 134 (I) and f (J), and morphotype 2 m (K) and f (L); S. mexicanum: morphotype 1 m (M) and f (N), and morphotype 2 m (O) and f (P); S. histrio: morphotype 1 m (Q and S) and f (R and T) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC2B3B452CBED000FF5FACE1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804198/files/figure.png	https://doi.org/10.5281/zenodo.804198	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC2A3B462CBED653FB46AA03.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804214/files/figure.png	https://doi.org/10.5281/zenodo.804214	FIGURE 14. Type specimens of S. rugosum: lectotype m (A) and paralectotype f (B); S. barrettii: lectotype m (C); S. borrei: lectotype m (D) and paralectotype f (E); S. bruneri lectotype m (F) and paralectotype f (G); S. variabile: holotype m (H) (Scale bars = 1 cm).	FIGURE 14. Type specimens of S. rugosum: lectotype m (A) and paralectotype f (B); S. barrettii: lectotype m (C); S. borrei: lectotype m (D) and paralectotype f (E); S. bruneri lectotype m (F) and paralectotype f (G); S. variabile: holotype m (H) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC2A3B462CBED653FB46AA03.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804218/files/figure.png	https://doi.org/10.5281/zenodo.804218	FIGURE 16. Male genital structures of S. crypticum sp. n. holotype (A – C); S. borrei (D – F); S. variabile: morphotype 1 paratype # 234 (G – I) and morphotype 2 (J – l). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	FIGURE 16. Male genital structures of S. crypticum sp. n. holotype (A – C); S. borrei (D – F); S. variabile: morphotype 1 paratype # 234 (G – I) and morphotype 2 (J – l). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC2A3B462CBED653FB46AA03.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804216/files/figure.png	https://doi.org/10.5281/zenodo.804216	FIGURE 15. External morphology of S. rugosum: morphotype 2 m (A) and f (B), morphotype 3 m (C) and f (D); S. crypticum sp. n. holotype m (E) and paratype f # 45 (F); S. borrei m (G) and f (H); S. variabile: morphotype 1 paratype m # 134 (I) and f (J), and morphotype 2 m (K) and f (L); S. mexicanum: morphotype 1 m (M) and f (N), and morphotype 2 m (O) and f (P); S. histrio: morphotype 1 m (Q and S) and f (R and T) (Scale bars = 1 cm).	FIGURE 15. External morphology of S. rugosum: morphotype 2 m (A) and f (B), morphotype 3 m (C) and f (D); S. crypticum sp. n. holotype m (E) and paratype f # 45 (F); S. borrei m (G) and f (H); S. variabile: morphotype 1 paratype m # 134 (I) and f (J), and morphotype 2 m (K) and f (L); S. mexicanum: morphotype 1 m (M) and f (N), and morphotype 2 m (O) and f (P); S. histrio: morphotype 1 m (Q and S) and f (R and T) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC2A3B462CBED653FB46AA03.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804202/files/figure.png	https://doi.org/10.5281/zenodo.804202	FIGURE 9. External morphology of S. purpurascens: morphotype 1 m (A) and f (B), morphotype 2 m (C) and f (D), and morphotype 3 m (E) and f (F); S. zapotecum sp. n. holotype m (G) and paratype f # 23 (H); S. tarascum sp. n. holotype m (I) and paratype f # 2 (J,); S. planum m (K) and f (L); S. macrophallicum paratype m # 230 (M) and f (N); S. minimum m (O) and f (P); S. infernalis sp. n. holotype m (Q) and paratype f # 2 (R); S. rugosum morphotype 1 m (S) and f (T) (Scale bars = 1 cm).	FIGURE 9. External morphology of S. purpurascens: morphotype 1 m (A) and f (B), morphotype 2 m (C) and f (D), and morphotype 3 m (E) and f (F); S. zapotecum sp. n. holotype m (G) and paratype f # 23 (H); S. tarascum sp. n. holotype m (I) and paratype f # 2 (J,); S. planum m (K) and f (L); S. macrophallicum paratype m # 230 (M) and f (N); S. minimum m (O) and f (P); S. infernalis sp. n. holotype m (Q) and paratype f # 2 (R); S. rugosum morphotype 1 m (S) and f (T) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC2A3B462CBED653FB46AA03.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804198/files/figure.png	https://doi.org/10.5281/zenodo.804198	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC2A3B462CBED653FB46AA03.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804220/files/figure.png	https://doi.org/10.5281/zenodo.804220	FIGURE 17. Type specimens of S. variabile: allotype f (A); S. mexicanum lectotype f (B) and paralectotype m (C); O. crassipes holotype m (D); S. ictericum: lectotype m (E) and paralectotype f (F); S. marginatum lectotype m (G) and paralectotype f (H) (Scale bars = 1 cm).	FIGURE 17. Type specimens of S. variabile: allotype f (A); S. mexicanum lectotype f (B) and paralectotype m (C); O. crassipes holotype m (D); S. ictericum: lectotype m (E) and paralectotype f (F); S. marginatum lectotype m (G) and paralectotype f (H) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC2A3B462CBED653FB46AA03.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804210/files/figure.png	https://doi.org/10.5281/zenodo.804210	FIGURE 12. Male genital structures of S. tarascum sp. n. holotype (A – C); S. planum (D – F); S. macrophallicum paratype # 234 (G – I); and S. minimum (J – L). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	FIGURE 12. Male genital structures of S. tarascum sp. n. holotype (A – C); S. planum (D – F); S. macrophallicum paratype # 234 (G – I); and S. minimum (J – L). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC293B5A2CBED70EFEBDAE33.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804220/files/figure.png	https://doi.org/10.5281/zenodo.804220	FIGURE 17. Type specimens of S. variabile: allotype f (A); S. mexicanum lectotype f (B) and paralectotype m (C); O. crassipes holotype m (D); S. ictericum: lectotype m (E) and paralectotype f (F); S. marginatum lectotype m (G) and paralectotype f (H) (Scale bars = 1 cm).	FIGURE 17. Type specimens of S. variabile: allotype f (A); S. mexicanum lectotype f (B) and paralectotype m (C); O. crassipes holotype m (D); S. ictericum: lectotype m (E) and paralectotype f (F); S. marginatum lectotype m (G) and paralectotype f (H) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC293B5A2CBED70EFEBDAE33.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804224/files/figure.png	https://doi.org/10.5281/zenodo.804224	FIGURE 19. Male genital structures of S. mexicanum: morphotype 1 (A – C) and morphotype 2 (D – F); S. histrio: morphotype 1 (G – I) and morphotype 2 (J – l). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	FIGURE 19. Male genital structures of S. mexicanum: morphotype 1 (A – C) and morphotype 2 (D – F); S. histrio: morphotype 1 (G – I) and morphotype 2 (J – l). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC293B5A2CBED70EFEBDAE33.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804216/files/figure.png	https://doi.org/10.5281/zenodo.804216	FIGURE 15. External morphology of S. rugosum: morphotype 2 m (A) and f (B), morphotype 3 m (C) and f (D); S. crypticum sp. n. holotype m (E) and paratype f # 45 (F); S. borrei m (G) and f (H); S. variabile: morphotype 1 paratype m # 134 (I) and f (J), and morphotype 2 m (K) and f (L); S. mexicanum: morphotype 1 m (M) and f (N), and morphotype 2 m (O) and f (P); S. histrio: morphotype 1 m (Q and S) and f (R and T) (Scale bars = 1 cm).	FIGURE 15. External morphology of S. rugosum: morphotype 2 m (A) and f (B), morphotype 3 m (C) and f (D); S. crypticum sp. n. holotype m (E) and paratype f # 45 (F); S. borrei m (G) and f (H); S. variabile: morphotype 1 paratype m # 134 (I) and f (J), and morphotype 2 m (K) and f (L); S. mexicanum: morphotype 1 m (M) and f (N), and morphotype 2 m (O) and f (P); S. histrio: morphotype 1 m (Q and S) and f (R and T) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC293B5A2CBED70EFEBDAE33.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804198/files/figure.png	https://doi.org/10.5281/zenodo.804198	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC293B5A2CBED70EFEBDAE33.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804222/files/figure.png	https://doi.org/10.5281/zenodo.804222	FIGURE 18. Type specimens of S. bolivari: lectotype m (A) and paralectotype m (B); S. histrio holotype m (C) and S. carinatum holotype m (D) (Scale bars = 1 cm).	FIGURE 18. Type specimens of S. bolivari: lectotype m (A) and paralectotype m (B); S. histrio holotype m (C) and S. carinatum holotype m (D) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC353B5B2CBED3E5FCC8A90B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804216/files/figure.png	https://doi.org/10.5281/zenodo.804216	FIGURE 15. External morphology of S. rugosum: morphotype 2 m (A) and f (B), morphotype 3 m (C) and f (D); S. crypticum sp. n. holotype m (E) and paratype f # 45 (F); S. borrei m (G) and f (H); S. variabile: morphotype 1 paratype m # 134 (I) and f (J), and morphotype 2 m (K) and f (L); S. mexicanum: morphotype 1 m (M) and f (N), and morphotype 2 m (O) and f (P); S. histrio: morphotype 1 m (Q and S) and f (R and T) (Scale bars = 1 cm).	FIGURE 15. External morphology of S. rugosum: morphotype 2 m (A) and f (B), morphotype 3 m (C) and f (D); S. crypticum sp. n. holotype m (E) and paratype f # 45 (F); S. borrei m (G) and f (H); S. variabile: morphotype 1 paratype m # 134 (I) and f (J), and morphotype 2 m (K) and f (L); S. mexicanum: morphotype 1 m (M) and f (N), and morphotype 2 m (O) and f (P); S. histrio: morphotype 1 m (Q and S) and f (R and T) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC353B5B2CBED3E5FCC8A90B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804224/files/figure.png	https://doi.org/10.5281/zenodo.804224	FIGURE 19. Male genital structures of S. mexicanum: morphotype 1 (A – C) and morphotype 2 (D – F); S. histrio: morphotype 1 (G – I) and morphotype 2 (J – l). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	FIGURE 19. Male genital structures of S. mexicanum: morphotype 1 (A – C) and morphotype 2 (D – F); S. histrio: morphotype 1 (G – I) and morphotype 2 (J – l). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC353B5B2CBED3E5FCC8A90B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804228/files/figure.png	https://doi.org/10.5281/zenodo.804228	FIGURE 21. Male genital structures of S. histrio: morphotype 3 (A – C), morphotype 4 (D – F), and morphotype 5 (G – I); and S. occidentalis sp. n. holotype (J – l). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	FIGURE 21. Male genital structures of S. histrio: morphotype 3 (A – C), morphotype 4 (D – F), and morphotype 5 (G – I); and S. occidentalis sp. n. holotype (J – l). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC353B5B2CBED3E5FCC8A90B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804226/files/figure.png	https://doi.org/10.5281/zenodo.804226	FIGURE 20. External morphology of S. histrio: morphotype 2 m (A and C) and f (B and D), morphotype 3 m (E) and f (F), morphotype 4 m (G) and f (H), and morphotype 5 m (I) and f (J); S. occidentalis sp. n. holotype m (K) and paratypes m # 2 (M), f # 3 (L) and f # 4 (N); S. totonacum sp. n. holotype m (O) and paratype f # 1 (P); S. adelinae sp. n. holotype m (Q) and paratype f # 1 (R); S. miztecum sp. n. holotype m (S) and paratype f # 1 (T) (Scale bars = 1 cm).	FIGURE 20. External morphology of S. histrio: morphotype 2 m (A and C) and f (B and D), morphotype 3 m (E) and f (F), morphotype 4 m (G) and f (H), and morphotype 5 m (I) and f (J); S. occidentalis sp. n. holotype m (K) and paratypes m # 2 (M), f # 3 (L) and f # 4 (N); S. totonacum sp. n. holotype m (O) and paratype f # 1 (P); S. adelinae sp. n. holotype m (Q) and paratype f # 1 (R); S. miztecum sp. n. holotype m (S) and paratype f # 1 (T) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC353B5B2CBED3E5FCC8A90B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804198/files/figure.png	https://doi.org/10.5281/zenodo.804198	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC353B5B2CBED3E5FCC8A90B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804222/files/figure.png	https://doi.org/10.5281/zenodo.804222	FIGURE 18. Type specimens of S. bolivari: lectotype m (A) and paralectotype m (B); S. histrio holotype m (C) and S. carinatum holotype m (D) (Scale bars = 1 cm).	FIGURE 18. Type specimens of S. bolivari: lectotype m (A) and paralectotype m (B); S. histrio holotype m (C) and S. carinatum holotype m (D) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC353B5B2CBED3E5FCC8A90B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804206/files/figure.png	https://doi.org/10.5281/zenodo.804206	FIGURE 11. Bayesian phylogeny of Sphenarium species based on CO 1 sequences of Pedraza-Lara et al. (2015) and this study. The analysis was conducted using the substitution model estimated previously and under the same conditions above specified for the concatenated phylogenetic analysis in methods section. Terminals of different colours represent the 17 recognized species in this study. Black dots behind the nodes indicate PP values greater than 95 %. Black bottom bars are equal to 0.04 substitutions per site.	FIGURE 11. Bayesian phylogeny of Sphenarium species based on CO 1 sequences of Pedraza-Lara et al. (2015) and this study. The analysis was conducted using the substitution model estimated previously and under the same conditions above specified for the concatenated phylogenetic analysis in methods section. Terminals of different colours represent the 17 recognized species in this study. Black dots behind the nodes indicate PP values greater than 95 %. Black bottom bars are equal to 0.04 substitutions per site.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC343B5D2CBED43DFB1EAE4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804226/files/figure.png	https://doi.org/10.5281/zenodo.804226	FIGURE 20. External morphology of S. histrio: morphotype 2 m (A and C) and f (B and D), morphotype 3 m (E) and f (F), morphotype 4 m (G) and f (H), and morphotype 5 m (I) and f (J); S. occidentalis sp. n. holotype m (K) and paratypes m # 2 (M), f # 3 (L) and f # 4 (N); S. totonacum sp. n. holotype m (O) and paratype f # 1 (P); S. adelinae sp. n. holotype m (Q) and paratype f # 1 (R); S. miztecum sp. n. holotype m (S) and paratype f # 1 (T) (Scale bars = 1 cm).	FIGURE 20. External morphology of S. histrio: morphotype 2 m (A and C) and f (B and D), morphotype 3 m (E) and f (F), morphotype 4 m (G) and f (H), and morphotype 5 m (I) and f (J); S. occidentalis sp. n. holotype m (K) and paratypes m # 2 (M), f # 3 (L) and f # 4 (N); S. totonacum sp. n. holotype m (O) and paratype f # 1 (P); S. adelinae sp. n. holotype m (Q) and paratype f # 1 (R); S. miztecum sp. n. holotype m (S) and paratype f # 1 (T) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC343B5D2CBED43DFB1EAE4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804228/files/figure.png	https://doi.org/10.5281/zenodo.804228	FIGURE 21. Male genital structures of S. histrio: morphotype 3 (A – C), morphotype 4 (D – F), and morphotype 5 (G – I); and S. occidentalis sp. n. holotype (J – l). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	FIGURE 21. Male genital structures of S. histrio: morphotype 3 (A – C), morphotype 4 (D – F), and morphotype 5 (G – I); and S. occidentalis sp. n. holotype (J – l). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC343B5D2CBED43DFB1EAE4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804198/files/figure.png	https://doi.org/10.5281/zenodo.804198	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC343B5D2CBED43DFB1EAE4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804206/files/figure.png	https://doi.org/10.5281/zenodo.804206	FIGURE 11. Bayesian phylogeny of Sphenarium species based on CO 1 sequences of Pedraza-Lara et al. (2015) and this study. The analysis was conducted using the substitution model estimated previously and under the same conditions above specified for the concatenated phylogenetic analysis in methods section. Terminals of different colours represent the 17 recognized species in this study. Black dots behind the nodes indicate PP values greater than 95 %. Black bottom bars are equal to 0.04 substitutions per site.	FIGURE 11. Bayesian phylogeny of Sphenarium species based on CO 1 sequences of Pedraza-Lara et al. (2015) and this study. The analysis was conducted using the substitution model estimated previously and under the same conditions above specified for the concatenated phylogenetic analysis in methods section. Terminals of different colours represent the 17 recognized species in this study. Black dots behind the nodes indicate PP values greater than 95 %. Black bottom bars are equal to 0.04 substitutions per site.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC323B512CBED3E8FE45A988.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804226/files/figure.png	https://doi.org/10.5281/zenodo.804226	FIGURE 20. External morphology of S. histrio: morphotype 2 m (A and C) and f (B and D), morphotype 3 m (E) and f (F), morphotype 4 m (G) and f (H), and morphotype 5 m (I) and f (J); S. occidentalis sp. n. holotype m (K) and paratypes m # 2 (M), f # 3 (L) and f # 4 (N); S. totonacum sp. n. holotype m (O) and paratype f # 1 (P); S. adelinae sp. n. holotype m (Q) and paratype f # 1 (R); S. miztecum sp. n. holotype m (S) and paratype f # 1 (T) (Scale bars = 1 cm).	FIGURE 20. External morphology of S. histrio: morphotype 2 m (A and C) and f (B and D), morphotype 3 m (E) and f (F), morphotype 4 m (G) and f (H), and morphotype 5 m (I) and f (J); S. occidentalis sp. n. holotype m (K) and paratypes m # 2 (M), f # 3 (L) and f # 4 (N); S. totonacum sp. n. holotype m (O) and paratype f # 1 (P); S. adelinae sp. n. holotype m (Q) and paratype f # 1 (R); S. miztecum sp. n. holotype m (S) and paratype f # 1 (T) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC323B512CBED3E8FE45A988.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804232/files/figure.png	https://doi.org/10.5281/zenodo.804232	FIGURE 22. Male genital structures of S. totonacum sp. n. holotype (A – C), S. adelinae sp. n. holotype (D – F); and S. miztecum sp. n. holotype (G – I). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	FIGURE 22. Male genital structures of S. totonacum sp. n. holotype (A – C), S. adelinae sp. n. holotype (D – F); and S. miztecum sp. n. holotype (G – I). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC323B512CBED3E8FE45A988.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804198/files/figure.png	https://doi.org/10.5281/zenodo.804198	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC3E3B532CBED4BBFACCAEF8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804226/files/figure.png	https://doi.org/10.5281/zenodo.804226	FIGURE 20. External morphology of S. histrio: morphotype 2 m (A and C) and f (B and D), morphotype 3 m (E) and f (F), morphotype 4 m (G) and f (H), and morphotype 5 m (I) and f (J); S. occidentalis sp. n. holotype m (K) and paratypes m # 2 (M), f # 3 (L) and f # 4 (N); S. totonacum sp. n. holotype m (O) and paratype f # 1 (P); S. adelinae sp. n. holotype m (Q) and paratype f # 1 (R); S. miztecum sp. n. holotype m (S) and paratype f # 1 (T) (Scale bars = 1 cm).	FIGURE 20. External morphology of S. histrio: morphotype 2 m (A and C) and f (B and D), morphotype 3 m (E) and f (F), morphotype 4 m (G) and f (H), and morphotype 5 m (I) and f (J); S. occidentalis sp. n. holotype m (K) and paratypes m # 2 (M), f # 3 (L) and f # 4 (N); S. totonacum sp. n. holotype m (O) and paratype f # 1 (P); S. adelinae sp. n. holotype m (Q) and paratype f # 1 (R); S. miztecum sp. n. holotype m (S) and paratype f # 1 (T) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC3E3B532CBED4BBFACCAEF8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804232/files/figure.png	https://doi.org/10.5281/zenodo.804232	FIGURE 22. Male genital structures of S. totonacum sp. n. holotype (A – C), S. adelinae sp. n. holotype (D – F); and S. miztecum sp. n. holotype (G – I). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	FIGURE 22. Male genital structures of S. totonacum sp. n. holotype (A – C), S. adelinae sp. n. holotype (D – F); and S. miztecum sp. n. holotype (G – I). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC3E3B532CBED4BBFACCAEF8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804198/files/figure.png	https://doi.org/10.5281/zenodo.804198	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC3E3B532CBED4BBFACCAEF8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804206/files/figure.png	https://doi.org/10.5281/zenodo.804206	FIGURE 11. Bayesian phylogeny of Sphenarium species based on CO 1 sequences of Pedraza-Lara et al. (2015) and this study. The analysis was conducted using the substitution model estimated previously and under the same conditions above specified for the concatenated phylogenetic analysis in methods section. Terminals of different colours represent the 17 recognized species in this study. Black dots behind the nodes indicate PP values greater than 95 %. Black bottom bars are equal to 0.04 substitutions per site.	FIGURE 11. Bayesian phylogeny of Sphenarium species based on CO 1 sequences of Pedraza-Lara et al. (2015) and this study. The analysis was conducted using the substitution model estimated previously and under the same conditions above specified for the concatenated phylogenetic analysis in methods section. Terminals of different colours represent the 17 recognized species in this study. Black dots behind the nodes indicate PP values greater than 95 %. Black bottom bars are equal to 0.04 substitutions per site.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC3C3B542CBED02BFE45AE84.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804226/files/figure.png	https://doi.org/10.5281/zenodo.804226	FIGURE 20. External morphology of S. histrio: morphotype 2 m (A and C) and f (B and D), morphotype 3 m (E) and f (F), morphotype 4 m (G) and f (H), and morphotype 5 m (I) and f (J); S. occidentalis sp. n. holotype m (K) and paratypes m # 2 (M), f # 3 (L) and f # 4 (N); S. totonacum sp. n. holotype m (O) and paratype f # 1 (P); S. adelinae sp. n. holotype m (Q) and paratype f # 1 (R); S. miztecum sp. n. holotype m (S) and paratype f # 1 (T) (Scale bars = 1 cm).	FIGURE 20. External morphology of S. histrio: morphotype 2 m (A and C) and f (B and D), morphotype 3 m (E) and f (F), morphotype 4 m (G) and f (H), and morphotype 5 m (I) and f (J); S. occidentalis sp. n. holotype m (K) and paratypes m # 2 (M), f # 3 (L) and f # 4 (N); S. totonacum sp. n. holotype m (O) and paratype f # 1 (P); S. adelinae sp. n. holotype m (Q) and paratype f # 1 (R); S. miztecum sp. n. holotype m (S) and paratype f # 1 (T) (Scale bars = 1 cm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC3C3B542CBED02BFE45AE84.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804232/files/figure.png	https://doi.org/10.5281/zenodo.804232	FIGURE 22. Male genital structures of S. totonacum sp. n. holotype (A – C), S. adelinae sp. n. holotype (D – F); and S. miztecum sp. n. holotype (G – I). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	FIGURE 22. Male genital structures of S. totonacum sp. n. holotype (A – C), S. adelinae sp. n. holotype (D – F); and S. miztecum sp. n. holotype (G – I). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC3C3B542CBED02BFE45AE84.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804198/files/figure.png	https://doi.org/10.5281/zenodo.804198	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC453B2A2CBED7C4FA3CA8E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804192/files/figure.png	https://doi.org/10.5281/zenodo.804192	FIGURE 4. External morphologic characters of Sphenarium and Prosphena: antennae filiform (A) or weakly ensiform (B); head subtriangular-compresed (C), subtriangular-elongated (D) or conical (E, F); tegmina spatula-like (G), strap-like (H) or tongue-like (I); subgenital plate of males tapered (J) or rounded moderately (K) or notably (L) developed posteriorly; dorsal ovipositor valves rounded (M), moderately lanceolate (N) or notably elongated (O).	FIGURE 4. External morphologic characters of Sphenarium and Prosphena: antennae filiform (A) or weakly ensiform (B); head subtriangular-compresed (C), subtriangular-elongated (D) or conical (E, F); tegmina spatula-like (G), strap-like (H) or tongue-like (I); subgenital plate of males tapered (J) or rounded moderately (K) or notably (L) developed posteriorly; dorsal ovipositor valves rounded (M), moderately lanceolate (N) or notably elongated (O).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC453B2A2CBED7C4FA3CA8E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804204/files/figure.png	https://doi.org/10.5281/zenodo.804204	FIGURE 10. Male genital structures of S. purpurascens: morphotype 1 (A – C), morphotype 2 (D – F), morphotype 3 (G – I); and S. zapotecum sp. n. holotype (J – l). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	FIGURE 10. Male genital structures of S. purpurascens: morphotype 1 (A – C), morphotype 2 (D – F), morphotype 3 (G – I); and S. zapotecum sp. n. holotype (J – l). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC453B2A2CBED7C4FA3CA8E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804218/files/figure.png	https://doi.org/10.5281/zenodo.804218	FIGURE 16. Male genital structures of S. crypticum sp. n. holotype (A – C); S. borrei (D – F); S. variabile: morphotype 1 paratype # 234 (G – I) and morphotype 2 (J – l). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	FIGURE 16. Male genital structures of S. crypticum sp. n. holotype (A – C); S. borrei (D – F); S. variabile: morphotype 1 paratype # 234 (G – I) and morphotype 2 (J – l). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC453B2A2CBED7C4FA3CA8E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804210/files/figure.png	https://doi.org/10.5281/zenodo.804210	FIGURE 12. Male genital structures of S. tarascum sp. n. holotype (A – C); S. planum (D – F); S. macrophallicum paratype # 234 (G – I); and S. minimum (J – L). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	FIGURE 12. Male genital structures of S. tarascum sp. n. holotype (A – C); S. planum (D – F); S. macrophallicum paratype # 234 (G – I); and S. minimum (J – L). For all taxa left squares show epiphallus (I) and ectophallus (II) in dorsal view, and endophallus in lateral view (III); meddle squares show ectophallus in posterior view; and right squares show a close up of ectophallus in lateral view (Scale bars = 1 mm).	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
A12C6E25AC453B2A2CBED7C4FA3CA8E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/804198/files/figure.png	https://doi.org/10.5281/zenodo.804198	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	FIGURE 7. Geographic distribution of Sphenarium species. Numbers within parenthesis in front of taxa names indicate the number of identified morphotype within the species. White surrounded areas and upper case abbreviations denote the Mexican biogeographic provinces. AC, Altos de Chiapas; AL, Altiplano Sur; BRB, Balsas River Basin; GMC, Gulf of Mexico Coast; MVB, Mexican Volcanic Belt; PC, Pacific Coast; SMOC, Sierra Madre Occidental; SMOR, Sierra Madre Oriental; SMS, Sierra Madre Sur; SO, Soconusco; and SOX, Sierra de Oaxaca.	2017-12-31	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del		Zenodo	biologists	Sanabria-Urbán, Salomón;Song, Hojun;Oyama, Ken;González-Rodríguez, Antonio;Castillo, Raúl Cueva Del			
