taxonID	type	description	language	source
A703878CA949FFBCFF26FF759EF7087E.taxon	discussion	Discussion. Lycoriella (Hemineurina) dearmata Mohrig & Krivosheina, 1987 is known only from a few specimens from Russia, Tuva and Altai Republics and Germany, Schleswig-Holstein (Heller 2004) and Thuringia, Saxonia (unpublished). The species was placed in the L. vitticollis group by Menzel & Mohrig (2000) but is exceptional in having a narrow gonostylus, medially not impressed, with a group of subapical megasetae and 2 – 3 medial elongated setae, and in having a peculiar intergonocoxal lobe of the hypopygium with unusually strong setae. Mohrig et al. (1987: 94) and Menzel & Mohrig (2000: 382) showed that the placement of L. dearmata in the L. vitticollis group of Lycoriella (Hemineurina) was somewhat problematic. The sensillar patch of the 1 st palpal segment is distinctly bordered (not deepened and pit-like), the wing vein R 5 are apically both dorsally and ventrally setose, the scutellum has 4 long and strong setae, the fore tibia has weak spinose setae [1 – 3 setae in the basic vestiture], the legs are long and thick, the fore tibial organ has fine and dense setosity and bow-like borders, and the tarsal claws are untoothed. It is striking that that the setosity of mesonotum and abdomen is, unlike the species of Lycoriella, Hemineurina und Trichocoelina, dark brown and much longer and dense. Furthermore, the species has a slender, not impressed gonostylus (without medial margin), 2 or 3 elongated and nearly straight setae on its apical half (not homologous with the apically curved whiplash seta of Lycoriella s. l.) and a slender apical tooth and 2 to 3 strong and nearly straight subapical megasetae. These characters support the view that L. dearmata is closer to the species of Bradysiopsis than to the here proposed genera Hemineurina and Trichocoelina (compare here with, for example, Bradysiopsis vittigera (Zetterstedt, 1851) in Menzel & Mohrig (2000: 189, fig. 159) and Bradysiopsis sordida (Mohrig, 1999) comb. n. (see below). The two, very closely placed intergonocoxal lobes of L. dearmata are in no case homologous with with the intergonocoxal lobes of Lycoriella s. str., Hemineurina or Trichocoelina, because at the base they are united with the intergonocoxal area by a strongly sclerotized bridge. Based on the above characters, Lycoriella (Hemineurina) dearmata Mohrig & Krivosheina is transferred to Bradysiopsis Tuomikoski.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA949FFBCFF26FB619F090A9D.taxon	discussion	Discussion. Both Br. sordida (Mohrig) comb. n. and Br. dearmata (Mohrig & Krivosheina) comb. n. (see above) are placed in the genus Bradysiopsis, in the here newly established Br. dearmata group. Both species have, unlike the Br. vittata group and Br. disjuncta group, a distinctly bordered patch of sensilla on the first palpal segment, conically narrowed and apically roundish tegmen, elongate-ovale gonostylus with the apex which is curved to the medial side and — like in Pseudolycoriella — densely setose, and which reaches apicad from the apical tooth. With their long three-segmented palpus, four strong setae on scutellum, untoothed tarsal claws, partly bow-like bordered and densely setose tibial organ, the lack of a whiplash seta on the gonostylus, the 2 – 3 long setae on the medial side of the gonostylus and a slender apical tooth in combination with three subapical megasetae, both species possess typical Bradysiopsis characters [compare with the genus diagnosis in Menzel & Mohrig (2000: 185)].	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA949FFBDFF26F97999790D66.taxon	discussion	Discussion. Lycoriella complexa was described based on one male from Spain (Rudzinski & Baumjohann 2009). We have not seen the holotype, but based on the description we exclude it from Lycoriella and transfer it to Camptochaeta Hippa & Vilkamaa, 2004. The single bristle on the first palpal segment, the sclerotized tegmen, the impressed gonostylus (in the original description the impression shown as a dotted line), the strong gonostylar megasetae in the apical half, three of which on the dorsal (apical) side of the apical tooth and the two moderately long elongated setae on the apical third of the gonostylus differ from the characters of Lycoriella s. str. and fit better with Camptochaeta. By these characters, Camptochaeta complexa (Rudzinski & Baumjohann, 2009) comb. n. resembles Cam. subcamptochaeta (Mohrig, 1992) [= Cam. pentacantha Komarova, Hippa & Vilkamaa, 2007]. It is possible that these species are even synonymous [compare the descriptions and figures in Mohrig & Eckert (1992: 295, fig. 1 a – e) and Komarova et al. (2007: 7, figs 1 – 5)].	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA948FFBDFF26FE2D9B0D0AE2.taxon	discussion	Discussion. Sasakawa (1997: 171) described the distinctive Lycoriella codonopsivora Sasakawa, 1997 from Hokkaido (Japan) whose larvae feed on deodeok leaves (Codonopsis lanceolata, Campanulaceae). For this species, he erected the monotypic subgenus Chorizomma of Lycoriella (Sasakawa 1997: 174 [preocc., not Chorizomma Simon, 1872; Araneae, Dictynidae] and renamed the subgenus later as Merizomma (Sasakawa 2003: 119, 128). The subgenus and the type species are not included in the monograph of the Palaearctic Sciaridae (Menzel & Mohrig 2000). Therefore, these taxa are discussed here in detail on the basis of the original descriptions by Sasakawa (1997) and included in the identification key and in the checklist.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA948FFBDFF26FE2D9B0D0AE2.taxon	description	On the basis of the above characters, L. codonopsivora Sasakawa must be excluded from the Lycoriella group. With its short gonocoxa, lacking intergonocoxal lobe and apical megaseta, the species resembles some species of the K. nepalensis group of Keilbachia Mohrig, 1987 [compare here Menzel & Martens (1995: 107), Menzel & Mohrig (2000: 348), Vilkamaa et al. (2006: 40) and Vilkamaa et al. (2009: 3)]. However, a strongly reduced eye bridge, a 2 - segmented palpus, an unbordered patch of sensilla on 1 st segment of palpus, a long R 1 and the lack of a spiral-formed megaseta on the medial side of gonostylus differ from the characters of Keilbachia. These characters are typical for the M. hippai group of the genus Mohrigia Menzel, 1995. Speaking against the combination of L. codonopsivora Sasakawa with Mohrigia are the Keilbachia - like form of the gonostylus, the missing long medial setae on the apical half of the gonostylus, the missing intergonocoxal lobe of hypopygium and the much smaller, weakly setose fore tibial organ [compare here Menzel & Martens (1995: 102), Menzel & Mohrig (2000: 415) and Rudzinski (2006: 450)]. Furthermore, neither Keilbachia nor Mohrigia have the lateral setae of scutellum reduced [both have two long and strong setae] nor an unmodified tegmen [the tegmen of Mohrigia has a straight, strongly sclerotized dorsomedial structure; the tegmen of Keilbachia is sclerotized laterally and often with a narrow apical process]. On the above grounds, Merizomma Sasakawa, 2003 stat. n. with the type species Merizomma codonopsivora (Sasakawa, 1997) comb. n. is raised to generic rank and excluded from the Lycoriella group sensu Menzel & Mohrig (2000).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA948FFB2FF26F8A1981F0F19.taxon	discussion	Discussion. Sciara hoyti Hardy was described based on 7 males and 2 females from the Hawaii Islands (Keanakolu, Mauna Kea). Steffan (1973) included this species in Lycoriella sensu Tuomikoski (1960), in spite of the missing bow-like bordered tibial organ and the missing whiplash seta, and compared the species with L. pallidior Tuomikoski, 1960 although the structure of the genitalia of the latter is very different. Contrary to the known species of Lycoriella, the first palpal segment of S. hoyti has only one seta, the gonostylus is allegedly strongly extended in the middle (Hardy 1956: fig. 6 c), and Hardy described six strong megasetae on the impressed area (three apically, two in the middle and one more basally at the medial corner of the impression). On the basis of these characters, S. hoyti does not belong to Lycoriella Frey. Mohrig et al. (2019: 427) have studied the holotype of S. hoyti Hardy and mention two conspecific males from Maui Island (Hawaii), and combine this species into the genus Scatopsciara Edwards s. str., with Scatopsciara spiculata Vilkamaa, Hippa & Mohrig, 2012 from New Caledonia as junior synonym. They found that the gonostyli of holotype are deformed in fig. 6 c by Hardy (1956: 81), the apex of the gonostyli has a strong tooth and the inner sides of gonostyli have only 4 megasetae (1 megaseta subapical and 3 further mesial up to the middle). The different lenght of tibial spurs at the middle and hind tibia, the narrow bristle row at the front tibia, the trapezoid tegmen, the sensory pit on the first palpal segment and the very short R 1 support the position in the Sc. atomaria group of the genus Scatopsciara Edwards (compare here especially with the detailed description in Vilkamaa et al. (2012 b: 69).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA947FFB2FF26FCFD99AA090E.taxon	discussion	Discussion. Neosciara biarmata Lengersdorf was described based on five males from the Austrian Alps. The collection data were defined on the basis of Janetschek’s collection list in Lengersdorf’s bequest by Menzel & Mohrig (2000) (letters and documents in ZFMK). Tuomikoski (1959 a) saw an immature type specimen (male) and wrote that N. biarmata Lengersdorf is very close to ‘ L. modesta (Staeger) ’. However, belonging nowadays in this species complex are many species which are difficult to distinguish from each other without comparing the specimens. A reliable concept of the species is not possible until a type revision is undertaken. It is treated as a nomen dubium in Menzel & Mohrig (2000: 589), and not included in the present checklist.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA947FFB2FF26FAF5980E0AE2.taxon	discussion	Discussion. Sciara morosa Meunier was described from Baltic amber on the basis of seven females. Mohrig & Röschmann (1994: 82) concluded in their revisionary work that the type material is lost and that Frey’s (1942) placement of the species in Lycoriella s. str. must be strongly suspected because up till now no species of Lycoriella have been found in amber. We follow the decision of Mohrig & Röschmann (1994) and exclude the preoccupied name Sciara morosa Meunier, 1904 [not Sciara morosa Winnertz, 1867] from the Lycoriella group and place it in the unplaced taxa (species incerta sedis).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA947FFB3FF26F8A19EF90DD3.taxon	discussion	Discussion. Sciara solita Walker was described based on one female from Malaysia (Borneo: Sarawak). One additional sample was found by Edwards (1931) from the lowlands of Northern Borneo (1 male, 14 females from ‘ Bettotan’ near Sandakan, Sabah). Brunetti (1920) wrote that the holotype is in BMNH, and in good condition. The original description by Walker (1857) comprises only five lines and is so meagre that the species cannot be identified without studying the type. Of the male found later it is unclear whether it really is S. solita sensu Walker (1857) as there is no description nor figures of this specimen. On this account, Sciara solita Walker, 1857 is removed from the Lycoriella group and placed among the unplaced taxa of Sciaridae (species incertae sedis). Taxonomic corrections within the Lycoriella group sensu Menzel & Mohrig (2000)	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA946FFB3FF26FD9598620A3C.taxon	synonymic_list	Synonyms: = arctica (Holmgren, 1869) [as Sciara]; = conglomerata (Pettey, 1918) [as Neosciara]; = ecalcarata (Holmgren, 1869) [as Sciara]; = frigida (Holmgren, 1869) [as Sciara, preocc.]; = fumatella (Lundbeck, 1898) [as Sciara]; = globiceps (Becher, 1886) syn. n. [as Sciara]; = groenlandica (Holmgren, 1872) [as Sciara]; = holmgreni (Rübsaamen, 1894) [as Sciara, new name for Sciara frigida Holmgren, 1869].	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA946FFB3FF26FD9598620A3C.taxon	materials_examined	Material studied. Lectotype, male of Sciara modesta Staeger (designated by Menzel in Menzel & Mohrig (2000): 405, 755). DEnmark, without locality details [as ‘ Danmark’], specimen no. 239, in May, Staeger (in ZMUC). Lectotype, male of Sciara globiceps Becher (designated by Menzel in Menzel & Mohrig (2000): 402, 752). NORWAY, Jan Mayen, inventory no. 5 / 70 A / 12 / NMW, [August or September] 1882, F. Fischer (in NHMW). Literature (selection). Sciara globiceps Becher — Becher (1886): 62, pl. 5, figs 2, 2 a – d; Edwards (1923): 236. Lycoriella globiceps (Becher) — Thunes et al. (2004): 85. Lycoriella (Hemineurina) globiceps (Becher) — Menzel & Mohrig (2000): 402 [in part]. Sciara modesta Staeger — Staeger (1840): 286. Bradysia (Hemineurina) modesta (Staeger) — Frey (1948): 66, 84; pl. 18, fig. 104; Frey (1953): 458; Nielsen et al. (1954): 21. Lycoriella modesta (Staeger) — Krivosheina & Mohrig (1986): 157, 162; Röschmann & Mohrig (1993): 383; Röschmann & Mohrig (1994): 203; Hellrigl (1996): 633; Hennicke et al. (1997): 99; Mukkala et al. (2005): 16, 32; Seeber et al. (2012): 369; Salmela et al. (2015): 87. Lycoriella (Hemineurina) modesta (Staeger) — Frey (1942): 36; Tuomikoski (1960): 75, 77;? Gerbachevskaja (1963): 498; Tuomikoski (1967): 48; Freeman (1983 a): 168; Freeman (1983 b): 30, fig. 99; Gerbachevskaja-Pavluchenko (1986): 31; Franz (1989): 15; Menzel et al. (1990): 337; Jakovlev (1994): 76; Menzel & Mohrig (2000): 405; Menzel et al. (2003): 88, 101; Coulson & Refseth (2004): 103; Menzel et al. (2006): 108; Coulson (2008): 161; Coulson (2013): 154; Mohrig et al. (2013): 213, fig. 37 a – e; Vilkamaa (2015): 551.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA946FFB3FF26FD9598620A3C.taxon	discussion	Discussion. The only existing male of Sciara globiceps Becher (lectotype) is in poor condition. Because of the strongly deformed male genitalia Menzel & Mohrig (2000) did not realize that the lectotype of Sciara globiceps Becher is not identical with Hemineurina algida (Frey) [misidentification] but in reality represents Hemineurina modesta (Staeger), which has a longer and therefore a more slender-looking gonostylus. Consequently, some records of H. algida were erroneously published by Menzel & Mohrig (2000) under the name ‘ L. globiceps (Becher) ’ [see literature under H. algida (Frey)]. These errors are corrected here, by treating Sciara globiceps Becher, 1886 syn n. as a junior synonym of Hemineurina modesta (Staeger, 1840) comb. n. and H. algida (Frey, 1948) restit. et comb. n. as a separate species. The distinct differences between H. algida and the similar H. thuringiensis are discussed under H. algida (Frey).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA946FFB0FF26F91B982B0836.taxon	materials_examined	Material studied. Lectotype, male (here designated). RUSSIA, ‘ Regio kuusamoensis, Paanajärvi’ [= Republic of Karelia, Kemsky District, village Paanajärvi], specimen no. 739, type no. 8372 (ID GE. 250), 24. VI. 1937, R. Frey (without genitalia, in MZH). Paralectotypes. 4 males [all misidentification; = Hemineurina modesta (Staeger, 1840)]: FINLAND, Ab (Regio aboensis), Vichtis [= Vihti], Päivölä, wood pile, 28. VIII. 1943, R. Frey, 1 male (ID GE. 1769, in MZH); FINLAND, Ta (Tavastia australis), Kangasala [SE of Tampere], 11. VI. 1942, R. Frey, 1 male (ID GE. 1770, in MZH); FINLAND, LKem (Lapponia kemensis), Pallastunturi [= mountain Pallastunturi SW of Raattama], 8. VII. 1943, R. Frey, 1 male (ID GE. 1773, in MZH); FINLAND, Le (Lapponia enontekiensis), Kilpisjärvi, Malla [= Lapland, Enontekiö, Malla mountains near Kilpisjärvi], 18. VII. 1943, R. Frey, 1 male (ID GE. 1771, in MZH). The other type specimens of ‘ Bradysia (Hemineurina) algida ’ mentioned by Frey (1948: 84) were collected in ‘ N. Helsingfors, Kottby’ [= FINLAND, Nylandia, Helsinki, Kottby district] and ‘ Le. Saana’ [= FINLAND, Lapponia enontekiensis, Enontekiö, Saana mountain near Kilpisjärvi]. These paralectotypes could not be found in the MZH collection and were not revised. Literature. Lycoriella (Hemineurina) globiceps (Becher) — Menzel & Mohrig (2000): 403 [in part, misidentification]; Coulson & Refseth (2004): 103; Salmela & Vilkamaa (2005): 291 [both misidentifications]. Lycoriella globiceps (Becher) — Mukkala et al. (2005): 16, 32; Vilkamaa et al. (2007): 228 [all misidentifications]. Bradysia (Hemineurina) algida Frey — Frey (1948): 66, 84, pl. 18, fig. 107. Lycoriella algida (Frey) — Jakovlev (1994): 76; Hellrigl (1996): 633. Lycoriella (Hemineurina) algida (Frey) — Tuomikoski (1960): 75, 77; Pavluchenko (1984): 94; Gerbachevskaja-Pavluchenko (1986): 30; Röschmann & Mohrig (1994): 203.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA946FFB0FF26F91B982B0836.taxon	discussion	Discussion. Hemineurina algida (Frey) is a small, common, boreoalpine species (body length of the male 1.7 – 2.0 mm). Of Frey’s original type series, only one male remains, which corresponds with the description by Frey (1948: 66, fig. 107). Frey (1948) prepared his figure 107 based on the now designated lectotype specimen (checked before the loss of male genital during the slide re-mounting). All other revised specimens of the type series in MZH belong to H. modesta (Staeger), as was already noted by Tuomikoski. Hemineurina algida (Frey) is similar to H. thuringiensis (Menzel & Mohrig) comb. n. The gonostylus of the very variable H. algida is mostly much narrower than that in H. thuringiensis, less impressed medially, the medial megasetae narrower and the basoventral seta groups of the intergonocoxal area hypopygium are less distinct. The seta groups of the intergonocoxal area can be very variable even in material from one locality [e. g., 1488 males were studied from the Tyresta National Park near Stockholm, Sweden (in SDEI, SMNH) and ca. 50 specimens from other European countries, incl. Finland and Norway (in MZH, SDEI)]. A morphological study showed that the basoventral seta groups can vary from 8 – 10 closely spaced setae to a near absence of setae. Furthermore, H. algida has a longer gonocoxa and somewhat wider basal bodies of the antennal flagellomeres with yellowish-whitish setosity. H. thuringiensis has the gonocoxa shorter and more compact, the flagellomeres slightly longer and with dirty pale brown setosity, the gonostylus shorter and thickened, the medial megasetae mostly stronger and the seta groups of the intergonocoxal area more dense and separated from each other like islands.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA945FFB0FF26FB1D99D209E6.taxon	discussion	Discussion Sciara speciosissima Strobl, 1898 is known only from one male in very poor condition and four females. They belong to Lycoriella (Hemineurina) sensu Menzel & Mohrig (2000) and to the genus Hemineurina Frey (former L. inflata group) in the present sense, but we cannot completely clarify their species status without an examination of male genitalia (Menzel 1992 a, Menzel & Mohrig 2000).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA945FFB1FF26F9AD99D80DB1.taxon	discussion	Discussion. When the figure of the lectotype of Sciara riparia Holmgren in Menzel & Mohrig (2000: 407, fig. 375) is compared with the figure of the holotype of S. unguicauda Malloch by Mohrig et al. (2013: 215: fig. 38 a), it is clear that the species are distinct. Hemineurina riparia (Holmgren, 1883) comb. n. has a narrower gonostylus and a slightly impressed part on the basal side of the apical tooth, where there are three megasetae [two very close to the apical tooth, one of which on its dorsal, one on its ventral side and one megaseta medially at the apical third]. The medial part of the gonostylus between the two groups of megasetae has only short setae, the apical tooth is not on a lobe-like process, and the whiplash seta is on the middle of the medial side. Furthermore, all megasetae are narrow, hyalinous, and on short basal bodies. In comparison, Hemineurina unguicauda (Malloch, 1923) restit. et comb. n. has a more voluminous gonostylus, the medial side of which is strongly impressed ventrally, and the medial margin between the apex of gonostylus and the medial megasetae has strikingly long setae. Of the four megasetae, three are much stronger than those of H. riparia and two of the megasetae are close to each other on high basal bodies at the middle of the gonostylus. Furthermore, the whiplash seta of H. unguicauda is more basally placed (on the basal third of the gonostylus), one slender megaseta is on the margin of the impressed area and one, stronger, megaseta just on the basal side of the apical tooth. Strikingly, the long apical tooth and the subapical megaseta are at the apex of a long non-setose lobe, which is absent from H. riparia (Holmgren).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA944FFB7FF26FD9599890904.taxon	discussion	Discussion. Lycoriella piristylata was described in the former subgenus Hemineurina Frey based on the holotype and nine paratypes from Northern Finland, Norway and Sweden (Vilkamaa et al. 2013). The species was provisionally placed in L. vitticollis group of Lycoriella (Hemineurina) but because it has a sensory pit on the first palpal segment, Lycoriella piristylata belongs — in spite of the completely reduced whiplash seta on the medial side of the gonostylus — to the genus Lycoriella Frey s. str. sensu Menzel & Mohrig (2000: 380). Furthermore, the species has a bow-like bordered fore tibial organ, lacks spinose setae on the fore tibia, the tarsal claws are without teeth, the basal portion of antennal flagellomeres have long and appressed sensilla, the gonocoxae are long, ventrobasally separated in a v-shape, and the tegmen is membraneous and broadly roundish. Basoventrally in the intergonocoxal area — contrary to the original description by Vilkamaa et al. (2013: p. 53, fig. 3 A) — there is no medial lobe. At that place on the medial margin there are at most a few setae, as, for example, in L. lundstromi (Frey, 1948) and L. vanderwieli (Schmitz, 1920). Phylogeny and classification The molecular phylogeny by Shin et al. (2013) challenged the monophyly of Lycoriella s. str. + (Hemineurina + Coelostylina) (only the L. inflata group was included in the ingroup of their analysis) suggested by Menzel & Mohrig (2000) on the morphological evidence. Shin et al. (2013) suggested that Hemineurina and Lycoriella s. str. are not closely related, as they appeared in different main clades in the phylogenetic tree. The morphological characters uniting Lycoriella s. str. and Hemineurina are indeed few: the presence of the whiplash seta and the pale and weak body setosity. Coelostylina Tuomikoski, 1960 (preocc.) is a junior homonym of Coelostylina Kittl, 1894 (Gastropoda: Coelostylinidae), and is here renamed as Stenacanthella nom. et stat. n. On the basis of the corrected nomenclature, the species groups of ‘ Coelostylina Tuomikoski’ sensu Menzel & Mohrig (2000: 380, 384) are kept and re-named here as the St. freyi group and St. secundaria group. Hemineurina, Trichocoelina and Stenacanthella have several putative synapomorphies: the intergonocoxal area is long and the apicoventral margin of gonocoxa is therefore short, the intergonocoxal area is almost exclusively with setose lobe (s), the gonostylar megasetae usually have basal bodies, the megasetae are mostly slender and at least some of them are oblique in position, the gonostylus is impressed, the apex of gonostylus is densely setose, the tegmen is at least partly sclerotized, and the fore tibia has spine-like setae among the ordinary vestiture (lacking in Lycoriella s. str.). Furthermore, Hemineurina and Trichocoelina as sister groups share as synapomorphies the similar intergonocoxal area with two lobes (in some species a medial lobe), the whiplash seta medial or subbasal in position, and a well-developed apical tooth of gonostylus. Following our present morphological interpretation, we agree with Menzel & Mohrig (2000) with the phylogenetic relationships of these three taxa, but propose raising them to genus rank: Stenacanthella nom. et stat. n. + (Hemineurina stat. n. + Trichocoelina gen. n.). The genus Hemineurina here includes only the former L. inflata group with the type species Sciara conspicua Winnertz, 1867, and the new genus Trichocoelina is established for the former L. vitticollis group (type species Sciara vitticollis Holmgren, 1883). The maximum likelihood tree for the COI gene indicates that Trichocoelina gen. n. is monophyletic (Appendix 3). However, the genus Hemineurina Frey in the present sense represented by two morphotypes sensu Menzel & Mohrig (2000) — H. inflata and H. conspicua — appears at different positions in the ML tree. This result is of course preliminary and not representative because only one gene was included in our genetic analysis and the taxon sampling was very limited. But this taxonomic problem was not the focus of this study and will be discussed in a later revision of the Hemineurina species. Diagnostic characters of the Lycoriella group of taxa The traditional Lycoriella group of taxa (subgenera Lycoriella s. str., Lycoriella (Hemineurina) with Hemineurina stat. n. [former L. inflata group] and Trichocoelina gen. n. [former L. vitticollis group], Stenacanthella nom. et stat. n. [former Lycoriella (Coelostylina)] and Bradysiopsis, can be distinguished from other genera of Sciaridae by the following characters: Body vestiture weak and pale (dark and strong in Bradysiopsis and some Stenacanthella), fore tibia with a distinct tibial organ (densely setose and with arcuate boundary), gonostylus with two or more upcurved whiplash setae sub-basomedially (lacking in Bradysiopsis and Stenacanthella), apical tooth and subapical / mesial megasetae present, intergonocoxal area of hypopygium or basoventral parts of gonocoxa mostly with lobe (s) or seta group (s). Species of Mohrigia Menzel, 1995 also have one or more whiplash setae on the gonostylus and an intergonocoxal lobe, but differ in having a modified apical tooth with megasetae inside, the tegmen narrow and with a dorsomedial sclerotized rim, the tarsal claws with tiny teeth, and a very short and weak aedeagal apodeme (Menzel & Martens 1995; Menzel & Mohrig 2000). Camptochaeta Hippa & Vilkamaa, 1994 also has elongated seta (e) on their gonostylus, but these are strong and not whiplash-like (similar as in Mohrigia). Camptochaeta can also be distinguished by typical lambda-shaped basal sclerotization in the gonostylus, stronger gonostylar megasetae, and by generally stronger and darker body setosity. Eugnoriste Coquillett, 1896 and Pseudolycoriella Menzel & Mohrig, 1998 also have a whiplash-like seta on the gonostylus, but the seta is downcurved and sub-apicoventral in position (not medial or subbasal as in Lycoriella s. str., Hemineurina and Trichocoelina). Furthermore, Eugnoriste and Pseudolycoriella have teeth on the tarsal claws, and strong body setosity.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA944FFB7FF26FD9599890904.taxon	materials_examined	Type species: Sciara vitticollis Holmgren, 1883	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA944FFB7FF26FD9599890904.taxon	description	Description. Male. Head. Eye bridge 2 – 4 facets wide. Coloration of antenna brown or scapus, pedicellus and / or 1 st flagellomere yellow, flagellomeres with unicolorous necks, necks short. Maxillary palpus with 3 segments, 1 st segment with dorsal patch of sensilla and a few sharp setae (without sensory pit). Face with many setae. Thorax. Brown and sparsely setose, setae usually pale. Scutellum with 4 (rarely 3) strong setae and many short setae. Katepisternum high and triangular, postpronotum non-setose. Legs. Yellow. Fore femur slender. Fore tibia with some spinose setae among vestiture and some spinose setae at apex. Fore tibial organ large and distinct, usually clearly demarcated, setae strong or fine, tibial spurs long, mid and hind legs with 2 equally long ones. Tarsal claws untoothed. Wing. Fumose. Anal lobe small. Veins distinct. Fork of M very long and weakly arcuate; R and R 1 long, R 1 merging with c nearly at level of base of M-fork; apical part of R 5 with only dorsal macrotrichia. Abdomen. Pale brown and sparsely setose, setae usually pale and fine. Hypopygium. Apicoventral corner with only one long seta, intergonocoxal area of hypopygium long, with one medial, usually at least apically divided lobe, or two separate lobes at bases of gonocoxae. Gonocoxa normal to wide, as long as or longer than gonostylus. Gonostylus rather narrow to very voluminous, apically densely setose, medially impressed, with an apical tooth (missing only in one known species), with 5 to numerous, usually slender and slightly procurved megasetae, with one or a few subbasal whiplash setae. Tegmen as long as broad, or shorter or longer, weakly sclerotized, rounded or with different modifications apically, with very small and fine aedeagal teeth (or these apparently lacking), and with distinct aedeagal apodeme. Female. Without diagnostic characters at species level.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA944FFB7FF26FD9599890904.taxon	discussion	Discussion. Trichocoelina gen. n. differs from Hemineurina in having the gonostylus usually broader and more strongly impressed or excavated, the ventrolateral margin extending over the basal parts of the megasetae, in having the megasetae more numerous and slender, usually with distinct basal bodies, the whiplash setae subbasal in position, and the tegmen usually only weakly sclerotized. The intergonocoxal area of hypopygium is longer and the apical part of gonocoxa shorter than in Hemineurina. Trichocoelina differs from Stenacanthella in having a whiplash seta on the gonostylus, in having stronger apical tooth, and no tendency to have three lobes in the intergonocoxal area. Trichocoelina differs from both Hemineurina and Stenacanthella in having 4 or 3, not just two, long and strong setae on the scutellum.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA944FFB7FF26FD9599890904.taxon	distribution	Distribution. Northern Holarctic, south of 50 ° N only in mountains over 1000 meters high.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA944FFB7FF26FD9599890904.taxon	etymology	Etymology. The name is formed from the latinized Greek words trichos, hair, and koilos, hollow, referring to the narrow megasetae in the medially hollowed gonostylus of most species.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA942FFB5FF26FA349BB90FDD.taxon	distribution	T. aemula sp. n. (Finland; Russia: Krasnodarsk region) T. biplex sp. n. (Canada: Newfoundland and Labrador, Yukon) T. brevicubitalis (Lengersdorf, 1926) comb. n. (Norway: mainland;? Estonia) T. chentejensis (Menzel, 1992) comb. n. (Finland; Mongolia) T. cochleata (Rübsaamen, 1898) comb. n. (Czech Republic; Finland; Russia: Karelia, Siberia; Norway: mainland, Svalbard; Greenland) T. dicksoni sp. n. (Russia: Arkhangelsk oblast, Kemerovsk oblast, Krasnodarsk region) T. dispansa sp. n. (Russia: Krasnodarsk region) T. dividua sp. n. (Canada: Northwest Territories) T. hians sp. n. (Canada: Yukon) T. hiemalis (Mohrig & Mamaev, 1985) comb. n. (Finland; Germany; Russia: Krasnodarsk region) T. imitator sp. n. (Canada: Yukon) T. incrassata sp. n. (USA: Alaska) T. ithyspina sp. n. (Norway: mainland) T. janetscheki (Lengersdorf, 1953) comb. n. (Austria; Greenland; Canada: Nunavut; USA: Colorado) T. jukkai sp. n. (Finland; Norway: mainland) T. magnifica sp. n. (Canada: Yukon) T. nefrens sp. n. (Russia: Krasnodarsk region) T. obesula sp. n. (Norway: Svalbard) T. olschwangi (Mohrig & Mamaev, 1983) comb. n. (Russia: Krasnodarsk region; Canada: Yukon; USA: Colorado) T. oricillifera sp. n. (Finland; Norway: mainland; Sweden) T. planilobata sp. n. (Finland) T. quintula sp. n. (Finland) T. semisphaera sp. n. (Finland; Norway: Svalbard) T. semusta sp. n. (Italy; USA: Alaska) T. subcochleata (Komarov, 2009) comb. n. (Russia: Republic of Altai) T. subpermutata (Mohrig & Mamaev, 1990) comb. n. (Russia: Krasnodarsk region; Sweden) T. tecta sp. n. (Russia: Krasnodarsk region, Yamalo-Nenets Autonomous Okrug; Canada: Nunavut, Yukon; USA: Alaska) T. vitticollis (Holmgren, 1883) comb. n. (Finland; Norway: Svalbard; Russia: Novaya Zemlya; Sweden; Greenland;? Canada: Nunavut)	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA942FFB5FF26FA349BB90FDD.taxon	discussion	Key to species of Trichocoelina (males) Note. Trichocoelina brevicubitalis (Lengersdorf, 1926) comb. n., known only in the female sex, is not included in the identification key (redescription and discussion see under ‘ The Trichocoelina species’).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA940FFB5FF26FB9E9B160BF1.taxon	description	Figs 1 A, 3 B	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA940FFB5FF26FB9E9B160BF1.taxon	materials_examined	Material studied. Holotype male. RUSSIA, Krasnoyarsk region, Taimyr Peninsula, 12.5 km S of Dixon, 73 ° 24 ’ N, 80 ° 39 ’ E, on the river Lemberova, pan trap, 7 – 10. VII. 2012, A. Barkalov (in ISEA).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA940FFB5FF26FB9E9B160BF1.taxon	description	Description. Male. Head. Face and antenna concolorous brown, maxillary palpus pale yellow. Eye bridge 3 facets wide. Face with 18 fine setae. Clypeus with 3 setae. Maxillary palpus with 3 segments, 3 rd segment longer than 1 st segment, 2 nd segment shortest; 1 st segment with 2 or 3 setae, with a dorsal patch of sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 2.3 x as long as wide, the neck shorter than broad, the longest setae slightly shorter than the width of flagellomere. Thorax. In poor condition in the specimen studied. Brown, setae pale. Wing. Fumose. Length 2.6 mm. Width / length 0.40. Anal lobe moderate. Veins distinct. c / w 0.60. R 1 / R 0.70. stM and fork of M subequal in length. r-m 2 x as long as bM, r-m and bM non-setose, Halter yellow. Legs. Yellow, coxal setae pale. Fore tibial organ forming a large patch of fine setae in depression. Fore tibial spur slightly longer than the tibial width. Abdomen. Pale brown, setae pale, short and fine. Hypopygium (Fig. 1 A). Brown, as abdomen. Intergonocoxal area long, with two short but distinct setose lobes. Gonocoxa broad, longer than gonostylus, medial margin basally smoothly curved, with short sparse setosity. Gonostylus (Fig. 3 B) broad, apically slightly narrowed, medially strongly impressed; with short setosity, with a long apical tooth, with about 15 megasetae medially, megasetae rather long and slender; with 2 well-differentiated whiplash setae on ventromedial margin. Tegmen subconical, weakly sclerotized, with a small area of minute aedeagal teeth. Aedeagal apodeme long. BIN. Unknown.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA940FFB5FF26FB9E9B160BF1.taxon	discussion	Discussion. In the form of its gonostylus, Trichocoelina absidata sp. n. most resembles T. dicksoni sp. n. and T. hians sp. n. It differs from T. dicksoni in having a longer apical tooth of gonostylus and shorter and more numerous gonostylar megasetae, in its more conical and unsclerotized tegmen which is not subtriangular and laterally and apically sclerotized, and in having shorter intergonocoxal lobes of the hypopygium. It is similar to T. hians in having numerous short gonostylar megasetae, but differs in its basomedially less bulged gonostylus, in missing the narrowed apical part of the tegmen and in having short intergonocoxal lobes of hypopygium.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA940FFB5FF26FB9E9B160BF1.taxon	etymology	Etymology. The name is Latin, absidata, hollowed, referring to the strongly recurved gonostylus with an impressed medial side.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA95EFFABFF26FF0D9BF5090E.taxon	description	Figs 1 B, 3 A, 18 A	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA95EFFABFF26FF0D9BF5090E.taxon	materials_examined	Material studied. Holotype male. RUSSIA, Krasnodarsk region, Taimyr Nature Reserve, Aru-Mas, 72.50 ° N, 101.94 ° E, pan trap, 9 – 20. VII. 2010, A. Barkalov (in ISEA). Paratypes. RUSSIA, same data as holotype, 4 males (in ISEA, MZH, NHMO, SDEI); FINLAND, Li (Lapponia inarensis) (grid 7759292: 3539670), sandy river bank, pitfall trap, 13. VI – 13. VII. 2016, J. Salmela, 3 males (in MZH, PJSR, SDEI).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA95EFFABFF26FF0D9BF5090E.taxon	description	Description. Male. Head. Face and antenna concolorous brown, maxillary palpus pale yellowish. Eye bridge 2 – 3 facets wide. Face with 13 – 15 setae. Clypeus with 1 – 3 setae. Maxillary palpus with 3 segments, 1 st segment as long as or longer than 3 rd segment, 2 nd segment shortest; 1 st segment with 1 – 2 setae, with dorsal patch of sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 1.7 – 2.0 x as long as wide, the neck shorter than broad, the longest setae shorter than the width of flagellomere. Thorax. Brown, setae pale. Anterior pronotum with 4 – 6 setae. Proepisternum with 3 – 10 setae. Scutellum with 4 longer and some short and fine setae. Wing. Fumose. Length 2.0 – 2.2 mm. Width / length 0.40. Anal lobe weak. Veins distinct. c / w 0.65 – 0.75. R 1 / R 0.6 – 0.7. stM shorter than fork of M. r-m as long as bM or longer, bM non-setose, r-m with 1 – 4 setae. Halter yellow. Legs. Yellow, coxal setae pale. Fore tibial organ forming a rather small patch in depression. Fore tibial spur longer than the tibial width. Abdomen. Pale brown, setae pale, short and fine. Hypopygium (Fig. 1 B). Brown, as abdomen. Intergonocoxal area long, with two short, distinctly separate setose lobes (Fig. 18 A). Gonocoxa broad, longer than gonostylus, medial margin basally strongly curved, with sparse and short setosity. Gonostylus (Fig 3 A) broad, laterally roundish, strongly impressed; with short setosity, with a long apical tooth, with 6 – 7 megasetae in the medial impression, megasetae long and slender, with distinct basal bodies; with 1 well-differentiated whiplash seta on ventromedial margin. Tegmen as long as broad, long and laterally rounded, apically straight, weakly sclerotized, with a dorsal finger-like process, aedeagal teeth not detectable in the specimens studied. Aedeagal apodeme long. BIN. Unknown.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA95EFFABFF26FF0D9BF5090E.taxon	discussion	Discussion. The basally narrowed gonostylus with a long apical tooth of Trichocoelina aemula sp. n. is almost identical to that of T. biplex sp. n., but differs in having the intergonocoxal lobes of hypopygium much shorter, and the tegmen less strongly narrowed towards apex and without basolateral corners. Because of its rather long gonostylar megasetae, roundish tegmen and short intergonocoxal lobes of hypopygium, T. aemula also resembles T. cochleata (Rübsaamen, 1898) but is distinguished by having the gonostylar apex less strongly curved, paler and with a longer apical tooth of gonostylus, see Fig. 3 A and fig. 377 in Menzel & Mohrig (2000).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA95EFFABFF26FF0D9BF5090E.taxon	etymology	Etymology. The name is Latin, aemula, comparable, referring to the close resemblance to Trichocoelina biplex sp. n. and T. cochleata.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA95EFFA8FF26FAF59B270EAA.taxon	description	Figs 2 A, 3 C	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA95EFFA8FF26FAF59B270EAA.taxon	materials_examined	Material studied. Holotype male. CANADA, Newfoundland and Labrador, Torngat Mountains National Park, Saglek / Base Camp, 58.451 ° N, 62.798 ° W, 5 m, 9. VIII. 2013, D. Whitaker (BOLD Sample ID BIOUG 11125 - C 09, in CNC). Paratypes. CANADA, same data as holotype but 16. VIII. 2013, 1 male (BOLD Sample ID BIOUG 11632 - G 07, in CBG); same data as previous but 20. VII. 2014, 3 males (BOLD Sample IDs BIOUG 18960 - C 12, BI- OUG 18960 - E 11 and BIOUG 18962 - F 10, in CBG); Yukon, Ogilvie Mts. North Fork Pass, 7. VIII. 1963, P. J. Skitsko, 1 male (in MZH).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA95EFFA8FF26FAF59B270EAA.taxon	description	Description. Male. Head. Face and antenna concolorous brown, maxillary palpus pale yellowish. Eye bridge 2 – 3 facets wide. Face with 22 – 32 setae. Clypeus with 1 seta. Maxillary palpus with 3 segments, 1 st segment as long as 3 rd segment, 2 nd segment shortest; 1 st segment with 1 seta, with dorsal patch of sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 2.2 x as long as wide, the neck shorter than broad, the longest setae shorter than the width of flagellomere. Thorax. Brown, setae pale. Anterior pronotum with 4 – 9 setae. Proepisternum with 6 – 9 setae. Scutellum with 3 – 4 longer and some short and fine setae. Wing. Fumose. Length 2.0 – 2.1 mm. Width / length 0.40. Anal lobe weak. Veins distinct. c / w 0.55 – 0.65. R 1 / R 0.55 – 0.70. stM shorter than fork of M. r-m longer than bM, bM non-setose, r-m with 1 – 6 setae. Halter yellow. Legs. Yellow, coxal setae pale. Fore tibial organ forming a sparse patch in depression. Fore tibial spur longer than the tibial width. Abdomen. Pale brown, setae pale, rather long and fine. Hypopygium (Fig. 2 A). Brown, as abdomen. Intergonocoxal area long, basally with strongly sclerotized medial stripe, with two long, subtriangular elongated setose lobes. Gonocoxa pale brown, narrow, longer than gonostylus, medial margin basally smoothly curved, with rather long but sparse setosity. Gonostylus (Fig. 3 C) yellow, rather long, apically slightly narrowed, strongly impressed; with normal setosity, with a long apical tooth, with 6 – 7 megasetae at middle, megasetae slightly curved, long and slender; with 1 well-differentiated whiplash seta on ventromedial margin. Tegmen broader than long, basolaterally broadened, apically roundish, with subapical semicircular sclerotization, aedeagal teeth not detectable in the specimens studied. Aedeagal apodeme short. BIN. BOLD: ACG 3979.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA95EFFA8FF26FAF59B270EAA.taxon	discussion	Discussion. By the general structure of its hypopygium, Trichocoelina biplex sp. n. resembles T. vitticollis (Holmgren) but differs in having the intergonocoxal lobes distinctly separate and shorter, the tegmen more modified with basolateral shoulders and a sclerotized apical rim, the gonostylus less curved and with a shorter apical tooth, the megasetae in a less compact group, and in its darker antennal flagellomeres. See also under T. aemula.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA95EFFA8FF26FAF59B270EAA.taxon	etymology	Etymology. The name is Latin, biplex, divided, referring to the distinctly separate, long intergonocoxal lobes of hypopygium.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA95DFFA8FF26FD699E800A0A.taxon	description	Redescription. Female. Head. Dark brown. Eye bridge united, 2 – 3 facets wide. Maxillary palpus long, with 3 segments, all segments narrow, 1 st segment with 4 sharp setae, with a dorsal unbordered patch of sensilla. Scapus and pedicellus dark bown and roundish; antennal flagellomers missing in the specimen studied. Thorax. Postpronotum non-setose. Scutellum with 8 long and strong setae. Legs. Strong, with long tibial spurs; tibial spurs of mid and hind tibiae unequally long [one spur is about 1 / 4 shorter than the other]; tarsal claws without teeth. Wing. Large, anal lobe strong, hind veins distinct, stCuA short, 1 / 3 of bM; bM = r-m, bM non-setose, r-m with 4 – 5 setae distally, c / w 0.50. Body length 2,7 mm, wing length 2,5 mm. BIN. Unknown.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA95DFFA8FF26FD699E800A0A.taxon	discussion	Discussion. The species was described from females from Northern Norway by Lengersdorf (1926): for the lectotype designation and literature, see Menzel & Mohrig (2000). After a study of the lectotype, which so far is the only specimen of the species (in ZFMK) available for study, Menzel & Mohrig (2000) stated that the scutellum has many long and strong setae, the 1 st palpal segment has a unbordered patch of sensilla, the fore tibia has spine-like setae, and that R 1 / R complex is long (R 1 ends near the level of the base of the M-fork). We have studied the lectotype again and have found characters that might be helpful in identification of the male of this species and for supporting its placement in the genus Trichocoelina. Of the characters given above, only the somewhat unequally long spurs of the mid and hind tibiae as well as the high number of long scutellars are unusual for Trichocoelina.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA95DFFA8FF26F9C9983E0B72.taxon	description	BIN. Unknown.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA95DFFA8FF26F9C9983E0B72.taxon	discussion	Discussion. The species was described from the holotype male from Mongolia (Menzel 1992 b). For discussion, see under Trichocoelina dispansa sp. n.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA95BFFAFFF26F8B09BAE0812.taxon	synonymic_list	Synonym: = haemorrhoidalis (Lundbeck, 1898) [as Sciara].	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA95BFFAFFF26F8B09BAE0812.taxon	discussion	Discussion. The species was described from Greenland (Rübsaamen, 1898), and redescribed and illustrated by Menzel & Mohrig (2000), with the discussion of the type material. Trichocoelina cochleata is characterized by having a medially impressed gonostylus, with a strongly curved apical part. The antennal scapus and pedicellus as well as the hypopygium and legs are yellow, but the apex of gonostylus has strikingly dark setosity. The gonostylus bears 6 – 7 rather long and slender gonostylar megasetae, the tegmen is broadly subconical with a sclerotized dorsoapical process. The intergonocoxal lobes of the hypopygium are short. For similar species, see under Trichocoelina aemula sp. n. and T. semusta sp. n.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA95AFFACFF26FBF199DC0CD6.taxon	description	Figs 2 B, 3 D	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA95AFFACFF26FBF199DC0CD6.taxon	materials_examined	Material studied. Holotype male. RUSSIA, Krasnoyarsk region, Dikson Island, Konus Islet, Korotkevich, 31. VII. 1948 (in ZIN). Paratypes. RUSSIA, same data as holotype, 3 males (1 in SDEI, 2 in ZIN). Other material. RUSSIA, Arkhangelsk oblast, Novaya Zemlya, Tuylenya Zaliv, 10. VII. 1901, Timofeev, 1 male (in ZIN); Kemerovsk oblast, river Mrassu on upper river Tom, 15. VII. 1908, Hvorov, 1 male (in ZIN).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA95AFFACFF26FBF199DC0CD6.taxon	description	Description. Male. Head. Face brown, maxillary palpus pale yellowish. Eye bridge and face not well visible in the specimens studied. Clypeus with 2 – 3 setae. Maxillary palpus with 3 segments, 1 st segment longer than 3 rd segment, 2 nd segment shortest; 1 st segment with 5 – 6 setae, with dorsal patch of sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 2.25 – 2.35 x as long as wide, the neck shorter than broad, the longest setae shorter than the width of flagellomere. Thorax. Dark brown, setae pale. Anterior pronotum with 7 – 12 setae. Proepisternum with 8 – 17 setae. Scutellum with 4 longer and some short and fine setae. Wing. In poor condition in the specimens studied. Fumose. Length 2.2 – 2.3 mm. Anal lobe weak. Veins distinct. c / w 0.55. Halter yellow. Legs. Yellow, coxal setae pale. Fore tibial organ forming a patch in depression. Fore tibial spur longer than the tibial width. Abdomen. Pale brown, setae pale and fine. Hypopygium (Fig. 2 B). Brown, as abdomen. Intergonocoxal area moderately long, with 2 long separate setose lobes. Gonocoxa moderately broad, slightly longer than gonostylus, medial margin basally strongly curved, with sparse setosity. Gonostylus (Fig. 3 D) long, apically truncate, strongly impressed; with normal setosity, a short apical tooth, and 7 rather short and slender megasetae; with 1 well-differentiated whiplash seta on ventromedial margin of the apical fourth of gonostylus. Tegmen subtriangular, laterally slightly curved, apically roundish, laterally and apically sclerotized; aedeagal teeth not detectable in the specimens studied. Aedeagal apodeme long. BIN. Unknown.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA95AFFACFF26FBF199DC0CD6.taxon	discussion	Discussion. The above description is based only on the type material. In its rather tumid gonostylus, Trichocoelina dicksoni sp. n. resembles most T. absidata sp. n. For distinguishing characters, see under the latter.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA95AFFACFF26FBF199DC0CD6.taxon	etymology	Etymology. The species is named after the Swedish merchant and philanthropist Oskar Dickson (1823 – 1897), after whom the type locality of the species, Dikson Island, was named.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA959FFACFF26FEBD9AB40AE2.taxon	description	Figs 4 A, 6 A, 18 B	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA959FFACFF26FEBD9AB40AE2.taxon	materials_examined	Material studied. Holotype male. RUSSIA, Krasnoyarsk region, Taimyr Peninsula, 12.5 km S of Dixon, 73 ° 24 ’ N, 80 ° 39 ’ E, on the river Lemberova, pan trap, 7 – 10. VII. 2012, A. Barkalov (in ISEA). Paratype. RUSSIA, same data as holotype (in MZH).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA959FFACFF26FEBD9AB40AE2.taxon	description	Description. Male. Head. Face and antenna concolorous darkish brown, maxillary palpus pale yellowish. Eye bridge 2 – 3 facets wide. Face with 21 – 22 setae. Clypeus with 1 – 2 setae. Maxillary palpus with 3 segments, 1 st segment as long as 3 rd segment, 2 nd segment shortest; 1 st segment with 1 – 2 setae, with a dorsal patch of sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 1.8 – 1.9 x as long as wide, the neck shorter than broad, the longest setae slightly shorter than the width of flagellomere. Thorax. Brown, setae pale. Anterior pronotum with 8 setae. Proepisternum with 10 setae. Scutellum with 3 longer and some short and fine setae. Wing. Fumose. Length 1.9 – 2.0 mm. Width / length 0.45. Anal lobe weak. Veins distinct. c / w 0.60. R 1 / R 0.70 – 0.85. stM shorter than fork of M. bM non-setose, r-m with 3 setae or non-setose. Halter yellow. Legs. Yellow, coxal setae pale. Fore tibial organ forming a large patch of setae in depression. Fore tibial spur longer than the tibial width. Abdomen. Pale brown, setae pale, fine and moderately long. Hypopygium (Fig. 4 A). Brown, as abdomen. Intergonocoxal area rather long, basally with strongly sclerotized medial stripe, with a large subtriangular apically bifid setose lobe (Fig. 18 B). Gonocoxa narrow, longer than gonostylus, medial margin basally smoothly curved. Gonostylus (Fig. 6 A) long, apically tumid, strongly impressed; densely setose on apical third; with a rather long apical tooth, with 2 megasetae subapically near the apical tooth and 2 or 3 at a medial position, megasetae long and slender, with distinct basal bodies; with 1 well-differentiated whiplash setae on ventromedial margin. Tegmen broader than long, apically and laterally roundish, sclerotized, aedeagal teeth not detectable in the specimens studied. Aedeagal apodeme long. BIN. Unknown.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA959FFACFF26FEBD9AB40AE2.taxon	discussion	Discussion. Trichocoelina dispansa sp. n. resembles T. chentejensis (Menzel, 1992), T. dividua sp. n., T. ithyspina sp. n., T. jukkai sp. n. and T. planilobata sp. n. in having a long apical tooth and 5 – 6 rather long gonostylar megasetae, some of which are positioned apically and / or subapically near the apical tooth, directed nearly perpendicularly, and partly more basal in position, directed more obliquely. Trichocoelina dispansa differs from all others in having a broader gonostylus, a distinctly broader tegmen and a shorter intergonocoxal area. Trichocoelina planilobata can be distinguished from other species of this group in having one very broad intergonocoxal lobe, medially only slightly notched, one gonostylar megaseta distinctly on the apical side of the tooth and the gonostylus impressed only on the apical half. Of the four other species in this group, T. chentejensis and T. ithyspina have long bodies of antennal flagellomeres (4 th flagellomere about 2.8 – 2.9 x as long as broad versus 2.0 x and ca 2.1 – 2.3 x as long as wide in T. dividua and T. jukkai). T. jukkai differs from the other remaining species in its at most very slightly apically divided medial intergonocoxal lobe and its apically roundish, not truncate, tegmen. T. jukkai differs from T. dividua further in having a more slender and more sparsely setose gonostylus, with weaker megasetae. T. dividua differs further from T. chentejensis in its longer gonostylar megasetae, and its longer and narrower tegmen and intergonocoxal lobes. T. ithyspina differs from T. chentejensis in having a long, apically bifid intergonocoxal lobe instead of two separate lobes. By the arrangement of the gonostylar megasetae, Trichocoelina obesula resembles all the species above, but differs distinctly in having a much broader gonostylus.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA959FFACFF26FEBD9AB40AE2.taxon	etymology	Etymology. The name is Latin, dispansa, spread, referring to the broad intergonocoxal lobe of the hypopygium.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA959FFADFF26F8A19B270812.taxon	description	Figs 4 B, 6 B	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA959FFADFF26F8A19B270812.taxon	materials_examined	Material studied. Holotype male. CANADA, Northwest Territories, Nahanni National Park, Reserve Nailicho (Virginia Falls), 61.606 ° N, 125.758 ° W, 578 m, 1. VIII. 2014, Parks Canada (BOLD Sample ID BIOUG 17502 - A 06, in CNC). Paratypes. CANADA, same data as holotype, 1 male (BOLD Sample ID BIOUG 17506 - C 03, in CBG); same data as previous but 4. VII. 2014, 3 males (BOLD Sample IDs BIOUG 17107 - F 02, BIOUG 17109 - F 06 and BIOUG 17129 - B 09, in CBG); same data but 12. VII. 2014, 1 male (BOLD Sample ID BIOUG 17213 - C 11, in CBG); same data but 25. VII. 2014, 1 male (BOLD Sample ID BIOUG 17423 - E 03, in CBG).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA959FFADFF26F8A19B270812.taxon	description	Description. Male. Head. Face brown, maxillary palpus pale yellowish. Eye bridge 2 – 3 facets wide. Face with 10 – 20 setae. Clypeus with 1 – 2 setae. Maxillary palpus with 3 segments, 1 st segment longer than 3 rd segment, 2 nd segment shortest; 1 st segment with 1 seta, with a large dorsal patch of sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 2.0 x as long as wide, the neck shorter than broad, the longest setae shorter than the width of flagellomere. Thorax. Brown, setae pale. Anterior pronotum with 5 – 9 setae. Proepisternum with 5 – 10 setae. Scutellum with 3 – 4 longer and some short and fine setae. Wing. Fumose. Length 1.5 – 1.6 mm. Width / length 0.35 – 0.40. Anal lobe weak. Veins distinct. c / w 0.65. R 1 / R 0.45 – 0.7. stM as long as fork of M. r-m as long as bM, bM non-setose, r-m non-setose or with 1 seta. Halter yellow. Legs. Yellow, coxal setae pale. Fore tibial organ forming a rather small patch in depression. Fore tibial spur longer than the tibial width. Abdomen. Pale brown, setae pale, long and fine. Hypopygium (Fig. 4 B). Brown, as abdomen. Intergonocoxal area moderately long, with long, apically bifid setose lobe. Gonocoxa moderately broad, longer than gonostylus, medial margin basally strongly curved, with sparse setosity. Gonostylus (Fig. 6 B) long, apically tumid, impressed; with normal setosity, a long apical tooth, and one megaseta on the dorsal side of apical tooth, one subapical and 3 – 4 megasetae more basally, megasetae long and slender; with one well-differentiated whiplash seta on ventromedial margin. Tegmen as long as broad, laterally roundish, apically straight, weakly sclerotized; aedeagal teeth not detectable in the specimens studied. Aedeagal apodeme long. BIN. BOLD: AAL 7893.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA959FFADFF26F8A19B270812.taxon	discussion	Discussion. Trichocoelina dividua sp. n. is especially similar to T. jukkai sp. n. For further discussion of these and similar species, see under Trichocoelina dispansa sp. n.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA959FFADFF26F8A19B270812.taxon	etymology	Etymology. The name is Latin, dividua, parted, referring to the apically distinctly divided intergonocoxal lobe of hypopygium.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA958FFA1FF26FBCE9F2F0D42.taxon	description	Figs 5 A, 6 C, 18 C	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA958FFA1FF26FBCE9F2F0D42.taxon	materials_examined	Material studied. Holotype male. CANADA, Yukon, Ogilvie Mountains, North Fork Pass, 4100 ft, 20. VI. 1962, R. E. Leech (in CNC).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA958FFA1FF26FBCE9F2F0D42.taxon	description	Description. Male. Head. Colours faded in the specimen studied. Eye bridge 3 facets wide. Setae of face not detectable in the specimen studied. Clypeus with 3 setae. Maxillary palpus with 3 segments, 3 rd segment longer than 1 st segment, 2 nd segment shortest; 1 st segment with 9 setae, with a dorsal patch of sensilla; antennae missing in the specimen studied. Thorax. Colours faded in the specimen studied. Anterior pronotum with 9 setae. Proepisternum with 18 setae. Scutellum with 5 longer and some short and fine setae. Wing. Fumose. Length 2.8 mm. Width / length 0.40. Anal lobe weak. Veins distinct. c / w 0.55. R 1 / R 0.60. stM shorter than fork of M. r-m 2 x as long as bM, bM and r-m non-setose. Legs. Colours faded in the specimen studied. Fore tibial organ forming a large patch in depression. Fore tibial spur slightly longer than the tibial width. Abdomen. Colours faded in the specimen studied. Hypopygium (Fig. 5 A). Colours faded in the specimen studied. Intergonocoxal area long, with two setose lobes (Fig. 18 C). Gonocoxa broad, slightly longer than gonostylus, medial margin basally rather strongly curved, with short sparse setosity. Gonostylus (Fig. 6 C) broad, apically tumid, strongly impressed; with short dense setosity, with a rather long apical tooth, with about 15 slender and rather short perpendicular megasetae; with 1 well-differentiated whiplash seta on ventromedial margin. Tegmen apically narrowed and truncated, laterally roundish, weakly sclerotized, aedeagal teeth not detectable in the specimen studied. Aedeagal apodeme long. BIN. Unknown.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA958FFA1FF26FBCE9F2F0D42.taxon	discussion	Discussion. By its broad and apically tumid gonostylus which is subbasally strongly broadened and which has numerous short megasetae, Trichocoelina hians sp. n. resembles T. hiemalis (Mohrig & Mamaev), T. incrassata sp. n. and T. magnifica sp. n. Trichocoelina hians differs from T. hiemalis in having the gonostylar apex broader and the apical tooth and megasetae shorter but intergonocoxal lobes of hypopygium longer, see Fig. 5 A and fig. 4 in Mohrig et al. (1985 b). Trichocoelina hians differs from T. incrassata and T. magnifica in having its gonostylus slightly narrower and lacking a subbasal lobe with a group of megasetae. From T. incrassata it differs also in having its tegmen less modified, laterally smoothly curved. Furthermore, Trichocoelina hians differs from T. magnifica in having all gonostylar megasetae of the same size, perpendicular and not arranged in groups, and in having shorter intergonocoxal lobes. See also under T. absidata sp. n.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA958FFA1FF26FBCE9F2F0D42.taxon	etymology	Etymology. The name is Latin, hians, opened, referring to its medially strongly excavated gonostylus.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA954FFA1FF26FE019E110ECE.taxon	description	BIN. Unknown.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA954FFA1FF26FE019E110ECE.taxon	discussion	Discussion. The species was described from three males from Southern Yamal, Northern Russia (Mohrig et al. 1985). By its broad, strongly impressed, basally broad gonostylus with numerous straight megasetae it resembles most the Nearctic Trichocoelina hians sp. n. (see above and under T. semisphaera sp. n.).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA954FFA1FF26FCB59B3F0ABE.taxon	description	Figs 5 B, 6 D, 18 A	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA954FFA1FF26FCB59B3F0ABE.taxon	materials_examined	Material studied. Holotype male. CANADA, Yukon, Ogilvie Mountains, North Fork Pass, 4100 ft, 20. VI. 1962, R. E. Leech (in CNC).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA954FFA1FF26FCB59B3F0ABE.taxon	description	Description. Male. Head. Face brown, maxillary palpus pale yellowish. Eye bridge 3 facets wide. Face with 21 fine setae. Clypeus with 2 setae. Maxillary palpus with 3 segments, 1 st segment as long as 3 rd segment, 2 nd segment shortest; 1 st segment with 3 setae, with a dorsal patch of sensilla; antennae missing in the specimen studied. Thorax. Brown, setae pale. Anterior pronotum with 5 setae. Proepisternum with 19 setae. Scutellum with 4 longer and some short and fine setae. Wing. In poor condition in the specimen studied. Fumose. Length about 2.8. Veins distinct. R 1 / R 0.70. r-m much longer than bM, bM non-setose, r-m with 4 setae. Halter yellow. Legs. Yellow, coxal setae pale. Fore tibial organ forming a large patch in depression. Fore tibial spur slightly longer than the tibial width. Abdomen. Pale brown, setae pale, fine and moderately long. Hypopygium (Fig. 5 B). Brown, as abdomen. Intergonocoxal area long, basally with strongly sclerotized medial stripe, with 2 short setose lobes (Fig. 18 A). Gonocoxa broad, as long as gonostylus, medial margin basally strongly curved, with short sparse setosity. Gonostylus (Fig. 18 A) long and broad, apically truncate, strongly impressed; with short and dense setosity, with a short dorsal apical tooth in subapical position, with 10 – 12 megasetae medially, megasetae long and slender; with one long and 2 – 3 shorter whiplash setae basad of the megasetae. Tegmen longer than broad, conical, with sharp apicolateral corners, basolaterally broadened, apically and laterally sclerotized; aedeagal teeth not detectable in the specimen studied. Aedeagal apodeme long. BIN. Unknown.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA954FFA1FF26FCB59B3F0ABE.taxon	discussion	Discussion. Trichocoelina imitator sp. n. is very similar to T. oricillifera sp. n. in the form of the large gonostylus with numerous perpendicular megasetae but differs from the latter in the form of the tegmen, which is basally broader and has a longer apical part, and in having shorter intergonocoxal lobes of hypopygium.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA954FFA1FF26FCB59B3F0ABE.taxon	etymology	Etymology. The name is Latin, imitator, referring to the close resemblance of the species to Trichocoelina oricillifera sp. n.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA954FFA6FF26F8859EF70F82.taxon	description	Figs 7 A, 9 A	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA954FFA6FF26F8859EF70F82.taxon	materials_examined	Material studied. Holotype male. USA, Alaska, 11 mi S Anderson Jct, Rte 3, mi 270, alder-poplar-spruce, Malaise trap, 23. VI – 11. VIII. 1984, S. & J. Peck (in MZH).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA954FFA6FF26F8859EF70F82.taxon	description	Description. Male. Head. Face and antenna concolorous brown, maxillary palpus pale yellowish. Eye bridge 2 – 3 facets wide. Face with 20 fine setae. Clypeus with 1 seta. Maxillary palpus with 3 segments, 3 rd segment longer than 1 st segment, 2 nd segment shortest; 1 st segment with 1 seta, with a dorsal patch of sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 2.2 x as long as wide, the neck shorter than broad, the longest setae as long as the width of flagellomere. Thorax. Brown, setae pale. Anterior pronotum with 6 setae. Proepisternum with 15 setae. Scutellum with 4 longer and some short and fine setae. Wing. Fumose. Length 2.6 mm. Width / length 0.40. Anal lobe weak. Veins distinct. c / w 0.70. R 1 / R 0.55. stM and fork of M subequal in length. r-m 2 x as long as bM. bM and r-m non-setose. Halter yellow. Legs. Yellow, coxal setae pale. Fore tibial organ forming a large patch in depression. Fore tibial spur as long as the tibial width. Abdomen. Pale brown, setae pale, rather short. Hypopygium (Fig. 7 A). Brown, as abdomen. Intergonocoxal area long, with two small setose lobes. Gonocoxa broad, about as long as gonostylus, medial margin basally smoothly curved, with short sparse setosity. Gonostylus (Fig. 9 A) semiglobular, strongly impressed; subbasally with a dorsal lobe, with short setosity, with a rather long and narrow apical tooth, with numerous megasetae medially, the ventral megasetae oblique, the dorsal ones perpendicular, megasetae slender and rather short; with 1 weakly differentiated whiplash seta on ventromedial margin. Tegmen as long as broad, apically truncate, laterally with distinct shoulders, sclerotized, with an area of small aedeagal teeth. Aedeagal apodeme long and strong. BIN. Unknown.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA954FFA6FF26F8859EF70F82.taxon	discussion	Discussion. By its large semigobular gonostylus with numerous short megasetae Trichocoelina incrassata sp. n. resembles most T. janetcheki (Lengersdorf, 1953) but differs in having groups of gonostylar megasetae pointing in two directions, in having shorter intergonocoxal lobes and in having the tegmen with lateral corners but without the apical process (see also under T. hians sp. n.).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA954FFA6FF26F8859EF70F82.taxon	etymology	Etymology. The name is Latin, incrassata, thickened, referring to the thick gonostylus.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA953FFA6FF26FC4198100B96.taxon	description	Figs 8 A, 9 B, 18 C	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA953FFA6FF26FC4198100B96.taxon	materials_examined	Material studied. Holotype male. NORWAY, Hedmark, Stor-Elvdal, river Atna Solbakken, 61.746 ° N, 10.746 ° E, 380 m, Malaise trap, 30. VI – 7. VII. 2008, E. Stur (NTNU-VM 201775, BOLD Sample ID TRD-Sci 029, in NTNU- VM).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA953FFA6FF26FC4198100B96.taxon	description	Description. Male. Head. Face and antenna concolorous brown but pedicellus yellow, maxillary palpus pale yellowish. Eye bridge, face and clypeus poorly visible in the specimen studied. Maxillary palpus with 3 segments, 1 st segment longer than 3 rd segment, 2 nd segment shortest; 1 st segment with 1 seta, with dorsal patch of sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 2.9 x as long as wide, the neck shorter than broad, the longest setae longer than the width of flagellomere. Thorax. Brown, setae pale. Anterior pronotum with 9 setae. Proepisternum with 11 setae. Scutellum with 4 longer and some short and fine setae. Wing. Fumose. Length 2.3 mm. Width / length 0.45. Anal lobe weak. Veins distinct. c / w 0.65. R 1 / R 0.80. stM shorter than fork of M. r-m longer than bM, bM non-setose, r-m with 1 seta. Halter yellow. Legs. Yellow, coxal setae pale. Fore tibial organ forming a large indistinct patch in depression. Fore tibial spur longer than the tibial width. Abdomen. Pale brown, setae pale, moderately long and fine. Hypopygium (Fig. 8 A). Brown, as abdomen. Intergonocoxal area moderately long, with long, apically bifid setose lobe (Fig. 18 C). Gonocoxa moderately broad, longer than gonostylus, medial margin basally smoothly curved, with rather long but sparse setosity. Gonostylus (Fig. 9 B) long, apically truncate, strongly impressed; with normal setosity, with a rather long apical tooth, with three megasetae subapically near the tooth and two more basally, megasetae long and slender, with distinct basal bodies; with 1 well-differentiated whiplash seta on ventromedial margin. Tegmen broader than long, laterally roundish, apically straight, weakly sclerotized, aedeagal teeth not detectable in the specimens studied. Aedeagal apodeme long. BIN. BOLD: ACX 5966.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA953FFA6FF26FC4198100B96.taxon	discussion	Discussion. The gonostylus shown in Figs 8 A and 9 B was slightly distorted in preparation. Trichocoelina ithyspina sp. n. resembles most T. jukkai sp. n. For discussion, see under T. dispansa sp. n.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA953FFA6FF26FC4198100B96.taxon	etymology	Etymology. The name is derived from the Greek word ithys, straight, and the Latin word spina, spine, referring to the straight megasetae of the gonostylus.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA96FFF9AFF26FF0D996409E6.taxon	description	Figs 10 A, 10 B, 17 D Literature. Neosciara janetscheki Lengersdorf — Lengersdorf (1953): 167, fig. 1; Janetschek (1956): 471. Lycoriella (Hemineurina) janetscheki (Lengersdorf) — Tuomikoski (1959 a): 35; Gerbachevskaja-Pavluchenko (1986): 31; Franz (1989): 14; Menzel & Mohrig (2000): 410; Wirta et al. (2016): appendix, unpaginated p. 21 (table S 1) and unpaginated p. 39 (cladogram).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA96FFF9AFF26FF0D996409E6.taxon	materials_examined	Material studied. CANADA, Nunavut, Sverdrup Islands, Elles Ringnes Island, Isachsen, 70.79 ° N, 103.55 ° W, lemming burrow, 17. VI. 1960, J. R. Vockeroth, 4 males (3 in CNC, 1 in MZH); same locality but moss near very small stream, 14. VII. 1960, J. F. McAlpine, 2 males (in CNC); Nunavut, Ellesmere Island, Fosheim Peninsula, Hot Weather Creek, 79 ° 58 ’ N, 84 ° 28 ’ W, 2. VII. 1990, F. Brodo, 1 male (in MZH); NE Greenland, Mestersvig, 72.24 ° N, 23,92 ° W, C. Vibe (in ZMUC); NE GREENLAND, Zackenberg, 74 ° 28 ’ N, 20 ° 34 ’ W (UTM 8265500: 0513778), 44 m, 2 – 11. VII. 2011, T. Roslin & G. Várkonyi, 1 male (in MZH); same locality but UTM 8265758: 0513786, 37 m, Malaise trap, 11 – 20. VII. 2011, T. Roslin & G. Várkonyi, 2 males (in MZH, SDEI); same locality but 74.50 ° N, 21.00 ° W, 44 m, 7. VII. 2011, T. Roslin & G. Várkonyi, 1 male (BOLD Sample ID GRPV 16, in DAUH); same locality but 74.4667 ° N, 20.5667 ° W, 48 m, 24. VII. 2011, T. Roslin & G. Várkonyi, 3 males (BOLD Sample IDs ZA 2012 - 50010, ZA 2012 - 50011 and ZA 2012 - 50012, in DAUH); USA, Colorado, Mt. Evans, 14 000 ft, 25. VII. 1961, W. R. M. Mason, 1 male (in USNM).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA96FFF9AFF26FF0D996409E6.taxon	description	Redescription. Male. Head. Face brown, antenna dark brown maxillary palpus pale yellowish. Eye bridge 2 – 3 facets wide. Face with 15 – 21 setae. Clypeus with 1 – 4 setae. Maxillary palpus with 3 segments, 1 st segment as long as or longer than 3 rd segment, 2 nd segment shortest; 1 st segment with 3 – 9 setae, with dorsal patch of sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 1.95 – 2.75 x as long as wide, the neck shorter than broad, the longest setae shorter than the width of flagellomere. Thorax. Dark brown, setae pale. Anterior pronotum with 5 – 11 setae. Proepisternum with 7 – 24 setae. Scutellum with 4 longer and some short and fine setae. Wing. Fumose. Length 2.2 – 3.1 mm. Width / length 0.35 – 0.40. Anal lobe weak. Veins distinct. c / w 0.50 – 0.60. R 1 / R 0.50 – 0.90. stM shorter than fork of M. r-m longer than bM, bM non-setose, r-m non-setose or with 1 – 2 setae. Halter yellow. Legs. Yellow, coxal setae pale. Fore tibial organ forming a large patch in shallow depression. Fore tibial spur as long as the tibial width. Abdomen. Pale brown, setae pale, short and fine. Hypopygium (Fig. 10 B). Brown, as abdomen. Intergonocoxal area long, with two short setose lobes. Gonocoxa broad, as long as gonostylus, medial margin basally smoothly curved, with short and fine setosity. Gonostylus (Fig. 10 A) voluminous, apically truncate, impressed; with short setosity, a short apical tooth, and numerous megasetae in the medial impression; megasetae slightly curved or straight, short and slender; with 1 well-differentiated whiplash seta basally at ventromedial margin. Tegmen (Fig. 17 D) longer than broad, laterally straight, apically with a narrow hyalinous acuminate process, weakly sclerotized, aedeagal teeth not detectable in the specimens studied. Aedeagal apodeme long. BIN. BOLD: ACK 5495.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA96FFF9AFF26FF0D996409E6.taxon	discussion	Discussion. In Fig. 10 A the gonostylus is flattened and distorted to show the arrangement of the megasetae. In its extremely voluminous gonostylus, Trichocoelina janetscheki (Lengersdorf, 1953) resembles most T. incrassata sp. n., for distinguishing characters, see under the latter.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA96FFF9BFF26F9AD9B6D0ECE.taxon	description	Figs 8 B, 9 D, 18 D	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA96FFF9BFF26F9AD9B6D0ECE.taxon	materials_examined	Material studied. Holotype male. FINLAND, Li (Lapponia inarensis), Utsjoki, Pummankijoki (grid 7759292: 539670), sandy shore, pitfall trap, 13. VI – 13. VII. 2016, J. Salmela (in MZH). Paratypes. FINLAND, same data as holotype, 10 males (6 in MZH, 2 in SDEI, 2 in PJSR); NORWAY, Troms, Tromsø, Nakkedalen, 300 m S of Estengammen, 69.612 ° N, 19.592 ° E, 150 m, Malaise trap, 16 – 19. VII. 2012, T. E. Barstad, 2 males (BOLD Sample IDs TSZD-JKJ- 103937 and TSZD-JKJ- 104088, in TMU).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA96FFF9BFF26F9AD9B6D0ECE.taxon	description	Description. Male. Head. Face and antenna brown, antenna darker, maxillary palpus pale yellowish. Eye bridge 2 facets wide. Face with 14 – 20 setae. Clypeus with 1 – 2 setae. Maxillary palpus with 3 segments, 1 st segment longer than 3 rd segment, 2 nd segment shortest; 1 st segment with 1 – 2 setae, with a large dorsal patch of sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 2.15 – 2.25 x as long as wide, the neck shorter than broad, the longest setae longer than the width of flagellomere. Thorax. Dark brown, setae pale. Anterior pronotum with 3 – 8 setae. Proepisternum with 7 – 9 setae. Scutellum with 4 longer and some short and fine setae. Wing. Fumose. Length 1.6 – 1.7 mm. Width / length 0.35 – 0.40. Anal lobe weak. Veins distinct. c / w 0.55 – 0.70. R 1 / R 0.45 – 0.55. stM longer or shorter than fork of M. r-m longer than bM, both non-setose. Halter yellow. Legs. Yellow, coxal setae pale. Fore tibial organ forming a rather small patch in depression. Fore tibial spur longer than the tibial width. Abdomen. Pale brown, setae pale and long. Hypopygium (Fig. 8 B). Brown, as abdomen. Intergonocoxal area moderately long, with long, apically bifid setose lobe (Fig. 18 D). Gonocoxa moderately broad, longer than gonostylus, medial margin basally strongly curved, with sparse setosity. Gonostylus (Fig. 9 D) long, apically truncate, impressed; with normal setosity, a long apical tooth, 1 megaseta on the apical side of the tooth, a pair of megaseta subapically near the tooth and another pair more basally; megasetae long and slender; with 1 well-differentiated whiplash seta at ventromedial margin. Tegmen broader than long, laterally and apically roundish, weakly sclerotized, aedeagal teeth not detectable in the specimens studied. Aedeagal apodeme long. BIN. BOLD: ADL 0130.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA96FFF9BFF26F9AD9B6D0ECE.taxon	discussion	Discussion. See under Trichocoelina dividua sp. n.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA96FFF9BFF26F9AD9B6D0ECE.taxon	etymology	Etymology. The species is named after the dipterist Jukka Salmela, Rovaniemi, Finland, who collected the holotype.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA96EFF9BFF26FCB59F420AE2.taxon	description	Figs 7 B, 9 C, 17 F	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA96EFF9BFF26FCB59F420AE2.taxon	materials_examined	Material studied. Holotype male. CANADA, Yukon, North Fork Crossing, Mi 43, Peel Plt. Rd., 3500 ft, 4. VII. 1962, R. J. Leech, 1 male (in CNC). Paratypes. CANADA, same data as holotype but 26. VI. 1962, 1 male (in MZH); Yukon, Ogilvie Mountains, 7. VII. 1962, P. J. Skitsko, 1 male (in MZH).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA96EFF9BFF26FCB59F420AE2.taxon	description	Description. Male. Head. Colours faded in the specimens studied. Eye bridge 2 facets wide. Face with 10 – 19 fine setae. Clypeus with 2 setae. Maxillary palpus with 3 segments, 1 st segment longer than 3 rd segment, 2 nd segment shortest; 1 st segment with 1 – 2 setae, with a dorsal patch of sensilla; antennae missing in the specimens studied. Thorax. Colours faded. Anterior pronotum with 6 – 9 setae. Proepisternum with 13 – 15 setae. Scutellum with 3 longer and some short and fine setae. Wing. Fumose. Length 2.0 – 2.2 mm. Width / length 0.40. Anal lobe weak. Veins distinct. c / w 0.55. R 1 / R 0.60 – 0.65. stM and fork of M subequal in length. r-m and bm subequal in length. bM non-setose, r-m with 3 – 4 setae non-setose. Legs. Colours faded. Fore tibial organ forming a small patch of strong setae in depression. Fore tibial spur longer than the tibial width. Abdomen. Colours faded. Hypopygium (Fig. 7 B). Intergonocoxal area long, basally with strongly sclerotized medial stripe, with two distinct setose lobes. Gonocoxa broad, as long as gonostylus, medial margin basally smoothly curved, with normal setosity. Gonostylus (Fig. 9 C) large, laterally roundish, apically slightly narrowed, medially impressed; with short setosity, with a short apical tooth, one megaseta with strong basal body at the apical third and a group of 3 megasetae on common basal body in a more basal position, on the dorsal side of the latter a group of 10 – 12 megasetae, megasetae rather straight, longer than apical tooth; with a long whiplash seta on ventromedial margin near the basalmost megasetae. Tegmen (Fig. 17 F) longer than broad, apically narrowed, with sharp corners at middle, sclerotized, with large area of small aedeagal teeth. Aedeagal apodeme moderate. BIN. Unknown.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA96EFF9BFF26FCB59F420AE2.taxon	discussion	Discussion. Trichocoelina magnifica sp. n. resembles T. nefrens sp. n. in having a highly modified tegmen and one outstanding medial megaseta, but differs from the latter in its broader tegmen, in having an apical tooth and lacking any subapical megasetae on the gonostylus. See also under T. hians sp. n.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA96EFF9BFF26FCB59F420AE2.taxon	etymology	Etymology. The name is Latin, magnifica, referring to the magnificent structure of the hypopygium.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA96EFF98FF26F8A19FEF0812.taxon	description	Figs 11 A, 13 B, 17 E	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA96EFF98FF26F8A19FEF0812.taxon	materials_examined	Material studied. Holotype male. RUSSIA, Krasnoyarsk region, Taimyr Nature Reserve, Aru-Mas, 72.50 ° N, 101.94 ° E, pan trap, 9 – 20. VII. 2010, A. Barkalov (in ISEA). Paratypes. RUSSIA, same data as holotype, 2 males (in MZH); Krasnoyarsk region, Taimyr Peninsula, on Zakharova River, Rassokha, 72.70 ° N, 101.08 ° E, pan trap, 1 – 10. VII. 2011, A. Barkalov, 2 males (in MZH, SDEI); Krasnoyarsk region, Taimyr Nature Reserve, VII. 2010, A. Barkalov, 1 male (in ISEA).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA96EFF98FF26F8A19FEF0812.taxon	description	Description. Male. Head. Face brown, antenna darker brown, maxillary palpus pale yellowish. Eye bridge 2 – 3 facets wide. Face with 24 – 26 fine setae. Clypeus with 2 – 3 setae. Maxillary palpus with 3 segments, 1 st segment longer than 3 rd segment, 2 nd segment shortest; 1 st segment with 1 – 2 setae, with a dorsal patch of sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 1.95 – 2 x as long as wide, the neck shorter than broad, the longest setae slightly shorter than the width of flagellomere. Thorax. Brown, setae pale. Anterior pronotum with 4 – 7 setae. Proepisternum with 6 – 8 setae. Scutellum with 4 longer and some short and fine setae. Wing. Fumose. Length 2.0 – 2.2 mm. Width / length 0.40. Anal lobe weak. Veins distinct. c / w 0.65 – 0.75. R 1 / R 0.50 – 0.60. stM slightly shorter than fork of M. r-m longer than bM, bM non-setose, r-m with 3 – 4 setae. Halter yellow. Legs. Yellow, coxal setae pale. Fore tibial organ forming a dense patch of fine setae in depression. Fore tibial spur slightly longer than the tibial width. Abdomen. Pale brown, setae pale and fine. Hypopygium (Fig. 11 A). Brown, as abdomen. Intergonocoxal area long, with 2 distinct narrow setose lobes. Gonocoxa broad, as long as gonostylus, basally with strongly sclerotized medial stripe, medial margin basally smoothly curved, with short sparse setosity. Gonostylus (13 B) broad, laterally roundish, apically strongly curved and narrowed, dorsally forming a lobe; strongly impressed; with short setosity, without apical tooth, with one dorsal and two ventral subapical megasetae and a group of three ventral and six dorsal megasetae subbasally, megasetae slightly curved and on distinct basal bodies; with 1 weakly differentiated whiplash seta on ventromedial margin. Tegmen (Fig. 17 E) much longer than broad, conical with sharp apex, with sharp lateral corners, sclerotized, with area of very tiny aedeagal teeth. Aedeagal apodeme long. BIN. Unknown.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA96EFF98FF26F8A19FEF0812.taxon	discussion	Discussion. Trichocoelina nefrens sp. n. differs from all other known species of the genus in lacking the apical tooth and in the peculiar form of the gonostylus. See also under T. magnifica sp. n.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA96EFF98FF26F8A19FEF0812.taxon	etymology	Etymology. The name is Latin, nefrens, toothless, referring to the lack of the apical tooth on the gonostylus.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA96DFF9CFF26FBF19B1F0BC6.taxon	description	Figs 11 B, 13 A	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA96DFF9CFF26FBF19B1F0BC6.taxon	materials_examined	Material studied. Holotype male. NORWAY, ‘ Svalbard, Engelskelva’ [= Svalbard, Bjørnøya, river Engelskelva in the NE part of island], 74.4768 ° N, 19.1779 ° E, 30 m, 30. VII. 2009, T. Ekrem (NTNU-VM 50594 and BOLD Sample ID BJ 244, in NTNU-VM). Paratypes. NORWAY, same data as holotype, 1 male (NTNU-VM 50595 and BOLD Sample ID BJ 245, in NTNU-VM); ‘ Svalbard, Lakselva’ [= Svalbard, Bjørnøya, river Lakselva], 74.4994 ° N, 18.9776 ° E, 15 m, 3. VIII. 2009, T. Ekrem, 1 male (NTNU-VM 50605 and BOLD Sample ID BJ 255, in NTNU- VM).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA96DFF9CFF26FBF19B1F0BC6.taxon	description	Description. Male. Head. Face and antenna concolorous brown, maxillary palpus pale yellowish. Eye bridge 3 facets wide. Face with 9 setae. Clypeus with 1 seta. Maxillary palpus with 3 segments, 1 st segment as long as 3 rd segment, 2 nd segment shortest; 1 st segment with 1 – 3 setae, with dorsal patch of sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 1.5 – 1.75 x as long as wide, the neck shorter than broad, the longest setae as long as the width of flagellomere. Thorax. Brown, setae pale. Anterior pronotum with 8 – 9 setae. Proepisternum with 10 – 13 setae. Scutellum with 4 longer and some short and fine setae. Wing. Fumose. Length 1.8 – 2.0 mm. Width / length 0.40. Anal lobe weak. Veins distinct. c / w 0.65 – 0.75. R 1 / R 0.55 – 0.60. stM shorter than fork of M. r-m as long as bM or longer, both non-setose. Halter yellow. Legs. Yellow, coxal setae pale. Fore tibial organ forming an indistinct sparse patch in depression. Fore tibial spur longer than the tibial width. Abdomen. Pale brown, setae pale, moderately long and fine. Hypopygium (Fig. 11 B). Brown, as abdomen. Intergonocoxal area moderately long, with two short, triangular setose lobes. Gonocoxa broad, longer than gonostylus, medial margin basally smoothly curved, with rather long but sparse setosity. Gonostylus (Fig. 13 A) broad, apically slightly narrowed, strongly impressed; with short and dense setosity, a short apical tooth, a megaseta on the ventral side of apical tooth, some subapical and some medial megasetae more basally, megasetae slightly curved, long and slender; with 1 welldifferentiated whiplash seta on ventromedial margin. Tegmen broader than long, laterally slightly curved, apically straight, weakly sclerotized, aedeagal teeth not detectable in the specimens studied. Aedeagal apodeme long. BIN. BOLD: ABA 5289.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA96DFF9CFF26FBF19B1F0BC6.taxon	discussion	Discussion. By its gonostylus and tegmen, Trichocoelina obesula sp. n. resembles T. dispansa sp. n. and reminiscent species but differs in having the gonostylus broader and more richly setose. See under T. dispansa.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA96DFF9CFF26FBF19B1F0BC6.taxon	etymology	Etymology. The name is derived from the Latin word obesa, fat, referring to the broad gonostylus and gonocoxa of the species.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA968FF9DFF26FF0D9EAB0DF0.taxon	description	New records. CANADA, Yukon, Ogilvie Mts., North Fork Pass, 4100 ft, 21. VI. 1962, P. J. Skitsko, 1 male (in CNC); USA, Colorado, Mt. Evans, Summit L. (Lake), Flats, 24. VII. 1961, C. W. Mann, 1 male (in MZH). BIN. Unknown.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA968FF9DFF26FF0D9EAB0DF0.taxon	discussion	Discussion. The species was described from one male from Southern Yamal by Mohrig & Mamaev in Mohrig et al. (1983 a) and redescribed by Menzel & Mohrig (2000). The species differs from all other Trichocoelina in in its gonostylus having a peculiar median lobe with a few megasetae pointing in different directions.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA968FF9DFF26FDA19BA40B10.taxon	description	Figs 12 B, 13 D, 17 B Literature. Lycoriella hiemalis Mohrig & Mamaev — Heller et al. (2009): 42 [misidentification].	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA968FF9DFF26FDA19BA40B10.taxon	materials_examined	Material studied. Holotype male. FINLAND, Lkor (Lapponia kemensis orientalis) Salla, Värriö, Kuntasjoki (grid 7520483: 3610905), spruce mire, brook, Malaise trap, 29. VI – 29. VII. 2013, J. Salmela (in MZH). Paratypes. FINLAND, same data as holotype, 1 male (in SDEI); Le (Lapponia enontekiensis), Kilpisjärvi, Saana, SW-slope, Malaise trap, 2006, J. Jakovlev & J. Penttinen, 1 male (in MZH); Lkor (Lapponia kemensis orientalis), Kittilä, Pallas-Yllästunturi National Park, SE of Lehtirova, 67 º 59 ’ N, 24 º 05 ’ E, mixed taiga (spruce, birch, pine), sweep-net, 18. VII. 2005, M. Jaschhof, 1 male (in SDEI); NORWAY, Finnmark, Tana, Nedre Storfoss, 4. VII. 1964, R. Tuomikoski, 1 male (in MZH); Finnmark, Karasjok, 12. VIII. 1985, J. Tuiskunen, 1 male (in MZH); SWEDEN, Lapland, Norrbottens län, Arjeplog, lake Sädvajaure (northern end), 66.33 ° N, 16.20 ° E, 500 m, subalpine birch forest, Malaise trap, 7. VII – 12. VIII. 2005, M. Jaschhof & C. Jaschhof, 5 males (no. 5846, 5848 and 5860, in PKHH; no. 898 and 980 in SMNH) [locality published in Heller et al. (2009) as ‘ Pite lappmark, Arjeplog 2 ’].	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA968FF9DFF26FDA19BA40B10.taxon	description	Description. Male. Head. Face brown, antenna slightly darker brown, maxillary palpus yellowish. Eye bridge 2 facets wide. Face with 17 – 28 fine setae. Clypeus with 2 – 3 setae. Maxillary palpus with 3 segments, 1 st segment either longer or shorter than 3 rd segment, 2 nd segment shortest; 1 st segment with 3 – 6 setae, with a dorsal patch of sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 2.3 – 2.6 x as long as wide, the neck shorter than broad, the longest setae shorter than the width of flagellomere. Thorax. Brown, setae pale. Anterior pronotum with 8 – 13 setae. Proepisternum with 10 – 13 setae. Scutellum with 3 – 4 longer and some short and fine setae. Wing. Fumose. Length 2.5 – 2.9 mm. Width / length 0.40. Anal lobe moderate. Veins distinct. c / w 0.60 – 0.85. R 1 / R 0.70 – 0.85. stM longer than fork of M. r-m and bM of variable lengths, bM non-setose, r-m with 2 – 4 setae. Halter yellow. Legs. Yellow, coxal setae pale. Fore tibial organ forming a large patch in depression. Fore tibial spur slightly longer than the tibial width. Abdomen. Pale brown, setae pale, fine and moderately long. Hypopygium (Fig. 12 B). Brown, as abdomen. Intergonocoxal area long, basally with strongly sclerotized medial stripe, with 2 distinct setose lobes (Fig. 17 B). Gonocoxa broad, as long as gonostylus, medial margin basally strongly curved, with short sparse setosity. Gonostylus (Fig. 13 D) long, apically truncate, strongly impressed; with short setosity, with a short dorsal apical tooth in subapical position, with 8 – 12 medial megasetae, megasetae long and slender, straight or slightly curved; with one long and 2 – 3 shorter whiplash setae basad of the megasetae. Tegmen longer than broad, conical, with sharp apicolateral corners, basolaterally broadened, apically and laterally sclerotized; aedeagal teeth not detectable in the specimens studied. Aedeagal apodeme long. BIN. Unknown.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA968FF9DFF26FDA19BA40B10.taxon	discussion	Discussion. See under Trichocoelina imitator sp. n.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA968FF9DFF26FDA19BA40B10.taxon	etymology	Etymology. The name is derived from the Latin words oricilla, small ear, and - fer, bearing, referring to the lateral lobes of the tegmen.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA968FF92FF26F8C1984B0F83.taxon	description	Figs 12 A, 13 C, 18 E	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA968FF92FF26F8C1984B0F83.taxon	materials_examined	Material studied. Holotype male. FINLAND, Li (Lapponia inarensis), Utsjoki, Pummankijoki (grid 7759292: 539670), sandy river bank, pitfall trap, 13. VI – 13. VII. 2016, J. Salmela (in MZH).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA968FF92FF26F8C1984B0F83.taxon	description	Description. Male. Head. Face and antenna pale brown, maxillary palpus pale yellowish. Eye bridge 3 – 4 facets wide. Face with 20 setae. Clypeus with 1 seta. Maxillary palpus with 3 segments, 1 st segment longer than 3 rd segment, 2 nd segment shortest; 1 st segment with 1 seta, with a large dorsal patch of sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 2.3 x as long as wide, the neck shorter than broad, the longest setae longer than the width of flagellomere. Thorax. Dark brown, setae pale. Anterior pronotum with 5 setae. Proepisternum with 12 setae. Scutellum with 4 longer and some short and fine setae. Wing. Fumose. Length 1.7 mm. Width / length 0.40. Anal lobe rather weak. Veins distinct. c / w 0.70. R 1 / R 0.50. stM shorter than fork of M. r-m longer than bM, both non-setose. Halter yellow. Legs. Yellow, coxal setae pale. Fore tibial organ forming a rather large patch in shallow depression. Fore tibial spur longer than the tibial width. Abdomen. Pale brown, setae pale and rather long. Hypopygium (Fig. 12 A). Brown, as abdomen. Intergonocoxal area moderately long, with very broad, apically divided setose lobe (Fig. 18 E). Gonocoxa moderately broad, longer than gonostylus, medial margin strongly curved, with sparse setosity, ventroapically with denser setosity. Gonostylus (Fig. 13 C) long, broadest medially, apically truncate; impressed; with normal setosity, a long apical tooth, and one dorsal, two subapical and two medial megasetae; megasetae longer than tooth, basally angulate; with one well-differentiated whiplash seta on ventromedial margin. Tegmen broader than long, strongly sclerotized, aedeagal teeth not detectable in the specimen studied. Aedeagal apodeme rather long and strong. BIN. Unknown.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA968FF92FF26F8C1984B0F83.taxon	discussion	Discussion. Trichocoelina planilobata sp. n. can be distinguished from all other congeneric species by its very broad intergonocoxal lobe of hypopygium. For further discussion, see under T. dispansa sp. n.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA968FF92FF26F8C1984B0F83.taxon	etymology	Etymology. The name is derived from the Latin words plana, flat, and lobus, lobe, referring to the broad intergonocoxal lobe of the hypopygium.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA967FF92FF26FC409B740BFD.taxon	description	Figs 14 A, 16 A	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA967FF92FF26FC409B740BFD.taxon	materials_examined	Material studied. Holotype male. FINLAND, Lkor (Lapponia kemensis orientalis) Salla, Värriö, Kuntasjoki (grid 7520406: 3610772), Malaise trap, 29. VII – 19. IX. 2013, J. Salmela (in MZH).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA967FF92FF26FC409B740BFD.taxon	description	Description. Male. Head. Face brown, antenna darker brown, maxillary palpus pale yellowish. Eye bridge 3 facets wide. Face with 20 fine setae. Clypeus with 2 setae. Maxillary palpus with 3 segments, lengths of segments poorly comparable in the specimen studied; 3 rd segment longer than 2 nd segment; 1 st segment with 1 seta, with a dorsal patch of sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 2.1 x as long as wide, the neck shorter than broad, the longest setae shorter than the width of flagellomere. Thorax. Brown, setae pale. Anterior pronotum with 7 setae. Proepisternum with 8 setae. Scutellum with 3 longer and some short and fine setae. Wing. Fumose. Length 2.0 mm. Width / length 0.40. Anal lobe moderate. Veins distinct. c / w 0.75. R 1 / R 0.90. stM and fork of M subequal in length. bM slightly longer than r-m, bM and r-m non-setose. Halter yellow. Legs. Yellow, coxal setae pale. Fore tibial organ forming a large distinct patch of pale setae in depression. Fore tibial spur longer than the tibial width. Abdomen. Pale brown, setae pale, moderately long. Hypopygium (Fig. 14 A). Brown, as abdomen. Intergonocoxal area long, with 2 small separate setose lobes. Gonocoxa narrow, longer than gonostylus, with short sparse setosity, medial margin basally smoothly curved. Gonostylus (Fig. 16 A) elongated, apically slightly narrowed, strongly impressed; with short setosity, a long and narrow apical tooth, and 5 medial megasetae; megasetae strong and slightly curved; with a long whiplash seta basad from the megasetae. Tegmen subtriangular, apically roundish, laterally straight, weakly sclerotized, with area of small aedeagal teeth. Aedeagal apodeme narrow and long. BIN. Unknown.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA967FF92FF26FC409B740BFD.taxon	discussion	Discussion. Trichocoelina quintula sp. n. resembles somewhat T. hiemalis (Mohrig & Mamaev) in having small intergonocoxal lobes and at most slightly curved, oblique gonostylar megasetae but differs in its much narrower gonostylus and in having only five, not more than ten, megasetae and in its narrower tegmen, see Figs 14 A, 16 A and fig. 4 in Mohrig et al. (1985). Trichocoelina quintula resembles also T. cochleata (Rübsaamen) but differs in having the gonostylus darker, not paler than gonocoxa, the gonostylus less strongly curved, the gonostylar megasetae shorter and stronger, smaller intergonocoxal lobes, and in lacking a semicircular rim on the tegmen.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA967FF92FF26FC409B740BFD.taxon	etymology	Etymology. The name is derived from the Latin word quintus, fifth, referring to the five megasetae of the gonostylus.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA965FF90FF26FF0D986E090E.taxon	description	Figs 14 B, 16 B	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA965FF90FF26FF0D986E090E.taxon	materials_examined	Material studied. Holotype male. FINLAND, Lkor (Lapponia kemensis orientalis), Salla, Värriö, Kuntasjoki (grid 7520406: 3610772), over a brook, Malaise trap, 29. VII – 19. IX. 2013, J. Salmela (in MZH). Paratypes. FINLAND, Ks (Regio kuusamoensis), Taivalkoski, Kylmäoja (grid 7275293: 3554865), by a brook, Malaise trap, 3. VII – 1. VIII. 2006, J. Salmela, 1 male (in PJSR); NORWAY, ‘ Svalbard, Lakselva’ [= Svalbard, Bjørnøya, river Lakselva], 74.4994 ° N, 18.9776 ° E, 15 m, 28. VII. 2009, T. Ekrem, 1 male (NTNU-VM 50613 and BOLD Sample ID BJ 263, in NTNU-VM).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA965FF90FF26FF0D986E090E.taxon	description	Description. Male. Head. Face brown, antenna darker brown, maxillary palpus pale yellowish. Eye bridge 2 – 3 facets wide. Face with 24 – 26 fine setae. Clypeus with 2 – 3 setae. Maxillary palpus with 3 segments, 1 st segment longer than 3 rd segment, 2 nd segment shortest; 1 st segment with 1 – 2 setae, with a dorsal patch of sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 1.95 – 2.0 x as long as wide, the neck shorter than broad, the longest setae slightly shorter than the width of flagellomere. Thorax. Brown, setae dark. Anterior pronotum with 4 – 7 setae. Proepisternum with 6 – 8 setae. Scutellum with 4 longer and some short and fine setae. Wing. Fumose. Length 2.0 – 2.2 mm. Width / length 0.40. Anal lobe weak. Veins distinct. c / w 0.65 – 0.75. R 1 / R 0.50 – 0.60. stM slightly shorter than fork of M. r-m longer than bM, bM non-setose, r-m with 3 – 4 setae. Halter yellow. Legs. Yellow, coxal setae pale. Fore tibial organ forming a large indistinct patch of fine setae in depression. Fore tibial spur longer than the tibial width. Abdomen. Pale brown, setae dark, moderately long. Hypopygium (Fig. 14 B). Brown, as abdomen. Intergonocoxal area long, with large conical setose lobe. Gonocoxa broad, as long as gonostylus, medial margin basally strongly curved, with short sparse setosity. Gonostylus (Fig. 16 B) broad, laterally roundish, apically narrowed, strongly impressed; with short setosity, a short curved apical tooth, and a dorsal and a ventral group of 6 – 7 megasetae medially; megasetae long and slender, directed slightly differently in both groups; with 1 – 2 weakly differentiated whiplash setae on ventromedial margin. Tegmen subconical and sclerotized, with area of minute aedeagal teeth. Aedeagal apodeme narrow and rather short. BIN. BOLD: ABA 5290.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA965FF90FF26FF0D986E090E.taxon	discussion	Discussion. By the subtriangular form of its gonostylus, Trichocoelina semisphaera sp. n. resembles T. hiemalis (Mohrig & Mamaev) but differs in having two groups of gonostylar megasetae, pointing in slightly different directions (all similarly directed in T. hiemalis), and one long intergonocoxal lobe of hypogium (two small ones in T. hiemalis) and in having the tegmen subconical (roundish in T. hiemalis).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA965FF90FF26FF0D986E090E.taxon	etymology	Etymology. The name is derived from the Latin words semi -, half, and sphaera, ball, referring to the roundish lateral margin of the gonostylus.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA965FF91FF26FAF59B5B0E3E.taxon	description	Figs 15 A, 16 C, 18 F	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA965FF91FF26FAF59B5B0E3E.taxon	materials_examined	Material studied. Holotype male. ITALY, Trentino-Alto Adige, Bolzano, Parco nazionale dello Stelvio (NP Stilfser Joch), Suldental E of Gomagoi, 46 ° 34 ’ 33 ’’ N, 10 ° 32 ’ 51 ’’ E, 1220 m, mountainous area, spruce forest, Malaise trap, 18 – 21. V. 2005, C. Lange & J. Ziegler (in ZMHB). Paratypes. ITALY, same data as holotype, 1 male (in SDEI); USA, Alaska, Kingfisher Creek, Grande Denali Lodge, 63.743024 ° N, 145.883416 ° W, 14. VI. 2016, J. Fitzgerald, 1 male (in MZH).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA965FF91FF26FAF59B5B0E3E.taxon	description	Description. Male. Head. Face and antenna concolorous brown, maxillary palpus pale yellowish. Eye bridge 3 facets wide. Face with 19 setae. Clypeus with 1 seta. Maxillary palpus with 3 segments, 3 rd segment longer than 1 st segment, 2 nd segment shortest; 1 st segment with 1 seta, with dorsal patch of sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 2.9 x as long as wide, the neck shorter than broad, the longest setae as long as the width of flagellomere. Thorax. Dark brown, setae pale. Anterior pronotum with 3 – 5 setae. Proepisternum with 6 – 9 setae. Scutellum with 4 longer and some short and fine setae. Wing. Fumose. Length 2.2 – 2.4 mm. Width / length 0.40. Anal lobe weak. Veins distinct. c / w 0.60. R 1 / R 0.65 – 0.80. stM shorter than fork of M. r-m as long as bM, both non-setose. Halter yellow. Legs. Yellow, coxal setae pale. Fore tibial organ forming a weakly demarcated patch. Fore tibial spur longer than the tibial width. Abdomen. Pale brown, setae pale, moderately long and fine. Hypopygium (Fig. 15 A). Brown, as abdomen. Intergonocoxal area long, with distinctly bifid setose lobe (Fig. 18 F). Gonocoxa moderately broad, longer than gonostylus, medial margin smoothly curved, with sparse setosity. Gonostylus (Fig. 16 C) long, narrow, strongly impressed; with dense setosity, a long apical tooth, and 5 – 6 megasetae at middle; megasetae long and slender; with 1 well-differentiated whiplash seta at ventromedial margin. Tegmen as long as broad, laterally and apically roundish, with a semicircular subapical sclerotized rim, otherwise weakly sclerotized, aedeagal teeth not detectable in the specimens studied. Aedeagal apodeme long. BIN. Unknown.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA965FF91FF26FAF59B5B0E3E.taxon	discussion	Discussion. By its strikingly dark setosity on the gonostylar apex, Trichocoelina semusta sp. n. resembles T. cochleata (Rübsaamen) but differs in having the apex less strongly curved, intergonocoxal lobes longer and closer to each other, the sclerotized rim of the tegmen broader and not protruding, and scapus and pedicellus concolorous brown with the flagellomeres, not yellow as in T. cochleata.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA965FF91FF26FAF59B5B0E3E.taxon	etymology	Etymology. The name is Latin, semusta, half-burned, referring to the dark setosity of the apex of the gonostylus of the species.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA964FF91FF26FD179BAB0846.taxon	description	BIN. Unknown.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA964FF91FF26FD179BAB0846.taxon	discussion	Discussion. Trichocoelina subcochleata (Komarov, 2009) was described based on one male from Russia (locus typicus: Republic of Altai, Kosh-Agachsky District, near Kosh-Agach, 50 ° 09 ’ N, 88 ° 19 ’ E). In its gonostylus with two groups of megasetae and its tegmen with sharp apicolateral corners it resembles most T. subpermutata (Mohrig & Mamaev) but differs in having two separate short interxoxal lobes instead of a long, apically bifid intergonocoxal lobe of hypopygium, and in its longer tegmen. By its tegmen and intergonocoxal lobes, T. subcochleata resembles also T. imitator and T. oricillifera, but these species have a large number of perpendicular megasetae in their gonostylus. We have not seen the holotype of T. subcochleata but suggest that the mention in the original description that the species lacks the whiplash setae from its gonostyli, is presumably a mistake, or that they have been broken off from the specimen studied.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA964FF91FF26FB1F9E550A2A.taxon	description	BIN. Unknown.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA964FF91FF26FB1F9E550A2A.taxon	discussion	Discussion. The species was described based on one male from Southern Yamal, Northern Russia (Mohrig et al. 1990). By its very large, apically bifid intergonocoxal lobe the species most resembles T. vitticollis, but differs in having its gonostylus broader, with megasetae pointing differently (subapical and subbasal megasetae perpendicular, medial oblique, whereas in T. vitticollis only a medial oblique group) and in having its tegmen with sharp apicolateral corners, absent in all other Trichocoelina. See also under T. tecta sp. n.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA964FF94FF26F9E298090ECE.taxon	description	Figs 15 B, 16 D	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA964FF94FF26F9E298090ECE.taxon	materials_examined	Material studied. Holotype male. USA, Alaska, 11 mi S Anderson Jct, Rte 3, mi 270, alder-poplar-spruce, Malaise trap, 23. VI – 11. VIII. 1984, S. & J. Peck (in MZH). Paratypes. USA, same data as holotype, 1 male (MZH); CANADA, Nunavut, Qikitaaluk, Western Bylot Island, 73.157 ° N, 79.950 ° W, 12 m, 3. VIII. 2007, L. Jolicoeur & L. McKinnon, 1 male (BOLD Sample ID 08 WOLVES- 01202, in CBG); Yukon, North Fork Crossing, Mi 42 Peel Plt. Road, 3500 ’, 24. VI. 1962, P. J. Skitsko, 1 male (in CNC); RUSSIA, Krasnoyarsk region, Taimyr Peninsula, river Zakharova, Rassokha, 72.70 ° N, 101.08 ° E, pan trap, 1 – 10. VII. 2011, A. Barkalov, 1 male (in ISEA); Yamalo-Nenets Autonomous Okrug, near Seyakh, 70.7788 ° N, 72.0750 ° E, nival meadow, Malaise trap, 13 – 29. VII. 2014, N. Zubryi, 5 males (1 in MZH, 1 in NHMO, 2 in SDEI, 1 in ZIN).	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA964FF94FF26F9E298090ECE.taxon	description	Description. Male. Head. Face and antenna concolorous brown, maxillary palpus pale yellowish. Eye bridge 2 – 3 facets wide. Face with 13 – 20 fine setae. Clypeus with 2 – 5 setae. Maxillary palpus with 3 segments, 1 st segment longer or shorter than 3 rd segment, 2 nd segment shortest; 1 st segment with 1 – 2 setae, with a dorsal patch of sensilla; surface of antennal flagellomeres smooth, body of 4 th antennal flagellomere 1.7 – 2.25 x as long as wide, the neck shorter than broad, the longest setae slightly shorter than the width of flagellomere. Thorax. Brown, setae pale. Anterior pronotum with 2 – 6 setae. Proepisternum with 3 – 10 setae. Scutellum with 4 longer and some short and fine setae. Wing. Fumose. Length 1.9 – 2.1 mm. Width / length 0.40 – 0.45. Anal lobe weak. Veins distinct. c / w 0.60 – 0.65. R 1 / R 0.60 – 0.85. stM and fork of M subequal in length. r-m and bM of variable length, bM non-setose, r-m with 2 – 4 setae or non-setose. Halter yellow. Legs. Yellow, coxal setae pale. Fore tibial organ forming a small patch of dark strong setae in depression. Fore tibial spur slightly longer than the tibial width. Abdomen. Pale brown, setae pale, moderately long. Hypopygium (Fig. 15 B). Brown, as abdomen. Intergonocoxal area long, basally with strongly sclerotized medial stripe, with large subtriangular lobe with setae at margin. Gonocoxa broad, longer than gonostylus, medial margin basally strongly curved, with long sparse setosity. Gonostylus (Fig. 16 D) long, apically narrowed, strongly impressed; with short setosity, with a long apical tooth, with 6 – 7 medial megasetae in dorsal and ventral groups, megasetae long and slender, directed slightly differently in both groups; with 1 well-differentiated whiplash seta on ventromedial margin. Tegmen conical, apically and laterally straight or slightly curved, weakly sclerotized, with an indistinct area of minute aedeagal teeth. Aedeagal apodeme long. BIN. BOLD: AAM 9263.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA964FF94FF26F9E298090ECE.taxon	discussion	Discussion. In having a long intergonocoxal lobe of hypopygium and 6 – 7 medial megasetae on the gonostylus, Trichocoelina tecta sp. n. resembles T. vitticollis (Rübsaamen) but differs in the intergonocoxal lobe being setose only on its margins, not on its ventral side, in its conical tegmen, not apically roundish, and in having the gonostylar megasetae in groups pointed at two directions, not all in a row pointing in one direction.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA964FF94FF26F9E298090ECE.taxon	etymology	Etymology. The name is Latin, tecta, covered, referring to the ventromedial part of the gonostylus covering almost completely the medial megasetae.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA961FF8BFF26FC879EF30F39.taxon	synonymic_list	Synonyms: = glacialis (Lundbeck, 1898) [as Sciara; preocc., not Sciara glacialis Rübsaamen, 1898]; = permutata (Lundbeck, 1900) [as Sciara; new name for Sciara glacialis Lundbeck, 1898].	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA961FF8BFF26FC879EF30F39.taxon	discussion	Discussion. The species was described from one female from Novaya Zemlya by Holmgren (1883) and later described again based on some males and females from Greenland as Sciara glacialis (Lundbeck 1898) and from Spitsbergen as Sciara permutata (Lundbeck 1900). The species was redescribed with a list of literature, synonyms and combinations by Menzel & Mohrig (2000). Trichocoelina vitticollis (Holmgren) can be distinguished from all other species of the genus by its very large, setose intergonocoxal lobe of hypopygium and in having the gonostylus with a group of slender, oblique medial megasetae. See also under Trichocoelina tecta sp. n.	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA97CFF89FF26FF0D99EC0BDE.taxon	description	= debilis (Winnertz, 1867) — PAL	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA97CFF89FF26FF0D99EC0BDE.taxon	description	= humilis (Winnertz, 1867) — PAL	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
A703878CA97BFF8FFF26FC1898D00CD6.taxon	distribution	absidata Vilkamaa & Menzel sp. n. — PAL aemula Vilkamaa & Menzel sp. n. — PAL biplex Vilkamaa & Menzel sp. n. — NEA brevicubitalis (Lengersdorf, 1926) comb. n. — PAL chentejensis (Menzel, 1992) comb. n. in Menzel (1992 b) — PAL cochleata (Rübsaamen, 1898) comb. n. — NEA / PAL = haemorrhoidalis (Lundbeck, 1898) — NEA dicksoni Vilkamaa & Menzel sp. n. — PAL dispansa Vilkamaa & Menzel sp. n. — PAL dividua Vilkamaa & Menzel sp. n. — NEA hians Vilkamaa & Menzel sp. n. — NEA hiemalis (Mohrig & Mamaev, 1985) comb. n. in Mohrig et al. (1985 b) — PAL imitator Vilkamaa & Menzel sp. n. — NEA incrassata Vilkamaa & Menzel sp. n. — NEA ithyspina Vilkamaa & Menzel sp. n. — PAL janetscheki (Lengersdorf, 1953) comb. n. — NEA / PAL jukkai Vilkamaa & Menzel sp. n. — PAL magnifica Vilkamaa & Menzel sp. n. — NEA nefrens Vilkamaa & Menzel sp. n. — PAL obesula Vilkamaa & Menzel sp. n. — PAL olschwangi (Mohrig & Mamaev, 1983) comb. n. in Mohrig et al. (1983 a) — NEA / PAL oricillifera Vilkamaa & Menzel sp. n. — PAL planilobata Vilkamaa & Menzel sp. n. — PAL quintula Vilkamaa & Menzel sp. n. — PAL semisphaera Vilkamaa & Menzel sp. n. — PAL semusta Vilkamaa & Menzel sp. n. — NEA / PAL subcochleata (Komarov, 2009) comb. n. — PAL subpermutata (Mohrig & Mamaev, 1990) comb. n. in Mohrig et al. (1990) — PAL tecta Vilkamaa & Menzel sp. n. — NEA / PAL vitticollis (Holmgren, 1883) comb. n. — NEA / PAL = glacialis (Lundbeck, 1898) [preocc.] — NEA = permutata (Lundbeck, 1900) — NEA / PAL	en	Vilkamaa, Pekka, Menzel, Frank (2019): Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species. Zootaxa 4665 (1): 1-67, DOI: 10.11646/zootaxa.4665.1.1
