taxonID	type	description	language	source
A6418788BC677057D098FCC7FDD3DCFB.taxon	description	(Figs. 2 A – B, 3 A – E)	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC677057D098FCC7FDD3DCFB.taxon	materials_examined	Material examined. Iran, Sistan va Baluchestan Province, IR 5 - 2011, Gerdoig village N of Chabahar, 5 km of Koch village, pool, 25 º 46.084 N, 60 º 54.968 E, 13. vii. 2011 5 / 11 / 5 (1 / 2 / 1 mounted). Morphology. Male — Idiosoma L / W 2260 / 2060; integument papillose; postocularia particularly distant from preocularia, together with the lateral eyes forming the edges of a longitudinally rectangular field (Fig. 2 B); coxal field: L 866, W 1700, medial margin of Cx-III + IV concave, Cx-IV bearing one seta, posteromedial corner of Cx- IV extended with cleaver-shaped posterolaterally directed secondary sclerotization; genital field L / W 441 / 413, posteriorly extending beyond posterior margin of Cx-IV, deeply indented anteriorly, extending over the anterior third of the plate, gonopore long, extending over the posterior third of the plate (Fig. 2 A); Ac small and numerous; ejaculatory complex L 375. Gnathosoma L 881, rostrum L 500, relatively long and curved (Fig. 3 C); chelicera L 1100; palp (Fig. 3 D): total L 773; dL: P- 1, 206; P- 2, 181; P- 3, 238; P- 4, 93; P- 5, 55; palp stout, P- 1 bearing one dorsodistal seta, P- 2 ventral margin slightly concave or straight. Legs: dL of I-L- 2 - 6: 194, 138, 178, 194, 222; dL of IV-L: 246, 281, 222, 503, 441, 231. Female (n = 2) — Idiosoma L / W 1810 / 1625; coxal field: L 770 – 825, W 1590 – 1720; genital field (Fig. 3 A) L / W 256 – 269 / 334 – 373, deeply indented anteriorly extending anterior half of the plate, the “ modified acetabulum ” not anterior to but between posterior setal fields; gnathosoma (Fig. 3 F) L 640 – 680, rostrum L 330 – 370; chelicera L 1000; palp (Fig. 3 E): total L 704 – 707; dL: P- 1, 184 – 194; P- 2, 150; P- 3, 219 – 225; P- 4, 91 – 95; P- 5, 47 – 56; palp stout, P- 4 more slender than in male (age dependant?). Legs: dL of I-L- 2 - 6: 181, 113, 141, 156, 185; dL of IV-L: 231 – 234, 256 – 281, 200 – 223, 419 – 422, 350 – 363, 209 – 225. Deutonymph (collected together with adults of Hydrachna cf. vaillanti at IR 5 - 2011 and suspected to represent this species). — Idiosoma L / W 1450 / 1280; coxal field: L 556, W 1025; individual provisional genital plate L / W 206 – 212 / 113 – 119; gnathosoma L 570, rostrum L 300; chelicera L 750; palp: total L 534; dL: P- 1, 141; P- 2, 128; P- 3, 153; P- 4, 71; P- 5, 41. Legs: dL of I-L- 2 - 6: 131, 94, 125, 136, 162; dL of IV-L: 163, 169, 153, 300, 288, 188.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC677057D098FCC7FDD3DCFB.taxon	discussion	Remarks. The population from Sistan va Baluchestan belongs to the Hydrachna processifera group of species which includes H. processifera Koenike, 1903 and several little documented species (e. g. H. danubialis Láska, 1964, H. vaillanti and H. sepasgozariani Bader, 1981). Due to the similar shape of the male genital field (minor in dimensions, anterior margin more elongated, extending over 1 / 3 of the genital field) the specimens from Sistan va Baluchestan most resemble Hydrachna vaillanti, a species known from a single male from Algeria (central Sahara, K. Viets 1951). However, as mentioned by Davids et al. (2005), these features in the type specimen of H. vaillanti might be an individual aberration. The morphological homogenity of specimens from Sistan va Baluchestan suggests that they represent a distinct taxon. Without information on the population variability of H. vaillanti, and in view of the large geographical distance of our records, this assignment is based mainly on non-identity with alternative species. Habitat. Standing water (Fig. 8 A).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC64705AD098FBBBFB2EDCC3.taxon	description	(Figs. 4, 5 A – F)	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC64705AD098FBBBFB2EDCC3.taxon	materials_examined	Material examined. Holotype male (idiosoma in fluid, gnathosoma, palp and I / IV-legs dissected and slide mounted), Iran, Sistan va Baluchestan Province, IR 5 - 2011, Gerdoig village N of Chabahar, 5 km of Koch village, pool, 25 º 46.084 N, 60 º 54.968 E, 13. vii. 2011 (RMNH). Paratypes: 5 / 0 / 0, same data as holotype (RMNH)	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC64705AD098FBBBFB2EDCC3.taxon	diagnosis	Diagnosis. Idiosoma dorsally and ventrally sclerotized except for a narrow strip of papillose integument extending around coxal and genital field, and two median strips extending posteriorly from the posteromedial corner of Cx-IV to about two-third of the postgenital length; gonopore surrounded by a row of small setae and located on a raised high ridge.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC64705AD098FBBBFB2EDCC3.taxon	description	Description. Male (holotype, in parentheses measurements of paratype specimens) — Idiosoma L / W 1280 (1040 – 1410) / 1100 (950 – 1250); idiosoma dorsally and ventrally sclerotized except for a narrow strip of papillose integument extending around coxal and genital field, and two median strips extending posteriorly from the posteromedial corner of Cx-IV to about two-third of the postgenital length (Fig. 4); coxal field: L 581, W 925; Cx-IV bearing one seta, posteromedial corner extending, with a cleaver-shaped extension of secondary sclerotization. Genital field L / W 252 / 216, heart-shaped, posteriorly extending beyond posterior margin of Cx-IV, anteriorly with a small indentation, gonopore surrounded by a row of small setae and located on a raised high ridge as shown in Figs. 5 A – B; genital field hinged at anterior end but movable posteriorly, this movement leads to apparent differences in placement of the gonopore (see Figs. 5 A – B); Ac small, but one larger Ac on each side located in the posterolateral part of the acetabular field. Gnathosoma stout, L 491, rostrum L 228, shorter than base, bent ventrally with a distinct kink (Fig. 5 C); chelicera L 572; palp (Figs. 5 D – E): total L 584; dL: P- 1, 166; P- 2, 138; P- 3, 163; P- 4, 81; P- 5, 36; P- 1 bearing one dorsodistal seta, P- 2 strongly convex dorsally, ventral margin concave. Legs: dL of I-L- 2 - 6 (Fig. 5 F): 144, 98, 113, 141, 168; dL of IV-L: 193, 175, 140, 244, 263, 242.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC64705AD098FBBBFB2EDCC3.taxon	etymology	Etymology. The species is named for its occurrence in Sistan va Baluchestan Province.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC64705AD098FBBBFB2EDCC3.taxon	discussion	Remarks. Due to the characteristic shape of the genital field, with its raised ridge surrounding the gonopore, and an extensive dorsal sclerotization, Hydrachna sistanica sp. nov. is most similar to H. mysorensis Cook, 1967, a species described from a pond in Mysore State in India (Cook 1967). Hydrachna mysorensis can be distinguished from the new species by the lack of ventral secondary sclerotization (see Cook 1967, Fig. 25). Furthermore, our species does not show the extreme humped condition of the dorsal side of P- 2 which is typical for H. mysorensis. Habitat. Standing water (Fig. 8 A).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC64705AD098FBBBFB2EDCC3.taxon	distribution	Distribution. Iran; only known from the type locality in Sistan va Baluchestan Province.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC697059D098FF4DFDD8D8C7.taxon	description	(Fig. 6 A – C)	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC697059D098FF4DFDD8D8C7.taxon	materials_examined	Material examined. Iran, Sistan va Baluchestan Province, IR 5 - 2011, Gerdoig village N of Chabahar, 5 km of Koch village, pool, 25 º 46.084 N, 60 º 54.968 E, 13. vii. 2011 0 / 6 / 1 (0 / 1 / 1 mounted). Morphology. Female — Idiosoma L 1809 – 2440; paired frontal plates (Fig. 6 C), L / W 500 / 288, separated from each other by membranous integument; coxal field: L 619, W 1187, medial margin of Cx-III + IV concave, Cx-IV bearing one seta, posteromedial corner of Cx-IV extended with cleaver-shaped posterolaterally directed secondary sclerotization; genital field (Fig. 6 A) L / W 197 / 263. Gnathosomal rostrum L 356, relatively long and curved; chelicera L 744; palp (Fig. 6 B): total L 663; dL (in parentheses dL / H ratio): P- 1, 181 (1.0); P- 2, 150 (1.4); P- 3, 213 (3.1); P- 4, 88 (2.1); P- 5, 31 (1.7); palp slender, P- 1 bearing one dorsodistal seta, P- 2 ventral margin slightly concave or straight. Legs: dL of I-L: 70, 175, 105, 119, 144, 172; dL of IV-L: 216, 216, 186, 297, 283, 222. Deutonymph (collected together with adults of Hydrachna globosa lacerata at IR 5 - 2011 and suspected to represent this species). — Idiosoma L / W 1138 / 925; frontal plate L / W 241 / 122; coxal field L 384; individual provisional genital plate L / W 128 / 84; gnathosomal rostrum L 167; chelicera L 441; palp: total L; dL: P- 1, 102; P- 2, 92; P- 3, 108; P- 4, 55; P- 5, 30. Legs: dL of I-L- 2 - 6: 98, 62, 72, 94, 126; dL of IV-L: 125, 117, 97, 166, 184, 175.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC697059D098FF4DFDD8D8C7.taxon	discussion	Remarks. The third Hydrachna species collected from a pool near Gerdoig village resembles Hydrachna globosa (De Geer, 1778). Our specimens are assigned to the Oriental subspecies, H. globosa lacerata Lundblad, 1969, known only from a female from Burma (Lundblad 1969). Compared with the stem species, this subspecies has more broad and relatively short frontal plates, a more slender P- 3 (L / H ratio 3.1) and extended posteromedial extensions of the fourth coxal plates. Additional information on the population variability of H. globosa lacerata, and the description of the male, are necessary to resolve the taxonomy of this taxon. Habitat. Standing waters (Fig. 8 A).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC69705ED098FB70FDCADA89.taxon	description	(Figs. 7 A – C)	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC69705ED098FB70FDCADA89.taxon	materials_examined	Material examined. Iran, Sistan va Baluchestan Province, IR 5 - 2011, Gerdoig village N of Chabahar, 5 km of Koch village, pool, 25 º 46.084 N, 60 º 54.968 E, 13. vii. 2011 25 / 2 (1 / 1 / 0 mounted). Morphology. Male — Idiosoma L / W 1270 / 1080; integument without sclerotized muscle insertions; Coxal field: medial subcutaneous extensions of Cx-I + II in their distal parts bent caudally (Fig. 7 A); L Cx-I + II 222, Cx- III + IV 381. Genital field: L / W 188 / 294, with triangular genital plates, maximum W at posterior margin, medial setae on genital plates regularly arranged along medial margin, a few longer setae concentrated near the posteromedial margin. Palp (Fig. 7 C) stout: total L 332; dL (in parentheses dL / H ratio): P- 1, 39 (0.63); P- 2, 85 (1.32); P- 3, 60 (0.95); P- 4, 117 (2.66); P- 5, 31 (2.0); P- 2 with three to five long and strong medial setae; L chelicera 366, basal segment 253, claw 134, L ratio basal segment / claw 1.89. Legs: dL of IV-L: 120, 111, 175, 231, 239, 206; swimming setae numbers: II-L- 5, 10; III-L- 4, 12; III-L- 5, 15; IV-L- 3, 1; IV-L- 4, 17; IV-L- 5, 13. Female — Idiosoma L / W 1450 / 1200; coxal field: Cx-I + II 259, Cx-III + IV 415. Genital field L / W 222 / 334. Palp: total L 416; dL (in parentheses dL / H ratio): P- 1, 52 (0.68); P- 2, 101 (1.34); P- 3, 78 (0.98); P- 4, 149 (2.85); P- 5, 36 (1.8); L chelicera (Fig. 7 B) 459, basal segment 316, claw 170, L ratio basal segment / claw 1.86. Legs: dL of IV-L: 153, 150, 222, 303, 303, 234; swimming setae numbers: II-L- 5, 10; III-L- 4, 8; III-L- 5, 15; IV-L- 3, 1; IV-L- 4, 16; IV-L- 5, 13 - 15.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC69705ED098FB70FDCADA89.taxon	discussion	Remarks. The specimens from Sistan va Baluchestan are provisionally assigned to the Oriental species Diplodontus silvestrii. However, they resemble both Diplodontus silvestrii and the Afrotropical D. schaubi (Koenike, 1893). According to Gerecke (2004 a), with the present state of knowledge, D. silvestrii differs from D. schaubi in a shorter cheliceral claw (L ratio basal segment / claw> 1.9, D. schaubi <1.9). The specimens from Sistan va Baluchestan agree in most dimensions with the measurements of D. silvestrii given by Cook (1967) for a single male specimen from India, as well with D. schaubi, a species widespread in standing waters in the Afrotropical region (Gerecke 2004 a). Our specimens differ from both D. schaubi and D. silvestrii in a stouter P- 4 (L / H 2.7 – 2.9, in the compared species 2.9 – 3.3). As Gerecke (2004 a) noted, the diagnostic features and taxonomic relationship of the Asian species require further revision. This should include the relationship with species from other zoogeographical regions, with emphasis to populations from the transitional area, what probably will require applying molecular techniques. Habitat. Standing water (Fig. 8 A)	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC6E705ED098F9ABFDB4DC51.taxon	materials_examined	Material examined. Iran, Hormozgan Province, Bandar Abbass: IR 18 - 2011, Taleguerdoo village, Poshtekeno spring, upper part of stream, 27 º 48.783 N, 56 º 24.459 E, 836 m asl, 18. vii. 2011 0 / 1 / 0; IR 22 - 2011, Rudan city, 5 km after Rudan, Rudan stream, 27 º 28.748 N, 57 º 15.648 E, 210 m asl, 21. vii. 2011 1 / 0 / 0. Habitat. Sandy / bouldery streams, often with strong exposure to sunlight.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC6E705ED098F9ABFDB4DC51.taxon	distribution	Distribution. India, Iran, Oman.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC6C705CD098FEC5FD1ADCFF.taxon	description	(Figs. 9 A – H, 10 A – C)	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC6C705CD098FEC5FD1ADCFF.taxon	materials_examined	Material examined. Holotype male, dissected and slide mounted, Iran, Hormozgan Province, IR 16 - 2011, shallow stream N of Bandar Abbass, 27 º 27.622 N, 56 º 20.590 E, 130 m asl, 16. vii. 2011 (RMNH). Paratypes: 6 / 14 / 0 (0 / 1 / 0 mounted), same data as holotype (RMNH). Further records. Iran, Hormozgan Province, IR 19 - 2011, Banglayan village N of Bandar Abbass, stream, 27 º 45.695 N, 56 º 32.402 E, 577 m asl, 18. vii. 2011 0 / 1 / 0; IR 21 - 2011, Rudan city, 5 km after Rudan, Rudan stream, 27 º 28.226 N, 57 º 15.153 E, 200 m asl, 21. vii. 2011 2 / 0 / 0 (mounted); IR 22 - 2011, 5 km N of Rudan city, Rudan stream, 27 º 28.748 N, 57 º 15.648 E, 210 m asl, 21. vii. 2011 1 / 0 / 0.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC6C705CD098FEC5FD1ADCFF.taxon	diagnosis	Diagnosis. Setae of D 2 and D 3 long; posterior dorsal plate moderately large (L / W <115 / 100, in both sexes); postgenital sclerite in male with moderately developed secondary sclerotization; postgenital sclerite in female smaller and not extending anteriorly as in male; pregenital sclerite of female moderate, bowed and not extending anteriorly.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC6C705CD098FEC5FD1ADCFF.taxon	description	Description. Male (holotype, in parentheses measurements of paratype specimens from IR 16 - 2011, n = 4; in square brackets specimens from IR 21 - 2011, n = 2) — Idiosoma soft, L / W 725 (690 – 741) [594 – 681] / 556 (500 – 572) [403 – 438]. Dorsum with one large posterior platelet (Fig. 9 A), L / W 100 (83 – 100) [109] / 91 (75 – 91) [99]; setae of glandularia D 2 and D 3 (sensu Wiles 1997 a) long. Coxal field L 419 [385 – 394], Cx-III W 422 [347 – 353]; Cx-I separated medially; Cx-III medially with a widely rounded corner anteriorly of genital field. Genital field (Figs. 9 D – E) L / W 103 (109 – 117) [102 – 108] / 89 (83 – 88) [83 – 84], with three pairs of elongated acetabula; pregenital sclerite moderate in size, bowed; posteriorly of genital field a somewhat triangular postgenital sclerite, L / W 93 (84 – 89) [87] / 47 (36 – 38) [39 – 49]. Ejaculatory complex L 186. Palp (Fig. 9 F): total L 386 [317 – 336]; dL: P- 1, 25 [22 – 24]; P- 2, 122 [102 – 106]; P- 3, 71 [59 – 65]; P- 4, 137 [109 – 115]; P- 5, 31 [25 – 26]; L P- 2 / P- 4 ratio, 0.89 [0.92 – 0.94]; P- 2 with a long ventral seta, L 72 [65 – 66]; ventral margin of P- 2 distally with small denticles; P- 3 with a long medial seta which extends beyond posterior margin of P- 4; P- 4 with a short setal tubercle, ventral margin with four setae of various length, the seta on the setal tubercle being the longest; gnathosoma vL 159 [150 – 151]; chelicera total L 309 [262 – 263], claw L 94 [78]. Legs: L of I-L- 2 - 6: [75 – 78], [84 – 87], [105 – 108], [127 – 133], [125 – 134]; I-L- 4 with four heavy distal setae, two longer, two shorter (Fig. 9 C); L of IV-L- 3 - 6: 124 [112 – 116], 208 [172 – 175], 252 [200 – 211], 268 [212 – 216]; IV-L- 4 with five heavy distal setae, some of these serrated; IV-L- 5 with three heavy distal setae, at least the longest of these serrated; IV-L- 6 with a long subterminal seta, L 92 [94 – 95]; claws of legs I – III with a comb and a clawlet. Female (paratype, in parentheses some measurements of paratype specimens from IR 16 - 2011, n = 4) — Idiosoma soft, L / W 813 (694 – 969) / 594 (431 – 687). Dorsum with one large posterior platelet, L / W 110 (88 – 112) / 98 (75 – 92); setae of glandularia D 2 and D 3 (sensu Wiles 1997 a) long. Coxal field L 438, Cx-III W 467; Cx-I separated medially; Cx-III with a blunt corner medially, anteriorly of genital field. Genital field L / W 155 (150 – 153) / 113 (109 – 114), with three pairs of elongated acetabula; pregenital sclerite as in male, but smaller (Fig. 10 B); postgenital sclerite in female smaller and not extending anteriorly as in male; maximum egg diameter L 144. Palp: total L 426; dL: P- 1, 31; P- 2, 134; P- 3, 75; P- 4, 152; P- 5, 34; L P- 2 / P- 4 ratio, 0.88; P- 2 ventral seta L 77; palp as in male; gnathosoma vL 180; chelicera total L 353, claw L 113. Legs: as in male; L of I-L- 3 - 6: 94, 122, 141, 148; L of IV-L- 2 - 6: 123, 140, 228, 270, 284; subterminal seta of IV-L- 6 L 91.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC6C705CD098FEC5FD1ADCFF.taxon	etymology	Etymology. The species is named for its occurrence in Hormozgan Province.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC6C705CD098FEC5FD1ADCFF.taxon	discussion	Remarks. Due to the similar general shape of the coxal field, palp and legs, as well the presence of long setae of D 2 and D 3, the specimens from Iran closely resemble Nilotonia longiseta Smit & Pešić, 2010 from northern Oman (Smit & Pešić 2010). The species from Oman differs from Nilotonia hormozgana sp. nov. in much larger dimensions of the dorsal plate (L / W 160 – 218 / 112 – 148 in male, 150 – 196 / 106 – 128 in female) and postgenital sclerite (L / W 106 / 104 in the holotype male) in both sexes, and in the shape of the female pregenital sclerite which is strongly extended anteriorly (see Smit & Pešić 2010). Habitat. Sandy / bouldery streams, often with strong exposure to sunlight (Fig. 44 B, D).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC6C705CD098FEC5FD1ADCFF.taxon	distribution	Distribution. Iran, Hormozgan Province.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC737041D098FEFFFBB9D839.taxon	description	(Figs. 11 A – C, 12 A – B)	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC737041D098FEFFFBB9D839.taxon	materials_examined	Material examined. Holotype female, dissected and slide mounted, Iran, Hormozgan Province, IR 19 - 2011, Banglayan village N of Bandar Abbass, unnamed stream, 27 º 45.695 N, 56 º 32.402 E, 577 m asl, 18. vii. 2011 (RMNH).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC737041D098FEFFFBB9D839.taxon	diagnosis	Diagnosis. Female (male unknown). Cxgl- 4 anterior to Cxgl- 2 and located adjacent to Leg-II socket; Cx-IV extending strongly posteriorly to genital field; P- 2 long (L ratio P- 2 / P- 4 1.65) and with small ventral denticles distally to strongly pronounced ventral projection, P- 3 with serrulate margin distally from small ventral projection; gnathosoma with long rostrum; cheliceral claw relatively short (basal segment / claw ratio 6.6); I-L- 6 relatively stout (L / H ratio 2.5).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC737041D098FEFFFBB9D839.taxon	description	Description. Female — Idiosoma (ventral view: Fig. 11 B) L 747, W 525; dorsal shield (Fig. 11 A) L 653, W 412, L / W ratio 1.59; dorsal plate 625; shoulder plate L 175 – 178, W 56 – 58, L / W ratio 3.1; frontal plate L 119 – 122, W 53 – 55, L / W ratio 2.2 – 2.3; shoulder / frontal plate L ratio 1.46 – 1.47. Gnathosomal bay L 131, Cx-I total L 250, Cx-I mL 119, Cx-II + III mL 61; ratio Cx-I L / Cx-II + III mL 4.1; Cx-I mL / Cx-II + III mL 1.95; Cx- IV extending posteriorly to genital field; Cxgl- 4 anteriorly to Cxgl- 2 and located adjacent to Leg-II socket; genital field L / W 159 / 145, L / W ratio 1.1; distance genital field-excretory pore 236, genital field-caudal idiosoma margin 272. Gnathosoma (Fig. 11 C) with long rostrum, vL 373; chelicera L 409, basal segment L 365, claw L 57, basal segment / claw ratio L 6.6; palp (Fig. 12 A) total L 384, dL: P- 1, 57; P- 2, 155; P- 3, 59; P- 4, 94; P- 5, 19; P- 2 / P- 4 ratio, 1.56; P- 2 with small ventral denticles distally to ventral projection bearing long seta, P- 3 with serrulate margin distally from small pointed ventral projection bearing seta; dL of I-L- 4 - 6 (Fig. 12 B): 112, 109, 112; I-L- 6 L / H ratio 2.5.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC737041D098FEFFFBB9D839.taxon	etymology	Etymology. Named after Dr Mahdieh Asadi (Kerman, Iran) in appreciation of her acarological studies in Iran.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC737041D098FEFFFBB9D839.taxon	discussion	Remarks. In the shape of the palp (P- 2 with ventral denticles distally to ventral projection, P- 3 with serrulate margin distally from small pointed ventral projection), Cxgl- 4 anterior to Cxgl- 2 and located adjacent to Leg-II socket, gnathosoma with long rostrum and Cx-IV greatly extended posterior to the genital field, Torrenticola asadiae sp. nov. is most similar to T. omanensis Smit & Pešić, 2010, a species known only from males from northern Oman (Smit & Pešić 2010). The latter species (in parentheses data taken from Smit & Pešić 2010) differs from T. asadiae sp. nov. in a longer P- 2 (L ratio P- 2 / P- 4 2.5) with much more developed ventral denticles, and in a slightly pronounced ventral projection, more elongated gnathosomal rostrum and a longer chelicera with a relatively shorter cheliceral claw (basal segment / claw ratio 12.0). Habitat. Sandy / bouldery stream, with strong exposure to sunlight and without vegetation (Fig. 44 C).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC737041D098FEFFFBB9D839.taxon	distribution	Distribution. Iran; only known from the type locality in Hormozgan Province.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC717047D098FC56FD1BDA6A.taxon	description	(Figs. 13 A – F, 14 A – B)	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC717047D098FC56FD1BDA6A.taxon	materials_examined	Material examined. Holotype male, dissected and slide mounted, Iran, Hormozgan Province, IR 19 - 2011, Banglayan village N of Bandar Abbass, unnamed stream, 27 º 45.695 N, 56 º 32.402 E, 577 m asl, 18. vii. 2011 (RMNH). Paratypes: 9 / 11 / 0 (1 / 2 / 0 mounted), same data as holotype (RMNH).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC717047D098FC56FD1BDA6A.taxon	diagnosis	Diagnosis. Cx-IV extending posterior to genital field; excretory pore posterior to line of primary sclerotization, Vgl- 2 a little posterior to excretory pore; gnathosoma with a long rostrum, ventral margin curved; P- 2 longer than P- 4 (ratio P- 2 / P- 4 1.18 - 1.22), ventral projection of P- 2 strongly pronounced, but not curved distally, distal margins of P- 2 and P- 3 without denticles, ventral protuberance of P- 4 forming an unique tip bearing one long, and three short hairs; postgenital area large. Males: Median suture line of Cx-II + III short; genital field roundish.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC717047D098FC56FD1BDA6A.taxon	description	Description. Male (holotype, in parentheses measurements of paratype, n = 1) — Idiosoma (ventral view: Fig. 13 B) L 738 (753), W 581 (538); dorsal shield (Fig. 13 A) L 634 (638), W 450 (438), L / W ratio 1.41 (1.46); dorsal plate 660 (606); shoulder plate L 177 – 181 (173 – 177), W 65 – 66 (56 – 58), L / W ratio 2.72 – 2.74 (3.05 – 3.09); frontal plate L 128 (125 – 130), W 55 – 56 (51), L / W ratio 2.29 – 2.33 (2.45 – 2.55); shoulder / frontal plate L ratio 1.38 – 1.41 (1.33 – 1.42). Gnathosomal bay L 139 (150), Cx-I total L 272 (290), Cx-I mL 132 (139), Cx-II + III mL 83 (84); ratio Cx-I L / Cx-II + III mL 3.28 (3.45); Cx-I mL / Cx-II + III mL 1.59 (1.65). Genital field L / W 131 (133) / 121 (125), L / W ratio 1.08 (1.06); ejaculatory complex normal in shape (Fig. 13 D), L 180 (197); distance genital field-excretory pore 163 (169), genital field-caudal idiosoma margin 238 (244). Gnathosoma (Fig. 13 C) ventral L 318 (321); chelicera total L 390 (373); palp (Fig. 13 E) total L 341 (346), dL: P- 1, 35 (37); P- 2, 122 (123); P- 3, 65 (63); P- 4, 100 (103); P- 5, 19 (20); P- 2 / P- 4 ratio 1.22 (1.19); dL of I-L- 4 - 6: 109, 109 (113), 103 (106); I-L- 6 L / H ratio, 2.9 (3.0). Female (n = 2) — Idiosoma (ventral view: Fig. 14 B) L 797 – 825, W 575 – 587; dorsal shield (Fig. 14 A) L 696 – 716, W 453 – 478, L / W ratio 1.50 – 1.54; dorsal plate 663 – 675; shoulder plate L 188 – 203, W 56 – 63, L / W ratio 3.17 – 3.45; frontal plate L 130 – 133, W 53 – 56, L / W ratio 2.32 – 2.47; shoulder / frontal plate L ratio 1.47 – 1.53. Gnathosomal bay L 173 – 186, Cx-I total L 304 – 316, Cx-I mL 128 – 130, Cx-II + III mL 43 – 50; ratio Cx-I L / Cx-II + III mL 6.3 – 7.07; Cx-I mL / Cx-II + III mL 2.56 – 3.02. Genital field L / W 156 – 166 / 153 – 156, L / W ratio 1.0 – 1.08; distance genital field-excretory pore 194, genital field-caudal idiosoma margin 278. Gnathosoma ventral L 352 – 357; chelicera total L 423 – 425; palp (Fig. 13 F) total L 368 – 378, dL: P- 1, 38 – 39; P- 2, 131 – 134; P- 3, 67 – 71; P- 4, 111 – 114; P- 5, 20 – 21; P- 2 / P- 4 ratio, 1.18; palp as in male; dL of I-L- 4 - 6: 115 – 119, 114 – 118, 106 – 107; I-L- 6 L / H ratio 2.9 – 3.0.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC717047D098FC56FD1BDA6A.taxon	etymology	Etymology. Named for its occurrence in Hormozgan Province.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC717047D098FC56FD1BDA6A.taxon	discussion	Remarks. Torrenticola hormozganensis sp. nov. is rather similar to T. arabica Smit & Pešić, 2010 from northern Oman (Smit & Pešić 2010), in respect to the general morphology of the idiosoma (relatively short median suture line of Cx-II + III and roundish genital field in males, Cx-IV posteriorly extending well beyond genital field in both sexes) and the gnathosoma with a long rostrum. The main difference is found in the ventral projection on P- 2 which distinctly curves distally in T. arabica (see Smit & Pešić 2010). Further differences are the excretory pore and Vgl- 2 shifted from the line of primary sclerotization in both sexes of T. hormozganensis. Habitat. Sandy / bouldery streams, with strong exposure to sunlight and without vegetation (Fig. 44 C).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC717047D098FC56FD1BDA6A.taxon	distribution	Distribution. Iran; Hormozgan Province.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC777044D098FA00FAA3DA12.taxon	description	(Fig. 15 A – B)	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC777044D098FA00FAA3DA12.taxon	materials_examined	Material examined. Iran, Hormozgan Province, IR 19 - 2011, Banglayan village N of Bandar Abbass, unnamed stream, 27 º 45.695 N, 56 º 32.402 E, 577 m asl, 18. vii. 2011 0 / 4 / 0 (0 / 2 / 0 mounted). Morphology. Female — Idiosoma (ventral view: Fig. 15 B) L 753, W 423, dorsal shield (Fig. 15 A) L 625, W 319, L / W ratio 1.96; shoulder plate L 138 – 144, W 47, L / W ratio 2.94 – 3.06; frontal plate L 109, W 53, L / W ratio 2.06; shoulder / frontal plate L ratio 1.32; gnathosomal bay L 133, Cx-I total L 240, Cx-I mL 106, Cx-II + III mL 63; ratio Cx-I L / Cx-II + III mL 3.8; Cx-I mL / Cx-II + III mL 1.68; genital field L / W 142 / 130, L / W ratio 1.09; distance genital field — excretory pore 217, genital field — caudal idiosoma margin 300; gnathosoma vL 185, chelicera L 215; palp total L 187, dL: P- 1, 26; P- 2, 58; P- 3, 37; P- 4, 46; P- 5, 20; P- 2 / P- 4 ratio, 1.26; distal margins of P- 2 and P- 3 without denticles; L I-L- 4 - 6: 92, 95, 103; I-L- 6 L / H ratio 2.48.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC777044D098FA00FAA3DA12.taxon	discussion	Remarks. The specimens from Hormozgan Province fit the description of Monatractides kermanshahus Pešić et al. 2008, a species known only and previously recorded only from females from Kermanshah Province (Pešić et al. 2008). Habitat. Sandy / bouldery streams with strong exposure to sunlight and without vegetation (Fig. 44 C).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC777044D098FA00FAA3DA12.taxon	distribution	Distribution. Iran: Kermanshah Province (Pešić et al. 2008); Hormozgan Province (present study).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC74704BD098FA3AFD26DD7A.taxon	description	(Fig. 16 A – F, 17 A – F)	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC74704BD098FA3AFD26DD7A.taxon	materials_examined	Type material. Holotype male, Hormozgan Province, Bandar Abbass, IR 21 - 2011, Rudan stream 5 km N of Rudan, lotic part, 27 º 28.226 N, 57 º 15.153 E, 200 m asl, 21. vii. 2011 (RMNH). Paratypes: 22 / 14 / 0 (4 / 2 / 0 mounted), same data as holotype; ibid., lentic part, 0 / 2 / 0 (RMNH); Rudan stream, upstream, IR 22 - 2011, 27 º 28.748 N, 57 º 15.648 E, 210 m asl, interstitial dig, 3 / 2 / 0 (RMNH). Further records. Iran: Hormozgan Province, IR 18 - 2011, Taleguerdoo village N of Bandar Abbas, Poshtekeno spring, upper part of stream, 27 º 48.783 N, 56 º 24.459 E, 836 m asl, 18. vii. 2011 13 / 4 / 2 (1 / 0 / 0 mounted); IR 19 - 2011, Banglayan village, stream, 27 º 45.695 N, 56 º 32.402 E, 577 m asl, 18. vii. 2011 4 / 4 / 0 (1 / 0 / 0 mounted); Sistan va Baluchestan Province, IR 11 - 2011, Hootgat Bala river near Dargas village NE of Chabahar, upstream, 25 º 49.539 N, 61 º 31.033 E, 65 m asl, 14. vii. 2011 0 / 2 / 0 (0 / 1 / 0 mounted).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC74704BD098FA3AFD26DD7A.taxon	diagnosis	Diagnosis. Shoulder plate elongated (shoulder / frontal plate L ratio 1.5 – 1.8); three pairs of knob-like protrusions on the lateral margin of the capitular bay: one pair at the apical tip of Cx-I, two further pairs distally and proximally from the subapical seta; suture line of Cx-IV partially distinct and curved; excretory pore well away from the line of primary sclerotization, Vgl- 2 posterior to excretory pore; distal margins of P- 2 without denticles, P- 3 with one denticle or without denticles at the mediodistal margin, P- 4 with a well visible denticle near the insertion of the ventral hairs.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC74704BD098FA3AFD26DD7A.taxon	description	Description. Male (holotype, in parentheses paratypes n = 3, in square brackets specimens from IR 19 - 2011) — Idiosoma (ventral view: Fig. 16 B) L 856 (788 – 813) [875], W 681 (556 – 613) [613], dorsal shield (Fig. 16 A) L 706 (606 – 634) [694], W 494 (400 – 472) [494], L / W ratio 1.43 (1.34 – 1.52) [1.40]; dorsal plate L 628 (541 – 566) [618]; shoulder plate L 222 – 223 (181 – 209) [234 – 241], W 80 – 84 (72 – 82) [84 – 88], L / W ratio 2.65 – 2.78 (2.48 – 2.68) [2.74 – 2.79]; frontal plate L 131 (114 – 128) [131 – 133], W 106 (90 – 103) [106 – 109], L / W ratio 1.24 (1.14 – 1.32) [1.20 – 1.26]; shoulder / frontal plate L ratio 1.7 (1.54 – 1.75) [1.76 – 1.84]; gnathosomal bay L 148 (134 – 177) [184], Cx-I total L 266 (233 – 294) [303], Cx-I mL 117 (97 – 116) [119], Cx-II + III mL 141 (126 – 136) [127]; ratio Cx-I L / Cx-II + III mL 1.89 (1.78 – 2.33) [2.39]; Cx-I mL / Cx-II + III mL 0.83 (0.74 – 0.92) [0.94]; genital field L / W 136 (122 – 127) [137] / 123 (112 – 122) [128], L / W ratio 1.1 (1.03 – 1.09) [1.07]; ejaculatory complex (Fig. 16 F), L 188 (180 – 205) [185]; distance genital field — excretory pore 175 (147 – 178) [189], genital field — caudal idiosoma margin 297 (242 – 259) [289]; gnathosoma vL 198 (178 – 201) [211]; chelicera L 232 (220 – 229) [234], H 32 (30 – 32) [27], ratio L / H 7.18 (7.09 – 7.4) [8.7]; palp (Figs. 16 D – E) total L (204 – 228) [217], dL: P- 1, 26 (23 – 29) [25]; P- 2, 72 (66 – 75) [69]; P- 3, 48 (42 – 45) [44]; P- 4, 58.5 (52 – 56) [57]; P- 5, 23 (21 – 23) [22]; P- 2 / P- 4 ratio, 1.23 (1.22 – 1.29) [1.21]; L I-L- 4 - 6: 123 (109 – 122) [119], 118 (100 – 109) [112], 105 (99 – 103) [100]; I-L- 6 L / H ratio 2.9 (2.68 – 2.82) [2.62]. Female — Idiosoma (ventral view: Fig. 17 B) L 863 – 882, W 610 – 650, dorsal shield (Fig. 17 A) L 675 – 700, W 450 – 473, L / W ratio 1.48 – 1.5; shoulder plate L 195 – 209, W 73 – 78, L / W ratio 2.63 – 2.68; frontal plate L 116 – 127, W 89 – 99, L / W ratio 1.23 – 1.32; shoulder / frontal plate L ratio 1.62 – 1.71; gnathosomal bay L 144 – 150, Cx-I total L 246 – 267, Cx-I mL 102 – 117, Cx-II + III mL 91 – 100; ratio Cx-I L / Cx-II + III mL 2.46 – 2.9; Cx-I mL / Cx-II + III mL 1.02 – 1.29; genital field L / W 154 – 163 / 153 – 163, L / W ratio 1.0 – 1.01; distance genital field — excretory pore 175 – 195, genital field — caudal idiosoma margin 312 – 319; gnathosoma (Fig. 17 E) vL 185 – 208, chelicera L 217 – 235, H 31 – 33, ratio L / H 6.78 – 7.4; palp (Fig. 17 F) total L 213 – 236, dL: P- 1, 25 – 29; P- 2, 69 – 74; P- 3, 42 – 59; P- 4, 54 – 59; P- 5, 23 – 25; P- 2 / P- 4 ratio, 1.25 – 1.28; L I-L- 4 - 6: 112 – 115, 109 – 119, 100 – 108; I-L- 6 L / H ratio 2.6 – 2.8.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC74704BD098FA3AFD26DD7A.taxon	etymology	Etymology. Named after Dr Peter Martin (Kiel) in appreciation of his acarological work and continuous effort to find referees for our papers.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC74704BD098FA3AFD26DD7A.taxon	discussion	Remarks. Monatractides martini sp. nov. belongs to a group of species characterized by a long median suture line of Cx-II + III in males and the presence of three pairs of knob-like protrusion on the lateral margin of the capitular bay. This group includes M. algeriensis (Lundblad, 1941) (Mediterranean — Lundblad 1942, Di Sabatino et al. 1992), M. balneatoris Di Sabatino & Gerecke, 2003 (Dead Sea area in Israel — Di Sabatino et al. 2003), M. oman Gerecke, 2004 (Oman — Gerecke 2004 b, Smit & Pešić 2010), M. vafaei Pešić, 2004 (Iran — Pešić & Saboori 2004, Turkey — Pešić et al. 2006). Differences between these species regard the more or less elongated idiosoma and gnathosoma, the denticulation on the mediodistal margin of P- 2 and P- 3, the shape of Cx-IV, the extension of postgenital area, length / height ratio of chelicera and length of ejaculatory complex. From all the aforementioned species M. martini sp. nov. can be distinguished by a much wider frontal plate, L / W ratio 1.1 – 1.3. Further diagnostic characters of M. martini sp. nov. are a moderately elongated idiosoma, shoulder plates relatively long (shoulder / frontal plate L ratio 1.5 – 1.8), distal margins of P- 3 bearing at most one pointed denticle, occasionally without a denticle, a slender chelicera (L / H ratio 7.1 – 8.7), a relatively small ejaculatory complex (length 180 – 205) and Cx-IV posteriorly moderately extending beyond the genital field. Pešić & Saboori (2004) reported M. algeriensis from the first order stream running from a thermal spring in Kerman Province, and mentioned that the single collected female differed from typical M. algeriensis in remarkably wider frontal plates. This female generally agrees well with our specimens, but the male differs in major dimensions and shorter frontal plates. Further material is needed for understanding the taxonomic placement of this population. Habitat. Most specimens were collected in lotic parts of sandy / bouldery streams, with strong exposure to sunlight (Figs. 44 C – D), only a few specimens in interstitial waters or in a lentic habitat.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC74704BD098FA3AFD26DD7A.taxon	distribution	Distribution. Iran (Hormozgan Province).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC7B704ED098F8D4FBB9DD1E.taxon	description	(Fig. 18 A – H, 19 A – B, 20 A – D)	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC7B704ED098F8D4FBB9DD1E.taxon	materials_examined	Material examined. Holotype female, dissected and slide mounted, Iran, Hormozgan Province, IR 19 - 2011, Banglayan village N of Bandar Abbass, unnamed stream, 27 º 45.695 N, 56 º 32.402 E, 577 m asl, 18. vii. 2011 (RMNH). Paratypes: 1 / 1 / 0 (1 / 0 / 0 mounted), same data as holotype (RMNH).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC7B704ED098F8D4FBB9DD1E.taxon	diagnosis	Diagnosis. Frontal platelet elongated and relatively narrow (L / W ratio 3.0 – 3.3); Cx-IV posteriorly extending well beyond genital field; excretory pore and Vgl- 2 on the margin of primary sclerotization, near posterior margin of idiosoma; P- 1 separate from P- 2, P- 2 relatively long and slender, P- 3 longer than P- 4, P- 2 and - 3 each with relatively long and slender ventral seta but without ventral projections, L P- 2 / P- 4 ratio 3.27. Gnathosoma with short posterodorsal projections and with inconspicuous oral papillae, rostrum narrower than remainder of the capitulum; chelicera long with relatively short cheliceral claw (L basal segment / claw ratio 4.3 – 4.4).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC7B704ED098F8D4FBB9DD1E.taxon	description	Description. Female — Idiosoma (ventral view: Fig. 18 B) L 633, W 509, dorsal shield (Fig. 18 A) L 544, W 397, L / W ratio 1.37; shoulder plate L 159 – 162, W 54 – 56, L / W ratio 2.8 – 3.0; frontal plate L 138 – 142, W 44 – 45, L / W ratio 3.06 – 3.24; shoulder / frontal plate L ratio 1.14 – 1.16; gnathosomal bay broad and very shallow, L 106, Cx-I total L 250, Cx-I mL 144, Cx-II + III mL 15; ratio Cx-I L / Cx-II + III mL 16.7; Cx-I mL / Cx-II + III mL 9.6; genital field L / W 143 / 128, L / W ratio 1.12; distance genital field — excretory pore 207, genital field — caudal idiosoma margin 219; gnathosoma (Fig. 18 D) vL 325; chelicera total L 402, claw L 75, basal segment L 330, L basal segment / claw ratio 4.4; palp (Figs. 18 E – F) total L 173, dL: P- 1, 33; P- 2, 72; P- 3, 32; P- 4, 22; P- 5, 14; L P- 2 / P- 4 ratio, 3.27; dL of I-L- 4 - 6 (Fig. 18 G): 72, 84, 96; I-L- 6 L / H ratio 2.45; dL of IV-L (Fig. 18 H): 134, 89, 103, 128, 150, 178. Male — Idiosoma (ventral view: Fig. 19 B) L 575, W 459, dorsal shield (Fig. 19 A) L 496, W 372, L / W ratio 1.33; shoulder plate L 147 – 148, W 56, L / W ratio 2.6 – 2.7; frontal plate L 125 – 134, W 41 – 42, L / W ratio 3.0 – 3.27; shoulder / frontal plate L ratio 1.2 – 1.18; gnathosomal bay L 94, Cx-I total L 238, Cx-I mL 144, Cx-II + III mL 62; ratio Cx-I L / Cx-II + III mL 3.8; Cx-I mL / Cx-II + III mL 2.3; genital field L / W 122 / 97, L / W ratio 1.26; ejaculatory complex (Fig. 20 C) L 168; distance genital field — excretory pore 136, genital field — caudal idiosoma margin 150; gnathosoma (Fig. 20 B) vL 295; chelicera total L 362, claw L 69, basal segment L 297, L basal segment / claw ratio 4.3; palp (Fig. 20 A) total L 159, dL: P- 1, 30; P- 2, 63; P- 3, 31; P- 4, 21; P- 5, 14; L P- 2 / P- 4 ratio, 3.0; dL of I-L- 4 - 6 (Fig. 20 D): 69, 78, 96; I-L- 6 L / H ratio 2.34; dL of IV-L: 122, 88, 97, 120, 146, 179.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC7B704ED098F8D4FBB9DD1E.taxon	etymology	Etymology. The name refers to the very hot conditions (up to 49 ° C) in which we had to collect water mites in southern Iran.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC7B704ED098F8D4FBB9DD1E.taxon	discussion	Remarks. Other species of the subgenus Allotorrenticola are Neoatractides abnormipalpis (Lundblad, 1941) (Burma, Malaysia, Brunei), N. bahtilli (Wiles, 1991) (Malaysia), N. malayensis (Wiles, 1991) (Malaysia) and N. suvarna (Cook, 1967) (India) (see Wiles 1997 b). Due to the absence of projections on P- 2 and P- 3, the female of the new species most resembles N. abnormipalpis and N. suvarna. Neoatractides abnormipalpis can be easily distinguished by P- 1 partially fused with P- 2 and in a very reduced P- 4 (shorter than P- 3) and P- 5 (see Wiles 1991). Neoatractides suvarna, a species similar in the capitulum with inconspicuous oral papillae and P- 1 separate from P- 2, differs in similarly shaped, broad frontal and shoulder platelets, a very short ventral seta on P- 2 and P- 3, P- 2 relatively short and stout, chelicera with a relatively much longer cheliceral claw (basal segment / claw ratio 2.06, calculated from Cook 1967) and the excretory pore located on a tubercle (see Cook 1967). Habitat. Sandy / bouldery stream, with strong exposure to sunlight (Fig. 44 C).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC7B704ED098F8D4FBB9DD1E.taxon	distribution	Distribution. Iran; only known from the type locality in Hormozgan Province.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC7E704ED098F970FAF0DCEE.taxon	materials_examined	Material. Iran, Hormozgan Province, Bandar Abbass, IR 21 - 2011 Rudan stream, 5 km N of Rudan, lentic part, 27 º 28.226 N, 57 º 15.153 E, 200 m asl, 21. vii. 2011 22 / 0 (1 / 1 / 0 mounted); ibid., lotic part, 2 / 0 / 0. Habitat. Most specimens were collected in the lentic part of sandy / bouldery streams, with strong exposure to sunlight (Fig. 46 D).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC7E704ED098F970FAF0DCEE.taxon	distribution	Distribution. Oman, Iran (Kerman Province — Asadi et al. 2010; Hormozgan Province — present study).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC7F704FD098FDF8FCE9D8E7.taxon	diagnosis	Diagnosis. Sixth segment of I- and II-L modified in male, distally tapering, and with multiple irregular constrictions at the ventral margin. Ac relatively small, Ac- 2 smaller than Ac- 1 and Ac- 3, the distance between Ac- 1 and Ac- 2 much larger than between Ac- 2 and Ac- 3; genital plates in male separate; numbers of genital setae low (5 - 6 in male, 3 in female). Ventral seta of P- 2 heavy, peg-shaped, inserted slightly laterally.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC7F704FD098FDF8FCE9D8E7.taxon	type_taxon	Type species. Limnesia diversipes sp. nov.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC7F704FD098FDF8FCE9D8E7.taxon	etymology	Etymology. The new subgenus is named for its occurrence in saline environment.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC7F704FD098FDF8FCE9D8E7.taxon	discussion	Remarks. The slightly modified first and especially the second leg, with the sixth segment distally tapering and with multiple irregular constrictions at the ventral margin is very distinctive and will separate members of the subgenus from representatives of all other other subgenera of this genus. Sexual dimorphism in I-L has not been reported in other limnesiids and a modified second leg is found only in Xenolimnesia Smit from Cameroon (Smit 1994), a genus having the three distal segments highly modified.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC7F704FD098FF4DFD52DAC0.taxon	materials_examined	Material. Iran, Sistan va Baluchestan Province, IR 5 - 2011, Gerdoig village N of Chabahar, 5 km from Koch village, pool, 25 º 46.084 N, 60 º 54.968 E, 13. vii. 2011 6 / 12 / 0; IR 6 - 2011, Kombedan Dam, 25 º 53.302 N, 60 º 58.912 E, 84 m asl, 13. vii. 2011 4 / 3 / 0; IR 7 - 2011, Sheikh Kallag Dam, 25 º 58.236 N, 61 º 01.344 E, 100 m asl, 13. vii. 2011 1 / 1 / 0; IR 9 - 2011, Shirgovaz — Machkor stream, 25 º 46.618 N, 61 º 27.632 E, 45 m asl, 14. vii. 2011 5 / 15 / 0. Habitat. Lenitobiont.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC7F704FD098FF4DFD52DAC0.taxon	distribution	Distribution. India, Oman (Smit & Pešić 2010), Iran (Kerman Province — Pešić et al. 2011; Hormozgan and Sistan va Baluchestan provinces — present study).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC7F7072D098FBE7FBB9DBF7.taxon	description	(Figs. 21 A – E, 22 A – E)	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC7F7072D098FBE7FBB9DBF7.taxon	materials_examined	Material examined. Holotype male, dissected and slide mounted, Iran, Hormozgan Province, IR 15 - 2011, Khorgoo village N of Bandar Abbass, upstream hot water spring, small saline pool near IR 14 - 2011, 27 º 29.155 N, 56 º 27.531 E, 113 m asl, 16. vii. 2011. Paratypes: 14 / 16 / 0 (0 / 1 / 0 mounted), same data as holotype.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC7F7072D098FBE7FBB9DBF7.taxon	description	Description. Male (holotype) — Idiosoma L / W 488 / 352; integument very thin; dorsum with a circular postero-medial platelet. Coxal field corresponding to other species of the genus, L 228, Cx-III W 239; Cx-I medially separated, subcutaneous projections at caudal margin of Cx-I, medial margin of Cx-III and posterior part of medial margin of Cx-IV; Glandulae Limnesiae at centre of the medial margin of Cx-III; Cx-IV with one seta in area of insertion of IV-L. Genital field nearly circular, L / W 103 / 102, genital plates separated, Ac relatively small, Ac- 2 smaller than Ac- 1 and Ac- 3, distance between Ac- 1 and Ac- 2 four to five times larger than between Ac- 2 and Ac- 3, each genital plate with 5 - 6 setae, pre- and postgenital sclerite moderately developed, postgenital sclerite caudally not extending beyond genital plates. Palp (Fig. 21 C): total L, 241; P- 1, 14; P- 2, 66; P- 3, 45; P- 4, 91; P- 5, 25; L P- 2 / P- 4 ratio, 0.73; ventral seta of P- 2 heavy, peg-shaped, inserting slightly laterally, ventral margin of P- 4 with a long seta accompanied by a flat tubercle. Gnathosoma vL 92; chelicera total L 147. Legs: dL of I-L- 3 - 6: 65, 66, 74, 92; dL of II-L- 2 - 6: 55, 81, 87, 102, 112; II-L- 6 dL / H ratio 3.1; sixth segment of I- and II-L sligtly modified (Figs. 22 B and - D), distally tapering, ventral margin with multiple irregular constrictions; dL of IV-L: 85, 71, 91, 120, 129, 128; IV-L- 6 (Fig. 21 B) with a strong sub-terminal seta, L 73. Female — Idiosoma L / W 553 / 442. Coxal field more slender than in male, L 273, Cx-III W 238. Genital field (Fig. 22 A) L / W 139 / 108, genital flap L 123, Ac- 2 close to Ac- 3, distance between Ac- 1 and Ac- 2 five to six times larger than between Ac- 2 and Ac- 3, each genital flap bearing three setae. Palp (Fig. 21 D): dL: P- 2, 77; P- 3, 48; P- 4, 108; P- 5, 28; L P- 2 / P- 4 ratio, 0.71; palp as in male, but P- 4 longer. Gnathosoma vL 124; chelicera (Fig. 19 E) total L 179. Legs: dL of I-L- 3 - 6: 62, 69, 77, 91; dL of II-L- 2 - 6: 59, 69, 87, 108, 109; II-L- 6 dL / H ratio 3.8; distal segments of I- and II-L unmodified, more slender than in male (Figs. 22 C and - E); dL of IV-L: 92, 75, 96, 121, 140, 129; IV-L- 6 subterminal seta L 76.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC7F7072D098FBE7FBB9DBF7.taxon	etymology	Etymology. Named for the modified legs of the male. Habitat. Remnant pool of a saline summer-dry stream (Fig. 8 B).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC7F7072D098FBE7FBB9DBF7.taxon	distribution	Distribution. Iran; only known from the type locality in Hormozgan Province.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC427072D098FCE0FBADDDBF.taxon	description	(Fig. 23 A – C)	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC427072D098FCE0FBADDDBF.taxon	materials_examined	Material. Iran, Hormozgan Province, IR 22 - 2011, 5 km N of Rudan N of Bandar Abbass, Rudan stream, 27 º 28.748 N, 57 º 15.648 E, 210 m asl, 21. vii. 2011 0 / 1 / 0 (mounted). Morphology. Female — Idiosoma L / W 931 / 669; dorsum without plates. Coxal field: Cx-I very long. Gnathosoma relatively small and broadly fused at base. Posterior apodemes of Cx-I + II moderately developed, suture line between Cx-III + IV complete, ending at bluntly-pointed medial margin of Cx-III + IV; posterior margin of Cx-IV with moderately developed projections; coxal field L 372, Cx-III W 416. Genital field L / W 136 / 164; genital plate narrow, L 87 - 92; egg (n = 2) maximum diameter 153. Palp (Fig. 23 C): total L 520, dL: P- 1, 30; P- 2, 142; P- 3, 131; P- 4, 188; P- 5, 29; L P- 2 / P- 4 ratio, 0.76; ventral margin of P- 2 and P- 3 with numerous small papillae, P- 2 and P- 3 swollen. Chelicera total L 287. Legs: dL of I-L- 2 - 6 (Fig. 23 B): 108, 136, 172, 185, 156; dL of IV-L: 141, 144, 231, 286, 325, 265.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC427072D098FCE0FBADDDBF.taxon	discussion	Remarks. The single specimen from Hormozgan province is provisionally assigned to the Oriental Hygrobates hamatus. It resembles both Hygrobates hamatus and the Afrotropical H. soari K. Viets, 1911. Cook (1967) described H. hamatus bharatensis Cook, 1967, from India and stated that this subspecies differs from the stem species in having much smaller projections on the posterior margin of Cx-IV. Later on, Gledhill & Wiles (1997) synonymized this subspecies with the stem species which is widespread in the Oriental region (Pešić et al. 2010) and furthermore reported from New Guinea and Australia (Wiles 1997 c, Cook 1986). Hygrobates hamatus is very similar to H. soari K. Viets, 1911, a species widspread in the Afrotropical region, reaching in its distribution to northern Oman (Gerecke 2004 b, Smit & Pešić 2010). The diagnostic differences separating these two species have been discussed only by Cook (1967) who mentioned that the anterior coxal group is much shorter in H. hamatus bharatensis. More material should be studied in order to get an insight into further diagnostic differences. Similar to Diplodontus silvestrii and D. schaubi, this probably will require applying molecular techniques. Habitat. Sandy / bouldery stream, with strong exposure to sunlight (Fig. 44 D).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC427072D098FE9DFA9AD991.taxon	description	Iranobates hesabii Peši ć, Smit & Asadi, 2011	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC427072D098FE9DFA9AD991.taxon	materials_examined	Material. Iran, Hormozgan Province, Bandar Abbass, IR 21 - 2011 Rudan city, 5 km after Rudan, Rudan stream, 27 º 28.226 N, 57 º 15.153 E, 200 m asl, 21. vii. 2011 3 / 9 / 0. Habitat. Lentic part of a sandy / bouldery stream, with strong exposure to sunlight; lenitophilous.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC427072D098FE9DFA9AD991.taxon	distribution	Distribution. Iran (Kerman Province — Pešić et al. 2011; Hormozgan Province — present study).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC427071D098F896FC8FD9A7.taxon	description	(Fig. 24 A – E)	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC427071D098F896FC8FD9A7.taxon	materials_examined	Material examined. Iran, Sistan va Baluchestan Province, Chabahar, IR 9 - 2011, Pools Shirgovaz River, 25 ° 46.618 N, 61 ° 27.632 E, 45 m asl, 14. vii. 2011 1 / 0 / 0 (mounted). Morphology. Male — Muscle attachment insertions sclerotized; dorsal and ventral shield present; dorsal shield divided into anterior and posterior plate (Fig. 24 A), L / W 466 / 349, anterior plate 128 / 263, posterior plate 341 / 349; coxal field: ventral surface covered by an extended shield (Fig. 24 E), L / W 450 / 434, including coxae, Vgl- 3 / 4 and genital field; genital field with 4 pairs of Ac, distance between most lateral pair of Ac 113, gonopore L 52; ejaculatory complex L 112; excretory pore sclerotized, Vgl- 1 fused to Vgl- 2. Palp (Fig. 24 C): palp total L 235, dL: P- 1, 26; P- 2, 49; P- 3, 48; P- 4, 89; P- 5, 23; L P- 2 / P- 4 ratio, 0.55; gnathosoma vL 105; chelicera total L 165. I-L (Fig. 24 D): I-L- 5 S- 1 and - 2 slender, close to each other, with obtuse tips, I-L- 6 stout, narrowed in the centre, claws with ventral and dorsal clawlets; I-L- 5 dL 114, vL 85, dL / vL 1.35, HB 34, dL / HB 3.36, S- 1 L 48, L / W 8.8, S- 2 L 55, L / W 8.9, S- 1 - 2 distance 7, L ratio S- 1 / 2 0.86; I-L- 6 L 86, HB 20, L / HB 4.3; L I-L- 5 / 6 1.33.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC427071D098F896FC8FD9A7.taxon	discussion	Remarks. The single male examined from the Sistan va Baluchestan Province fits well the description of Atractides biscutatus Cook, 1967, a species known only from India (Maharashtra and Kerala States — Cook 1967, Gujarat — Pešić & Ranga Reddy 2009) and Sri Lanka (Gledhill & Wiles 1997). Habitat. Sandy / muddy stream, with strong exposure to sunlight.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC427071D098F896FC8FD9A7.taxon	distribution	Distribution. India, Sri Lanka, Iran (present study).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC417075D098FD28FD24D8E2.taxon	description	(Figs. 25 A – F, 26 A – D, 27 A – D)	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC417075D098FD28FD24D8E2.taxon	materials_examined	Material examined. Holotype male, dissected and slide mounted, Iran, Hormozgan Province, IR 21 - 2011, Rudan city N of Bandar Abbass, 5 km N of Rudan, Rudan stream, lotic part, 27 º 28.226 N, 57 º 15.153 E, 200 m asl, 21. vii. 2011 (RMNH). Paratypes: 19 / 15 / 2, same data as holotype, 2 / 1 / 1 mounted; 1 / 1 / 0, IR 22 - 2011, Rudan stream, interstitial dig, 27 º 28.748 N, 57 º 15.648 E, 210 m asl, 21. vii. 2011 (all RMNH). Further records. Iran, Hormozgan Province, IR 18 - 2011, Taleguerdoo village N of Bandar Abbass, Poshtekeno spring, upper part of stream, 27 º 48.783 N, 56 º 24.459 E, 836 m asl, 18. vii. 2011 3 / 3 / 0 (1 / 1 / 0 mounted); IR 19 - 2011, Banglayan village, stream, 27 º 45.695 N, 56 º 32.402 E, 577 m asl, 18. vii. 2011 2 / 3 / 0; Sistan Province, IR 11 - 2011, Hootgat Bala river near Dargas village, NE of Chabahar, upstream, 25 º 49.539 N, 61 º 31.033 E, 65 m asl, 14. vii. 2011 1 / 1 / 0 (1 / 0 / 0 mounted).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC417075D098FD28FD24D8E2.taxon	diagnosis	Diagnosis. Dorsal integument striated; muscle attachment plates smooth; genital field in male with anterior margin medially with more or less protruding border of secondary sclerotization; palp slender (L / H ratio P- 2 1.4 – 1.6, P- 4 3.8 – 4.1 in both sexes, P- 3 ♂ 1.7 – 1.9, ♀ 2.3 – 2.4), P- 4 ventral margin divided by hair insertions into sectors 2: 3: 2; I-L- 5 smaller (L ♂ 146 – 171, ♀ 204 – 220, L ratio I-L- 5 / 6 1.1 – 1.2). General features. Dorsal integument striated; muscle attachment plates smooth. Coxal field: mediocaudal margin of Cx-I + II with a large, slightly concave area between the laterally directed apodemes of Cx-II. Genital field: Ac in a weakly curved line. Excretory pore smooth; Vgl- 1 not fused to Vgl- 2. Palp: weak sexual dimorphism, P- 2 ventral margin weakly S-shaped, convex in distal part, P- 4 ventral hairs long, sword seta close to distoventral hair. I-L- 5: S- 1 and - 2 strongly heteromorphic and distanced, ventral seta close to insertion of S- 1, I-L- 6 curved and slender, strongly narrowed in the centre, slightly thickened distally.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC417075D098FD28FD24D8E2.taxon	description	Description. Male (holotype, in parentheses measurements of paratypes, n = 2; in square brackets specimen from IR 18 - 2011) — Idiosoma L / W 538 (480 – 494) [506] / 380 (341 – 366) [375]. Coxal field (Fig. 25 A): L 278 (252 – 259) [288], Cx-III W 305 (278 – 283) [322], Cx-I + II mL 77 (66 – 69) [80], Cx-I + 2 lL 179 (156 – 162) [177]. Genital field (Figs. 25 A, C – D): L / W 74 (71 – 75) [80] / 84 (73 – 80) [85], anterior margin medially with more or less protruding border of secondary sclerotization, rarely straight, posterior margin straight, not indented, lateral margin weakly indented between Ac- 2 and Ac- 3, Ac- 3 distanced from anterior two acetabula on each side, L Ac- 1 - 3: 18 – 19 (15 – 20) [19 – 22], 17 – 20 (19 – 22) [21 – 23], 17 – 19 (17 – 19) [16 – 17]. Ejaculatory complex (Fig. 25 B) L 94 (91). Palp (Figs. 25 E – F): palp total L 270 (267) [268], dL: P- 1, 27 (24 – 26) [28]; P- 2, 62 (55 – 61) [63]; P- 3, 66 (61 – 66) [64]; P- 4, 89 (81 – 87) [87]; P- 5, 26 (27 – 28) [26]; L P- 2 / P- 4 ratio, 0.7 (0.67 – 0.7) [0.72]. I-L (Fig. 26 A – B): I-L- 5 dL 168 (146 – 157) [171], vL 94.6 (84 – 88) [100], dL / vL 1.78 (1.74 – 1.78) [1.71], HB 42.2 (35 – 41.5) [41], dL / HB 3.98 (3.78 – 4.2) [4.17], S- 1 L 95 (82 – 88) [85], L / W 8.8 (9.6 – 9.7) [10.0], S- 2 L 64 (60 – 63) [60], L / W 5.0 (5.7 – 6.5) [5.2], S- 1 - 2 distance 29 – 30 (26 – 28) [28 – 29], L ratio S- 1 / 2 1.49 (1.37 – 1.39) [1.41]; I-L- 6 L 147 (129 – 136) [144], HB 16 (15) [16], L / HB 9.04 (8.5 – 8.9) [9.06]; L I-L- 5 / 6 1.15 (1.13 – 1.15) [1.17]. Female (paratype, n = 1; in parentheses specimen from IR 18 - 2011) — Idiosoma L / W 766 (747) / 550 (503). Coxal field (Fig. 27 A): L 308 (334), Cx-III W 355 (394), Cx-I + II mL 79 (83), Cx-I + II lL 184 (197). Genital field (Fig. 27 B): L / W 142 / 137, genital plates slightly indented between Ac, genital plate L 77 – 82 (84), L Ac- 1 - 3: 22 (20 – 21), 22 – 23 (22), 23 – 25 (28). Palp (Fig. 27 D): palp total L 316 (335), dL: P- 1, 33 (34); P- 2, 71 (79); P- 3, 82 (85); P- 4, 100 (106); P- 5, 30 (31); L P- 2 / P- 4 ratio, 0.71 (0.75). I-L (Fig. 27 C): I-L- 5 dL 204 (220), vL 102 (112), dL / vL 2.0 (2.0), HB 55 (61), dL / HB 3.72 (3.62), S- 1 L 108 (123), L / W 8.8 (8.9), S- 2 L 68 (71), L / W 3.7 (3.23), S- 1 - 2 distance 53 (52), L ratio S- 1 / 2 1.6 (1.73); I-L- 6 L 170 (188), HB 16 (16), L / HB 10.6 (11.5); L I-L- 5 / 6 1.2 (1.17). Deutonymph (collected together with adults of Atractides hormozganus at IR 21 - 2011 and suspected to represent this species) — Idiosoma L / W 406 / 253; integument dorsally striated; muscle attachments unsclerotized. Coxal field: L 181, Cx-III W 215, Cx-I + II mL 45, Cx-I + II lL; provisional genital field with two pairs of Ac, L / W 37 / 48; excretory pore smooth; Vgl- 1 not fused to Vgl- 2. Palp (Fig. 26 C): palp total L 175, dL: P- 1, 15; P- 2, 42; P- 3, 43; P- 4, 57; P- 5, 18; L P- 2 / P- 4 ratio, 0.74. I-L (Fig. 26 D): I-L- 5 dL 94, vL 69, dL / vL 1.36, HB 29, dL / HB 3.24, S- 1 L 52, L / W ratio 8.4, S- 2 L 51, L / W ratio 6.6, S- 1 - 2 distance 0, L ratio S- 1 / 2 1.02; I-L- 6 L 79, HB 15, L / HB 5.4; L I-L- 5 / 6 1.2.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC417075D098FD28FD24D8E2.taxon	etymology	Etymology. Named for its occurrence in Hormozgan Province.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC417075D098FD28FD24D8E2.taxon	discussion	Remarks. Due to the similar morphology of the genital field (relatively small Ac arranged in a weakly curved line, male genital field apple-shaped with Ac- 3 distanced from Ac- 1 and - 2), I-L- 5 and - 6 (S- 1 and S- 2 with relatively wide setal interspace, I-L- 6 strongly curved and slender) and palp (without sexual dimorphism, P- 2 ventral margin slightly S-shaped in the both sexes), A. hormozganus sp. nov. is most similar to A. nikooae Pešić, 2004, a species known from both sexes from the Markazi Province (Pešić et al. 2004). From the latter species, A. hormozganus differs in both sexes in smaller dimensions of I-L- 5 (dL / H ♂ 196 / 50, ♀ 252 / 77 in A. nikooae), and more slender palp segments (L / H ratio: P- 2, 1.4 – 1.6, P- 4, 3.8 – 4.1 in both sexes, P- 3 ♂ 1.7 – 1.9, ♀ 2.3 – 2.4 vs. P- 2, 1.2 – 1.3, P- 4, 3.3 – 3.5 in both sexes, P- 3 ♂ 1.5, ♀ 2.0 in A. nikooae, measurements taken from the type material). Moreover, the ventral margin of P- 4 is divided by hair insertions in three sections with a ratio of 2: 3: 2 (2: 2: 3 in A. nikooae). Further differences are found in a more slender S- 2 (L / W ratio 4.4) in the female of A. nikooae, a shorter medial suture line of Cx-I + II in A. hormozganus (in A. nikooae L ♂ 100, ♀ 135) and the genital field anterior margin medially with a more or less protruding border of secondary sclerotization in the male of A. hormozganus (concave in A. nikooae). Habitat. Most specimens were collected in the lotic part of sandy / bouldery streams, with strong exposure to sunlight (Figs. 44 C – D); only a few specimens were found in an interstitial dig.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC417075D098FD28FD24D8E2.taxon	distribution	Distribution. Iran (Hormozgan Province).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC45707AD098FB1FFE1DDBF9.taxon	description	(Figs. 28 A – B, 29 A – B)	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC45707AD098FB1FFE1DDBF9.taxon	materials_examined	Material. Iran, Hormozgan Province, IR 21 - 2011, Rudan city N of Bandar Abbass, 5 km N of Rudan, Rudan stream, lentic part, 27 º 28.226 N, 57 º 15.153 E, 200 m asl, 21. vii. 2011 1 / 2 / 0 (0 / 1 / 0 mounted). Sistan va Baluchestan Province, IR 9 - 2011, pools Shirgovaz River, 25 ° 46.618 N, 61 ° 27.632 E, 45 m asl, 14. vii. 2011 0 / 2 / 0. Morphology. Female (from IR 21 - 2011) — Idiosoma L / W 731 / 622; L between anterior end of Cx-I and posterior end of Cx-IV 478, Cx-III W 464; apodemes of Cx-I long, extending posteriorly beyond the suture line between Cx-III and – IV. Fig. 28 A shows the coxal area. Genital field W 242; acetabular plate L / W 85 / 63, 10 - 11 acetabula on each side; egg maximum diameter 111. Palp: total L 306, dL: P- 1, 29; P- 2, 94; P- 3, 57; P- 4, 94; P- 5, 32; P- 2 / P- 4 ratio, 1.0; P- 4 with a short anteroventral tubercle (Fig. 28 B). Chelicera total L 121; dL of I-L- 2 - 6: 125, 147, 184, 178, 161; dL of IV-L: 81, 106, 122, 169, 195, 193; first (Fig. 29 A) and second leg with long fluted or grooved setae. Swimming setae numbers: II-L- 4 one, II-L- 5 one, III-L- 3 one, III-L- 4 five, III-L- 5 three, IV-L- 3 one to two, IV-L- 4 three and IV-L- 5 three.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC45707AD098FB1FFE1DDBF9.taxon	discussion	Remarks. Neumania maharashtris most closely resembles N. falcipes africana K. Viets, 1925, from which it can be distinguished by the pronounced concavity near the distal end of the ventral side of IV-L- 5, and proportionally much longer and less curved IV-L- 6 (Cook 1967). Neumania maharashtris was originally described from India (Maharashtra State), and recently reported from southern Iran (Pešić et al. 2011). The male specimens from Iran agree well with specimens from India, and the only differences are found in the presence of pinnate setae on IV-L- 5 in the specimen from Iran versus smooth setae in Cook’s drawings of the specimen from India. Checking the holotype of N. maharashtris is necessary before we can assess the taxonomic status of the population from Iran. Unfortunately, this was not possible at the time of preparing this paper. Thus far, only male specimens of this species were known, and this is the first description of the female. Habitat. Lentic part of a sandy / bouldery stream, with strong exposure to sunlight.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC45707AD098FB1FFE1DDBF9.taxon	distribution	Distribution. India; Iran (Kerman Province — Pešić et al. 2011; Hormozgan and Sistan va Baluchestan provinces — present study).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC4A707ED098FA30FD26D871.taxon	description	(Figs. 30 A – F, 31 A – F)	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC4A707ED098FA30FD26D871.taxon	materials_examined	Material examined. Holotype male, dissected and slide mounted, Iran, Hormozgan Province, IR 21 - 2011, Rudan city N of Bandar Abbass, 5 km N of Rudan, Rudan stream, lotic part, 27 º 28.226 N, 57 º 15.153 E, 200 m asl, 21. vii. 2011 (RMNH). Paratypes: 5 / 28 / 0 (1 / 1 / 0 mounted), same data as holotype; 0 / 2 / 0 (0 / 1 / 0 mounted), same data as holotype, lentic part; 5 / 4 / 0, same data as holotype, interstitial dig; IR 22 - 2011, Rudan city, 5 km N of Rudan, Rudan stream, upstream (lotic), 27 º 28.748 N, 57 º 15.648 E, 210 m asl, 21. vii. 2011 5 / 4 / 0 (all RMNH). Further records. Iran, Hormozgan Province, IR 18 - 2011, Taleguerdoo village N of Bandar Abbass, Poshtekeno spring, upper part of stream, 27 º 48.783 N 56 º 24.459 E, 836 m asl, 18. vii. 2011 0 / 1 / 0; IR 20 - 2011, Bandar Khamir to Bandar Lengeh road, ca 80 km from Bandar Abbass, saline stream near Dezhgas, 26 º 52.873 N, 55 º 16.354 E, 20 m asl, 20. vii. 2011 1 / 0 / 0.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC4A707ED098FA30FD26D871.taxon	diagnosis	Diagnosis. Dorsal furrow on each side with three short glandular platelets and three slit organs, slit organs free in integument; extreme posterior end of dorsal shield bearing the excretory pore and a pair of glandularia; IV-L showing sexual dimorphism. Male: IV-L- 5 curved (dorsal margin convex, ventral margin slightly concave) and anteriorly slightly enlarged. Female: acetabular plates separated from postgenital sclerite; excretory pore fused with postgenital sclerite; a pair of glandularia platelets located immediately lateral to the anterior end of acetabular plates.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC4A707ED098FA30FD26D871.taxon	description	Description. Male (holotype, in parentheses paratype, n = 1) — Dorsal shield (Fig. 30 A) L / W 359 (349) / 236 (224); extreme posterior end of dorsal shield bearing the excretory pore and a pair of glandularia; dorsal furrow on each side, from anterior to posterior, bearing: one slit organ, three short glandularium platelets, two slit organs. Ventral shield (Fig. 30 B) (including genital field) L / W 366 (344) / 298 (297); gnathosomal bay L 68 (65); anterior coxae not extending to anterior end of ventral shield; Cx-III W 194 (193). Genital field fused into the ventral shield; distance between the most lateral pair of Ac 75 (75); gonopore L 32 (29); genital field with three pairs of Ac, not flanked posteriorly by glandularia platelets; ejaculatory complex (Fig. 30 D) L 84 (80). Palp (Fig. 30 C) total L 153 (147), dL: P- 1, 22 (20); P- 2, 35 (36); P- 3, 26 (26); P- 4, 51 (46); P- 5, 19 (19); L P- 2 / P- 4 ratio, 0.69 (0.78); gnathosoma vL 66; chelicera total L 100. Legs: dL of I-L- 4 - 6 (Fig. 30 E): 50, 54 (52), 43 (39); dL of IV-L: 77 (79), 48 (48), 70 (66), 77 (72), 95 (85), 77 (72); III-L- 5 and IV-L- 5 each with one swimming seta; IV-L- 5 curved (dorsal margin convex, ventral margin slightly concave) and anteriorly slightly enlarged (Fig. 30 F). Female (paratypes, n = 2) — Similar to male except for the shape of ventral shield, genital field and IV-L- 5. Dorsal shield (Fig. 31 A) L / W 390 – 409 / 256 – 264. Ventral shield (Fig. 31 B) L / W 348 – 362 / 325. Gnathosomal bay L 68; Cx-III W 228 – 239. Genital field not fused with the ventral shield, with three pairs of Ac, arranged in an arc; acetabular plates L 44 – 45; distance between outer margins of acetabular plates 159; acetabular plates separated from postgenital sclerite. Excretory pore fused with postgenital sclerite. A pair of glandularia platelets located immediately lateral to the anterior end of acetabular plates (Fig. 31 C). Palp (Fig. 31 D) total L 156 – 162, dL: P- 1, 21 – 22; P- 2, 39 – 40; P- 3, 26 – 28; P- 4, 49 – 50; P- 5, 20 – 23; L P- 2 / P- 4 ratio, 0.78 – 0.82; palp as in male. Gnathosoma vL 66 – 74; chelicera total L 128. Legs: dL of I-L (Fig. 31 F): 34, 26, 37 – 40, 47 – 49, 52 – 54, 41 – 42; dL of IV-L (Fig. 31 G): 75 – 78, 46, 62 – 63, 70, 81 – 85, 68 – 76; IV-L- 5 not curved, distally not enlarged.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC4A707ED098FA30FD26D871.taxon	etymology	Etymology. Named for its occurrence in Iran (Persia)	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC4A707ED098FA30FD26D871.taxon	discussion	Remarks. The subgenus Javalbiopsis Cook, 1967, includes only one species, Javalbia africana Cook, 1966, taken from streams in West Africa (Liberia, Cook 1966). This subgenus is characterized by the excretory pore being fused with the dorsal shield in both sexes, and a postgenital sclerite fused with dorsal shield in the female (Cook 1974). Javalbia africana can be distinguished from the new species by the glandularia platelets in the dorsal furrow being greatly enlarged, the glandularia platelets associated with the genital field are fused with the anterior end of the acetabular plates, the shape of the palp (P- 4 more stout, ventral setae similar in length and closer to each other, P- 5 longer, see Fig. 626, Cook 1966), the shape of IV-L (all segments much shorter and stockier, see Cook 1966: Fig. 623) and I-L- 6 (distally strongly tapering, see Cook 1966: Fig. 633). Habitat. Most specimens were collected in the lotic part, a few specimens in the lentic part of sandy / bouldery streams with strong exposure to sunlight (Fig. 44 D).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC4A707ED098FA30FD26D871.taxon	distribution	Distribution. Iran (Hormozgan Province).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC4E707ED098FC09FA9DDDD2.taxon	description	(Fig. 32 A – D) New records. Iran: Sistan va Baluchestan Province, IR 11 - 2011, Hootgat Bala river near Dargas village, NE of Chabahar, upstream, 25 º 49.539 N, 61 º 31.033 E, 65 m asl, 14. vii. 2011 0 / 2 / 0 (0 / 1 / 0 used for SEM). Hormozgan Province, IR 22 - 2011, Rudan city N of Bandar Abbass, 5 km N of Rudan, Rudan stream, upstream, interstitial dig, 27 º 28.748 N, 57 º 15.648 E, 210 m asl, 21. vii. 2011 1 / 0 / 0 (used for SEM).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC4E707ED098FC09FA9DDDD2.taxon	discussion	Remarks. This species was recently described from a pool in a stream, where it was found to be relatively abundant (Asadi et al. 2010). The presence of this species in interstitial habitats was probably accidental.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC4E707ED098FC09FA9DDDD2.taxon	distribution	Distribution. Iran: (Kerman Province — Asadi et al. 2010; Hormozgan and Sistan va Baluchestan provinces — present study). Axonopsis (Hexaxonopsis) manoojani Peši ć & Asadi, 2010 New records. Iran, Hormozgan Province, IR 22 - 2011, Rudan city N of Bandar Abbass, 5 km N of Rudan, Rudan stream, interstitial dig, 27 º 28.748 N, 57 º 15.648 E, 210 m asl, 21. vii. 2011 0 / 1 / 0.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC4E707ED098FC09FA9DDDD2.taxon	discussion	Remarks. This species was recently described from a pool of a stream in South Iran (Asadi et al. 2010), and probably occurs only occasionally in the hyporheic habitat	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC4E707ED098FC09FA9DDDD2.taxon	distribution	Distribution. Iran (Kerman Province — Asadi et al. 2010; Hormozgan Province — present study).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC4E7061D098F8F5FBCAD98A.taxon	description	(Figs. 33 A – F, 34, 35 A – F)	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC4E7061D098F8F5FBCAD98A.taxon	materials_examined	Material examined. Holotype male, dissected and slide mounted, Iran, Hormozgan Province, IR 16 - 2011, shallow stream, N of Bandar Abbass, 27 º 27.622 N, 56 º 20.590 E, 130 m asl, 16. vii. 2011 (RMNH). Paratypes: 9 / 10 / 0 (0 / 1 / 0 mounted), same data as holotype (RMNH); 1 / 0 / 0, IR 15 - 2011, Khorgoo village before hot water spring, small pool in dry river bed near IR 14 - 2011, N of Bandar Abbass, 27 º 29.155 N, 56 º 27.531 E, 113 m asl, 16. vii. 2011.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC4E7061D098F8F5FBCAD98A.taxon	diagnosis	Diagnosis. Tips of Cx-I and - II with pointed, recurved, hook-like projections; two pairs of glandularia lying between the genital field and insertions of IV-L, the posterior one closely approaching the genital field; venter without a ridge extending posteriorly from the region of insertion of the IV-L; middle of ventral side of P- 4 expanded, bearing a hair-like seta lying on a small pointed tubercle; II-, III- and IV-L with swimming setae.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC4E7061D098F8F5FBCAD98A.taxon	description	Description. Male (holotype) — Dorsal and ventral shields anteriorly fused; dorsal shield (Fig. 33 A) L / W 363 / 297, with seven pairs of glandularia (seventh pair inconspicuous, flanking the excretory pore); postocularia well distanced from anterior margin; colour pattern indistinct; eye pigment well developed; excretory pore located at posterior end of dorsal shield. Ventral shield (Fig. 33 B) L / W 372 / 359, convex at anterior end, lateral margins rounded; tips of Cx-I and - II with pointed, recurved, hook-like projections. Gnathosomal bay L 101. Venter without a ridge on each side extending posteriorly from the region of insertion of the IV-L. Three pairs of Ac, arranged in an arc; distance between most lateral pair of Ac 95; gonopore W 23. Ejaculatory complex (Fig. 33 F) L 104. Palp (Figs. 33 C-D): total L 207, dL: P- 1, 34; P- 2, 50; P- 3, 29; P- 4, 73; P- 5, 21; L P- 2 / P- 4 ratio, 0.69; ventral margin of P- 2 convex, distal margin of P- 3 with hyaline extensions, middle of ventral side of P- 4 expanded, bearing a hairlike seta lying on a small pointed tubercle. Gnathosoma vL 79, chelicera total L 95. Legs: dL of I-L- 5 - 6 (Fig. 33 E): 67, 72; dL of IV-L: 47, 77, 57, 70, 89, 80; swimming setae numbers: II-L- 4, 2; II-L- 5, 3; III-L- 4, 2; III-L- 5, 4; IV- L- 4, 2; IV-L- 5 3. Female — similar to male except for the shape of the genital field (Fig. 34). Dorsal shield L / W 391 / 309; ventral shield L / W 395 / 391. Gnathosomal bay L 106. Distance between most lateral pair of Ac 129; gonopore relatively large, W 79. Palp: total L 207, dL: P- 1, 34; P- 2, 48; P- 3, 29; P- 4, 75; P- 5, 21; L P- 2 / P- 4 ratio, 0.64. Gnathosoma vL 79. Legs: dL of I-L- 3 – 6: 49, 65, 68; dL of IV-L- 2 - 6: 74, 60, 69, 87, 86. Palp chaetotaxy and swimming setae numbers as in male.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC4E7061D098F8F5FBCAD98A.taxon	etymology	Etymology. The species is named after Dr Peter Glöer (Hetlingen) in appreciation of his work on Iranian freshwater molluscs.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC4E7061D098F8F5FBCAD98A.taxon	discussion	Remarks. Axonopsis gloeeri sp. nov. is similar to A. kermanica Pešić & Asadi, 2010, with respect to the general morphology of the idiosoma and palps. The latter species differs from A. gloeeri sp. nov. in the presence of a semicircular ridge forming a sunken, elliptical area in the anterior part of the dorsal shield (see Fig. 32 A – B) and in the distinct colour pattern of the dorsum (see Pešić et al. 2010). Habitat. Sandy / bouldery stream, partly shaded by riparian vegetation (Fig. 44 B).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC4E7061D098F8F5FBCAD98A.taxon	distribution	Distribution. Iran; only known from the type locality in Hormozgan Province. Axonopsis (Brachypodopsis) interstitialis sp. nov. (Figs. 36 A – F, 37)	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC4E7061D098F8F5FBCAD98A.taxon	materials_examined	Material examined. Holotype male, dissected and slide mounted, Iran, Hormozgan Province, IR 22 - 2011, Rudan city N of Bandar Abbass, 5 km N of Rudan, Rudan stream, interstitial (dig), 27 º 28.748 N, 57 º 15.648 E, 210 m asl, 21. vii. 2011 (RMNH). Paratypes: 3 / 1 / 0, same data as holotype, 0 / 1 / 0 mounted; 4 / 2 / 0, same data as holotype, interstitial (pump) (all RMNH).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC4E7061D098F8F5FBCAD98A.taxon	diagnosis	Diagnosis. Idiosoma slender (L / W ratio 1.45 – 1.55 in both sexes) with subparallel lateral margins; dorsal shield colour pattern indistinct. Two pairs of glandularia lying between the genital field and insertions of the IV-L, these relatively close together. Venter with a short ridge on each side extending posteriorly from the region of insertion of the IV-L. P- 2 stout with a convex ventral margin, ventral margin of P- 3 with a kink, ventral margin of P- 4 convex, with two long hair-like setae, inserting side by side; II-, III- and IV-L with swimming setae (II-L- 5, 2; III-L- 4, 1; III-L- 5, 2; IV-L- 4, 1; IV-L- 5, 2).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC4E7061D098F8F5FBCAD98A.taxon	description	Description. Male (holotype, in parentheses measurements of paratypes, n = 2) — Dorsal and ventral shields fused anteriorly; dorsal shield (Fig. 36 A) L / W 450 / 264, with seven pairs of glandularia (the seventh pair inconspicuous, flanking the excretory pore); postocularia well distanced from anterior margin; colour pattern indistinct; eye pigment developed; excretory pore located at posterior end of dorsal shield. Ventral shield (Fig. 36 B) L / W 455 (447 – 481) / 313 (290 – 325), with subparallel lateral margins; lateral margins of Cx-I and - II without hook-like projections, forming a slightly undulating line, suture line of Cx-II / III indistinct. Gnathosomal bay L 101. Venter with a short ridge on each side extending posteriorly from the region of insertion of the IV-L. Three pairs of Ac, arranged in triangle; distance between most lateral pair of Ac 120; gonopore small, W 29. Ejaculatory complex (Fig. 36 C) L 78. Palp (Figs. 36 D-E): total L 192, dL: P- 1, 25; P- 2, 47; P- 3, 25; P- 4, 72; P- 5, 23; L P- 2 / P- 4 ratio, 0.65; P- 1 slender, with a blunt kink; ventral margin of P- 2 convex; dorsal margin of P- 3 convex, ventral margin of P- 3 with a nearly right-angled kink in its distal half; ventral margin of P- 4 convex, with two long hair-like setae, inserting side by side; P- 5 relatively long, inserted between weakly projecting convex lateral and medial projections of P- 4. Gnathosoma vL 49, chelicera total L 95. Legs: dL of I-L- 4 - 6: 45, 57, 60; dL of IV-L: 57, 63, 62, 74, 83, 79; IV-L- 2 / 3 distally swollen with pointed sheaths covering the base of the following segment (Fig. 36 F); swimming setae numbers: II-L- 5, 2; III-L- 4, 1; III-L- 5, 2; IV-L- 4, 1; IV-L- 5 2. Female (slide mounted paratype, in parentheses some measurements of other paratypes, n = 2). — Similar to male except for the shape of the genital field (Fig. 37). Dorsal shield L / W 466 / 289; ventral shield L / W 475 (472 – 522) / 333 (306 – 334). Gnathosomal bay L 103. Distance between most lateral pair of Ac 131; gonopore W 51. Palp: total L 193, dL: P- 1, 24; P- 2, 48; P- 3, 25; P- 4, 72; P- 5, 24; L P- 2 / P- 4 ratio, 0.67. Gnathosoma vL 55. Legs: dL of IV-L: 57, 66, 63, 72, 82, 83; palp chaetotaxy and swimming setae numbers as in male.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC4E7061D098F8F5FBCAD98A.taxon	etymology	Etymology. The species is named for its occurrence in interstitial waters.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC4E7061D098F8F5FBCAD98A.taxon	discussion	Remarks. The new species can be distinguished from other member of this subgenus by the more slender idiosoma with subparallel lateral margins, in combination with the following further diagnostic characters: (1) glandularia between the genital field and the insertions of the fourth legs lying close together; (2) venter with a short ridge on each side extending posteriorly from the region of insertion of the IV-L, and (3) the morphology of the palp (P- 2 stout, convex, ventral margin of P- 3 forming a kink, ventral margin of P- 4 convex, with two long hair-like setae, inserted side by side). Habitat. Interstitial waters associated with sand and gravel deposits of stream.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC4E7061D098F8F5FBCAD98A.taxon	distribution	Distribution. Iran; only known from interstitial waters in Hormozgan Province.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC517067D098FD0DFBCADAFA.taxon	description	(Figs. 38 A – H)	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC517067D098FD0DFBCADAFA.taxon	materials_examined	Material examined. Holotype male, dissected and slide mounted, Iran, Hormozgan Province, IR 22 - 2011, Rudan city N of Bandar Abbass, 5 km N of Rudan, Rudan stream, upstream, interstitial dig, 27 º 28.748 N, 57 º 15.648 E, 210 m asl, 21. vii. 2011 (RMNH). Paratypes: 2 / 3 / 0 (0 / 2 / 0 mounted), same data as holotype, interstitial dig; 3 / 2 / 0, same data as holotype, interstitial (pump); 0 / 1 / 0, same data as holotype, surface water sample (all RMNH).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC517067D098FD0DFBCADAFA.taxon	diagnosis	Diagnosis. Idiosoma relatively elongated (L / W ratio 1.7 – 1.9); dorsal furrow on each side in the posterior part with one short and one elongate glandular platelet and two free slit organs not fused with the neighbouring glandular platelets. Palp: P- 4 relatively long (L P- 2 / P- 4 ratio 0.65), proximally and distally narrowed, in the middle of ventral margin expanded, with two short setae, inserted side by side; II-, III- and IV-L with swimming setae.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC517067D098FD0DFBCADAFA.taxon	description	Description. Male (holotype, in parentheses measurements of paratypes) — Dorsal and ventral shield anteriorly fused; dorsal shield (Fig. 38 A) L / W 384 (379 – 386) / 213 (189 – 192), with four pairs of small glandularia (the fourth pair inconspicuous, flanking the excretory pore); postocularia well distanced from anterior margin; colour pattern indistinct; eye pigment developed. Dorsal furrow on each side, from anterior to posterior, bearing: one slit organ slightly posterior to the centre of idiosoma, a short glandularium platelet, a second pair of slit organs and an elongate glandularium platelet. Excretory pore located at posterior end of dorsal shield. Ventral shield (Fig. 38 B) L / W 415 (412 – 413) / 219 (213 – 217), with subparallel lateral margins; distal margin of Cx-I projecting beyond the strong concave frontal margin; lateral margins of Cx-I and - II without hook-like projections, forming a slightly undulating line, Cx-II / III not prominent. Gnathosomal bay L 105. Venter without a ridge on each side extending posteriorly from the region of insertion of the IV-L. Genital field distinctly set off from the remainder of the ventral shield. Four pairs of Ac arranged in two anterior and two posterior pairs; distance between most lateral pair of Ac 95; gonopore narrow W 13. A pair of slit organs and a pair of seta-bearing platelets lying in the soft integument posterior to the ventral shield. Ejaculatory complex (Fig. 38 E) L 155. Palp (Fig. 38 F): total L 189, dL: P- 1, 21; P- 2, 46; P- 3, 26; P- 4, 71; P- 5, 25; L P- 2 / P- 4 ratio, 0.65; P- 2 stout, ventral margin nearly straight, P- 3 with dorsal margin equally convex, distal margins forming sheets flanking the insertion of the subsequent segment; P- 4 proximally and distally narrowed, middle of ventral margin expanded, with two short setae, inserting side by side; P- 5 relatively long, inserted between weakly convex lateral and medial projections of P- 4. Gnathosoma vL 70, chelicera total L 101. Legs: dL of I-L- 5 - 6 (Fig. 38 G): 50, 72; dL of IV-L: 40, 55, 46, 68, 75, 80; IV-L- 2 / 3 distally with pointed sheaths covering the base of the following segment (Fig. 38 H); swimming setae numbers: II-L- 5, 2; III-L- 4, 1; III- L- 5, 2; IV-L- 4, 1; IV-L- 5, 2. Female (paratypes, n = 2) — Similar to male except for the shape of the genital field (Fig. 38 D). Dorsal shield L / W 394 / 209; ventral shield L / W 408 – 409 / 242; gnathosomal bay L 100 – 107. Distance between most lateral pair of Ac 103 – 112; gonopore W 23 – 27. Palp: total L 200, dL: P- 1, 21; P- 2, 48; P- 3, 29; P- 4, 74; P- 5, 28; L P- 2 / P- 4 ratio, 0.65. Gnathosoma vL 75 – 79, chelicera total L 97. Legs: dL of I-L- 3 – 6: 43 – 46, 46 – 49, 63 – 66; dL of IV-L: 39 – 42, 51 – 56, 47 – 48, 64 – 65, 69 – 73, 74; palp chaetotaxy and swimming setae numbers as in male.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC517067D098FD0DFBCADAFA.taxon	etymology	Etymology. The species is named for its occurrence in hyporheic habitat.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC517067D098FD0DFBCADAFA.taxon	discussion	Remarks. Due to the comparatively long and slender idiosoma and the lack of colour pattern, Axonopsis hyporheica sp. nov., most closely resembles A. phreaticola Cook, 1967, a hyporheobiontic species known from Maharashtra State in India (Cook 1967, Pešić & Ranga Reddy 2009). The new species differs from A. phreaticola (in the following, in parentheses measurements taken from Cook 1967) in its slightly smaller size, the second pair of slit-organs not fused (fused to posterior margin of anterior glandular platelets in the species from India, see Cook 1967: Fig. 631), and in a more slender and relatively longer P- 4 (stouter and relatively shorter, see Cook, 1967: Fig. 629; L P- 4 59 - 65 µm in both sexes, L P- 2 / P- 4 ratio 0.84 - 0.85). Habitat. Interstitial waters associated with sand and gravel deposits of a stream.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC517067D098FD0DFBCADAFA.taxon	distribution	Distribution. Iran; only known from interstitial waters in Hormozgan Province.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC577067D098FD9DFDCCDFCC.taxon	diagnosis	Diagnosis. Characters of the Aturidae and the subfamily Axonopsinae. Dorsal and ventral shields anteriorly fused; dorsal shield with seven pairs of glandularia and a pair of postocularia. Venter without a ridge on each side extending posteriorly from the region of insertion of the IV-L. Two pairs of glandularia lying between the genital field and insertions of the IV-L, relatively close together in male; posterior pair halfway between the fist pair of glandularia and genital field in female. Genital field with three pairs of so-called wheel-like acetabula; II-, III- and IV-L with swimming setae.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC577067D098FD9DFDCCDFCC.taxon	type_taxon	Type species. Haloaxonopsis salina sp. nov.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC577067D098FD9DFDCCDFCC.taxon	etymology	Etymology. Named for the salty environment where the type species was collected. The gender of the genus is feminine.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC577067D098FD9DFDCCDFCC.taxon	discussion	Remarks. The presence of wheel-like acetabula is unique in Aturidae and will separate the new genus from all other members of this family. Until now, so-called wheel-like acetabula were found only in Pontarachnidae, a family of marine water mites. The term ‘ wheel-like acetabula’ coined by Cook (1996) refers to the cuticular structure which is characterized by small radiating folds surrounding a cuticular depression, and as shown by Smit & Alberti (2009) these structures take part in osmoregulation. However, it was uncertain whether these structures are homologues of the acetabula of other freshwater mites (or Actinotrichida). Our findings support the supposed homology of acetabula and ' wheel-like acetabula'.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC577064D098FACCFB17DC87.taxon	description	(Figs. 39 A – B, 40 A – C, 41 A – E, 42 A – C, 43 A – F)	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC577064D098FACCFB17DC87.taxon	materials_examined	Material examined. Holotype male, dissected and slide mounted, Iran, Hormozgan Province, IR 15 - 2011, Khorgoo village before hot water spring, Bandar Abbass, saline stream, 27 º 29.347 N, 56 º 27.802 E, 125 m asl, 16. vii. 2011 (RMNH). Paratypes: 31 / 47 / 1, same data as holotype, 1 / 2 / 0 mounted. Further records. Iran, Hormozgan Province, IR 15 - 2011, Khorgoo village before hot water spring, small pool near IR 14 - 2011, N of Bandar Abbass, 27 º 29.155 N, 56 º 27.531 E, 113 m asl, 16. vii. 2011 1 / 7 / 1; IR 20 - 2011, Bandar Khamir to Bandar Lengeh road, ca 80 km from Bandar Abbass, saline stream near Dezhgas, 26 º 52.873 N, 55 º 16.354 E, 20 m asl, 20. vii. 2011 6 / 5 / 0 (1 / 0 / 0 mounted). Sistan va Baluchestan Province, IR 11 - 2011, Hootgat Bala river near Dorgas village, NE of Chabahar, upstream, 25 º 49.539 N, 61 º 31.03 E, 65 m asl, 14. vii. 2011 1 / 0 / 0 (mounted).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC577064D098FACCFB17DC87.taxon	diagnosis	Diagnosis. As for genus.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC577064D098FACCFB17DC87.taxon	description	Description. Male (holotype; in parentheses paratype, n = 1; in square brackets specimen from IR 20 - 2011, n = 1) — Dorsal and ventral shields anteriorly fused; dorsal shield (Fig. 39 A) L / W 513 (538) [470] / 389 (322) [345], with seven pairs of glandularia (the seventh pair inconspicuous, flanking the excretory pore); postocularia well distanced from anterior margin; colour pattern indistinct; extending beyond posterior margin of the ventral shield; eye pigment well developed; excretory pore located at posterior end of dorsal shield. Ventral shield (Fig. 39 B) L / W 519 (503) [494] / 459 (413) [469], slightly truncate at anterior end, lateral margins rounded. Gnathosomal bay L 134 (136) [125]. Venter without a ridge on each side extending posteriorly from the region of insertion of the IV-L. Three pairs of ' wheel-like' acetabula (Figs. 43 A – D), arranged in an arc; distance between most lateral pair of wheel-like Ac 107 (103) [113]. Gonopore extending beyond the anterior pair of weel-like acetabula, L / W 45 (44) [30] / 34 (30) [26]; gonopore flanked by several small setae. Ejaculatory complex (Fig. 41 D) L 142 (143) [135]. Palp (Figs. 41 A-B): total L 257 (256) [244], dL: P- 1, 42 (42) [37]; P- 2, 72 (69) [67]; P- 3, 32 (32) [31]; P- 4, 82 (83) [80]; P- 5, 29 (30) [29]; L P- 2 / P- 4 ratio, 0.88 (0.83) [0.84]; ventral margin of P- 2 convex, middle of ventral side of P- 4 expanded, bearing a hair-like seta lying on a small pointed tubercle. Gnathosoma vL 120 (122) [103]. Legs: dL of I-L- 4 - 6 (Fig. 41 C): 72 [65], 88 (88) [84], 94 (96) [85]; dL of IV-L (Fig. 41 E): 85 [83], 112 (115) [104], 85 (90) [79], 104 (108) [98], 122 (119) [112], 115 (117) [110]; swimming setae numbers: II-L- 5, 3; III-L- 4, 2; III-L- 5, 3; IV-L- 4, 2; IV-L- 5 3. Female (paratypes, n = 2) — Similar to male except for the shape of the genital field (Figs. 40 A, C). Dorsal shield L / W 497 – 544 / 353 – 381; ventral shield L / W 525 – 528 / 438; gnathosomal bay L 130 – 133. Distance between most lateral pair of weel-like Ac 118 – 124. Gonopore W 69 – 72; maximum egg diameter 212; palp (Fig. 42 A): total L 244 – 256, dL: P- 1, 39 – 43; P- 2, 68 – 69; P- 3, 31 – 33; P- 4, 78 – 82; P- 5, 28 – 29; L P- 2 / P- 4 ratio, 0.84 – 0.87; gnathosoma (Fig. 42 C) vL 109 – 128, chelicera (Fig. 42 B) total L 179 – 200. Legs: dL of I-L- 3 – 6: 57, 76 – 84, 82 – 86; dL of IV-L: 72 – 79, 88 – 97, 69 – 76, 89 – 95, 106 – 113, 93 – 112; palp chaetotaxy and swimming setae numbers as in male.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC577064D098FACCFB17DC87.taxon	etymology	Etymology. The species is named for its occurrence in saline habitat. Habitat. Apparently restricted to saline habitats. This is the only water mite and one of the most commonly collected invertebrate species in saline streams along the Persian Gulf. The associated fauna includes the snail species Cerithidaea cingulata (Gmelin) and Ecrobia sp., which are characteristic for brackish waters (Glöer & Pešić in prep.). We did not measure the salinity during our field work but most probably this species has a high salinity tolerance, as it is able to inhabit saturated brine (Fig. 44 A). The weel-like acetabula should be considered as an adaptation to the salty environment.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC577064D098FACCFB17DC87.taxon	distribution	Distribution. Iran; only known from saline streams along Persian Gulf and Oman Sea.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC55706BD098FB08FBF0DDD3.taxon	description	(Fig. 45 A – H)	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC55706BD098FB08FBF0DDD3.taxon	materials_examined	Material examined. Holotype male, dissected and slide mounted, Iran, Hormozgan Province, IR 22 - 2011, Rudan city N of Bandar Abbass, 5 km N of Rudan, Rudan stream, interstitial (pump), 27 º 28.748 N, 57 º 15.648 E, 210 m asl, 21. vii. 2011 (RMNH). Paratypes: 0 / 1 / 0 (mounted), same data as holotype, interstitial (pump); 1 / 0 / 0, same data as holotype, interstitial waters (dig); 2 / 4 / 0, same data as holotype, surface waters (all RMNH).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC55706BD098FB08FBF0DDD3.taxon	diagnosis	Diagnosis. Idiosoma dimensions small (L dorsal shield 414 – 423, ventral shield 400 – 409); glandularia on Cx- III shifted closer to D 2 than E 2; genital field with 6 - 11 Ac on each side.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC55706BD098FB08FBF0DDD3.taxon	description	Description. Male (holotype) — Dorsal and ventral shields present; dorsal shield (Fig. 45 A) L / W 414 / 247, oval, lateral margins indented posteriorly in the region of one of the posterior pair of glandularia; a ridge present on anterior and posterior portions of dorsal shield; dorsal shield colour indistinct. Ventral shield (Fig. 45 B) L / W 409 / 263, oval, notched on lateral margins at level of posterior end of Cx-III. Gnathosomal bay L 71. Suture line of Cx- IV well developed. Genital field completely fused with the ventral shield; 6 – 7 Ac on each side; distance between most lateral pair of Ac 106; gonopore small L / W 27 / 4. Excretory pore and a pair of flanking glandularia fused with the ventral shield. Ejaculatory complex (Fig. 45 D) L 65. Palp (Fig. 45 F): total L 123, dL: P- 1, 14; P- 2, 22; P- 3, 23; P- 4, 47; P- 5, 17; L P- 2 / P- 4 ratio, 0.47. Palps without sexual dimorphism: P- 2 with two enlarged setae ventrodistally, P- 4 medially with a thickened seta, near proximoventral hair, P- 5 abruptly tapering at distal end. Gnathosoma (Fig. 45 E) vL 106, chelicera total L 116. Legs: dL of I-L- 4 - 6 (Fig. 45 H): 39, 40, 59; dL of IV-L: 74, 42, 51, 72, 72, 61; IV-L- 2 / 3 distally with pointed sheaths covering the base of the following segment; swimming setae numbers: III-L- 5, 3; IV-L- 5, 3. Female (paratype) — Similar to male except for the shape of the genital field (Fig. 45 C). Dorsal shield L / W 423 / 263; ventral shield L / W 400 / 281; gnathosomal bay L 71. Genital field not fused with ventral shield; 9 – 10 Ac on each side; acetabular plates L 52 – 54; distance between outer margins of acetabular plates 128. Excretory pore and a pair of flanking glandularia lying free in the integument posterior to the genital field. Palp (Fig. 42 G): total L 120, dL: P- 1, 16; P- 2, 24; P- 3, 23; P- 4, 41; P- 5, 16; L P- 2 / P- 4 ratio, 0.59. Gnathosoma vL 77, chelicera total L 113. Legs: dL of I-L- 3 – 6: 40, 42, 62; dL of IV-L: 72, 38, 51, 72, 72, 59; palp chaetotaxy and swimming setae numbers as in male.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC55706BD098FB08FBF0DDD3.taxon	etymology	Etymology. The species is named for its occurrence in hyporheic waters.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC55706BD098FB08FBF0DDD3.taxon	discussion	Remarks. The subgenus Dentalbia Cook, 1974 is characterized (after Wiles 1992) by: 1) two glandularia (medially displaced L 1 and L 2 [sensu Wiles, 1996]) located on Cx-III; 2) P- 2 ventrodistally with 1 - 3 greatly enlarged setae; and 3) suture line between Cx-III and Cx-IV greatly reduced or absent. This subgenus includes: Albia paucipora Viets, 1935 (Java, Indonesia), A. dentipalpis Cook, 1967 (India), A. phreatica Cook, 1967 (India) and A. minutissima Lundblad, 1971 (Java, Indonesia). Due to P- 2 with two stout setae ventrodistally and the enlarged leg claws, the new species from Iran most closely resembles Albia phreatica Cook, 1967, a hyporheobiontic species known from Maharashtra State in India (Cook 1967, Pešić & Ranga Reddy, 2009). The new species differs from A. phreaticola (in the following, in parentheses, data taken from Cook 1967) in its smaller size (L dorsal shield 460 - 517, ventral shield 464 - 517, in the both sexes), the lower number of Ac on each side (13 - 25 Ac, in the both sexes), and in the glandularia on Cx-III shifted closer to D 2 than E 2 (closer to E 2 than to D 2). Habitat. Interstitial waters associated with sand and gravel deposits of stream.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC55706BD098FB08FBF0DDD3.taxon	distribution	Distribution. Iran; only known from the interstitial waters in Hormozgan Province.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC587068D098FEFFFBBADF5D.taxon	description	(Figs. 46 A – E)	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC587068D098FEFFFBBADF5D.taxon	materials_examined	Material examined. Iran, Hormozgan Province, IR 22 - 2011, Rudan city N of Bandar Abbass, 5 km N of Rudan, Rudan stream, interstitial dig, 27 º 28.748 N, 57 º 15.648 E, 210 m asl, 21. vii. 2011 1 / 0 / 0 (mounted); same data as holotype, surface water, 0 / 0 / 1 (mounted). Morphology. Male — Idiosoma (Fig. 46 A) L / H 491 / 256 (ratio 1.92). Genital field fused with the secondary sclerotization at posterior ventral shield margin. Three pairs of Ac, distance between outer margins of Ac- 1 and Ac- 3, 63. Ejaculatory complex (Fig. 46 C) L 86. Palp (Fig. 46 B): total L 178, dL: P- 1, 20; P- 2, 51; P- 3, 29; P- 4, 52; P- 5, 26; L P- 2 / P- 4 ratio 0.98; palp morphology as illustrated in Fig. 46 B. Legs: dL of I-L- 4 - 6 (Fig. 43 D): 43, 59, 75; segments of IV-L expanded and flattened (Fig. 46 E); dL of IV-L: 109, 38, 84, 88, 95, 75.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC587068D098FEFFFBBADF5D.taxon	discussion	Remarks. In having the idiosoma proportionally longer and lower (L / H ratio> 1.5), and the proportionally shorter segments of the fourth leg, especially IV-L- 5 and - 6 (proportionally longer in Indian subspecies F. reticulatifrons indicus, see Cook 1967), the specimen reported here is most similar to the European subspecies F. reticulatifrons reticulatifrons, but differs from typical specimens in the IV-L segments being more expanded and flattened. A further subspecies, F. reticulatifrons persicus Schwoerbel & Serpasgozarian, 1980, decribed from a single female from northern Iran (Mazandaran Province), is similar in having more expanded segments of the fourth leg (especially IV-L- 4 - 6), but differs in having the idiosoma proportionally much higher (idiosoma L / H ratio 1.4, data taken from Schwoerbel & Sepasgosarian 1980). Information about the variability of diagnostic characters of these subspecies is lacking. For this reason, we are convinced that it is best to postpone a decision on the taxonomic state of this specimen, probably a representative of an undescribed (sub) species, until the variability in additional specimens is known. Habitat. Interstitial waters associated with sand and gravel deposits of stream.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC587069D098FAEFFB19DBDA.taxon	description	(Fig. 47 A – D)	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC587069D098FAEFFB19DBDA.taxon	materials_examined	Material examined. Iran, Sistan va Baluchestan Province, IR 6 - 2011, Komb Dam NE of Chabahar, 25 º 53.302 N, 60 º 58.912 E, 84 m asl, 13. vii. 2011, 18 / 7 / 1; Reservoir Sheikh Kalag, IR 7 - 2011, 25 ° 58.236 N, 61 ° 01.344 E, 100 m a. s. l, 13. vii. 201 1 / 0 / 0; IR 9 - 2011, pools Shirgovaz River, 25 ° 46.618 N, 61 ° 27.632 E, 45 m asl, 14. vii. 2011 1 / 1 / 0. Morphology. Male — In addition to the description of Cook (1967) the following characters can be added. Males have a large hyaline membrane, with a concave posterior margin. The petiole is in ventral view tongueshaped (Fig. 47 D), in dorsal view four curved lobes can be seen (Fig. 47 C); medially the petiole has a sclerotized ridge. Female — Idiosoma greenish, truncated anteriorly, L / W 1532 / 1407. Anterior margin concave; dorsal shield (Fig. 47 A) complete, L 965, W 721. D 1 on small humps, L 4 on large humps. First coxal plates not extending to anterior idiosoma margin. Medial margin of fourth coxal plates much larger than medial margin of third coxal plates (Fig. 47 B). Medial distance of fourth coxal plates slightly larger than width of one genital valve. Genital field W 810. Gonopore without sclerotized patches, L / W 144 / 175. Genital plates about three times as long as wide, slightly enlarged laterally, sloping a little bit posterolaterally. V 2 on small humps. Palp: total L 412, dL: P- 1, 44; P- 2, 100; P- 3, 80; P- 4, 124; P- 5, 64; palp as in male, but P- 2 with three setae in anteroventral corner. Legs: dL of I-L- 4 - 6: 211, 203, 219; dL of IV-L- 4 - 6: 284, 227, 211; II-IV-Legs with numerous swimming setae.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC587069D098FAEFFB19DBDA.taxon	discussion	Remarks. Only the male of this species was known, and this is the first description of the female. Habitat. Standing waters.	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
A6418788BC587069D098FAEFFB19DBDA.taxon	distribution	Distribution. India (Cook 1967), Sulawesi, Indonesia (Smit 1996), Iran (present study).	en	Pešić, Vladimir, Smit, Harry, Saboori, Alireza (2012): 3330. Zootaxa 3330: 1-67
