taxonID	type	description	language	source
A62987A0360EFFCE3395F9F5FB2D4A13.taxon	description	(Figs. 2 – 9)	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A0360EFFCE3395F9F5FB2D4A13.taxon	materials_examined	Type material. — Arcastes sumatrensis: Holotype: Female, “ Spj, 4.77 / Arcastes sumatrensis Jac. “ (NNML; Fig. 8). Type locality: Indonesia, Soepajang 0 ° 27 ' S / 100 ° 54 ' E. Holotype by original indication “ single specimen from Soepajang (Sumatra- Expedition) ”. Neolepta biplagiata: Lectotype, Male, “ Dr. B. Hagen. Tandjong Morawa, Serdang, N. O. Sumatra / Neolepta biplagiata Jac. “ (NNML; Fig. 9). Type locality: Indonesia, Tandjong Morawa, 0 ° 35 ' S / 101 ° 18 ' E. Jacoby gave no number on the specimens he studied, but there are 14 specimens with labels from the type locality available. Thus, we herein designate a lectotype to fix the name on single specimen. Paralectotypes: 13 ex., same data as lectotype (7 ex. BMNH, 6 ex. NNML). – Invalid types: 1 ex. from Sumatra, Si-Rambé, 6 ° 11 ' S / 106 ° 48 ' E, Dec. 1990 – Mar. 1991, E. Modigliani in MCZ and 2 ex. from Dohrn, Sumatra, Soekaranda, 0 ° 37 ' S / 104 ° 29 ' E, 71484 in MNHU are labelled as co-type, but are not from the type series. Further material examined. — Indonesia. 3 ex., Sumatra, Collect. Duvivier, Arcastes sumatrensis Jacoby (IRSN; NNML; BMNH); 15 ex., Sumatra, Manna, 4 ° 27 ' S / 102 ° 59 ' E, M. Knappert, coll. Veth (NNML); 2 ex., Sumatra, Palembang, 2 ° 59 ' S / 104 ° 45 ' E, M. Knappert, coll. Veth (NNML); 1 ex., SVL, Deli, 3 ° 35 ' N / 98 ° 39 ' E, coll. Veth (NNML); 1 ex., N. Sumatra, Toba Plateau Tiga Dolok, Holzweg Eins, ca. 950 m, 2 ° 55 ' N / 99 ° 03 ' E, 20 Jun. 1972, J. Krikken, no. 20 (NNML); 1 ex., N. E. Sumatra, Deli, Penatangsiantar Balimbingan Est., forest 600 m, 3 ° 35 ' N / 98 ° 39 ' E, Aug. 1953, A. Sollaart (NNML); 2 ex., Sumatra, Fort cte kock, 0 ° 17 ' S / 100 ° 22 ' E, Oct. 1913, Edward Jacobson (BMNH, NNML); 1 ex., Soekaranda, 0 ° 37 ' S / 104 ° 29 ' E, Jan. 1894, Dohrn (BMNH); 1 ex., West Sumatra, Pulau Tello, 0 ° 04 ' S / 98 ° 13 ' E, Nov. 1924, C. B. K. and N. S. (BMNH); 1 ex., Sumatra, Siolak Daras, Korinchi Valley, 3100 ft, 2 ° 08 ' S / 101 ° 29 ' E, Mar. 1914 (BMNH); 1 ex., Sumatra, Sandaran Agong, Korinchi Lake, 2450 ft, 2 ° 08 ' S / 101 ° 29 ' E, May – Jun. 1914 (BMNH); 1 ex., Sumatra, Pagherang Pisang, 0 ° 35 ' S / 101 ° 20 ' E, Oct. 1890, Mar. 1891, E. Modigliani (BMNH); 1 ex., Sumatra, Sungei-Bulu, Sep. 1878, G. Beccari (MCGD); 2 ex., Sumatra, Ajer-Mantcior, Aug. 1878, G. Beccari (MCGD); 3 ex., Sumatra, Pangherang-Pisang, Oct. 1890 – Mar. 1891, G. Beccari (MCGD); 1 ex., Brastagi, Sumatra, 3 ° 11 ’ N / 98 ° 28 ' E, Mjoberg (NHRS); 1 ex., N. O. Sumatra, Deli, 3 ° 35 ' N / 98 ° 39 ' E, L. Martin S., 86977 (MNHU); 1 ex., W. Sumatra, Liman Manis b. Padang, 0 ° 25 ' S / 101 ° 34 ' E, 8 Jan. 1909, Schoede S. G (MNHU); 1 ex., West Sumatra, Batu Insel Mentawei Gruppe, 0 ° 25 ' S / 101 ° 34 ' E, 16 Jan. 1909, H. Schoede (MNHU). – Malaysia. 9 ex., Malay Penin., Selangor, Bukit Kutu, 3300 ft, 3 ° 33 ' N / 101 ° 43 ' E, 19 Mar. 1931, 27 Sep. 1932, H. M. Pendlebury (BMNH); 4 ex., Semangko pass Selangor-Pahang, 2700 ’, 3 ° 35 ' N / 103 ° 24 ' E, Mar. 1912, Ex. F. M. S. Museum, B. M. 1955 – 354 (BMNH); 1 ex., Pahang, F. M. S. Cameron Highland, Gunung Kial, 5000 ft, 4 ° 30 ' N / 101 ° 23 ' E, 27 Jul. 1938 (BMNH); 1 ex., Perak, Gunong Kledang, 4 ° 35 ' N / 101 ° 01 ' E, 2646, Nov. 1916 (BMNH); 3 ex., Perak, 4 ° 48 ' N / 101 ° 09 ' E, Doherty, Fry coll. 1905.100 (BMNH); 1 ex., West Sumatra, 0 ° 44 ' S / 100 ° 48 ' E, Jacoby coll. 1909 – 28 a (BMNH); 1 ex., Kampong, 1450 m, 21 Jan. 1981 (NHRS); 1 ex., Pahang, Genting Highland, 2 km top, 3 ° 22 ' N / 102 ° 06 ' E, 2 Aug. 1992, C. W. & L. B. O’ Brein (UMNH); 1 ex., Borneo, Sabah, W Crocker Range E, West of Apin Apin, 5 ° 34 ' N / 116 ° 05 ' E, leg. Snizek, Feb. 2000 (CJB). Total length. — 4.85 – 5.75 mm (mean: 5.23 mm, n = 15) Head. — Reddish-brown, impunctate, first and second antennomeres contrasting reddish-brown, third to outer antennomere black (Figs. 2, 8, 9). Ratio length of second to third antennomeres 0.75 – 0.80 (mean: 0.76); median antennomeres insignificantly widened, ratio length of third to fourth antennomeres 0.40 – 0.50 (mean: 0.44), terminal are more slender (Fig. 3). Thorax. — Pronotum reddish-brown, pronotal width 1.50 – 1.75 mm (mean: 1.62), ratio length to width 0.52 – 0.60 (mean: 0.56 mm). Elytra black with transverse yellowish spot on the disc of each elytron, not quite extending to either margin (Figs. 2, 8, 9). Elytral length 3.80 – 4.50 mm (mean: 4.15 mm), maximal width of both elytra together 2.00 – 3.30 mm (mean: 2.93 mm), ratio of maximal width of both elytra together to length of elytra 0.69 – 0.76 (mean: 0.72). Legs reddish-brown. Abdomen. — Pale yellow to reddish-brown. Male genitalia. — Median lobe parallel-sided, rounded at apex, not incised, apex carinated ventrally in the middle. Tectum broad and became lanceolate at apex. Endophallus with one distinct type of spiculae, median spiculae long, slender and straight (Fig. 4). Female genitalia. — Like description of the genus (Figs. 5, 6).	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A0360EFFCE3395F9F5FB2D4A13.taxon	distribution	Distribution. — Neolepta sumatrensis is up to now known from Sumatra and some part of Peninsular Malaysia and Borneo (Fig. 7).	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A0360EFFCE3395F9F5FB2D4A13.taxon	diagnosis	Diagnosis. — Neolepta sumatrensis can be differentiated from N. quadriplagiata by the dorsal colouration. Neolepta sumatrensis are often with one transverse yellowish oval spot on each elytron while N. quadriplagiata has two yellowish oval spots on each elytron. As written by Jacoby (1884) in his original description, N. sumatrensis has a more transverse prothorax than other species. It is true that ratio pronotal length to width of N. sumatrensis is on average larger (0.52 – 0.60) than for N. quadriplagiata is (0.59 – 0.61). Characteristic for N. sumatrensis are also the widened median antennomeres in both sexes (Fig. 3), while N. quadriplagiata are only slender antennomeres (Fig. 11). The median lobe and the tectum of N. sumatrensis (Fig. 4) are broader than in N. quadriplagiata (Fig. 12).	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A0360CFFC030B7FED4FA104A31.taxon	description	(Figs. 10 – 15)	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A0360CFFC030B7FED4FA104A31.taxon	materials_examined	Type material. — Lectotype: “ Sumatra Mt. Singgalang, Luglio 1878. O. Beccari / Neolepta 4 plagiata Jac. ” (MCGD; Fig. 15). Type locality: Indonesia, Singgalang, 0 ° 24 ' S / 100 ° 21 ' E. Paralectotypes: 3 ex., same data as lectotype (MCGD); 1 ex., additionally with “ 1 st Jacoby coll. / Type 18314 ” (MCZH). Jacoby gave no number on the specimens he studied, and we herein designate a lectotype to fix the name on single specimen. Further material examined. — Indonesia. 2 ex., Sumatra, Ajer Mantcior, Aug. 1878, G. Beccari (MCGD); 2 ex., Sumatra, Pangherang-Pisang, Oct. 1890 – Mar. 1891, E. Modigliani (MCGD); 1 ex., Sumatra, St. Rambe, Dec. 1890 – Mar. 1891, E. Modigliani (MCGD); 2 ex., Sumatra, Korinchi Peak, 7300 ft, 1 ° 35 ' S / 103 ° 36 ' E, May 1914, Ex. F. M. S. Museum, B. M. 1955 – 354 (BMNH); 7 ex., Sumatra, Si-Rambe, 6 ° 11 ’ S / 106 ° 48 ’ E, Dec. 90 – Mar. 91, E. Modigliani (BMNH; IRSN, MCGD); 2 ex., Sumatra, Manna, 4 ° 27 ' S / 102 ° 59 ' E, 1902, M. Knappert, coll. Veth (NNML); 1 ex., N. Sumatra, Bivouac Two, Mt. Bandahara, no 24 ca 1430 m, submontane multistratal evergreen forest, 3 ° 44 ' N / 97 ° 43 ' E, 5 – 10 Jul. 1972, J. Krikken, at light (NNML); 1 ex., Sumatra Barat, Mt. Singgalang, ca 15 km Bukit Tinggi, 2100 m, 0 ° 24 ' S / 100 ° 21 ' E, 12 Feb. 1996, leg. C. Zorn (CJB); 1 ex., W. Sumatra, Bengkulu prov., nr. Curup, Bukit Kaba Mt., 1000 – 1500 m, 3 ° 29 ' S / 102 ° 36 ' E, 30 Jan. – 3 Feb. 2000, J. Bezdek leg. (CJB). Total length. — 5.00 – 5.25 mm (mean: 5.11 mm, n = 8) Head. — Reddish-brown, impunctate. First and second basal antennomeres contrasting brown-reddish, third to outer antennomeres black (Figs. 10, 15). Ratio length of second to third antennomeres 0.75 – 0.88 (mean: 0.79); median to terminal antennomeres are slender, ratio length of third to fourth antennomere 0.40 – 0.44 (mean: 0.41; Fig. 11). Thorax. — Pronotum reddish-brown. Pronotal width 1.30 – 1.45 mm (mean: 1.40 mm), ratio length to width 0.59 – 0.62 (mean: 0.61). Elytra black at base, a broad yellowish spot occupies the anterior half of the disc, extending nearly to either the outer margin, the anterior edge of this spot is obliquely rounded near the suture, another smaller and triangular spot is placed near the apex of each elytron (Figs. 10, 15). Elytral length 4.00 – 4.25 mm (mean: 4.15 mm), maximal width of both elytra together 2.60 – 3.00 mm (mean: 2.84 mm), ratio of maximal width of both elytra together to length of elytra 0.63 – 0.70 (mean: 0.68). Coxa and femur reddish-brown, tibia and tarsus black. Abdomen. — Pale yellow to reddish brown. Male genitalia. — Median lobe not incised, apex carinated ventrally in the middle. Tectum narrow, became pointed at apex. Endophallus consists of only median spiculae; long, slender and straight. Stronger sclerotized ventral spur with apical hook occurred at the apical first of the median lobe. Several basal endophallus structure at the bottom, close to sacculus (Fig. 12). Female genitalia. — Like description of the genus (Figs. 13, 14).	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A0360CFFC030B7FED4FA104A31.taxon	diagnosis	Diagnosis. — Median antennomeres of N. quadriplagiata are rather slender while N. sumatrensis are insignificantly widened. The median lobe of N. quadriplagiata (Fig. 12) is quite similar to N. sumatrensis, but the latter has a broadened tectum (Fig. 4). Elytron always with two yellowish spots.	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A03602FFC130F1FEF5FB244A10.taxon	type_taxon	Type species. — Ochralea marginata Jacoby, 1884: 55 – 56; herein designated.	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A03602FFC130F1FEF5FB244A10.taxon	etymology	Etymology. — Combination of para (latin: next to) and Neolepta; gender: feminine. Total length. — 6.65 – 10.10 mm (mean: 7.43 mm) Head. — Pale yellow to reddish-brown. Impunctate, with significant transverse impression between posterior third of eyes. Eyes large, convex. Labial palpi and maxillary palpi slender (Fig. 16 a). Labrum yellow to brown and black at apex. Mandibulae brownish. Antennae slender, entirely black and only the first basal antennomeres contrasting reddishbrown, extended to the apical third of the elytra (Figs. 17, 25, 31). Third to terminal antennomere with very dense and rough cover of partly bristle-like setae. First antennomere club shaped; second antennomere a bit shorter than third antennomere; ratio length of second to third antennomere 0.75 – 0.86 (mean: 0.81); ratio length of third to fourth antennomere 0.40 – 0.50 (mean: 0.45; Figs. 18, 26, 32). Thorax. — Pronotum transverse, lateral and posterior margin evenly rounded, the anterior angles slightly thickened (Figs. 17, 25, 31), often with transverse depression beyond the middle of the base. Pronotum pale yellow to reddishbrown. Pronotal width 1.65 – 2.70 mm (mean: 2.14 mm), ratio length to width 0.60 – 0.63 (mean: 0.62). Scutellum triangular, impunctate, yellow to black. Procoxal cavities nearly closed (Fig. 16 b). Meso- and metathorax yellowish to blackish. Mesosternum broad (Fig. 16 c). Elytra strongly punctuated, punctuation slightly coarser and denser than that of pronotum, entirely yellow to brown, sutural and lateral margin narrowly black. Elytral length 4.60 – 8.10 mm (mean: 5.90 mm), maximal width of both elytra together 3.20 – 5.40 mm (mean: 4.14 mm), ratio of maximal width of both elytra together to length of elytra 0.65 – 0.74 (mean: 0.71). Alae fully developed (Fig. 16 d). Legs long and slender, basi-metatarsus elongated (Fig. 16 e – g), yellow to brown, tibiae and tarsi blackish. In particular tibia with dense and fine setae. Abdomen. — Pale yellow to brown. Last visible sternite in females rounded at apex (Fig. 16 h) and in males with two deep, parallel sided incisions (Fig. 16 i). Male genitalia. — The median lobe symmetrical and usually insignificantly narrowed towards apex, not incised apically. Tectum broad at base, vary from long to short and became lanceolate at apex (Fig. 19 a). The endophallus consist of a bundle of laterally arranged long, slender and straight median spiculae. At the bottom of median spiculae, with distinct endophallus structure. Sacculus clearly visible (Figs. 19, 27, 33). Female genitalia. — Spermatheca with slender and elongated nodulus. Middle part long and slightly curved, cornu long, curved, arrow-like widened beyond apex (Figs. 20, 28, 34), with a pair of sclerotized bursa sclerites (Figs. 21, 29, 35).	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A03602FFC130F1FEF5FB244A10.taxon	distribution	Distribution. — The genus is restricted to South-East Asia and up to now only recorded from Brunei, Malaysia (Peninsular) and Indonesia (Borneo, Sumatra), southwards to Java (Fig. 22).	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A03602FFC130F1FEF5FB244A10.taxon	diagnosis	Diagnosis. — Paraneolepta, new genus are relatively large galerucines with a strongly elongated basi-metatarsus, pale yellow to reddish-brown head and pronotum and usually contrasting black antennae with slender antennomeres. The pronotum is coarsely punctuated and a transverse depression beyond the middle of the base occurred. The dorsal colouration is without spots and bands, entirely yellow to brown, but the elytral suture and the elytral lateral margin are often narrowly black lined (Figs. 17, 25, 31). Paraneolepta, new genus resembles Neolepta and Orthoneolepta, new genus. Among these three groups, Paraneolepta, new genus is largest in term of the body length (6.65 – 10.10 mm; Neolepta 4.85 – 5.75 mm; Orthoneolepta, new genus 4.50 – 5.75 mm). The antennomeres of Paraneolepta, new genus are slender while in Neolepta, the median antennomeres are insignificantly widened, and they are strongly widened in Orthoneolepta, new genus. The second antennomere of Paraneolepta, new genus is somewhat shorter than third antennomere (ratio length of second to third antennomere: 0.75 – 0.86; Figs. 18, 26, 32), similar to Neolepta (0.75 – 0.80; Figs. 3, 11) while in Orthoneolepta, new genus varied from short to more or less the same length (0.60 – 1.00; Figs. 39, 46). Third to terminal antennomere densely covered by bristle-like setae and extended to the apical third of elytra in these three groups. The pronotum of Paraneolepta, new genus is transverse and comparatively less broad (0.60 – 0.63), like in Orthoneolepta, new genus (0.59 – 0.65) compared to Neolepta (0.52 – 0.60). The depression beyond the middle of the base occurrs in these three groups, but the punctuation is coarsest in Paraneolepta, new genus. Besides many similarities among these three groups, they can be clearly distinguished by the genital characters in both sexes. The median lobe of Paraneolepta, new genus are without apical incision (Figs. 19, 27, 33) as in Neolepta, but the latter is with stronger sclerotized ventral spur and carinated ventrally (Figs. 4, 12). In comparison with Orthoneolepta, new genus, the median lobe has a deep incision apically (Figs. 40, 47). The endophallus consist of a bundle of laterally arranged long, slender and straight median spiculae in these three groups. The median lobe of Paraneolepta, new genus is bigger and longer than in Neolepta, and narrower towards the apex. The spermatheca is the same shape for Paraneolepta, new genus (Figs. 20, 28, 34) and Orthoneolepta, new genus (Figs. 41, 48). Spermatheca with small, slender and elongated nodulus and middle part slightly curved. For Neolepta, the nodulus is rounded and usually possess a strongly curved cornu (Figs. 5, 13). Bursa sclerites showed a different shape (Figs. 6, 14, 21, 29, 35, 42, 49).	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A03603FFC4308BFE15FDB24A30.taxon	description	(Figs. 16 – 24)	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A03603FFC4308BFE15FDB24A30.taxon	materials_examined	Type material. — Ochralea marginata: Lectotype, Male, “ Soerol., 4.18 / Ochralea marginata Jac. “ (NNML; Fig. 23). Type locality: Indonesia, Soerol, 0 ° 35 ’ S / 101 ° 20 ’ E. Jacoby (1884) stated that there are at least two specimens when he described the species. Thus we designate a lectotype to fix the specimen that available to us. Luperodes marginata: Holotype, “ Suma. / Luperodes marginata / Type / Luperodes marginata Baly, Type “ (BMNH; Fig. 24). Type locality: Indonesia, Sumatra. Baly gave no number on the specimens he studied, but there are three specimens available, and only one of them carries a label with “ type ”, which can be treated as holotype by indication. – Paratypes, 2 ex., Sumatra, Baly coll. (BMNH) Further material examined. — Indonesia. 16 ex., Sumatra, Museum Leiden, ex. collection, J. J. de Vos tot, Nederveen Cappel (NNML); 15 ex., Sumatra, Palembang, 2 ° 59 ' S / 104 ° 45 ' E, M. Knappert, coll. Vth. (NNML); 5 ex., Dr. B. Hagen, Tandjong Morawa Serdang (N. O. Sumatra), 0 ° 35 ' S / 101 ° 18 ' E (NNML); 4 ex., N. E. Sumatra, Deli, Seleleh, Kuala Limpang, Medang Ara State, 3 ° 34 ' N / 98 ° 40 ' E, Mar. 1954, A. Sollaart, Lowland Forest (NNML); 2 ex., Sumatra, S. E. coast, Laut Tador, 90 m, 3 ° 32 ' N / 99 ° 04 ' E, 2 / 5 Aug. 1950, E. Straatman leg. (NNML); 1 ex., Java, Wallace, 6 ° 23 ' S / 106 ° 48 ' E, 67.56 (NNML); 1 ex., marginata Jac., Jacoby coll. 1909 - 28 a (BMNH). Total length. — 7.10 – 10.10 mm (mean: 8.45 mm; n = 10) Head. — Reddish-brown, impunctate. Antennae pale brown (Fig. 17). Ratio length of second to third antennomere 0.83 – 0.86 (mean: 0.85); slender, ratio length of third to fourth antennomere 0.43 – 0.50 (mean: 0.47; Fig. 18). Thorax. — Pronotum pale brown to reddish-brown, finely punctuated. Pronotal width 2.25 – 2.70 mm (mean: 2.46 mm), ratio length to width 0.60 – 0.63 (mean: 0.61). Scutellum black. Elytra entirely yellow to brownish, sutural and lateral margin narrowly black. Elytral length 1.40 – 1.65 mm (mean: 1.51 mm), maximal width of both elytra together 2.25 – 2.70 mm (mean: 2.46 mm), ratio of maximal width of both elytra together to length of elytra 0.65 – 0.70 (mean: 0.67). Legs entirely pale yellow to brownish. Abdomen. — Pale yellow to brown. Male genitalia. — Median lobe is broad at middle and slightly narrowed towards apex and not incised apically. Tectum broadened at apex and with a fine, pointed tip (Fig. 19 a). Endophallus consist of a bundle of laterally arranged long, slender and straight median spiculae. At the bottom of the median spiculae, several basal endophallus structures. Sacculus clearly visible (Fig. 19). Female genitalia. — Spermatheca like description of the genus (Fig. 20), two pairs of bursa sclerites like Fig. 21.	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A03603FFC4308BFE15FDB24A30.taxon	distribution	Distribution. — This species was often collected at Sumatra Island, and occurred also in Java (Fig. 22).	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A03603FFC4308BFE15FDB24A30.taxon	diagnosis	Diagnosis. — Paraneolepta marginata is on average the largest species of this group (7.10 – 10.10 mm). The antennae are slender and entirely pale yellow to brown. The body colouration is brownish with narrowly black suture, and the punctuation of the pronotum is coarse. The median lobe is larger and broad from base to apex, the tectum is also broadened at the tip. The spermatheca is big and two pairs of bursa sclerites evenly sclerotised.	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A03606FFC5339AFEF5FCF44A10.taxon	description	(Figs. 25 – 30)	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A03606FFC5339AFEF5FCF44A10.taxon	materials_examined	Type material. — Holotype, “ Luperodes limbella / Singa. / Type / Luperodes limbella Baly type / Baly coll. “ (BMNH; Fig. 30). Type locality: Singapore, 2 ° 38 ' N / 103 ° 39 ' E. Baly gave no number on the specimens he studied, but there are three specimens available, and only one of them carries a label with “ type ”, which can be treated as holotype by indication. Further material examined. — Brunei. 5 ex., Temburong District, Ridge NE of Kuala Belalong, 4 ° 37 ’ N / 115 ° 8 ’ E, 300 m, Oct. 1992, 125 W MV Light Trap, J. H. Martin (BMNH); 1 ex., Labi, Bukit Teraja 60 m, Mxt. Dipt. Forest, 4 ° 25 ' N / 114 ° 27 ' E, B. M. 1983 – 39, Light trap 3, 25 m above ground, 23 Aug. 1979, S. L. Suttor (BMNH). – Malaysia. 6 ex., Sarawak, Kapit dist., Sebong, Baleh riv., 1 ° 54 ' N / 113 ° 38 ' E, 9 – 21 Mar. 1994, J. Horak leg. (CJB); 3 ex., Sarawak, Mt. Matang, 1.55 ° N / 110.35 ° E, 13 Dec. – 14 Jan. 1914, G. E. Bryant coll. 1919 - 147 (BMNH); 7 ex., Quop, W. Sarawak, 1 ° 33 ' N / 101 ° 24 ' E, G. E. Bryant, Mar. – Apr. 1914 (BMNH); 8 ex., N. Borneo, Samawang, Nr. Sandakan, jungle, 5 ° 55 ' N / 117 ° 46 ' E, 13 Jul. 1927, C. B. K & H. M. P., F. M. S. Museum (BMNH); 2 ex., N. Borneo, Bettotan, Nr. Sandakan, 5 ° 47 ' N / 117 ° 52 ' E, 2 Aug. 1927, C. B. K. & H. M. P., F. M. S. Museum (BMNH); 6 ex., W. Sarawak, Lundu, 1 ° 40 ' N / 109 ° 48 ' E, Jan. 1914, G. E. Bryant (BMNH); 2 ex., Sarawak, 2 ° 30 ' N / 113 ° 15 ' E, 1907 – 1909, C. J. Brooks, B. M. 1936 - 681, Wallace, Baly coll., (BMNH); 1 ex., Sarawak, Bau, 1 ° 25 ' N / 110 ° 9 ' E, 18 Jul. – 1 Aug. 1909, C. J. Brooks, B. M. 1936 - 681 (BMNH); 1 ex., Fed. Malay State, 1909, C. J. Brooks, B. M. 1931 - 570 (BMNH); 5 ex., Sabah, Lembah Danum, 5 ° 08 ' N / 117 ° 24 ' E, 3 – 5 Dec. 1916, 8 Nov. 1994, 27 – 31 Aug. 1991, Salleh, Zaidi, Mail, Lan (BMNH); 1 ex., N. Sembilan, Gemencheh, 2 ° 35 ' N / 102 ° 24 ' E, 4 – 6 Aug. 1990, Zabidi (UKM); 4 ex., Sabah, Danum Valley, 5 ° 08 ' N / 117 ° 24 ' E, 6 – 15 May 2007, B. H. Izfa leg. (UKM); 1 ex., Sarawak, Lanjak Entimau, 1 ° 11 ' N / 111 ° 51 ' E, 28 – 29 Feb. 1992, Zaidi (UKM); 1 ex., Sandakan, Borneo, 5 ° 50 ' N / 118 ° 3 ' E, Baker (NHRS). Total length. — 6.00 – 7.50 mm (mean: 6.81 mm; n = 10) Head. — Pale yellow to brown, impunctate. Antennae entirely black and only the first basal antennomeres usually contrasting pale yellow to brown (Fig. 25). Ratio length of second to third antennomere 0.75 – 0.83 (mean: 0.79); ratio length of third to fourth antennomere 0.40 – 0.50 (mean: 0.44; Fig. 26). Thorax. — Pronotum pale yellow to brown. Pronotal width 1.65 – 2.35 mm (mean: 1.99 mm), ratio length to width 0.60 – 0.63 (mean: 0.62). Scutellum usually black, but at least in 30 % of the specimens examined, scutellum are yellowish and in that case, the elytra are entirely yellow to brown without further narrowly black sutural as in the other specimens, with sutural and lateral margin narrowly black. Elytral length 4.60 – 5.90 mm (mean: 5.25 mm), maximal width of both elytra together 3.20 – 4.30 mm (mean: 3.79 mm), ratio of maximal width of both elytra together to length of elytra 0.70 – 0.74 (mean: 0.72). Legs brownish, tibiae and tarsi blackish. Abdomen. — Yellow to brown. Male genitalia. — Median lobe symmetrical, parallel sided, insignificantly narrow towards apex, not incised apically. Tectum short, broad at base, and became lanceolate at apex (Fig. 27 a). Endophallus consist of a bundle of laterally arranged long, slender and straight median spiculae. At the bottom of median spiculae, several basal endophallus structure. Sacculus clearly visible (Fig. 27). Female genitalia. — Spermatheca like description of the genus (Fig. 28), one pair of bursa sclerites like Fig. 29.	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A03606FFC5339AFEF5FCF44A10.taxon	distribution	Distribution. — Recorded from Borneo, Brunei, Peninsular Malaysia (Fig. 22).	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A03606FFC5339AFEF5FCF44A10.taxon	diagnosis	Diagnosis. — Paraneolepta limbella is the most similar in body outline and general colouration to P. imitans. The differences on both species can be obtained by genital structures. The median lobe of P. limbella is much narrowed towards the apex (Fig. 27), while for P. imitans, the median lobe is much broader at the apex (Fig. 33). The spermatheca of both species are similar, but only one pair of bursa sclerites in P. limbella (Fig. 29), while two pairs in P. imitans (Fig. 35).	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A03607FFC73031FED5FA224BD3.taxon	description	(Figs. 31 – 36)	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A03607FFC73031FED5FA224BD3.taxon	discussion	Remarks. — Monolepta aemula is a replacement name. Weise (1922) transferred the species to Monolepta and named it new to avoid the homonymy with Monolepta imitans Jacoby, 1888 from Central America.	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A03607FFC73031FED5FA224BD3.taxon	materials_examined	Type material. — Lectotype, female, “ Type H. T / Perak / Ochralea imitans Jac. / Jacoby coll. 1909 - 28 a, ” (BMNH; Fig. 36). Type locality: Malaysia, Perak, 4 ° 48 ' N / 101 ° 09 ' E. Jacoby gave no number on the specimens he studied, but there must be more than one concerning the original description, and we herein designate a lectotype to fix the name on single specimen. Further material examined. — Indonesia. 1 ex., N. O. Sumatra, Tebing Tinggi, 3 ° 19 ' N / 99 ° 09 ' E, Dr. Schulteiss, coll. Kraatz (MNHU); 1 ex., Dinding Island, 2 ° 30 ' N / 107 ° 35 ' E, H. N. Ridley, 1900 – 111 (BMNH); 1 ex., Sipora Island, West Sumatra, 2 ° 13 ' S / 99 ° 40 ' E, Oct. 1924, C. B. K. and N. S. (BMNH); 4 ex., North East Sumatra Deli, Kuala Simpang, Medang Ara Estate, lowland forest, 3 ° 34 ' N / 98 ° 40 ' E, Feb. – Mar. 1954, A. Sollaart (NNML). – Malaysia. 6 ex., Malay Penin., Selangor, Bukit Kutu, Foot of hill, 3500 ft., 5500 ft., 3 ° 33 ' N / 101 ° 43 ' E, Apr. 1915, 5 – 10 Sep. 1929, H. M. Pendlebury (BMNH); 6 ex., Malaya, Doherty, Perak, 4 ° 48 ' N / 101 ° 09 ' E, Fry coll. 1905 – 100 (BMNH); 1 ex., Malay Penin., Pahang, F. M. S, Batu Balai Jerantut, 3 ° 56 ' N / 102 ° 22 ' E, 19 Mar. 1927 (BMNH); 1 ex., Sarawak, Bau, 1 ° 25 ' N / 110 ° 09 ' E, 27 Sep. – 15 Oct. 1909, C. J. Brooks, B. M. 1936 – 681 (BMNH); 1 ex., Perak, F. M. S. Larut Hills, 3700 – 4000 ft., 5 ° N / 100 ° 53 ' E, 11 Feb. 1932, H. M. Pendlebury (BMNH); 1 ex., Baly coll. (BMNH); 1 ex., Lundu, West Sarawak, 1 ° 40 ' N / 109 ° 48 ' E, Jan. 1914, G. E. Bryant (BMNH); 2 ex., Malay Penin., Pahang, F. M. S., Fraser Hill, 4200 ft., 5 ° N' 100 ° 53 ' E, 22 Jul. 1936 (BMNH); 1 ex., Sandakan, Borneo, 5 ° 50 ' N / 118 ° 03 ' E, Baker (NHRS); 1 ex., Perak, Temenggor, 5 ° 19 ' N / 101 ° 22 ' E, Ekspedisi MNS-Belum, 29 – 30 Jan. 1994, Salleh & Ismail (UKM); 2 ex., Kelantan, Pasir Putih, J. Linang, 5 ° 49 ' N / 102 ° 22 ' E, 28 May 1994, Ismail & Zabidi (UKM); 1 ex., N. Sembilan, K. Kelawang, 2 ° 56 ' N / 102 ° 05 ' E, 16 Jun. 1987, Salleh & Ismail (UKM); 1 ex., Kedah, Sik, Hutan Lipur Lata Mengkuang, 5 ° 48 ' N / 100 ° 44 ' E, 15 Jun. 1994, Ismail, Ruslan, Yusof (UKM). Total length. — 6.65 – 8.00 mm (mean: 7.03 mm; n = 10) Head. — Reddish-brown, impunctate. Antennae entirely black and only the first basal antennomeres usually contrasting reddish-brown (Fig. 31). Ratio length of second to third antennomere 0.80 – 0.83 (mean: 0.81); ratio length of third to fourth antennomere 0.42 – 0.46 (mean: 0.44; Fig. 32). Thorax. — Pronotum reddish-brown, coarsely punctuated. Pronotal width 1.80 – 2.25 mm (mean: 1.98 mm), ratio length to width 0.60 – 0.63 (mean: 0.61). Scutellum black. Elytra entirely yellow to brown, sutural and lateral margin narrowly black. Elytral length 5.00 – 6.25 mm (mean: 5.51 mm), maximal width of both elytra together 3.60 – 4.50 mm (mean: 3.95 mm), ratio of maximal width of both elytra together to length of elytra 0.69 – 0.74 (mean: 0.72). Legs yellow to brown, black line along dorsal part of femur, tibiae and tarsi blackish. Abdomen. — Yellow to pale brown. Male genitalia. — Median lobe insignificantly narrowed towards apex, where it is blunt (Fig. 33 a). Tectum parallelsided, with short, pointed tip, much shorter than the ventral part (Fig. 33 b). Female genitalia. — Spermatheca like description of the genus (Fig. 34), two pairs of bursa sclerites like Fig. 35.	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A03607FFC73031FED5FA224BD3.taxon	distribution	Distribution. — Recorded from Peninsular Malaysia, Borneo and Sumatra (Fig. 22).	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A03607FFC73031FED5FA224BD3.taxon	diagnosis	Diagnosis. — The body size and colouration of P. imitans is most similar to P. limbella. The differences on both species can be obtained clearly by median lobe and bursa sclerites. The tectum of median lobe of P. limbella is shorter and narrowed towards the pointed tip (Fig. 27) while in P. imitans is broader at base, and pointed at the tip of it (Fig. 33). Two pairs of bursa sclerites present in P. imitans (Fig. 35) and only one pair of bursa sclerites in P. limbella (Fig. 29).	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A03605FFD930DDFE94FAD14AD0.taxon	type_taxon	Type species. — Neolepta fulvipennis Jacoby, 1884: 222 – 223; herein designated.	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A03605FFD930DDFE94FAD14AD0.taxon	etymology	Etymology. — Combination of ortho (latin: straight to) and Neolepta; gender: feminine. Total length. — 4.50 – 5.75 mm (mean: 5.12 mm) Head. — Brown to reddish-brown. Impunctate, with significant transverse impression between posterior third of eyes. Eyes small, convex. Labial palpi slender and maxillary palpi enlarged (Fig. 37 a), occasionally dark brown. Labrum and mandibulae contrasting dark brown. Antennae elongated, extended to the apical third of the elytra (Figs. 38, 45), the median antennomeres significantly broadened. Antennae entirely black and only the first basal antennomeres usually contrasting reddish-brown. First antennomere club shaped; second and third antennomere varied from a bit shorter to more or less the same length; ratio length of second to third antennomere 0.60 – 1.00 (mean: 0.79); ratio length of third to fourth antennomere 0.30 – 0.50 (mean: 0.42; Figs. 39, 46). Thorax. — Pronotum transverse, broad, with a transverse depression beyond the middle of the base, anterior margin concave and posterior margin broadly rounded (Figs. 39, 46). Pronotum brown to reddish-brown. Pronotal width 1.30 – 1.80 mm (mean: 1.56 mm), ratio length to width 0.59 – 0.65 (mean: 0.62). Scutellum triangular, impunctate, reddish-brown to black. Procoxal cavities nearly closed (Fig. 37 b). Meso- and metathorax reddish-brown to black. Metasternum broad (Fig. 37 c). Elytral strongly punctuated, punctuation partly arranged in longitudinal rows, entirely yellow to reddish-brown and in N. banggiensis, sutural and lateral margin narrowly black. Elytral length 3.50 – 4.80 mm (mean: 4.16 mm), maximal width of both elytra together 2.50 – 3.40 mm (mean: 2.92 mm), ratio of maximal width of both elytra together to length of elytra 0.67 – 0.72 (mean: 0.70). Alae fully developed (Fig. 37 d). Legs long and slender, basi-metatarsus elongated (Fig. 37 e – g), blackish throughout and in O. fulvipennis, coxa and trochanter contrasting brown to reddish-brown. In particular tibia with dense and fine setae. Abdomen. — Pale yellow to red. Last visible sternite in females rounded at apex (Fig. 37 h), and in males with two deep, parallel-sided incisions (Fig. 37 i). Male genitalia. — The median lobe is symmetrical, parallelsided at base and usually narrowed towards apex. Apically incised. Tectum long, broad at base and became pointed at apex. As Neolepta, endophallus consist of bundle of laterally arranged long, slender and straight median spiculae. At the bottom of median spiculae, several basal endophallus structure. Sacculus clearly visible (Figs. 40, 47). Female genitalia. — Spermatheca with small, slender and elongated nodulus. Middle part long and slightly curved, cornu long and curved (Figs. 41, 48). Two pairs of bursa sclerites; one spiny and sclerotized and the other pair hooked shaped, attached together (Figs. 42, 49).	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A03605FFD930DDFE94FAD14AD0.taxon	distribution	Distribution. — Restricted to the Malaysian Peninsula, Sumatra and Borneo (Fig. 43).	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A03605FFD930DDFE94FAD14AD0.taxon	diagnosis	Diagnosis. — Orthoneolepta, new genus are medium sized (4.50 – 5.75 mm), dorsoventrally bulged galerucine with strongly elongated basi-metatarsus and transverse depression on pronotum. The punctuation on elytra is coarser than that of pronotum. The dorsal colouration is yellow to reddishbrown, and in certain species, sutural and lateral margin of elytra are narrowly black (Figs. 38, 45). As the genotype of this genus O. fulvipennis are transferred from Neolepta, most of the character of this genus resembles to Neolepta. The pronotum of both groups are with transverse depression, but it is comparatively less broad (0.59 – 0.65) in Orthoneolepta, new genus than in Neolepta (0.52 – 0.60). The second and third antennomere of Orthoneolepta, new genus varied from a bit shorter to more or less the same length (ratio length of second to third antennomere: 0.60 – 1.00; Figs. 39, 46), about similar to Neolepta, of which the second antennomere is a bit shorter (0.75 – 0.80; Figs. 3, 11), but in Orthoneolepta, new genus, the fourth to sixth antennomere are significantly widened. The procoxal cavities are nearly closed in both groups (Figs. 1 b, 37 b). Besides the similarities, the differences in both group can be obtained on the genital characters. The median lobe are incised apically in Orthoneolepta, new genus (Figs. 40, 47) and not incised in Neolepta and Paraneolepta, new genus (Figs. 4, 12, 19, 27, 33). The spermatheca are the same shape with Paraneolepta, new genus, but two pairs of bursa sclerites; one spiny and sclerotized and one other pair of hook-shaped occurred in Orthoneolepta, new genus and very characteristic for this group (Figs. 42, 49). At the first glance, O. fulvipennis looks very similar to Rubrarcastes sanguinea (Jacoby, 1892; Hazmi & Wagner, 2010 b) with the punctuation coarser and reddish-brown dorsal colouration. Both species can be misidentified at a first glance, but the crucial characters that can distinguish both species are the second and third antennomere of Rubrarcastes (0.43 – 0.57) are not of the same length as in Orthoneolepta, new genus (0.60 – 1.00). The median lobes of these two groups are also different — in Orthoneolepta, new genus, they are incised apically and spermathecae are also of different shape.	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A0361BFFDC3085FDD5FAA24A70.taxon	description	(Figs. 37 – 44)	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A0361BFFDC3085FDD5FAA24A70.taxon	materials_examined	Type material. — Lectotype, “ Neolepta fulvipennis Jac. / Dr. B. Hagen. Tandjong Morawa Serdang (N. O. Sumatra) / Baly coll. ” (BMNH; Fig. 44). Type locality: Indonesia, Sumatra 0 ° 35 ' S / 101 ° 18 ' E. Jacoby gave no number of the specimens he studied, but there are several specimens from the type locality available, and thus we herein designate a lectotype to fix the name on single specimen. – Paralectotypes, 5 ex., same data as lectotype (NNML). – Invalid types: 1 ex., “ Soekaranda, 0 ° 37 ' S / 104 ° 29 ' E, Jan. 1894, Dohrn, 71490 ”, and 1 ex., “ Dohrn, Sumatra, Lianggas, 0 ° 37 ' S / 94 ° 29 ' E, 71491, Arcastes sanguinea ” in MNHU are both labelled as co-type, but are not from the type series. Further material examined. — Indonesia. 1 ex., Medan, Sumatra, 3 ° 30 ' N / 98 ° 37 ' E, J. J. D. V. Museum Leiden ex. Collection J. J. de Vos tot Nederveen Cappel (NNML); 1 ex., N. E. Sumatra, Deli, Kuala Simpang, Semadan Estate, 3 ° 34 ’ N / 98 ° 40 ’ E, Nov. 1954, A. Sollaart, lowland forest (NNML); 1 ex., Soekaranda, 0 ° 37 ' S / 104 ° 29 ' E, J. V. Hasselt (NNML); 6 ex., Sumatra, Forides, Baly coll. (BMNH); 3 ex., Soekaranda, 0 ° 37 ' S / 104 ° 29 ' E, Jan. 1894, Dohrn, 380., Arcastes sanguinea Jac. (MNHU). – Malaysia. 2 ex., Borneo, Sarawak, 1865 – 1866, coll. G. Doria (MCGD); 6 ex., Mt. Matang, W. Sarawak, 1.55 ° N / 110.35 ° E, Dec. 1913, Jan. 1913, Mar. 1914, G. Bryant coll. 1919 – 147 (BMNH); 2 ex., Quop, West Sarawak, 1 ° 33 ' N / 101 ° 24 ' E, Mar. 1914, G. Bryant coll. 1919 – 147 (BMNH); 2 ex., Borneo, Mahakam (BMNH); 3 ex., Sarawak, 2 ° 30 ' N / 113 ° 15 ' E, C. J. Brooks, B. M. 1928 – 193 (BMNH); 3 ex., Fed. Malay States: 3 ° 08 ' N / 101 ° 42 ' E, 1909, C. J. Brooks, B. M. 1931 – 570 (BMNH); 1 ex., Sarawak, Matang, 4 1 / 2 miles, 1 ° 32 ' N / 110 ° 15 ' E, 6 Jun. 1909, J. E. A. Lewis 1910 – 116 (BMNH); 8 ex., Borneo, Banja, 2 ° 30 ' N / 113 ° 15 ' E, German mission, Fry coll. 1909.100 (BMNH); 1 ex., Long Navang, Borneo, 2 ° 30 ' N / 113 ° 15 ' E, Mjoberg (NHRS); 1 ex., Taman Negara NP Kuala Tahan, primaval forests, 4 ° 19 ' N / 102 ° 20 ' E, 5 – 9 Mar. 2007, V. Hula, Puchart, L. Ruzicka, F. (CJB). Total length. — 4.75 – 5.60 mm (mean: 5.20 mm; n = 10) Head. — Brown to reddish-brown. Impunctate. First basal antennomeres contrasting reddish-brown, second to outer antennomere usually black (Fig. 38). Ratio length of second to third antennomere 0.60 – 0.75 (mean: 0.73); widened, insignificantly enlarged, ratio length of third to fourth antennomere 0.38 – 0.50 (mean: 0.45; Fig. 39). Thorax. — Pronotum brown to reddish-brown. Pronotal width 1.50 – 1.70 mm (mean: 1.57 mm), ratio length to width 0.59 – 0.63 (mean: 0.61). Scutellum brown to reddish-brown. Elytral entirely brown to reddish-brown (Fig. 38), coarsely punctuated, punctuation partly arranged in longitudinal rows. Elytral length 3.90 – 4.60 mm (mean: 4.27 mm), maximal width of both elytra together 2.70 – 3.30 mm (mean: 2.95 mm), ratio of maximal width of both elytra together to length of elytra 0.67 – 0.71 (mean: 0.69). Legs blackish, except coxa and trochanter contrasting brown to reddish-brown. In particular tibia with dense and fine setae. Abdomen. — Brown to red. Male genitalia. — Median lobe broad, incised apically. Tectum long, broad and became pointed at apex. Median spiculae consist of laterally arranged slender structures. At the bottom of median lobe, several basal endophallus structure of plate like structured present, and sacculus can visibly seen (Fig. 40). Female genitalia. — Spermatheca with small, slender and elongated nodulus. Middle part long and slightly curved, cornu long and curved (Fig. 41). Two pairs of bursa sclerites; one spiny and sclerotized and one other pair of hook-shaped, bigger in size (Fig. 42).	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A0361BFFDC3085FDD5FAA24A70.taxon	distribution	Distribution. — Distributed throughout Borneo, Sumatra and Peninsular Malaysia (Fig. 43).	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A0361BFFDC3085FDD5FAA24A70.taxon	diagnosis	Diagnosis. — Orthoneolepta fulvipennis resembles Rubrarcastes sanguinea (Jacoby, 1892) from colouration and antennomeres (both possess widened medium antennomeres), but O. fulvipennis posses second and third antennomere almost the same length (ratio length of second to third antennomere 0.60 – 1.00) differed to R. sanguinea which third antennomere longer than second antennomere (0.43 – 0.57). Median lobe and spermatheca are different between these two species. From O. banggiensis, O. fulvipennis can be differentiated by the dorsal colouration. Elytral entirely brown to reddish-brown in O. fulvipennis (Fig. 38) while often the sutural and lateral margin narrowly black on yellow to dark brown elytra of O. banggiensis (Fig. 45).	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A0361EFFDD306CFE35FC3A4A51.taxon	description	(Figs. 45 – 50)	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A0361EFFDD306CFE35FC3A4A51.taxon	materials_examined	Type material. — Holotype, “ Holotype Neolepta banggiensis n. sp. des Mohamedsaid 1997 / Sabah: Pulau Banggi, 8 – 12 May 1996, Salleh, Zaidi, Ismail & Sham ” (UKM). – 5 Paratypes, same data as holotype (UKM; Fig. 50). Type locality: Malaysia, Sabah, 7 ° 16 ' N / 117 ° 09 ' E. Further material examined. — Malaysia. 2 ex., Sabah, Lembah Danum, 5 ° 08 ' N / 117 ° 24 ' E, 3 – 5 Dec. 1991, 17 – 20. Nov. 1994, Ismail, Sham & Ruslan (UKM); 3 ex., Sabah, Pulau Gaya, 6 ° 01 ' N / 116 ° 01 ' E, 26 – 30 Nov. 1991, Zaidi & S. Abin (UKM); 28 ex., N. Borneo, Kudat, 6 ° 49 ' N / 116 ° 43 ' E, 2 – 27 Aug. 1927, 1 – 18 Sep. 1927, Ex. F. M. S. Museum, B. M. 1955 – 354 (BMNH); 1 ex., Sabah, 5 m, S. Mt. Trus Madi, 1800 ft, 5 ° 33 ’ N / 116 ° 30 ’ E, 18 – 28 Aug. 1977, M. E. Bacchus, B. M. 1978 – 48 (BMNH); 6 ex., Borneo (BMNH); 3 ex., Sarawak, 2 ° 33 ' N / 113 ° E, C. J. Brooks, B. M. 1928 – 193 (BMNH); 1 ex., Lundu, West Sarawak, 1 ° 40 ' N / 109 ° 48 ' E, Jan. 1914, G. E. Bryant (BMNH); 3 ex., Borneo, Sandakan, 5 ° 50 ' N / 118 ° 03 ' E, Baker (NHRS). Total length. — 4.50 – 5.75 mm (mean: 5.03 mm; n = 10) Head. — Brown to reddish-brown. Impunctate. Antennae entirely black and only the first basal antennomeres usually contrasting brown (Fig. 45). Ratio of length of second to third antennomere 0.75 – 1.00 (mean: 0.86); ratio of length of third to fourth antennomere 0.30 – 0.50 (mean: 0.39; Fig. 46). Thorax. — Pronotum brown to reddish-brown. Pronotal width 1.30 – 1.80 mm (mean: 1.55), ratio length to width 0.61 – 0.65 (mean: 0.63). Scutellum black. Elytra entirely yellow to dark brown, sutural and lateral margin narrowly black (Fig. 45). Elytral length 3.50 – 4.80 mm (mean: 4.05 mm), maximal width of both elytra together 2.50 – 3.40 mm (mean: 2.88 mm), ratio of maximal width of both elytra together to length of elytra 0.68 – 0.72 (mean: 0.70). Legs black throughout. Abdomen. — Pale yellow to brown. Male genitalia. — Median lobe less broad than in O. fulvipennis and narrowed towards the apex. Apically incised. Tectum long and broadened at base and pointed at apex. Endophallus in O. fulvipennis consists of laterally arranged slender median spiculae. The basal endophallus structures are present (Fig. 47). Female genitalia. — Nodulus of spermatheca small, slender and elongated, middle part and cornu long and curved (Fig. 48). Two pairs of bursa sclerites; one spiny and sclerotized and one other pair of hook-shaped, bigger in size (Fig. 49).	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A0361EFFDD306CFE35FC3A4A51.taxon	distribution	Distribution. — Only known from northern Borneo (Fig. 43).	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
A62987A0361EFFDD306CFE35FC3A4A51.taxon	diagnosis	Diagnosis. — Orthoneolepta banggiensis can be differentiated from dorsal colouration with O. fulvipennis. The elytra of O. banggiensis are often with sutural and lateral margin narrowly black, and in O. banggiensis, the scutellum is black while brown to reddish-brown in O. fulvipennis (Figs. 38, 45). The median lobe is quite similar (Figs. 40, 47) and bursa sclerites of O. banggiensis are bit smaller than that of O. fulvipennis (Figs. 42, 49). The punctuation of O. fulvipennis is coarser.	en	Hazmi, Izfa Riza, Wagner, Thomas (2013): Revision Of Neolepta Jacoby, 1884 And Related Galerucines From The Oriental Region, Including Descriptions Of Two New Genera (Coleoptera: Chrysomelidae: Galerucinae). Raffles Bulletin of Zoology 61 (1): 73-95, DOI: 10.5281/zenodo.4508559
