taxonID	type	description	language	source
A554879758537E3AE78D2019FBDAFE6B.taxon	materials_examined	Type species: Myzocallis sawashibae (Matsumura, 1917), by original designation. Generic Diagnosis: According to Quednau (2003), Mesocallis belongs to the Pterocallis group of genera. This genus is similar to Pterocallis Passerini but can be distinguished by having two setae on ABD VIII, black-bordered fore tibiae and Cu 1 b of forewing distinctly dark bordered. Representatives of Mesocallis can be also distinguished from Pterocallis by oval to narrow elliptical secondary rhinaria on ANT III, epicranial suture weakly developed but present on the head of apterous viviparous females. Host plant: Species belonging to this genus are known to feed on Alnus spp., Carpinus spp., Corylus spp., Ostrya spp. (Betulaceae) and Fagus sp. (Fagaceae). Distribution: This genus is known from South- East Asian countries (China, India, Japan, Korea, Nepal, and Siberia).	en	Lee, Yerim, Kanturski, Mariusz, Lee, Seunghwan (2018): Agarna malayi Tiwari 1952. Zoological Studies 57 (17): 1-20, DOI: 10.6620/ZS.2018.57-17, URL: http://dx.doi.org/10.5281/zenodo.13316922
A554879758527E3BE7E423C8FB27FB0A.taxon	description	(Figs. 1, 2, 3 a-b, 5, 10; Table 1) urn: lsid: zoobank. org: act: 966 B 034 F- 6 ABD- 40 AA- 972 B- 4 FBDADB 1 ED 2 D Material examined: Holotype: 1 alate viviparous female, Jangsan-forest, Jangsan-ri, Maam-myeon, Goseong-gun, GN, South Korea, 35 ° 02 ' 48 " N, 128 ° 24 ' 44 " E, on Carpinus laxiflora, Yerim Lee and Hyoseok Lee leg., 3. v. 2015, no. 150503 YR- 11. Paratypes: 19 alate viviparous females, same data as the holotype, including DNA voucher no. Mes 1 - 2, Mes 1 - 3, Mes 1 - 4, Mes 1 - 5. Etymology: The species name, carpinicola, is derived from the genus name of the host plant (Carpinus) and the Latin suffix ‘ - cola’ (dweller, inhabitant). Diagnosis: This species is close to M. (M.) pteleae in having similar coloration and Cu 1 b of forewing dark bordered. However, the new species can be distinguished from M. (M.) pteleae by its shorter PT, 0.06 - 0.08 mm (0.08 - 0.10 mm in M. pteleae) and shorter URS length, 0.10 - 0.11 mm (0.11 - 0.14 mm in M. pteleae), ANT III with 4 - 5 setae (5 - 9 in M. pteleae), URS with 4 - 6 accessory setae (8 - 9 in M. pteleae), cauda with 10 - 15 setae (7 - 11 in M. pteleae). Description (Alate viviparous female): Color in life: Head pale yellow, head vertex and ANT I-III fuscous, distal 1 / 3 of ANT IV and ANT V darken, BASE and distal 1 / 2 of PT also darken. Thorax and abdomen pale yellow. Legs pale, fore and hind tibiae including 1 / 9 of femur and tarsus black, 1 / 9 of middle tibiae and tarsi dark. Wing vein Cu 1 b of forewing bordered with black pigment. SIPH pale. Cauda and anal plate pale. Morphology: Body oval, BL, 1.30 - 1.55 mm. Head smooth with a flat median protrusion on frons, head vertex with 0.01 mm pointed setae, epicranial suture weakly developed, head dorsum without tubercles. ANT 6 - segmented, 0.92 - 1.05 mm long, ANT 0.66 - 0.78 × BL, ANT III longest with 12 - 16 narrow transversely elliptical secondary rhinaria in a row on the whole segment, with 4 - 5 short and pointed setae, ANT III-VI imbricated with very short and inconspicuous setae, Ls ANT III × 0.50 BD III, ANT IV without secondary rhinaria, ANT V shorter than ANT IV, PT 0.60 - 0.80 × BASE. Rostrum not reaching to middle coxae, URS 0.10 - 0.11 mm long with 4 - 6 accessory setae, URS 1.00 - 1.22 × BASE, 1.22 - 1.38 × HT II. Thorax smooth, without tubercles. Fore coxae weakly enlarged, longest setae on TIBIAE III 0.67 - 1.00 × middle width of TIBIAE III, HT I with 7 chaetotaxy, HT II 0.08 - 0.09 mm long. Wing vein Rs of forewing weakly developed. Dorsal spinal setae sometimes on small elevations, conspicuous tubercles not developed, ABD VIII with a pair of spinal setae, ABD margin I-IV with a single seta on short cone-shaped marginal tubercles, 4 th marginal tubercle 0.03 mm. SIPH cylindrical, truncated, 0.05 - 0.06 mm long. Cauda knobbed, 0.07 - 0.09 mm long, bearing 10 - 15 setae. Anal plate bilobed, each lobe with 6 - 8 setae. Distribution: So far, the species is known from Goseong-gun in the southern part of South Korea. Host plant: The species feed on the underside of leaves of Carpinus laxiflora (Betulaceae). Remarks: This species was first referred to as Mesocallis sp. in Lee et al. (2017).	en	Lee, Yerim, Kanturski, Mariusz, Lee, Seunghwan (2018): Agarna malayi Tiwari 1952. Zoological Studies 57 (17): 1-20, DOI: 10.6620/ZS.2018.57-17, URL: http://dx.doi.org/10.5281/zenodo.13316922
A554879758527E3EE45A2598FB99FE0A.taxon	description	(Figs. 1, 2, 3 c-d, 6; Table 1)	en	Lee, Yerim, Kanturski, Mariusz, Lee, Seunghwan (2018): Agarna malayi Tiwari 1952. Zoological Studies 57 (17): 1-20, DOI: 10.6620/ZS.2018.57-17, URL: http://dx.doi.org/10.5281/zenodo.13316922
A554879758527E3EE45A2598FB99FE0A.taxon	materials_examined	Material examined: 3 alate viviparous females, Mt. Beagunsan, Gwangyang-si, JN, South Korea, 35 ° 06 ' 26 " N, 127 ° 37 ' 19 " E, on Carpinus sp., Yerim Lee leg., 25. vii. 2013, no. 130725 YR- 23, including DNA voucher no. Ca 2; 33 alate viviparous females, Mt. Chilgapsan, Cheongyang-gun, CN, South Korea, 36 ° 24 ' 48 " N, 126 ° 53 ' 03 " E, on Corylus sp., Yerim Lee and Hyoseok Lee leg., 10. v. 2014, no. 140510 YR- 1, including DNA voucher no. Co 10 - 1, Co 10 - 2, Co 10 - 3, Co 10 - 4, Co 20; 2 alate viviparous females, Mt. Chilgapsan, Cheongyang-gun, CN, South Korea, 36 ° 24 ' 48 " N, 126 ° 53 ' 03 " E, on Corylus sp., Yerim Lee and Hyoseok Lee leg., 6. v. 2016, no. 160506 YR- 2. Diagnosis: This species is similar to M. (M.) taoi Quednau by the color of ANT and black pigmented frons. However, it can be distinguished by its longer ANT III length, 0.33 - 0.43 mm (0.14 - 0.20 mm in M. taoi) and long and slender URS, 0.11 - 0.14 mm (0.06 - 0.07 mm in M. taoi). Redescription (Alate viviparous female): Color in life: Head pale, head vertex and ANT I-III fuscous, compound eye red, distal joint of ANT IV- V, BASE and distal 1 / 2 of PT also dark. Thorax and abdomen pale. Legs pale, fore tibiae including 1 / 9 of femur black, distal margin of FEMUR III dark, tarsus dark. Wing vein Cu 1 b of forewing dark bordered. SIPH and cauda pale. Morphology: Body oval, BL 1.51 - 1.76 mm long. Head smooth with a flat median protrusion on frons, head vertex with 0.01 mm pointed setae, epicranial suture developed, head dorsum without tubercles. ANT 6 - segmented, ANT 0.63 - 0.74 × BL, ANT III longest with 13 - 16 transversely elliptical secondary rhinaria in a row on the whole segment, ANT III with 8 - 10 short pointed setae, ANT III-VI imbricated, ANT IV without secondary rhinaria, ANT V shorter than ANT IV, BASE with a single inconspicuous seta, PT 0.71 - 0.91 × BASE. Rostrum not reaching to middle coxae, URS 0.11 - 0.14 mm long with 8 - 9 accessory setae, URS 0.86 - 1.27 × BASE, 1.22 - 1.50 × HT II. Thorax smooth, without tubercles. Fore coxae weakly enlarged, longest setae on TIBIAE III 1.00 - 1.05 × middle width of TIBIAE III, HT I with 7 chaetotaxy, HT II 0.08 - 0.10 mm long. Wing vein Rs of forewing weakly developed. Dorsal spinal setae sometimes on small elevations, conspicuous tubercles not developed, ABD VIII with a pair of spinal setae, rarely with 3 setae, ABD margin I-IV with a single seta on short cone-shaped tubercles, 4 th marginal tubercle 0.02 mm long. SIPH cylindrical, truncated, 0.03 - 0.06 mm long. Cauda knobbed, 0.09 - 0.11 mm long, bearing 8 - 11 setae. Anal plate bilobed, each lobe with 7 - 10 setae. Distribution: The species is known from Chungcheongnam-do in western Korea. (First record), China (Zhang and Zhong 1990), Japan (Higuchi 1972) Host plant: This species feeds on Alnus spp., Betula spp., Carpinus spp., Ostrya spp. (Holman 2009). In Korea, it was only found on Corylus spp. (Betulaceae). Matsumura (1919) gave Ptelea trifoliata (Rutaceae) as a host plant, however, it is unlikely to be a true host.	en	Lee, Yerim, Kanturski, Mariusz, Lee, Seunghwan (2018): Agarna malayi Tiwari 1952. Zoological Studies 57 (17): 1-20, DOI: 10.6620/ZS.2018.57-17, URL: http://dx.doi.org/10.5281/zenodo.13316922
A554879758567E3FE7A42198FAEEFA0A.taxon	description	(Figs. 1, 2, 3 e-f, 7; Table 1)	en	Lee, Yerim, Kanturski, Mariusz, Lee, Seunghwan (2018): Agarna malayi Tiwari 1952. Zoological Studies 57 (17): 1-20, DOI: 10.6620/ZS.2018.57-17, URL: http://dx.doi.org/10.5281/zenodo.13316922
A554879758567E3FE7A42198FAEEFA0A.taxon	materials_examined	Material examined: 4 alate viviparous females, Seoul, South Korea, 37 ° 33 ' 59.53 " N, 126 ° 58 ' 40.69 " E, on Carpinus erosa, Woon-Ha Paik leg., 3. xi. 1970, no. 6146; 1 alate viviparous female, Seoul, South Korea, 37 ° 33 ' 59.53 " N, 126 ° 58 ' 40.69 " E, on Carpinus koreana, Woon-Ha Paik leg., 6. v. 1971, no. 6241; 1 alate viviparous female, Seoul, South Korea, 37 ° 33 ' 59.53 " N, 126 ° 58 ' 40.69 " E, on Carpinus erosa, Woon-Ha Paik leg., 6. v. 1971, no. 6247; 1 alate viviparous female, Seoul, South Korea, 37 ° 33 ' 59.53 " N, 126 ° 58 ' 40.69 " E, on Carpinus cordata, Woon-Ha Paik leg., 21. x. 1971, no. 6837; 2 alate viviparous females, Manmulsang-area, Mt. Geumgangsan, North Korea, 38 ° 39 ' 24.10 " N, 128 ° 6 ' 17.85 " E, on Carpinus cordata, Jan Havelka, 24. v. 1988, no. 88 HA 2587; 1 alate viviparous female, Soholeub, Pocheon-si, GG, South Korea, 37 ° 49 ' 24 " N, 127 ° 08 ' 18 " E, on Carpinus laxiflora, Yerim Lee leg., 2. viii. 2013, no. 130802 YR- 3, DNA voucher no. Ca 4; 4 alate viviparous females, Girin-myeon, Inje-gun, GW, South Korea, 37 ° 57 ' 19 " N, 128 ° 19 ' 09 " E, on Carpinus sp., Duwal Keshari Ram leg., 26. ix. 2013, no. 130926 Ram- 6, including DNA voucher no. Ca 5 - 2, Ca 5 - 3, Ca 5 - 4; 10 alate viviparous females, Mt. Manisan, Is. Ganghwado, Incheon-si, GG, South Korea, 37 ° 36 ' 41 " N, 126 ° 26 ' 05 " E, on Carpinus turczaninowii, Yerim Lee and Hyoseok Lee leg., 17. v. 2014, no. 140517 YR- 1; 33 alate viviparous females, Yongdae-ri, Inje-gun, GW, South Korea, 38 ° 11 ' 02 " N, 128 ° 21 ' 43 " E, on Carpinus sp., Yerim Lee leg., 4. vi. 2014, no. 140604 YR- 42, including DNA voucher no. Ca 10; 13 alate viviparous females, Mt. Manisan, Is. Ganghwad, Incheon-si, GG, South Korea, 37 ° 36 ' 41 " N, 126 ° 26 ' 05 " E, on Carpinus turczaninowii, Yerim Lee and Hyoseok Lee leg., 21. vi. 2014, no. 140621 YR- 3, including DNA voucher no. Ca 9. Diagnosis: This species is morphologically close to M. (M.) obtusirostris Gosh by having pale head and body color, and partly pigmented ANT. However, it can be distinguished by having 6 - 12 secondary rhinaria in a row on basal 2 / 3 of ANT III (4 - 7 secondary rhinaria mostly restricted to basal half of ANT III in M. obtusirostris). Redescription (Alate viviparous female): Color in life: Head and antenna pale yellow, distal joint of Ant. III-VIb dark, compound eye red; Thorax and abdomen pale yellow; Legs pale; cu 1 b of forewing, slightly dark bordered; SIPH and cauda. Morphology: Body oval, BL 1.27 - 1.77 mm. Head smooth with a flat median protrusion on frons, head vertex with 0.01 mm pointed setae, epicranial suture weakly developed, head dorsum without tubercles. ANT 6 - segmented, ANT 0.64 - 0.76 × BL, ANT III longest with 6 - 12 transversely elliptical secondary rhinaria in a row on 2 / 3 of segment, bearing 6 - 11 short pointed setae, ANT IV-VI imbricated with inconspicuous hair-like setae, ANT IV without secondary rhinaria, ANT V shorter than ANT IV, BASE with a single inconspicuous seta, PT 0.80 - 1.25 × BASE. Rostrum not reaching to middle coxae, URS 0.06 - 0.08 mm long with 4 - 5 accessory setae, URS 0.60 - 0.89 × BASE, 0.67 - 1.00 × HT II. Fore coxae weakly enlarged, longest setae on TIBIAE III 0.67 - 1.00 × middle width of TIBIAE III, HT I with 7 chaetotaxy, HT II 0.08 - 0.10 mm long. Wing vein Rs of forewing weakly developed. Thorax smooth, without tubercles. Dorsal abdominal setae sometimes on small elevations, conspicuous tubercles not developed, ABD VIII with a pair of spinal setae, ABD margin I-IV with a single seta on short cone-shaped tubercles, 4 th marginal tubercle 0.02 - 0.04 mm long. SIPH cylindrical, truncated, 0.04 - 0.06 mm long. Cauda knobbed, 0.08 - 0.11 mm long, bearing 11 - 13 setae. Anal plate bilobed, each lobe with 5 - 10 setae. Distribution: This species is widely distributed in Korea (North and South). Also known from China (Zhang and Zhong 1990), Japan (Higuchi 1972), and East Siberia (Quednau and Shaposhnikov 1988). Host plant: This species was found on different Carpinus species (Betulaceae). In Korea also on Corylus sieboldiana (Paik 1972).	en	Lee, Yerim, Kanturski, Mariusz, Lee, Seunghwan (2018): Agarna malayi Tiwari 1952. Zoological Studies 57 (17): 1-20, DOI: 10.6620/ZS.2018.57-17, URL: http://dx.doi.org/10.5281/zenodo.13316922
A554879758567E32E4432498FAA5FDEA.taxon	description	(Figs. 1, 2, 4 a-b, 8; Table 1)	en	Lee, Yerim, Kanturski, Mariusz, Lee, Seunghwan (2018): Agarna malayi Tiwari 1952. Zoological Studies 57 (17): 1-20, DOI: 10.6620/ZS.2018.57-17, URL: http://dx.doi.org/10.5281/zenodo.13316922
A554879758567E32E4432498FAA5FDEA.taxon	materials_examined	Material examined: 1 alate viviparous female, Mt. Gyeryongsan, Gyeryong-si, CN, South Korea, 36 ° 20 ' 7 " N, 127 ° 12 ' 3 " E, on Alnus japonica, Woon-Ha Paik leg., 17. ix. 1963; 1 alate viviparous female, Daegwanryeong-myeon, Pyeongchang-gun, GW, South Korea, 37 ° 40 ' 24 " N, 128 ° 42 ' 22 " E, host plant unknown, Woon-Ha Paik leg., 11 - 20. viii. 1968; 3 alate viviparous females, Seoul, South Korea, 37 ° 34 ' 1 " N, 126 ° 57 ' 5 " E, on Physocarpus insularis, Woon-Ha Paik leg., 12. v. 1971; 1 alate viviparous female, Mt. Myohyangsan, PB, North Korea, 40 ° 1 ' 3 " N, 126 ° 19 ' 46 " E, on Corylus heterophylla, Jan Havelka leg., no. 85 HA 0789; 3 alate viviparous females, Mt. Baekdusan, Naegokri, YG, North Korea, 41 ° 7 ' 52 " N, 128 ° 15 ' 42 " E, on Corylus heterophylla, Jan Havelka leg., 18. vi. 1988, no. 88 HA 3031; 1 alate viviparous female, Danseong-myeon, Danyang-gun, CB, South Korea, 36 ° 56 ' 13 " N, 128 ° 19 ' 24 " E, on Corylus sp., Seunghwan Lee leg., 18. v. 2001, no. 011517 - SH 22; 4 alate viviparous females, Mt. Beagunsan, Gwangyang-si, JN, South Korea, 35 ° 06 ' 26 " N, 127 ° 37 ' 19 " E, on Corylus sieboldiana, Yerim Lee leg., 25. vii. 2013, no. 130725 YR- 22, including DNA voucher no. Co 2, Co 3; 1 alate viviparous females, Mt. Manwolsan, Yangyang-gun, GW, South Korea, 37 ° 57 ' 11 " N, 128 ° 41 ' 37 " Eon Corylus sieboldiana, Yerim Lee leg., 14. viii. 2013, no. 130814 YR- 4, DNA voucher no. Co 4; 15 alate viviparous females, Mt. Hwangbyeongsan, Chahang-ri, Pyeongchang-gun, GW, South Korea, 37 ° 42 ' 27 " N, 128 ° 41 ' 14 " E, on Corylus sieboldiana, Yerim Lee leg., 15. viii. 2013, no. 130815 YR- 11, including DNA voucher no. Co 6 - 1, Co 6 - 2, Co 6 - 3, Co 6 - 4, Co 6 - 5; 1 alate viviparous females, Silnae-pass, Hwacheon-gun, GW, 38 ° 09 ' 05 " N, 127 ° 31 ' 02 " E, on Corylus sieboldiana, Yerim Lee leg., 1. v. 2014, no. 140501 YR- 15, DNA voucher no. Co 8 - 1; 2 alate viviparous females, Sangdong-ri, Yeongwol-gun, GW, South Korea, 37 ° 47 ' 09 " N, 127 ° 17 ' 58 " E, on Corylus sieboldiana, Yerim Lee leg., 12. vi. 2014, no. 140612 YR- 23, DNA voucher no. Co 15 - 1, Co 15 - 2; 1 alate viviparous female, Is. Geojedo, GN, South Korea, 34 ° 53 ' 12 " N, 128 ° 37 ' 25 " E, on Corylus sp., Yerim Lee and Hyoseok Lee leg., 14. vii. 2014, no. 140714 YR- 7, DNA voucher no. Co 14. Diagnosis: This species is similar to M. (P.) yunanensis (Zhang) but it differs by having pointed setae on the dorsal abdomen, URS 0.80 - 1.00 × HT II (1.50 - 1.70 in M. yunanensis), and PT 0.90 - 1.10 × BASE (0.60 - 0.70 in M. yunanensis). Redescription (Alate viviparous female): Color in life: Head and antenna pale yellow, distal joint of ANT III-VIb dark. Thorax and abdomen pale yellow. Legs pale, distal 1 / 3 of fore tibiae and tarsi fuscous. Wing vein Cu 1 b of forewing dark bordered. SIPH and cauda. Morphology: Body oval, BL 1.24 - 1.61 mm. Head smooth with a flat median protrusion on frons, epicranial suture weakly indicated, head vertex with 0.01 mm pointed setae, head dorsum without tubercles. ANT 6 - segmented, ANT 1.02 - 1.29 × BL, ANT III with 7 - 12 transversely elliptical formed secondary rhinaria in a row on the whole segment, ANT III-VI imbricated with inconspicuous hair-like setae, ANT IV without secondary rhinaria, ANT V shorter than ANT IV, PT 0.92 - 1.10 × BASE. Rostrum not reaching to middle coxae, URS 0.08 - 0.09 mm long with 4 - 5 accessory setae, 0.80 - 1.00 × HT II. Thorax smooth, without tubercles. Fore coxae weakly enlarged, longest setae on TIBIAE III 1.00 - 1.50 × middle width of TIBIAE III, HT I with 7 - 8 chaetotaxy, HT II 0.08 - 0.11 mm long. Dorsal spinal setae sometimes on small elevations, conspicuous tubercles not developed, ABD VIII with a pair of setae, ABD margin I-IV with 2 setae on short cone-shaped tubercles, 4 th marginal tubercle 0.02 - 0.04 mm. SIPH cylindrical, truncated, 0.05 - 0.07 mm long. Cauda knobbed, 0.08 - 0.10 mm long, bearing 10 - 14 setae. Anal plate bilobed, each lobe with 6 - 11 setae. Distribution: The species is rather common in the Korean peninsula (both North and South Korea), China (Zhang et al. 1986), and Japan (Higuchi 1972). Host plant: The species feeds on several species of Corylus (Betulaceae) in almost whole distributional range, but in Russian Far East it was collected from Quercus dentata (Holman 2009). Remarks: In Paik’s collection, some specimens were collected on Alnus japonica and Physocarpus insularis. However, these do not seem to be a true host plants because M. (P.) corylicola is typically recorded as Corylus.	en	Lee, Yerim, Kanturski, Mariusz, Lee, Seunghwan (2018): Agarna malayi Tiwari 1952. Zoological Studies 57 (17): 1-20, DOI: 10.6620/ZS.2018.57-17, URL: http://dx.doi.org/10.5281/zenodo.13316922
A5548797585A7E37E7FF2198FB09FD4A.taxon	description	(Figs. 1, 2, 4 c-d, 9, 11; Table 1) urn: lsid: zoobank. org: act: 21873460 - 79 B 3 - 4 ECE-BD 7 B- BD 9100 F 727 A 6 Material examined: Holotype: 1 alate viviparous female, Sinwon-ri, Yongin-si, GG, South Korea, 37 ° 18 ' 56 " N, 127 ° 12 ' 56 " E, on Corylus sp., Yerim Lee and Hyoseok Lee leg., 16. v. 2014, no. 140516 YR- 12. Paratypes: 9 alate viviparous females, same data as holotype, including DNA voucher no. Co 12; 1 alate viviparous female, Silnae-pass, Hwacheon-gun, GW, 38 ° 09 ' 05 " N, 127 ° 31 ' 02 " E, on Corylus sieboldiana, Yerim Lee leg., 1. v. 2014, no. 140501 YR- 15, DNA voucher no. Co 9; 1 alate viviparous female, Mt. Jugeumsan, Gapyeonggun, GG, South Korea, 38 ° 08 ' 38 " N 126 ° 54 ' 17 " E, on Corylus heterophylla, Yerim Lee leg., 9. v. 2014, no. 140509 YR 10, DNA voucher no. Co 11; 30 alate viviparous females, Mt. Hwayasan, Gapyeonggun, GG, South Korea, 37 ° 40 ' 17 " N 127 ° 25 ' 40 " E, on Corylus heterophylla, Yerim Lee leg., 26. v. 2014, no. 140526 YR- 1, including DNA voucher no. Co 16 - 1; 2 alate viviparous females, Mt. Gyeryongsan, Gongju-si ,, CN, South Korea, 36 ° 20 ' 33 " N, 127 ° 12 ' 21 " E, on Corylus heterophylla, Yerim Lee and Hyoseok Lee leg., 24. iv. 2015, no. 150424 YR- 3, DNA voucher no. Mes 3 - 1, Mes 3 - 2; 1 alate viviparous females, Mt. Gwanggyosan, Suwonsi, GG, South Korea, 37 ° 19 ' 40 " N, 127 ° 00 ' 54 " E, on Corylus heterophylla, Yerim Lee and Hyoseok Lee leg., 10. v. 2015, no. 150510 YR- 5, DNA voucher no. Mes 5 - 1. Etymology: The species name, occulta is derived from Latin adjective, ‘ occulta’ (hidden). Diagnosis: This species is morphologically very similar to M. (P.) corylicola Higuchi in that it has a pale body and URS that is 0.07 - 0.09 mm in length. However, it can be distinguished by its shorter ANT IV length, 0.12 - 0.22 mm (0.23 - 0.32, in M. corylicola), shorter ANT V length, 0.13 - 0.18 mm (0.12 - 0.22, in M. corylicola), ANT 0.60 - 0.77 × BL (0.75 - 0.92, in M. corylicola) and URS 1.00 - 1.14 × HT II (0.80 - 1.00 in M. corylicola). Description (Alate viviparous female): Color in life: Head pale yellow, ANT pale with slight dark margin of ANT III and BASE. Thorax and abdomen pale yellow. Legs pale with 1 / 9 of fore tibiae and tarsi fuscous. Wing vein Cu 1 b of forewing bordered with brown pigment. SIPH pale. Cauda and anal plate pale. Morphology: Body oval, BL 0.95 - 1.44 mm. Head smooth with a flat median protrusion on frons, epicranial suture weakly developed, head dorsum without tubercles with 8 acuminate setae shorter than 0.01 mm long. ANT 6 - segmented, 0.73 - 1.07 mm long, ANT 0.60 - 0.77 × BL, ANT III longest with 5 - 9 narrow elliptical secondary rhinaria in a row on the whole segment, bearing 3 - 4 short pointed setae, ANT III-VI imbricated with inconspicuous pointed setae, Ls ANT III × 0.50 - 1.00 BD III, ANT IV without secondary rhinaria, ANT V shorter than ANT IV, BASE with a single inconspicuous seta, PT 0.83 - 1.10 × BASE. Rostrum not reaching to middle coxae, URS 0.07 - 0.09 mm long with 4 - 6 accessory setae, URS 0.75 - 1.00 × BASE, 1.00 - 1.14 × HT II. Thorax smooth without tubercles. Fore coxae slightly enlarged, longest setae on TIBIAE III 1.50 × middle width of TIBIAE III, HT I with 8 chaetotaxy, HT II 0.07 - 0.09 mm long. Wing vein Rs weakly developed. Dorsal spinal setae sometimes on small elevations, conspicuous tubercles not developed, ABD VIII bearing a pair of short setae, ABD margin I-IV with 0.01 - 0.02 mm long of 2 setae on short cone-shaped tubercle, 4 th marginal tubercle 0.01 - 0.02 mm. SIPH cylindrical, truncated, 0.04 - 0.06 mm long. Cauda knobbed, 0.07 - 0.09 mm with 9 - 13 setae. Anal plate bilobed, each lobe with 7 - 10 setae. (a) (b) (c) (e) (d) (f) (g) Distribution: So far, the new species is known from Gangwon-do, Gyeongsangnam-do, and Jeollanam-do in South Korea. Host plant: Representatives of the new species feed on the underside of leaves of Corylus sp. (Betulaceae). Remarks: This species was first recognized as a cryptic species of M. (P.) corylicola in the previous DNA barcoding study (Lee et al. 2017). Molecular identification The overall mean genetic distance was 7.33 % for the 45 partial COI sequences of the 5 Mesocallis species. All species showed very low intraspecific distance level, ranging from 0 % to 1.07 % (Table 2). Among the five species, intraspecific distance was most varied in M. corylicola, with ranges of up to 1.07 %; in contrast, 0 % of intraspecific distance was observed among M. carpinicola sp. nov. In the remaining species, intraspecific distances were below 0.61 %. Interspecific distance ranged from 6.21 % to 12.72 %. 11.81 % to 12.72 % of highest interspecific distances were observed between M. pteleae and M. corylicola (Table 2). M. corylicola and M. occulta sp. nov. showed 6.21 % to 7.40 % of lowest interspecific distances. The COI sequence NJ tree showed that the 45 sequences are clearly divided into five specific groups (Fig. 12). This result indicates that molecular identification based on COI barcoding region is effective for Mesocallis. Haplotype analysis results found 128 variable positions in the 658 bp COI region. Overall, 19 different haplotypes were observed, consisting of 1 haplotype for M. carpinicola sp. nov., 3 for M. pteleae, 4 for M. sawashibae, 9 for M. corylicola and 2 for M. occulta sp. nov. (Fig. 13).	en	Lee, Yerim, Kanturski, Mariusz, Lee, Seunghwan (2018): Agarna malayi Tiwari 1952. Zoological Studies 57 (17): 1-20, DOI: 10.6620/ZS.2018.57-17, URL: http://dx.doi.org/10.5281/zenodo.13316922
