identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
A50C0231FFAFFF9AFF14FABAFC82F986.text	A50C0231FFAFFF9AFF14FABAFC82F986.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gianius camachoae Martinez-Ansemil 2025	<div><p>Gianius camachoae Martínez-Ansemil sp. nov.</p><p>(Figs 1, 2)</p><p>Holotype. MHNUSC_10146: mature worm, incomplete (broken at segment XIV), unstained and whole-mounted in Canada balsam. Leg. Ana I. Camacho. Type locality. CAN002 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.19491&amp;materialsCitation.latitude=-3.58453" title="Search Plazi for locations around (long 43.19491/lat -3.58453)">De los Santos cave</a>, Cantabria, Spain, –3.58453, 43.19491 (4 June 2002) .</p><p>Paratypes. MHNUSC_10147_1–2: 1 mature and 1 immature specimen, unstained and whole-mounted in Canada balsam, from type locality (4 June 2002); MHNUSC_10147_3–5: 2 mature specimens (1 stained in haematoxylin and 1 unstained) and 1 immature stained in haematoxylin, whole-mounted in Canada balsam, CAN004 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.22852&amp;materialsCitation.latitude=-3.61173" title="Search Plazi for locations around (long 43.22852/lat -3.61173)">Covalagua cave (cueva del Agua)</a>, Cantabria, –3.61173, 43.22852 (7 June 2002); MHNUSC_10147_6–10: 2 mature specimens (1 stained in haematoxylin and 1 unstained) and 3 immature unstained, whole-mounted in Canada balsam, CAN022 Upwelling of the <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.27797&amp;materialsCitation.latitude=-3.61717" title="Search Plazi for locations around (long 43.27797/lat -3.61717)">Bustablado stream</a>, Cantabria, –3.61717, 43.27797 (14 Mars 2003); MHNUSC_10147_11: 1 mature specimen stained in haematoxylin and whole-mounted in Canada balsam, CAN034 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.27404&amp;materialsCitation.latitude=-3.60441" title="Search Plazi for locations around (long 43.27404/lat -3.60441)">Bustablado stream</a>, Arredondo, Cantabria, –3.60441, 43.27404 (8 June 2002); MHNUSC_10147_12: 1 mature specimen stained in haematoxylin and whole-mounted in Canada balsam, CAN008 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.27576&amp;materialsCitation.latitude=-3.62032" title="Search Plazi for locations around (long 43.27576/lat -3.62032)">Arduengo cave</a> (Cuviobramante), Cantabria, – 3.62032, 43.27576 (14 Mars 2003); MHNUSC_10147_13: 1 mature specimen stained in haematoxylin and whole-mounted in Canada balsam, CAN047 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.25108&amp;materialsCitation.latitude=-3.60577" title="Search Plazi for locations around (long 43.25108/lat -3.60577)">Asón stream</a>, La Cubera, Cantabria, –3.60577, 43.25108 (20 January 2002). All leg. Ana I. Camacho.</p><p>Etymology. Named in honor of Dr.Ana Isabel Camacho (MNCN, Madrid, Spain), the leader of the PASCALIS project in Spain, who contributes greatly to the knowledge and development of the biospeleology.</p><p>Description. More than 2.6 mm long (all mature specimens are broken and lack posterior segments). Maximum number of segments (observed in immature specimens), 31. Diameter at segment V (slightly compressed specimens) 90–130 μm, at segment XI 100–180 μm. Clitellum 1/2 X–XII. Prostomium rounded, 42–58 μm long and 40–70 μm wide. Secondary annulation indistinct or slightly marked, up to VI. Somatic setae bifid, 26–35 μm long, with upper tooth shorter and thinner than lower (Fig. 2A); (2)3–5 setae per bundle anteriorly and 2–3 in the clitellar and postclitellar segments. Ventral setae of XI modified into bifid penial setae with short teeth, in number of 6–13 per bundle, 28–40 μm long and 1–1.5 μm thick, arranged comb-like, forming an open arc towards the sagittal plane and placed close to the male pores on their inner side (Figs 1A, B, 2D [ps]). One pair of male pores located in the posterior third of segment XI, in line with somatic ventral setae (Fig. 1A). One pair of spermathecal pores in segment XI; four of the ten specimens studied have an additional pair in XII (Fig. 2H), and one (from upwelling of Bustablado) has even additional spermathecal pores in X and XIII. All spermathecal pores are placed in the most anterior part of their respective segments, in line with somatic ventral setae (Figs 1A, 2H [sp]).</p><p>Pharyngeal glands in segments IV–VI, poorly developed. Chloragogen cells beginning in VI. Oesophageal appendages and diverticula absent. One pair of testes in segment X. One pair of ovaries in segment XI. Sperm sacs extending by segments (IX) X–XI(XII); mature eggs in segments XI–XII(XIII). Vasa deferentia twisted, shorter than atria, 45–60 μm long and around 3 μm wide, entering the apical (ental) end of the atria (Fig. 1A vd). Atria tubular to spindle-shaped, curved, 78–95 μm long and 12–16 μm maximum diameter, with a thin muscular outer layer (1–2 μm thick); atrial lumen frequently filled by spermatozoids; ectally, each atrium ends in a dilated spherical ampulla which is surrounded by a rather thin muscular layer (atrial bulb), 16–26 μm in diameter, acting as a pseudopenis in extrusion (Figs 1A, 2B–D, F–H [a, b, pp]). Two prostate glands per atrium, the anterior one large and placed near the entrance of vas deferens, and the posterior one small and placed close to the atrial bulb (Figs 1A, 2C, D [pr1, pr2]). Spermathecae of XI and XII filled with spermatozoids (Fig. 1A, D); spermathecae of X and XIII empty (Fig. 1C). Spermathecal duct muscular, conical and long (up to 30 µm) in XI, globular and short in XII and XIII, tubular and short in X; spermathecal ampulla tubular (up to 80 µm long, 10–20 µm in diameter) and directed forward (Figs 1A, C–D, 2B, E–H [sd, sa]).</p><p>Distribution and habitat. Caves, springs, phreatic and hyporheic habitats at the upper and middle parts of the Asón River basin, Cantabria, Spain. Alt. 161– 710 m.</p><p>Remarks. Following Erséus (1992a), the presence of elongated and curved atria, each with two prostate glands attached to proximal and distal ends, respectively, vasa deferentia entering apical ends of atria, slender spermathecae with short ducts, and small straight penial setae allocate the new species described above in the genus Gianius Erséus.</p><p>From the twelve species of Gianius currently known, Gianius camachoae sp. nov. is clearly distinguished by the presence of spermathecae in the atrial segment (segment XI), a character only shared with the marine species G. eximius Erséus, 1997 and G. urgorrii Martínez-Ansemil &amp; Caramelo, 2025 . Among other characters, G. camachoae sp. nov. clearly differs from these two species by the shape and structure of the spermathecae, ampullae elongate and ducts simple in the former, and ampullae globular and ducts bipartite in the two latter. G. crypticus (Rodriguez &amp; Giani, 1989), described from the same geographical area (a cave in the upper part of the Clarin stream basin), appears as its most closely related species, although having exclusively a typical pair of spermathecae in the preatrial segment X. The variability in the number and location of the spermathecae observed in G. camachoae sp. nov. is high, but all specimens have, at least, one functional pair of spermathecae in the atrial segment. Furthermore, G. camachoae sp. nov. clearly differs from G. crypticus for at least two other characters, the atrial muscular layer (thin in the former; very thick in the later), and the atrial bulb (with a large lumen and surrounded by a relatively thin muscular layer in the former; heavily muscular in the later).</p></div>	https://treatment.plazi.org/id/A50C0231FFAFFF9AFF14FABAFC82F986	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFAAFF99FF14F946FAA4FE7A.text	A50C0231FFAAFF99FF14F946FAA4FE7A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gianius inermis Sambugar & Martinez-Ansemil 2025	<div><p>Gianius inermis Sambugar &amp; Martínez-Ansemil sp. nov.</p><p>(Figs 3, 4)</p><p>Holotype. MHNUSC_10149: mature worm, incomplete (broken at segment XVI), stained in paracarmine and whole-mounted in Canada balsam. Leg. Fabio Stoch.</p><p>Type locality. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.61771&amp;materialsCitation.latitude=10.98458" title="Search Plazi for locations around (long 45.61771/lat 10.98458)">LES050 Bus delle Piatte cave</a> (441 V/VR), Rocca, Lessinian Massif, Italy, 10.98458, 45.61771 (20 June 2006).</p><p>Paratypes. MHNUSC_10150_1: 1 mature specimen stained in paracarmine and whole mounted in Canada balsam, from type locality (20 June 2006). MHNUSC_10150_2: 1 mature specimen stained in paracarmine and whole-mounted in Canada balsam, LES107 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.60577&amp;materialsCitation.latitude=11.04847" title="Search Plazi for locations around (long 45.60577/lat 11.04847)">Regosse cave</a> (161 V/VR), Regosse, Lessinian Massif, 11.04847, 45.60577 (22 August 2001); MHNUSC_10150_3: 1 mature specimen stained in haematoxylin and whole-mounted in Canada balsam, LES036 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.6155&amp;materialsCitation.latitude=10.91867" title="Search Plazi for locations around (long 45.6155/lat 10.91867)">Fumane-Molina interstitial</a>, Lessinian Massif, 10.91867, 45.61550 (August 2001); MHNUSC_10150_4: 1 mature specimen stained in haematoxylin and whole-mounted in Canada balsam, LES124 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.4595&amp;materialsCitation.latitude=11.0665" title="Search Plazi for locations around (long 45.4595/lat 11.0665)">Verona–Montorio via del Lanificio interstitial</a>, Lessinian Massif, 11.06650, 45.45950 (August 2001). Leg. Fabio Stoch and Diana Galassi.</p><p>Etymology. The specific name refers to the lack of penial setae (Latin inermis, unarmed).</p><p>Description. Mature worms up to 5 mm long. Number of segments 28–33. Diameter at segment V (slightly compressed specimens) 105–135 μm, at segment XI 100–160 μm. Clitellum 1/2 X–XII. Prostomium rounded, 45– 65 μm long and 60–82 μm wide. Secondary annulation unnoticeable or slightly marked, up to V. Somatic setae bifid, 30–45 μm long, with upper tooth shorter and thinner than lower (Fig. 4A); (2)3–4 somatic setae per bundle anteriorly and 2(3) in the clitellar and postclitellar segments (ventral setae absent in XI). Penial setae absent. One pair of male pores located in the posterior third of segment XI, in line with somatic ventral setae. One pair of spermathecal pores in segment X, placed in the most anterior part of the segment, in line with somatic ventral setae.</p><p>Pharyngeal glands in segments IV–VI. Chloragogen cells beginning in VI. Oesophageal appendages and diverticula absent. One pair of testes in segment X. One pair of ovaries in segment XI. Sperm sacs extending by segments X–1/2XII. Egg sac extending to 13/14. Vasa deferentia about as long as atria, around 5 μm wide, entering the apical end of the atria. (Figs 3A, 4C [vd]). Atria tubular and curved, 115–140 μm long and 13–20 μm maximum diameter, with a thin muscular outer layer (1–2 μm thick); atrial lumen frequently containing spermatozoids; ectally, the atrium ends directly in a simple pore, surrounded by muscular strands, and can extrude somewhat as a small pseudopenis (Figs 3A, 4E [mp, ms]). Two roundish compact prostate glands per atrium, similar in size, the anterior one placed near the entrance of vas deferens, and the posterior one near the ectal end (Figs 3A, 4B–C). Spermathecae composed by a short conical muscular duct (13–20 μm long) dilated in a roundish vestibulum (16–30 μm in diameter) and followed by a long tubular ampulla (90–110 µm long, 13–22 µm wide) generally directed backwards in the segment (Figs 3B, 4D).</p><p>Distribution and habitat. Caves and interstitial habitats in the western and southern parts of the Lessinian Massif (Progno di Fumane basin, Progno di Valpantena basin and Vaio di Squaranto basin), southernmost part of the Venetian pre-Alps, Italy. Alt. 71– 700 m.</p><p>Remarks. Following Erséus (1992a), the presence of elongated and curved atria, each with two prostate glands attached to proximal and distal ends, respectively, vasa deferentia entering apical ends of atria, slender spermathecae with short ducts, and small straight penial setae allocate the new species described above in the genus Gianius Erséus.</p><p>The species most closed relative to Gianius inermis sp. nov. appears to be G. labouichensis (Rodriguez &amp; Giani, 1989), described from the underground stream of Labouiche (Ariège, France). G. labouichensis is the only species of Gianius known until now with somatic setae in the ventral bundles of XI at maturity, instead of differentiated penial setae; the mature specimens of G. inermis sp. nov. are completely devoid of ventral setae in XI. The male ducts of G. labouichensis and G. inermis sp. nov. are similar in shape, but clearly smaller in the former (87 μm long, 7.5 μm in diameter). The location of the male pores in about the middle of the segment in G. labouichensis and in the posterior third of the segment in G. inermis sp. nov., and the absence in G. inermis sp. nov. of a dorsal appendage in the gut, present in G. labouichensis, are two other characters that clearly separate these two species.</p></div>	https://treatment.plazi.org/id/A50C0231FFAAFF99FF14F946FAA4FE7A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFA9FF97FF14FE1AFABFFDEB.text	A50C0231FFA9FF97FF14FE1AFABFFDEB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gianius parvogenitalis Martinez-Ansemil 2025	<div><p>Gianius parvogenitalis Martínez-Ansemil sp. nov.</p><p>(Figs 5, 6)</p><p>Holotype. MHNUSC_10148: mature worm stained in paracarmine and whole-mounted in Canada balsam. Leg. Ana I. Camacho.</p><p>Type locality. CAN072 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.34713&amp;materialsCitation.latitude=-3.5413" title="Search Plazi for locations around (long 43.34713/lat -3.5413)">Upwelling of the Clarión stream</a>, Cantabria, Spain, –3,54130, 43.34713 (13 Mars 2003).</p><p>Etymology. The specific name refers to the small (Latin parvus) size of the genital organs (Latin genitalia).</p><p>Description. Small worm, 1.8 mm long. Number of segments 26. Diameter at segment V 125 μm, at segment XI 175 μm. Clitellum 1/2 X–XII. Prostomium rounded, 45 μm long. Secondary annulation up to VI. Somatic setae bifid, 18–32 μm long, with upper tooth equal or slightly thinner and longer than lower (Fig. 6B); 3–5 setae per bundle in segments II–VII and 2 thereafter. Penial setae bifid, with a minute upper tooth, 2–3 setae per bundle, about 22 μm long and 1.5 μm thick (Figs 5A–B, 6D [ps]). One pair of male pores located in the posterior third of segment XI, in line with somatic ventral setae. One pair of spermathecal pores in segment X, placed anteriorly in the segment, slightly lateral with respect to the alignment of the ventral somatic setae.</p><p>Pharynx with a well-developed pharyngeal pad. Pharyngeal glands in segments IV–VII. Chloragogen cells beginning in VI. Oesophageal appendages and diverticula absent. One pair of testes in segment X. One pair of ovaries in segment XI. Sperm sacs not observed. Egg sac extending to 12/13. Vasa deferentia coiled, 4–5 μm wide, about as long as the atria (Figs 5A, 6C, E [vd]). Atria cylindrical and curved (C-shaped), about 50 μm long and 7–8 μm in diameter, with a thin muscular outer layer (about 1 μm thick); ectally, the atrium ends directly in the tegument, surrounded by a thin circular muscular layer (Figs 5A, 6C–E [a]). Two prostate glands per atrium, the anterior one placed near the entrance of the vas deferens, and the posterior one at the ectal part of the atrium, close to the penial setae (Figs 5A, 6C–E [pr1, pr2]). Spermathecae composed of a short duct (22 μm long, 11 μm wide) and a thin-walled, pyriform, slender ampulla (48 µm long, up to 14 µm wide), filled with spermatozoa (Figs 5A, 6F [sd, sa]).</p><p>Distribution and habitat. Upwelling of the Clarión stream, Cantabria, Spain. Alt. 57 m.</p><p>Remarks. Following Erséus (1992a), the presence of elongated and curved atria, each with two prostate glands attached to proximal and distal ends, respectively, vasa deferentia entering apical ends of atria, slender spermathecae with short ducts, and small straight penial setae allocate the new species described above in the genus Gianius Erséus.</p><p>The narrow cylindrical C-shaped atria, only slightly tapering towards the ends, its small size (only 50 μm long and 7–8 μm in diameter), and the presence of bifid penial setae in low number (only 2–3 per bundle) are distinctive characters of Gianius parvogenitalis sp. nov. with respect to all the other known species in the genus.</p></div>	https://treatment.plazi.org/id/A50C0231FFA9FF97FF14FE1AFABFFDEB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFA7FF97FF14FD6BFA2BFC1E.text	A50C0231FFA7FF97FF14FD6BFA2BFC1E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cantabrodrilus Martinez-Ansemil 2025	<div><p>Cantabrodrilus Martínez-Ansemil gen. nov.</p><p>Type species. Cantabrodrilus pilarae Martínez-Ansemil sp. nov.</p><p>Diagnosis. Freshwater subterranean phallodrilines. Somatic setae bifid. Penial setae present, single pointed hook-shaped. Coelomocytes absent. Spermathecal pores in segment X. Vasa deferentia narrow, entering apical ends of atria. Atria tripartite, elongate; ental part strongly muscular, middle part tubular U-shaped, ectal part thin-walled. Two large prostate glands per atrium; the anterior one attached near entrance of vas deferens and the posterior one at ectal end of atrium.</p><p>Etymology. The name Cantabrodrilus means that the type species is a “worm” (Greek drilos) from Cantabria.</p></div>	https://treatment.plazi.org/id/A50C0231FFA7FF97FF14FD6BFA2BFC1E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFA7FF92FF14FBFFFBF2FE06.text	A50C0231FFA7FF92FF14FBFFFBF2FE06.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cantabrodrilus pilarae Martinez-Ansemil 2025	<div><p>Cantabrodrilus pilarae Martínez-Ansemil sp. nov.</p><p>(Figs 7, 8)</p><p>Holotype. MHNUSC_10151: mature worm, incomplete (broken after the setae of segment XII), unstained and whole-mounted in Canada balsam. Leg. Ana I. Camacho.</p><p>Type locality. CAN177 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=43.2803&amp;materialsCitation.latitude=-4.48694" title="Search Plazi for locations around (long 43.2803/lat -4.48694)">Latarmá stream near Toyu cave</a>, Venta Fresnedo, Cantabria, Spain, –4.48694, 43.28030 (3 September 2002).</p><p>Etymology. Named after Professor Pilar Rodriguez (University of the Basque Country, Spain), who contributes greatly to the knowledge of the oligochaetes in inland waters.</p><p>Description. Small worm, about 80 µm in diameter at segments V and XI (specimen broken after the setae of segment XII). Clitellum unnoticeable. Prostomium short (28 µm long), rounded. Secondary annulation not observed. Somatic setae bifid, 20–30 μm long, with short teeth, upper slightly shorter than lower, 2–3 somatic setae per bundle (Fig. 8A); 5 single pointed hook-shaped penial setae per bundle in segment XI, about 20 μm long (Figs 7B, 8E [ps]). One pair of male pores located in the posterior third of segment XI, in line with somatic ventral setae. One pair of spermathecal pores in segment X, surrounded by a glandular area, in the lateral anterior part of the segment (Fig. 8B, D [ga]).</p><p>Pharyngeal glands in segments III–V. Chloragogen cells beginning in VI. One pair of testes in segment X. One pair of ovaries in segment XI. Sperm sacs entering the posterior part of segment IX. Vasa deferentia short and twisted, 4–5 μm wide, entering apical ends of atria (Fig. 7A [vd]). Atria elongated, divided in three clearly differentiated parts. The ental part consist in a long, sigmoid, spindle shaped and heavily muscular structure (75–85 μm long, 13 μm in maximum diameter, muscular wall up to 4 μm thick) (Figs 7A, 8F [ma, m]), the middle part is a simple short curved U-tube (about 5 μm in diameter) connecting the ental and the ectal part (Fig. 7A [ta]), and the latter is a pyriform, thin-walled and granulated ampulla (60–75 μm long, 20–22 μm in maximum diameter, muscular wall about 2 μm thick), which ends in a slightly curved and gradually narrowing ejaculatory duct (about 15 μm long) provided with a thin layer of circular muscles (Figs 7A, 8C, E [pa, ad]). Two large prostate glands per atrium, one attached to the anterior end of its muscular ental part, and the other to the posterior end of the ectal ampulla (Figs 7A, 8B–C, F [pr1, pr2]). Spermathecae composed by a tubular duct (36–40 μm long), and a very long, thin-walled ampulla (115–130 μm long, 20 μm in maximum diameter) directed backwards all along the segment X (Figs 7C, 8B [sd, sa]).</p><p>Distribution and habitat. Hyporheic flow of the Latarmá stream, a tributary of the Lamasón stream at the left side of the Nansa stream basin, Cantabria, Spain. Alt. 180 m.</p><p>Remarks. The new species described here is a typical member of the Phallodrilinae, a characteristic subfamily of Naididae that, according to Erséus (1992a) has two important autapomorphies: the absence of coelomocytes and the presence of two compact prostate glands attached to each atrium, one to anterior face, another to posterior face of latter.</p><p>Cantabrodrilus pilarae gen. nov., sp. nov. is the only phallodriline with tripartite atria. The highest degree of complexity observed until know in the atria of the Phallodrilinae was the bipartite condition of the atria of Nootkadrilus Baker, 1982 (atria elongate, bipartite and folded at middle), with which C. pilarae sp. nov. also shares the shape of the penial setae.At first glance, the well set-off U-tube connecting the ental and the ectal part of the atria in Cantabrodrilus could be interpreted as an evolved character with respect to the bipartite atria of Nootkadrilus, but the latter is an atypical genus among the Phallodrilinae, living in the Northwest Pacific Ocean, and defined not only by the bipartite condition of the atria but also by the following apomorphies: absence of anterior prostate gland in the atria, presence of thickened and heavily granulated walls, resembling (and functioning as?) prostate glands, or diffuse prostate-like cells in the ental part of the vasa deferentia (see Erséus 1992a).</p></div>	https://treatment.plazi.org/id/A50C0231FFA7FF92FF14FBFFFBF2FE06	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFA2FF90FF14FDC7FD6AF96A.text	A50C0231FFA2FF90FF14FDC7FD6AF96A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aberrantidrilus sketi Martinez-Ansemil & Sambugar 2025	<div><p>Aberrantidrilus sketi Martínez-Ansemil &amp; Sambugar sp. nov.</p><p>(Figs 9, 10)</p><p>Abyssidrilus sp. 1 in Giani et al. (2011)</p><p>Holotype. MHNUSC_10144: mature worm, incomplete (broken at segment XVII), stained in paracarmine and whole-mounted in Canada balsam. Leg. Boris Sket.</p><p>Type locality. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=45.77766&amp;materialsCitation.latitude=13.8622" title="Search Plazi for locations around (long 45.77766/lat 13.8622)">Cave 4101</a>, Brje pri Koprivi, Sežana, Slovenia, 13.86220, 45.77766 (December 1979).</p><p>Paratypes. MHNUSC_10145_1–4: 3 mature specimens, unstained and whole-mounted in Canada balsam; 1 immature specimen, stained in paracarmine and whole-mouted in Canada balsam. All from type locality (December 1979). Leg. Boris Sket.</p><p>Etymology. Named in honor of the late Professor Boris Sket (University of Ljubljana, Slovenia), who made outstanding contributions to the knowledge and development of biospeleology.</p><p>Description. Small worms, 3.2 mm long and 32 segments (complete paratype specimen). Diameter at segment V (slightly compressed specimens) 150–185 μm, at segment XI 175–195 μm. Clitellum hardly noticeable, extending over 1/2 X–XII. Prostomium rounded, 60–62 μm long. Secondary annulation at segments II–V. Somatic setae 32–50 μm long, 3–5(6) in the preclitellar bundles and 2–5 per bundle in clitellar and postclitellar ones; ventral setae absent in XI in mature specimens; preclitellar dorsal setae and ventral setae until about segment XXV bifid, with upper tooth slightly longer than or as long as lower (Fig. 10A, C–D); a mixture of single-pointed and bifid setae with upper tooth reduced in the following one or two segments; thereafter the bundles are composed exclusively of single pointed sigmoid setae (Fig. 10B). Penial setae absent. One pair of male pores located in two conical protuberances (porophores) in the posterior third of segment XI, in line with somatic ventral setae (Fig. 10E–F, H [mp, po]); each porophore has a large ring of glandular cells surrounding the male pore (Fig. 10F g). One pair of spermathecal pores in segment XII, surrounded by a small circle of glandular cells, and placed anteriorly in the segment, in a lateral position slightly ventral with respect to the lateral lines (Figs 9, 10H–G [sp, g]).</p><p>Pharyngeal glands in (III) IV– VI. Chloragogen cells beginning in VI. One pair of testes in X. Anterior and posterior sperm sacs extending to IX and XI, respectively. One pair of ovaries in XI. Egg sac not developed, one mature egg present in the ovarian segment. Vasa deferentia shorter than atria, 5.5–7 μm in diameter, entering apical end of atria (Figs 9, 10J [vd]). Atria tubular, comma-shaped, 205–215 μm long, 16–20 μm wide, strongly muscular (muscular layer 5–7 μm thick) (Figs 9, 10H, K [a]). Two prostate glands per atrium. The anterior prostate is large, compact and lobed and is attached to the apical end of the atrium (Figs 9, 10J [pr1]), and the posterior prostate consists in a small group of cells, hardly visible, connected by a few strands with the ectal end of the atrium (or with the porophore?) near the male pore (Figs 9, 10I [pr2]). Spermathecae elongated, confined to XII, each consisting of a short conical duct (about 45 μm long), curved at base, and dilating in a club-shaped ampulla filled with spermatozoids; length of the ampulla about 75 μm long, maximum width 33 μm (Figs 9, 10E [sd, sa]).</p><p>Distribution and habitat. Cave in the Slovenian coastal karst, Sežana, Slovenia. Alt. 290 m.</p><p>Remarks. The genus Aberrantidrilus Martin was erected to include a new groundwater species of Phallodrilinae, A. stephaniae Martin, 2015, from the Mercantour National Park, France, and the only two groundwater species of Abyssidrilus Erséus known until then, A. cuspis (Erséus &amp; Dumnicka, 1988), from Umbria, Italy and A. subterraneus (Rodriguez &amp; Giani, 1989), from Cantabria, Spain (Martin et al. 2015).The decision to erect this genus was supported by “a unique combination of characters rather than formally proposed apomorphies”. Thus, these three species share the “aberrant” position of the spermathecal pores (in the postatrial segment XII), the presence of a single penial seta per bundle oriented in anterior direction, the atria covered by a thick muscular layer and opening to the exterior on conical male protuberances (porophores), and the small size of the posterior prostate glands. A fourth species of Aberrantidrilus, A. mihaljevici Vučković, Rodriguez &amp; Kerovec, 2020 was recently described from the tufa barrier of Roški slap, Croatia (Rodriguez et al. 2020); the authors remark that this species shows all the abovementioned characters except the thick muscular layer of the atria and, perhaps also, the presence of posterior prostate glands (“not seen, probably due to its small size”).</p><p>The new species described here is tentatively ascribed to the genus Aberrantidrilus . The only major character by which A. sketi sp. nov. stands out in the genus is the lack of penial setae, but this is probably an unstable character in the genus as the absence of penial setae was observed also in one mature specimen of A. stephaniae (Martin et al. 2015) and “maybe” also in the specimens of A. cf. cuspis cited from Trieste, Italy (Sambugar et al. 1999). The tubular, narrow, comma-shaped and strongly muscular atria, the characteristically lobed anterior prostate glands and the thin and long spermathecal ducts clearly separate A. sketi sp. nov. from all the other species of the genus; the presence of glandular cells surrounding the spermathecal pores is also noteworthy, as similar structures or true glandular tissues associated or close to spermathecal pores are also present in Abyssidrilus opulentus Erséus, 1992 and Abyssidrilus potens Erséus, 1992 (Erséus 1992a).As pointed out by Martin (in Martin et al. 2015), the specimens tentatively identify by Sambugar et al. (1999) as A. cuspis or A. cf. cuspis from Italy, France and Slovenia can be considered as a complex of cryptic species. Aberrantidrilus can be regarded now as a diversified groundwater genus of Phallodrilinae from the karst of southern Europe.</p></div>	https://treatment.plazi.org/id/A50C0231FFA2FF90FF14FDC7FD6AF96A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBAFF8AFF44FE3EFA2CFDCE.text	A50C0231FFBAFF8AFF44FE3EFA2CFDCE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aberrantidrilus cuspis (Erseus & Dumnicka 1988)	<div><p>Aberrantidrilus cf. cuspis</p><p>Romana Cave, Venezia Giulia, Italy and Škocjanske Cave, Slovenia (Sambugar et al. 1999) [see comments above]</p></div>	https://treatment.plazi.org/id/A50C0231FFBAFF8AFF44FE3EFA2CFDCE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBAFF8AFF44FEAEFD2CFE06.text	A50C0231FFBAFF8AFF44FEAEFD2CFE06.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aberrantidrilus cuspis (Erseus & Dumnicka 1988)	<div><p>Aberrantidrilus cuspis (Erséus &amp; Dumnicka, 1988)</p><p>Umbrian wells near Perugia, Italy (Erséus &amp; Dumnicka 1988; Dumnicka 1990); Bocca Lupara Cave, La Spezia, Liguria, Italy, Papipano Cave, Udine, Friuli, Italy and Aven de la Cuisinière Cave, Calcomier, Aveyron, France (Sambugar et al. 1999) [see comments above]</p></div>	https://treatment.plazi.org/id/A50C0231FFBAFF8AFF44FEAEFD2CFE06	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBAFF8AFF44FDC6FC93FDB6.text	A50C0231FFBAFF8AFF44FDC6FC93FDB6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aberrantidrilus mihaljevici Vuckovic, Rodriguez & Kerovec 2020	<div><p>Aberrantidrilus mihaljevici Vučković, Rodriguez &amp; Kerovec, 2020</p><p>Tufa barrier of Roški slap, Croatia (Rodriguez et al. 2020)</p></div>	https://treatment.plazi.org/id/A50C0231FFBAFF8AFF44FDC6FC93FDB6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBAFF8AFF44FD8EFC95FD7E.text	A50C0231FFBAFF8AFF44FD8EFC95FD7E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aberrantidrilus sketi Martinez-Ansemil & Sambugar 2025	<div><p>Aberrantidrilus sketi Martínez-Ansemil &amp; Sambugar sp. nov.</p><p>Cave 4101, Brje pri Koprivi, Sežana, Slovenia (this work)</p></div>	https://treatment.plazi.org/id/A50C0231FFBAFF8AFF44FD8EFC95FD7E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBAFF8AFF44FD56FA20FD25.text	A50C0231FFBAFF8AFF44FD56FA20FD25.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aberrantidrilus stephaniae Martin 2015	<div><p>Aberrantidrilus stephaniae Martin, 2015</p><p>Hyporheic zone in alluvial aquifer at Bévéra River, Sospel, Mercantour National Park, France (Martin et al. 2015)</p></div>	https://treatment.plazi.org/id/A50C0231FFBAFF8AFF44FD56FA20FD25	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBAFF8AFF44FD1EFDF3FC89.text	A50C0231FFBAFF8AFF44FD1EFDF3FC89.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aberrantidrilus subterraneus (Rodriguez & Giani 1989)	<div><p>Aberrantidrilus subterraneus (Rodriguez &amp; Giani, 1989)</p><p>Jivero 1 Cave at Ozana, Matienzo, Cantabria, Spain (Rodríguez &amp; Giani 1989); Alisas–Ramales karst, Matienzo, Cantabria, Spain (Achurra et al. 2015)</p></div>	https://treatment.plazi.org/id/A50C0231FFBAFF8AFF44FD1EFDF3FC89	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBAFF8AFF44FC4AFB91FBC5.text	A50C0231FFBAFF8AFF44FC4AFB91FBC5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aktedrilus argatxae Giani & Rodriguez 1988	<div><p>Aktedrilus argatxae Giani &amp; Rodriguez, 1988</p><p>Karstic spring of Argatxa, Bizkaia, Spain (Giani &amp; Rodriguez 1988); Santa Eufemia–Ereñozar, Bizkaia, Spain, Llanes–Ribadesella and Gijón–Villaviciosa, Asturias, Spain (Achurra et al. 2015)</p></div>	https://treatment.plazi.org/id/A50C0231FFBAFF8AFF44FC4AFB91FBC5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBAFF8AFF44FBFEFC92FBAA.text	A50C0231FFBAFF8AFF44FBFEFC92FBAA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aktedrilus cuneus Erseus 1984	<div><p>Aktedrilus cuneus Erséus, 1984</p><p>Mineral spring in Charco Verde, Isla de la Palma, Canary Islands (Erséus, 1992b). A. cuneus is a species commonly found in marine intertidal environments around the world.</p></div>	https://treatment.plazi.org/id/A50C0231FFBAFF8AFF44FBFEFC92FBAA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBAFF8AFF44FB62FF6FFB3E.text	A50C0231FFBAFF8AFF44FB62FF6FFB3E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aktedrilus leeuwinensis Pinder 2006	<div><p>Aktedrilus leeuwinensis Pinder, 2006</p><p>Budjur Mar Cave (pool area of a stream, 25 m asl), Leeuwin–Naturaliste Ridge, Western Australia (Pinder et al. 2006)</p></div>	https://treatment.plazi.org/id/A50C0231FFBAFF8AFF44FB62FF6FFB3E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBAFF8AFF44FB16FD53FAAE.text	A50C0231FFBAFF8AFF44FB16FD53FAAE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aktedrilus parvithecatus (Erseus 1978)	<div><p>Aktedrilus parvithecatus (Erséus, 1978)</p><p>Anchyaline karst window about 500 m from the sea at Cape Range, Western Australia (Pinder et al. 2006). A. parvithecatus is a circumtropical species, also found in brackish anchialine waters in the Cabo Verde Islands (Erséus 1992b) and frequent in littoral marine environments.</p></div>	https://treatment.plazi.org/id/A50C0231FFBAFF8AFF44FB16FD53FAAE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBAFF8AFF44FA66FEC7FA32.text	A50C0231FFBAFF8AFF44FA66FEC7FA32.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aktedrilus podeilema Pinder 2006	<div><p>Aktedrilus podeilema Pinder, 2006</p><p>Tombstone Rocks Cave (deep pool of freshwater about 9 km from the ocean), Nambung region, Western Australia (Pinder et al. 2006)</p></div>	https://treatment.plazi.org/id/A50C0231FFBAFF8AFF44FA66FEC7FA32	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBAFF8AFF44FA0AFCE5F9FA.text	A50C0231FFBAFF8AFF44FA0AFCE5F9FA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aktedrilus ruffoi Sambugar, Giani & Martinez-Ansemil 1999	<div><p>Aktedrilus ruffoi Sambugar, Giani &amp; Martínez-Ansemil, 1999</p><p>Spring of the river Tione, Verona, Italy (Sambugar et al. 1999)</p></div>	https://treatment.plazi.org/id/A50C0231FFBAFF8AFF44FA0AFCE5F9FA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBAFF8AFF44F9D2FC69F9A2.text	A50C0231FFBAFF8AFF44F9D2FC69F9A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aktedrilus yacoubii Martin & Boughrous 2012	<div><p>Aktedrilus yacoubii Martin &amp; Boughrous, 2012</p><p>Exsurgence Aïn Maarouf, El Hajeb, Morocco (Martin &amp; Boughrous, 2012)</p></div>	https://treatment.plazi.org/id/A50C0231FFBAFF8AFF44F9D2FC69F9A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBAFF8AFF44F922FC48F8D2.text	A50C0231FFBAFF8AFF44F922FC48F8D2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cantabrodrilus pilarae Martinez-Ansemil 2025	<div><p>Cantabrodrilus pilarae Martínez-Ansemil gen. nov., sp. nov.</p><p>Latarmá stream near Toyu cave, Venta Fresnedo, Cantabria, Spain (this work)</p></div>	https://treatment.plazi.org/id/A50C0231FFBAFF8AFF44F922FC48F8D2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBAFF8AFF44F8B2FCCEF842.text	A50C0231FFBAFF8AFF44F8B2FCCEF842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coralliodrilus amissus Arslan, Timm & Erseus 2007	<div><p>Coralliodrilus amissus Arslan, Timm &amp; Erséus, 2007</p><p>Göksu stream, West-Central Anatolia, Turkey (Arslan et al. 2007)</p></div>	https://treatment.plazi.org/id/A50C0231FFBAFF8AFF44F8B2FCCEF842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBBFF8BFF44FF5EFCCEFF2F.text	A50C0231FFBBFF8BFF44FF5EFCCEFF2F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gianius anatolicus Arslan, Timm & Erseus 2007	<div><p>Gianius anatolicus Arslan, Timm &amp; Erséus, 2007</p><p>Göksu stream, West-Central Anatolia, Turkey (Arslan et al. 2007)</p></div>	https://treatment.plazi.org/id/A50C0231FFBBFF8BFF44FF5EFCCEFF2F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBBFF8BFF44FEE6FC5DFE06.text	A50C0231FFBBFF8BFF44FEE6FC5DFE06.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gianius aquaedulcis (Hrabe 1960)	<div><p>Gianius aquaedulcis (Hrabĕ, 1960)</p><p>Several localities along the lower part of the River Weser, Germany (Hrabĕ 1960); Mairuelegorreta Cave at Zigoitia, Alava, Spain, Jivero 1 Cave at Ozana, Matienzo, Cantabria, Spain and Labouiche Cave, Vernajoul, Ariège, France (Rodriguez &amp; Giani 1989); Tonawanda Channel, Niagara River, North America (Farara &amp; Erséus 1991); Santa Eufemia–Ereñozar, Bizkaia (Achurra &amp; Rodriguez 2008 and Achurra et al. 2015); Itxina, Gorbeia, Bizkaia, Spain, and Alisas–Ramales karst, Matienzo, Cantabria, Spain (Achurra et al. 2015)</p></div>	https://treatment.plazi.org/id/A50C0231FFBBFF8BFF44FEE6FC5DFE06	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBBFF8BFF44FE3EFC37FDCF.text	A50C0231FFBBFF8BFF44FE3EFC37FDCF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gianius camachoae Martinez-Ansemil 2025	<div><p>Gianius camachoae Martínez-Ansemil sp. nov.</p><p>Several groundwater environments in Cantabria, Spain (this work)</p></div>	https://treatment.plazi.org/id/A50C0231FFBBFF8BFF44FE3EFC37FDCF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBBFF8BFF44FDC6FAB5FDB7.text	A50C0231FFBBFF8BFF44FDC6FAB5FDB7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gianius cavealis Juget & des Chatelliers 2001	<div><p>Gianius cavealis Juget &amp; des Chȃtelliers, 2001</p><p>Le Trou des Voleurs Cave, near Poncin, Southern Jura, France (Juget &amp; Creuzé des Chȃtelliers 2001)</p></div>	https://treatment.plazi.org/id/A50C0231FFBBFF8BFF44FDC6FAB5FDB7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBBFF8BFF44FD8EFBB4FD7F.text	A50C0231FFBBFF8BFF44FD8EFBB4FD7F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gianius crypticus (Rodriguez & Giani 1989)	<div><p>Gianius crypticus (Rodríguez &amp; Giani, 1989)</p><p>Orillon Cave at Ozana, Matienzo, Cantabria, Spain (Rodriguez &amp; Giani 1989)</p></div>	https://treatment.plazi.org/id/A50C0231FFBBFF8BFF44FD8EFBB4FD7F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBBFF8BFF44FD56FC59FD27.text	A50C0231FFBBFF8BFF44FD56FC59FD27.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gianius inermis Sambugar and Martinez-Ansemil 2025	<div><p>Gianius inermis Sambugar and Martínez-Ansemil sp. nov.</p><p>Several groundwater environments in the Lessinian Massif, Italy (this work)</p></div>	https://treatment.plazi.org/id/A50C0231FFBBFF8BFF44FD56FC59FD27	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBBFF8BFF44FD1EFC1CFCEF.text	A50C0231FFBBFF8BFF44FD1EFC1CFCEF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gianius labouichensis (Rodriguez & Giani 1989)	<div><p>Gianius labouichensis (Rodriguez &amp; Giani, 1989)</p><p>Labouiche Cave, Vernajoul, Ariège, France (Rodriguez &amp; Giani 1989)</p></div>	https://treatment.plazi.org/id/A50C0231FFBBFF8BFF44FD1EFC1CFCEF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBBFF8BFF44FCA6FCE9FC57.text	A50C0231FFBBFF8BFF44FCA6FCE9FC57.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gianius navarroi Rodriguez & Achurra 2010	<div><p>Gianius navarroi Rodriguez &amp; Achurra, 2010</p><p>Ojo Guareña Cave, Burgos, Spain (Rodriguez &amp; Achurra 2010)</p></div>	https://treatment.plazi.org/id/A50C0231FFBBFF8BFF44FCA6FCE9FC57	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBBFF8BFF44FC6EFCFBFC1F.text	A50C0231FFBBFF8BFF44FC6EFCFBFC1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gianius parvogenitalis Martinez-Ansemil 2025	<div><p>Gianius parvogenitalis Martínez-Ansemil sp. nov.</p><p>Upwelling of the Clarión stream, Cantabria, Spain (this work)</p></div>	https://treatment.plazi.org/id/A50C0231FFBBFF8BFF44FC6EFCFBFC1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBBFF8BFF44FC36FF6FFB8F.text	A50C0231FFBBFF8BFF44FC36FF6FFB8F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gianius riparius (Giani & Martinez-Ansemil 1981)	<div><p>Gianius riparius (Giani &amp; Martínez-Ansemil, 1981)</p><p>Hyporheic flow of the river Tambre, Galicia, Spain (Giani &amp; Martínez-Ansemil 1981); hyporheic flow of the river Dordogne, France (Lafont 1989); underground water at Ile du Grand Gravier, river Rhône (Lafont &amp; Durbec 1990)</p></div>	https://treatment.plazi.org/id/A50C0231FFBBFF8BFF44FC36FF6FFB8F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBBFF8BFF44FB43FA24FB33.text	A50C0231FFBBFF8BFF44FB43FA24FB33.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pectinodrilus ningaloo Pinder 2006	<div><p>Pectinodrilus ningaloo Pinder, 2006</p><p>Anchialine groundwater (well about 500 m from the ocean) at Cape Range, Western Australia (Pinder et al. 2006)</p></div>	https://treatment.plazi.org/id/A50C0231FFBBFF8BFF44FB43FA24FB33	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBBFF8BFF44FAD7FB6BFAA6.text	A50C0231FFBBFF8BFF44FAD7FB6BFAA6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phallobaikalus gladiiseta (Martin & Brinkhurst 1998)	<div><p>Phallobaikalus gladiiseta (Martin &amp; Brinkhurst, 1998)</p><p>Lake Baikal, southern Siberia, Russia (Martin &amp; Brinkhurst 1998), as Rhyacodriloides gladiiseta .</p></div>	https://treatment.plazi.org/id/A50C0231FFBBFF8BFF44FAD7FB6BFAA6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBBFF8BFF44FA27FBBCF9D7.text	A50C0231FFBBFF8BFF44FA27FBBCF9D7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phallodriloides macmasterae (Erseus 1986)	<div><p>Phallodriloides macmasterae (Erséus, 1986)</p><p>Inland anchialine Prospero Cave, near Castle Harbour, Bermuda (Erséus 1986)</p></div>	https://treatment.plazi.org/id/A50C0231FFBBFF8BFF44FA27FBBCF9D7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBBFF8BFF44F9B7FD4BF92B.text	A50C0231FFBBFF8BFF44F9B7FD4BF92B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spiridion modricensis (Hrabe 1973)	<div><p>Spiridion modricensis (Hrabĕ, 1973)</p><p>Salt spring near Zadar, Dalmatia, Croatia (Hrabĕ 1973); Jama pri ogradi, Gangarol island, Murter, Croatia (Martínez-Ansemil et al. 2016); freshwater (probably influenced by sea water) at the mouth of a small stream, Tjärnö, Sweden (Erséus et al. 1992). S. modricensis is also present in marine littoral sandy habitats, and is qualify as a probably oligohaline brackish-water species (Erséus et al. 1992)</p></div>	https://treatment.plazi.org/id/A50C0231FFBBFF8BFF44F9B7FD4BF92B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFBBFF8BFF44F8E3FC36F82F.text	A50C0231FFBBFF8BFF44F8E3FC36F82F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spiridion phreaticola (Juget 1987)	<div><p>Spiridion phreaticola (Juget, 1987)</p><p>Hyporheic and phreatic waters at the alluvial plain of the river Rhône, upstream from Lyon, France (Juget 1987); hyporheic flow of the river Neste d’Aure, Hautes-Pyrénées, middle reach of the river Dordogne, and Labouiche Cave, Vernajoul, Ariège, France (Erséus et al. 1992); hyporheos of the Lachein creek, Alas, Ariège (Route et al. 2004); alluvial aquifer at Forez plain, catchment of the river Loire, France (Paran et al. 2005) [more detailed data of the French sites are given in Artheau &amp; Giani (2006)]; spring v Dragi 1, spring at Resnik, spring Malence and hyporheal of the stream Borovniščica, Slovenia (Giani et al. 2011)</p></div>	https://treatment.plazi.org/id/A50C0231FFBBFF8BFF44F8E3FC36F82F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
A50C0231FFB8FF88FF44FF5EFC13FF2E.text	A50C0231FFB8FF88FF44FF5EFC13FF2E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thalassodrilus hallae (Cook & Hiltunen 1975)	<div><p>Thalassodrilus hallae (Cook &amp; Hiltunen, 1975)</p><p>Saint Lawrence Great Lakes, North America (Cook &amp; Hiltunen 1975)</p></div>	https://treatment.plazi.org/id/A50C0231FFB8FF88FF44FF5EFC13FF2E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Martínez-Ansemil, Enrique;Sambugar, Beatrice	Martínez-Ansemil, Enrique, Sambugar, Beatrice (2025): Inventory of the inland-water Phallodrilinae (Clitellata: Naididae), with descriptions of a new genus and five new species from groundwaters of southern Europe. Zootaxa 5692 (3): 446-468, DOI: 10.11646/zootaxa.5692.3.2, URL: https://doi.org/10.11646/zootaxa.5692.3.2
