identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
B508A3A24FAB59BF84BCFEC207B96689.text	B508A3A24FAB59BF84BCFEC207B96689.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolbelasmus Boucomont 1910	<div><p>Bolbelasmus Boucomont, 1910</p><p>Diagnosis.</p><p>Species of Bolbelasmus are small to medium-sized (5.6-15.2 mm in body length) and can be recognized by the glossy, unicolored, reddish-brown to black dorsal surface; presence of a conical frontal tubercle with a rounded or bifurcated tip in males; transverse frontal carina present in females; eyes protruding, divided by a canthus anteriorly, canthus with anterior margin smooth; antennal club with first segment glabrous on inner side; pronotum usually quadrituberculate in major males (vestigial or reduced to bituberculate in minor males), females with transverse carina only; first elytral stria terminated by scutellum; parameres usually weakly sclerotized.</p></div>	https://treatment.plazi.org/id/B508A3A24FAB59BF84BCFEC207B96689	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Chun-Lin;Wang, Chuan-Chan	Li, Chun-Lin, Wang, Chuan-Chan (2024): A systematic review of the genus Bolbelasmus Boucomont (Coleoptera, Geotrupidae, Bolboceratinae) from Indochina and surrounding areas. ZooKeys 1191: 287-305, DOI: http://dx.doi.org/10.3897/zookeys.1191.114021, URL: http://dx.doi.org/10.3897/zookeys.1191.114021
13FA9162948C5A5A92FF18A5591DFCB1.text	13FA9162948C5A5A92FF18A5591DFCB1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolbelasmus chifengi Wang & Li 2024	<div><p>Bolbelasmus chifengi Wang &amp; Li sp. nov.</p><p>Figs 3, 4, 12, 18, 24, 31, 32, 40, 46, 47</p><p>Type materials.</p><p>Holotype male. "China: Yunnan, Bangdashan (邦達山), 16.IX.2015. leg. Y.-T. Wang." (glued on label, TARI), Taichung, Taiwan. Paratypes. 3♂♂, 1♀(TARI). same collecting data as the holotype. 1♀ (TARI). "China: Yunnan, Wudian (武甸), 17.IX.2014. leg. Y.-T. Wang". 1♂, 2♀♀(TARI). "China: Yunnan, CCCC, Nabang (那邦), 21.VI.2017. leg. Y.-T. Wang". 1♀ (CCLI). "China: Yunnan, Banggunjianshan (邦棍尖山), 19.IX.2015. leg. Y.-T. Wang". 1♂ (CCLI). "China: Yunnan, Bangdashan (邦達山), 01.IX.2015. leg. Y.-T. Wang". 2♂♂, 1♀(CCLI). "China: Yunnan, Ruili (瑞麗), 01.IX.2014. leg. Y.-T. Wang".</p><p>Description.</p><p>Holotype male (Figs 3, 4). Body length 9.7 mm; width across humeri 6.0 mm. Dorsum moderately shiny. Head, pronotum and scutellum dark brown with elytra reddish brown. Head (Fig. 12): labrum with anterior margin crenulate, disc transversally rugose. Clypeus trapezoidal, surface densely rugopunctate; protrusion at basal angle moderately developed. Clypeofrontal suture well defined, distinctly curved in front of frontal tubercle. Frons with surface sparsely punctate, punctures fine, frontal tubercle vertically located in junction of suture, right-triangle in shape in lateral view. Eye prominent, canthus simple, not exceeding eye. Thorax: pronotum (Fig. 18) quadrituberculate, tubercles placed in a line, lateral tubercle greatly reduced in size; anterior face of median tubercles almost perpendicular to plane of pronotum; primary punctures coarse, moderately distributed on disc and intermixed with impunctate area, punctures between lateral margins of pronotum and fovea bigger and denser, posterior area between elytral humeri and suture impunctate except for four coarse punctures in front of scutellum, secondary punctures tiny, evenly scattered throughout surface of pronotum; frontal and lateral margins beaded, posterior margin beaded only in front of elytral humeri. Scutellum elongate, secondary punctures sparse throughout surface with a coarse puncture at center. Elytron (Fig. 24): elytral striae shallow, punctures mostly spaced 2-3 times diameters of punctures. Intervals slightly convex including sutural one, surface with scattered secondary punctures. Male genitalia (Figs 31, 32, 40).</p><p>Female (Figs 46, 47). Body length 7.7-10.0 mm; width across humeri 5.4-6.5 mm. Similar to male with minor differences in the form of strongly wrinkled surface of clypeus, transverse frontal carina trilobed, central lobe more prominent than lateral lobes, punctures on frons and vertex rugose, transverse pronotal carina feebly bilobed with lobes broadly developed, punctures on pronotal disc coarser and denser than males.</p><p>Variation in male. Dorsum brown, smaller body size, 6.6 mm in length and 5.2 mm in width across humeri, frontal tubercle less developed and not in junction of clypeofrontal suture, pronotal tubercles feebly convex, and number of coarse punctures arranged at pronotal posterior margin variable.</p><p>Diagnosis.</p><p>Bolbelasmus chifengi is morphologically similar to B. concavisuturalis, but can be distinguished from the latter by the coarser and denser primary punctures on the pronotal disc (finer and scattered in B. concavisuturalis), elytral intervals evenly convex (elytral intervals flat with sutural interval concave in B. concavisuturalis) and by the longer parameres (shorter in B. concavisuturalis).</p><p>Distribution.</p><p>Southern Yunnan, China (Fig. 52).</p><p>Etymology.</p><p>Bolbelasmus chifengi sp. nov. is named after Dr Chi-feng Lee, the curator of the Department of Applied Zoology, Taiwan Agriculture Research Institute, Taichung, Taiwan, who kindly provided materials used in this study.</p></div>	https://treatment.plazi.org/id/13FA9162948C5A5A92FF18A5591DFCB1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Chun-Lin;Wang, Chuan-Chan	Li, Chun-Lin, Wang, Chuan-Chan (2024): A systematic review of the genus Bolbelasmus Boucomont (Coleoptera, Geotrupidae, Bolboceratinae) from Indochina and surrounding areas. ZooKeys 1191: 287-305, DOI: http://dx.doi.org/10.3897/zookeys.1191.114021, URL: http://dx.doi.org/10.3897/zookeys.1191.114021
04B3E5DBEF5D5AC68E2205860EBA22D6.text	04B3E5DBEF5D5AC68E2205860EBA22D6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolbelasmus concavisuturalis Li & Wang 2024	<div><p>Bolbelasmus concavisuturalis Li &amp; Wang sp. nov.</p><p>Figs 1, 2, 11, 17, 23, 29, 30, 39, 44, 45</p><p>Type materials.</p><p>Holotype male. "Mon-Angget, near Chiangmai, North Thailand, 31-V-1990, K. Masumoto leg." (glued on label, NSMT). Paratypes. 1♂ (ZMUC) "Northern Thailand, Doi Sutep, 21.6.1958, B. Degerbøl leg., Pr. 548 (1.7.59)". 1♀ (NSMT) "Doi Suthep, Chiang Mai, Thailand, 15-VIII-1989, Y. MANIT leg."</p><p>Description.</p><p>Holotype male (Figs 1, 2). Body length 10.2 mm; width across humeri 6.3 mm. Dorsum distinctly shiny. Head, pronotum and scutellum reddish brown with elytra brown in color. Head (Fig. 11): labrum with anterior margin crenulate, disc transversally rugose. Clypeus trapezoidal, surface densely rugopunctate; protrusion at basal angle reduced. Clypeofrontal suture well defined, distinctly curved in front of frontal tubercle. Frons with surface sparsely punctate, punctures fine, frontal tubercle vertically located in junction of suture, triangular in lateral view. Eye prominent, canthus simple, not exceeding eye. Thorax: pronotum (Fig. 17) quadrituberculate, tubercles situated in a line, lateral tubercle smaller; anterior face of median tubercles almost perpendicular to surface of pronotum; primary punctures weakly defined, sparse on disc except between lateral margin of pronotum and fovea, these coarser and denser, line in front of scutellum with a coarse puncture, secondary punctures tiny, evenly scattered on surface of pronotum; frontal and lateral margins beaded, posterior margin beaded only anterior to humeri of elytra. Scutellum elongate, secondary punctures sparsely distributed. Elytron (Fig. 23): elytral striae shallowly impressed, punctures mostly spaced 2-3 times diameters of punctures. Intervals flat, with sutural interval weakly concave, surface scattered with secondary punctures. Male genitalia (Figs 29, 30, 39).</p><p>Female paratype (Figs 44, 45). Body length 9.8 mm; width across humeri 5.8 mm. Similar to male with minor differences in the form of strongly wrinkled surface of clypeus, transverse frontal carina trilobed, central lobe more prominent than lateral lobes, punctures on frons and vertex rugose, transverse pronotal carina feebly bilobed, lobes broad, punctures on pronotal disc coarser and denser than those of males.</p><p>Male paratype. The single male paratype is smaller in body size, 9.4 mm in length and 5.1 mm in width across humeri, frontal tubercle less developed and with three coarse punctures along pronotal posterior margin in front of scutellum.</p><p>Diagnosis.</p><p>Bolbelasmus concavisuturalis sp. nov. is morphologically similar to B. coreanus, but can be distinguished from the latter by having denser punctures along the midline of the pronotum (Fig. 17) (sparser punctures in B. coreanus (Fig. 22)), punctures in elytral striae moderately developed (Fig. 23) (punctures weakly developed in B. coreanus (Fig. 28)) and ventrally curved parameres (straight in B. coreanus).</p><p>Distribution.</p><p>Northern Thailand (Fig. 52).</p><p>Etymology.</p><p>Concavi - (L.) = concave, - suturalis (L.) = suture. In reference to the concave sutural intervals of the elytra.</p></div>	https://treatment.plazi.org/id/04B3E5DBEF5D5AC68E2205860EBA22D6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Chun-Lin;Wang, Chuan-Chan	Li, Chun-Lin, Wang, Chuan-Chan (2024): A systematic review of the genus Bolbelasmus Boucomont (Coleoptera, Geotrupidae, Bolboceratinae) from Indochina and surrounding areas. ZooKeys 1191: 287-305, DOI: http://dx.doi.org/10.3897/zookeys.1191.114021, URL: http://dx.doi.org/10.3897/zookeys.1191.114021
6A70B116BB565A2BA23B6323990FDA36.text	6A70B116BB565A2BA23B6323990FDA36.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolbelasmus coreanus (Kolbe 1886)	<div><p>Bolbelasmus coreanus (Kolbe, 1886)</p><p>Bolboceras coreanus Kolbe, 1886: 188. Original combination (type locality: Seoul, Korea, female type in the Museum für Naturkunde, Berlin, Germany).</p><p>Bolbelasmus coreanus (Kolbe, 1886): Cartwright 1953: 97 (generic combination); Krikken 1977: 288 (notes; diagnosis; illustration); Kim 2000: 45 (diagnosis; collecting records); Li et al. 2008: 480 (redescription, illustrations, collecting records, distribution, remarks); Král, Löbl and Nikolajev 2006: 83 (catalog, subgenus Bolbelasmus Kolbeus); Nikolajev, Král and Bezdӗk 2016: 33 (catalog, subgenus Bolbelasmus Kolbeus).</p><p>Kolbeus coreanus (Kolbe, 1886): Boucomont 1911: 335 (generic combination); Boucomont 1912: 17 (catalog); Boucomont and Gillet 1921: 72 (record to Taiwan; diagnosis); Miwa 1930: 164 (catalog); Miwa 1931: 276 (catalog); Miwa and Chûjô 1939: 30 (catalog); Paulian 1945: 42 (diagnosis; figures; distribution).</p><p>Bolbelasmus kurosawai Masumoto, 1984: 76; Li et al. 2008: 481 (junior synonym).</p><p>Bolboceras conicifrons Fairmaire, 1896: 82; Boucomont and Gillet 1921: 72 (junior synonym).</p><p>Distribution.</p><p>Korean Peninsula; China (Anhui, Zhejiang, Kweichow, Szechuan, Yunnan); Taiwan.</p><p>Remarks.</p><p>The detailed distribution of B. coreanus in China requires further investigation, particularly those from southwestern areas. Based on a large number of Bolbelasmus specimens recently collected from Yunnan and neighboring areas, we found no representatives of B. coreanus among them. Therefore, we reserve a decision about whether B. coreanus occurs in Yunnan, Thailand and India, as recorded by Krikken (1977). Voucher specimens from the areas mentioned above are required.</p></div>	https://treatment.plazi.org/id/6A70B116BB565A2BA23B6323990FDA36	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Chun-Lin;Wang, Chuan-Chan	Li, Chun-Lin, Wang, Chuan-Chan (2024): A systematic review of the genus Bolbelasmus Boucomont (Coleoptera, Geotrupidae, Bolboceratinae) from Indochina and surrounding areas. ZooKeys 1191: 287-305, DOI: http://dx.doi.org/10.3897/zookeys.1191.114021, URL: http://dx.doi.org/10.3897/zookeys.1191.114021
8409E5B68AF95ABB8545DE166EA5384F.text	8409E5B68AF95ABB8545DE166EA5384F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolbelasmus korshunovi Zinchenko 2016	<div><p>Bolbelasmus korshunovi Zinchenko, 2016</p><p>Figs 7, 8, 14, 20, 26, 35, 36, 42</p><p>Bolbelasmus korshunovi Zinchenko, 2016: 328. Original combination (type locality: Nong Bun Nak, Nakhon Prov., Thailand).</p><p>Material examined.</p><p>(5♂♂). 2♂♂ (ZMUC). Thailand: Loei Province, Phu Luang Wildlife Sanctuary, 8.-14.x.1984, 700-900 m, Karsholt, Lomboldt &amp; Nielsen leg., Pral Siaw, 1923-9-33, Paūl Fogh / Coll. Roseberg. 3♂♂ (NSMT). Sansai, Chiang Mai, Thailand, 17. VI. 1993 .</p><p>Diagnosis.</p><p>Body length, males, 6.8-9.0 mm, greatest width at pronotal base 4.4-5.7 mm; females, 7.1-8.7 mm in length, 4.3-5.6 mm in width (Zinchenko 2016). Bolbelasmus korshunovi is distinguished from the other Oriental Bolbelasmus species by elytral sutural intervals that are moderately convex, primary punctures sparsely distributed either side of the center of the pronotum, and shapes of the parameres.</p><p>Notes.</p><p>Thirteen type specimens were designated in the original description of the species (Zinchenko 2016), 12 of them collected from June to August, and the holotype during November. Accordingly, the temporal activity of adults is likely at least half the year during both rainy and dry seasons. This is identical to the sympatric species, B. meridionalis .</p><p>Distribution.</p><p>Northern Thailand.</p><p>Remarks.</p><p>Bolbelasmus korshunovi inhabits plains to low-elevational montane areas and occurs sympatrically with B. meridionalis in northern Thailand.</p></div>	https://treatment.plazi.org/id/8409E5B68AF95ABB8545DE166EA5384F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Chun-Lin;Wang, Chuan-Chan	Li, Chun-Lin, Wang, Chuan-Chan (2024): A systematic review of the genus Bolbelasmus Boucomont (Coleoptera, Geotrupidae, Bolboceratinae) from Indochina and surrounding areas. ZooKeys 1191: 287-305, DOI: http://dx.doi.org/10.3897/zookeys.1191.114021, URL: http://dx.doi.org/10.3897/zookeys.1191.114021
6B60480DF66550FFAC3D424931B0F1D0.text	6B60480DF66550FFAC3D424931B0F1D0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolbelasmus krikkeni Nikolajev 1979	<div><p>Bolbelasmus krikkeni Nikolajev, 1979</p><p>Bolbelasmus krikkeni Nikolajev, 1979: 225. Original combination (type locality: Gopaldhara, Sikkim, India); Král, Löbl and Nikolajev 2006: 83 (catalog, in subgenus Bolbelasmus Kolbeus); Nikolajev, Král and Bezdӗk 2016: 33 (catalog, subgenus Bolbelasmus Kolbeus).</p><p>Distribution.</p><p>Northern India.</p><p>Remarks.</p><p>Based on the collecting data from the monotypic specimen, B. krikkeni occurs in mid-elevation forests above 1000 m and is unique compared to its congeners that usually inhabit plains to low-elevation montane areas in the region. No additional specimens have been recorded since the publication of the original description.</p></div>	https://treatment.plazi.org/id/6B60480DF66550FFAC3D424931B0F1D0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Chun-Lin;Wang, Chuan-Chan	Li, Chun-Lin, Wang, Chuan-Chan (2024): A systematic review of the genus Bolbelasmus Boucomont (Coleoptera, Geotrupidae, Bolboceratinae) from Indochina and surrounding areas. ZooKeys 1191: 287-305, DOI: http://dx.doi.org/10.3897/zookeys.1191.114021, URL: http://dx.doi.org/10.3897/zookeys.1191.114021
E0B0EB6A0CC35F4CA382A1C954C295D7.text	E0B0EB6A0CC35F4CA382A1C954C295D7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolbelasmus meridionalis Krikken 1977	<div><p>Bolbelasmus meridionalis Krikken, 1977</p><p>Figs 5, 6, 13, 19, 25, 33, 34, 41, 48, 49</p><p>Bolbelasmus meridionalis Krikken, 1977: 285. Original combination (type locality: Java, Indonesia); Král, Löbl and Nikolajev 2006: 83 (catalog, subgenus Bolbelasmus Kolbeus); Nikolajev, Král and Bezdӗk 2016: 33 (catalog, subgenus Bolbelasmus Kolbeus).</p><p>Material examined.</p><p>(11♂♂, 8♀♀). 1♂ (NSMT). Thailand: Sansai, Chiang Mai, 17. VI. 1993 (1 male at NSMT) . 9♂♂, 7♀♀ (NSMT); same locality, 12-V-1996 . 1♀ (NSMT); near Chiang Mai, N Thailand, VII-1996, native collector . 1♂ (NSMT). Taiwan: Formosa, Heito, 10-VII-1941, H. Kondo / Sizumu Nomura Bequest, 1981 .</p><p>Diagnosis.</p><p>Body length, males, 6.1-8.2 mm, greatest width at pronotal base, 3.7-5.0 mm; females, 5.6-8.2 mm in length, 3.3-5.2 mm in width. Both B. meridionalis and B. minutus constitute a distinctive group among southeastern Asian congeners based on sharing the distinctly convex elytral sutural intervals and the tips of the parameres acute and curved ventrally in lateral view. Due to a lack of further material being available of the later species, B. meridionalis and B. minutus can only be separated by the shape of male genitalia and the lateral margin of the pronotum in B. meridionalis, which is more widely explanate than that of B. minutus .</p><p>Chinese name.</p><p>脊背厚角金龜</p><p>Notes.</p><p>Li et al. (2008) excluded B. meridionalis from the registered Taiwan fauna due to the lack of verified records. During the present study, we examined a male B. meridionalis specimen housed in NSMT bearing identical labels as the paratype of the species collected in Heito (now Pingtung) by the late Japanese coleopterist, Yushiro Miwa. We therefore confirm the record of B. meridionalis in Taiwan, though it has been lacking in reports of the genus for 90 years. Consequently, the conservation status of B. meridionalis in Taiwan is in urgent need of study, along with that of B. minutus Li &amp; Masumoto, 2008 and Bolbotrypes davidis (Fairmaire, 1891). These species are restricted to habitats in highly urbanized areas and/or intensively farmed plains of Taiwan.</p><p>Distribution.</p><p>Indonesia (Java); eastern China; Thailand; Vietnam; Taiwan.</p><p>Remarks.</p><p>Bolbelasmus meridionalis has the widest known distribution among congeners in the region in eastern and southeastern Asia. Also, the records from Java for the holotype and paratypes indicated that it is the only member from the Sunda Islands of the genus.</p></div>	https://treatment.plazi.org/id/E0B0EB6A0CC35F4CA382A1C954C295D7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Chun-Lin;Wang, Chuan-Chan	Li, Chun-Lin, Wang, Chuan-Chan (2024): A systematic review of the genus Bolbelasmus Boucomont (Coleoptera, Geotrupidae, Bolboceratinae) from Indochina and surrounding areas. ZooKeys 1191: 287-305, DOI: http://dx.doi.org/10.3897/zookeys.1191.114021, URL: http://dx.doi.org/10.3897/zookeys.1191.114021
96D9C6B638E45F6882FAFB32CD2C99DC.text	96D9C6B638E45F6882FAFB32CD2C99DC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolbelasmus minutus Li & Masumoto 2008	<div><p>Bolbelasmus minutus Li &amp; Masumoto, 2008</p><p>Bolbelasmus minutus Li &amp; Masumoto, 2008: 482. Original combination (type locality: Heito (presently Pingtung), Taiwan); Nikolajev, Král and Bezdӗk 2016: 33 (catalog, subgenus Bolbelasmus Kolbeus).</p><p>Distribution.</p><p>Taiwan.</p><p>Remarks.</p><p>Bolbelasmus minutus was described from a pair of specimens collected during 1931, and no further records of the species have been recorded. This species occurs sympatrically with B. meridionalis and B. nativus in the plains of southern Taiwan.</p></div>	https://treatment.plazi.org/id/96D9C6B638E45F6882FAFB32CD2C99DC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Chun-Lin;Wang, Chuan-Chan	Li, Chun-Lin, Wang, Chuan-Chan (2024): A systematic review of the genus Bolbelasmus Boucomont (Coleoptera, Geotrupidae, Bolboceratinae) from Indochina and surrounding areas. ZooKeys 1191: 287-305, DOI: http://dx.doi.org/10.3897/zookeys.1191.114021, URL: http://dx.doi.org/10.3897/zookeys.1191.114021
34EDE9E61640502AA9D47695608C0F8B.text	34EDE9E61640502AA9D47695608C0F8B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolbelasmus nativus subsp. nativus Krikken 1977	<div><p>Bolbelasmus nativus nativus Krikken, 1977</p><p>Bolbelasmus nativus Krikken, 1977: 287. Original combination (type locality: Heito (presently Pingtung), Taiwan); Nikolajev, Král and Bezdӗk 2016: 33 (catalog, in nominate subgenus Bolbelasmus Bolbelasmus).</p><p>B. n. ishigakiensis ssp. Masumoto, 1984.</p><p>Bolbelasmus ishigakiensis Masumoto, 1984: 73. Original combination (type locality: Ishigaki island, Okinawa, Japan); Král, Löbl and Nikolajev 2006: 83 (catalog, subgenus Bolbelasmus Kolbeus); Ochi and Masumoto 2005: 244 (as subspecies of B. nativus); Nikolajev, Král and Bezdӗk 2016: 33 (catalog, subgenus Bolbelasmus Kolbeus).</p><p>Distribution.</p><p>Taiwan (southern areas and Lanyu island); Japan (Iriomote, Ishigaki and Tarama islands, Okinawa Prefecture).</p><p>Remarks.</p><p>Bolbelasmus nativus was originally described based on a single male from Taiwan. Ochi and Masumoto (2005) treated the population distributed on a few small islands near Taiwan as a subspecies, B. nativus ishigakiensis .</p></div>	https://treatment.plazi.org/id/34EDE9E61640502AA9D47695608C0F8B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Chun-Lin;Wang, Chuan-Chan	Li, Chun-Lin, Wang, Chuan-Chan (2024): A systematic review of the genus Bolbelasmus Boucomont (Coleoptera, Geotrupidae, Bolboceratinae) from Indochina and surrounding areas. ZooKeys 1191: 287-305, DOI: http://dx.doi.org/10.3897/zookeys.1191.114021, URL: http://dx.doi.org/10.3897/zookeys.1191.114021
4E4E95D6287F52E9B77D452832EF4044.text	4E4E95D6287F52E9B77D452832EF4044.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolbelasmus orientalis Petrovitz 1968	<div><p>Bolbelasmus orientalis Petrovitz, 1968</p><p>Bolbelasmus orientalis Petrovitz, 1968: 185. Original combination (type locality: Wladiwostok [Vladivostok], Primorskii Territory, Russia); Krikken 1977: 289 (notes; diagnosis; illustration); Nikolajev, Král and Bezdӗk 2016: 33 (catalog).</p><p>Distribution.</p><p>Described from Vladivostok, Russian Far East.</p><p>Remarks.</p><p>Bolbelasmus orientalis was described from one male and one female. Krikken (1977) examined both type specimens and found that the female allotype to be a minor male. Also, he noted that the species has no direct affinity with three congeners, B. coreanus, B. meridionalis and B. nativus, which are geographically close to B. orientalis but are allied to the western Palaearctic B. unicornis . Bezborodov and Koshkin (2014) doubted the locality label attached to the type specimens because there were no additional records of the species documented in Russia or nearby territories other than that of type specimens. We, therefore, exclude B. orientalis from the Bolbelasmus fauna in the eastern Palaearctic and the Oriental regions.</p></div>	https://treatment.plazi.org/id/4E4E95D6287F52E9B77D452832EF4044	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Chun-Lin;Wang, Chuan-Chan	Li, Chun-Lin, Wang, Chuan-Chan (2024): A systematic review of the genus Bolbelasmus Boucomont (Coleoptera, Geotrupidae, Bolboceratinae) from Indochina and surrounding areas. ZooKeys 1191: 287-305, DOI: http://dx.doi.org/10.3897/zookeys.1191.114021, URL: http://dx.doi.org/10.3897/zookeys.1191.114021
EA0192B91149563696F487E9239620EA.text	EA0192B91149563696F487E9239620EA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolbelasmus shibatai Masumoto 1984	<div><p>Bolbelasmus shibatai Masumoto, 1984</p><p>Bolbelasmus shibatai Masumoto, 1984: 75. Original combination (type locality: Amami Oshima Island, Japan); Nikolajev, Král and Bezdӗk 2016: 33 (catalog).</p><p>Distribution.</p><p>Japan (Amami oshima and Okinawa island).</p><p>Remarks.</p><p>Populations of B. shibatai are restricted to a few small islands in the southwestern archipelagos of Japan. Males possess strongly sclerotized parts of the parameres that can be distinguished from the similar species, B. coreanus .</p></div>	https://treatment.plazi.org/id/EA0192B91149563696F487E9239620EA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Chun-Lin;Wang, Chuan-Chan	Li, Chun-Lin, Wang, Chuan-Chan (2024): A systematic review of the genus Bolbelasmus Boucomont (Coleoptera, Geotrupidae, Bolboceratinae) from Indochina and surrounding areas. ZooKeys 1191: 287-305, DOI: http://dx.doi.org/10.3897/zookeys.1191.114021, URL: http://dx.doi.org/10.3897/zookeys.1191.114021
4203E0EB4F025F3D935674590CF2005F.text	4203E0EB4F025F3D935674590CF2005F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bolbelasmus yutangi Li & Wang 2024	<div><p>Bolbelasmus yutangi Li &amp; Wang sp. nov.</p><p>Figs 9, 10, 15, 21, 27, 37, 38, 43, 50, 51</p><p>Type materials.</p><p>Holotype male. "Myanmar: Bago Region, Moe Yin Gyi, CCCC, 21.V.2017. leg. Y.-T. Wang." (glued on label, TARI). Paratypes. 1♀(TARI). data as the holotype. 5 ♂♂ (TARI). "China: Yunnan, Wudian (武甸), 17.IX.2014. leg. Y.-T. Wang". 1♀ (CCLI) "China: Yunnan, Banggunjianshan (邦棍尖山), 18.IX.2015. leg. Y.-T. Wang". 6♂♂ (CCLI). "China: Yunnan, Bangdashan (邦達山), 01.IX.2015. leg. Y.-T. Wang". 1♂ (CCLI). "China: Yunnan, Ruili (瑞麗), 15.IX.2014. leg. Y.-T. Wang". 1♂ (NSMT). "Doi Saket, Chiang Mai, Thailand, 12-X-1989, Y. MANIT leg". 1♂ (NSMT). "Doi Mon Unggate, Samoeng Distr., Chiang Mai Prov., Thailand, 18-VII-1989, Y. MANIT leg".</p><p>Description.</p><p>Holotype male (Figs 9, 10). Body length 9.3 mm; width across humeri 5.9 mm. Dorsum moderately shiny. Head, pronotum and scutellum dark brown with elytra reddish brown. Head (Fig. 15): labrum with anterior margin crenulate, disc transversally rugose. Clypeus trapezoidal, surface densely rugopunctate; protrusion at basal angle moderately developed. Clypeofrontal suture well defined, slightly curved in front of frontal tubercle. Frons with surface moderately punctate, punctures coarse, frontal tubercle vertically located at center of disc, triangular when viewed laterally. Eye prominent, canthus simple, not exceeding eye. Thorax: pronotum (Fig. 21) quadrituberculate, tubercles placed in a line, lateral tubercle greatly reduced in size; anterior face of median tubercles almost perpendicular to plane of pronotum; primary punctures coarse, dense on disc with small impunctate area in front of middle of posterior margin, punctures between lateral margins of pronotum and fovea bigger and denser, scattered coarse punctures distributed along posterior margin with seven punctures in front of scutellum, secondary punctures tiny, evenly scattered on surface of pronotum; frontal and lateral margins beaded, posterior margin beaded only in front of elytral humeri. Scutellum elongate, fine punctures sparsely distributed on surface. Elytron (Fig. 27): elytral striae shallowly impressed, punctures mostly spaced by 1-3 times diameters of punctures. Intervals slightly convex, including sutural interval, surface with scattered secondary punctures. Male genitalia. Figs 37, 38, 43.</p><p>Female (Figs 50, 51). Body length 8.4-8.7 mm; width across humeri 5.1-5.3 mm. Similar to male with minor differences in the form of strongly wrinkled surface of clypeus, transverse frontal carina trilobed, central lobe more prominent than lateral lobes, punctures on frons and vertex rugose, transverse pronotal carina feebly bilobed, with lobes broadly developed to reduced, bigger punctures on pronotal disc denser than those of males and scutellum with 1 or 2 bigger punctures.</p><p>Variation in male. Male paratypes differ from the holotype in the following respects: smaller body size, 6.6 mm in body length and 5.2 mm in width across humeri, frontal tubercle less developed and not in junction of clypeofrontal suture, pronotal tubercle feebly convex, reduced, and number of coarse punctures along pronotal posterior margin variable.</p><p>Diagnosis.</p><p>Bolbelasmus yutangi is similar to B. nativus in sharing the frontal tubercle location at the center of the frons as well as having smaller parameres. It can be distinguished from the latter by the weakly convex basal angle of the clypeus (distinctly bulging in B. nativus), primary punctures densely distributed on the disc (almost impunctate along the middle of disc in B. nativus), sutural interval convex, similar to discal intervals (distinctly more convex than discal intervals in B. nativus) and parameres with the tips tapered (parameres broader at tips in B. nativus).</p><p>Distribution.</p><p>Eastern Myanmar, northern Thailand and western Yunnan, China (Fig. 52).</p><p>Etymology.</p><p>Bolbelasmus yutangi sp. nov. is named after Mr Yu-tang Wang, a beetle enthusiast of Taiwan, who collected most of the material used in this study.</p><p>Species with doubtful locality record</p></div>	https://treatment.plazi.org/id/4203E0EB4F025F3D935674590CF2005F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Chun-Lin;Wang, Chuan-Chan	Li, Chun-Lin, Wang, Chuan-Chan (2024): A systematic review of the genus Bolbelasmus Boucomont (Coleoptera, Geotrupidae, Bolboceratinae) from Indochina and surrounding areas. ZooKeys 1191: 287-305, DOI: http://dx.doi.org/10.3897/zookeys.1191.114021, URL: http://dx.doi.org/10.3897/zookeys.1191.114021
