identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
FB9F285E51FB58E99A5B63824F522FAE.text	FB9F285E51FB58E99A5B63824F522FAE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sulawesidrobia lasoloensis Kadow & von Rintelen & Iqram & Bichain & Haase 2025	<div><p>Sulawesidrobia lasoloensis sp. nov.</p><p>Etymology.</p><p>This species is named after the Lasolo River as it is the first description of a Sulawesidrobia occurring in the system of this river.</p><p>Material examined.</p><p>Holotype (Fig. 2 A): Indonesia • Sulawesi, North Konawe Regency, Amorome, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=121.98289&amp;materialsCitation.latitude=-3.4265556" title="Search Plazi for locations around (long 121.98289/lat -3.4265556)">northeast of Tambua, in a tributary of the Lasolo River</a>. 3°25'35.6"S, 121°58'58.4"E. July 2010; Jean-Michel Bichain leg.; MZB Gst. 12193.</p><p>Paratypes (Fig. 3 A, B): Indonesia • N = 20; same collection data as for holotype; MZB Gst. 12194 . N = 20; same collection data as for holotype; ZMB 118374 . N = 10; same collection data as for holotype; MHNEC -MOLL 2025_04_001 . N = 8; Sulawesi, North Konawe Regency, Amorome, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=121.98256&amp;materialsCitation.latitude=-3.428639" title="Search Plazi for locations around (long 121.98256/lat -3.428639)">northeast of Tambua, in a tributary of the Lasolo River</a>. 3°25'43.1"S, 121°58'57.2"E. July 2010; Jean-Michel Bichain leg. MZB Gst. 12195 . N = 7; same collection data, ZMB 118375 .</p><p>Diagnosis.</p><p>This new species is characterized by the cylindro-conical shell shape, the anterodorsal insertion of the bursal duct in the ovoid bursa copulatrix, and the penis resting in U-shape with a swelling in the distal third on the left side and a small, round lobe on the right side behind the swelling.</p><p>Description.</p><p>Shell. Cylindro-conical to conical, slender, 1.7 to 2 times higher than wide, 3.5 to 4.125 whorls, sutures shallow, whorls moderately flat in profile, semitranslucent, goldish-brown, periostracum without color (Figs 2 A, 3 A, B); protoconch (N = 4) wrinkled comprising 0.8–1 whorls (Fig. 4 A), teleoconch with growth lines, otherwise without structure; umbilicus a narrow slit; outer lip straight.</p><p>Operculum (N = 9). Light orange to yellow, thin, paucispiral, nucleus eccentric,</p><p>External features (N = 9). General appearance fairly light, black pigmentation rather sparse, but variable; eyes black, epidermal black pigment spots almost like eyebrows behind and toward the inner side of the eyes, tentacles otherwise white, proximal region of snout and head pigmented, mirrored, sickle-shaped spots on pharynx; mantle more or less loosely pigmented, almost unpigmented over distal genital glands in both sexes, upper parts of whorls darker than lower ones.</p><p>Mantle cavity (N = 8). Gill with 10–13 filaments almost abutting to pericard; osphradium ovate to elongate-ovate, about 1 / 3 of length of gill, rather behind middle of gill.</p><p>Digestive system (N = 3 for radula, 9 for rectum and stomach). Radula formula R: 4-5 1 4-5 / 2 - 3 2-3; L: 3-4 1 4-5, M 1: 16-22, M 2: 20-25; lateral tooth with membranous junction between face and flank (Fig. 5); stomach without caecum, outer side brown to black; rectum makes a short loop in roof of mantle cavity underneath distal genital glands, more conspicuous in males than in females (Fig. 6 A – C).</p><p>Female genitalia (N = 5; Fig. 7 A-C). Ovary a simple sac, starting 1–1.5 whorls below apex, comprising 0.25 to almost 0.5 whorls, reaching stomach; renal oviduct with U-shaped loop followed by 270 ° - loop underneath ovoid bursa copulatrix lying largely behind albumen gland, bursal duct short emerging anterodorsally, no receptaculum seminis, sperm storage in distal part of second oviduct loop; albumen gland milky-white, capsule gland bipartite with a posterior opaque white and an anterior milky-white portion.</p><p>Male genitalia (N = 4). Testis lobate, starting 0.75 to 1.25 whorls below apex, comprising more than 1 whorl, reaching or overlapping proximal part of stomach; vesicula seminalis coiling underneath distal half of testis; penis in rest U-shaped, tip pointing forward, for most of the length with parallel sides, a swelling in the distal third on the left side and a small, round lobe on the right side behind the swelling (Fig. 8 A), penial base with black pigment.</p><p>Remarks.</p><p>The cylindro-conical form of this new species is rare among species of Sulawesidrobia and only seen in S. angusta Haase &amp; Bouchet, 2006 from Lake Poso (Haase and Bouchet 2006), which has a smaller aperture and less convex whorls, though. The shapes of bursa copulatrix and penis are unique. The morphological variation at the type locality was higher than in the other stream with the slender forms occurring only in the former (Fig. 9). The broader form approaches S. marmer sp. nov. described below but differs through the smaller size, the less translucent shell, and the gold-brown color. Both species were well supported as sister species in the phylogenetic analyses. Their relationship to a clade of the Malili Lake system including S. abreui Zielske, Glaubrecht &amp; Haase, 2011, received moderate support (Figs 10, 11).</p><p>This species is the only species from Sulawesi where a membranous junction between face and flank in the lateral radular teeth typical for the majority of Pacific tateids (e. g., Haase and Zielske 2015) has been observed so far.</p></div>	https://treatment.plazi.org/id/FB9F285E51FB58E99A5B63824F522FAE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Kadow, Laura;von Rintelen, Thomas;Iqram, Muhammad;Bichain, Jean-Michel;Haase, Martin	Kadow, Laura, von Rintelen, Thomas, Iqram, Muhammad, Bichain, Jean-Michel, Haase, Martin (2025): Three new non-lacustrine species of the endemic freshwater gastropod genus Sulawesidrobia (Caenogastropoda, Truncatelloidea, Tateidae) from Sulawesi, Indonesia, with additional records of species from ancient Lake Poso. Zoosystematics and Evolution 101 (4): 1585-1600, DOI: 10.3897/zse.101.161812
EF22A6C280BD52B1A6FFAAB7830C9DFF.text	EF22A6C280BD52B1A6FFAAB7830C9DFF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sulawesidrobia marmer Kadow & von Rintelen & Iqram & Bichain & Haase 2025	<div><p>Sulawesidrobia marmer sp. nov.</p><p>Etymology.</p><p>The word marmer is Indonesian and means marble. It refers to the shell’s appearance reminiscent of marble and is used as noun in apposition.</p><p>Material examined.</p><p>Holotype (Fig. 2 B): Indonesia • Sulawesi, North Konawe Regency, Lasolo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=122.235306&amp;materialsCitation.latitude=-3.7185001" title="Search Plazi for locations around (long 122.235306/lat -3.7185001)">south of Abola, in a small stream next to the Wawotobi Road</a>, on rocks. 3°43'06.6"S, 122°14'07.1"E. September 2023; Thomas von Rintelen, Muhammad Iqram, and Sara Nejad leg. MZB Gst. 12187 .</p><p>Paratypes (Fig. 3 C, D): Indonesia • N = 15; same collection data as for holotype; MZB Gst. 12188 . N = 15; same collection data as for holotype; ZMB 118347 .</p><p>DIAGNOSIS. This new species has the conical shell shape and female genitalia seen in several species of Sulawesidrobia . But the translucent shell with its light-brown to whitish-grey appearance is characteristic. The coiled penis with the hook-shaped appendage pointing backwards is unique.</p><p>Description.</p><p>Shell. Conical, 1.6 to 1.7 times higher than wide, 3.625 to 4.5 whorls, sutures shallow, whorls moderately convex, transparent. light-brown to greyish, periostracum without colour (Figs 2 B, 3 C, D); protoconch (N = 3) wrinkled with 0.8 to slightly more than 1 whorls (Fig. 4 B), teleoconch with growth lines, otherwise without structure; umbilicus narrow; outer lip straight.</p><p>Operculum (N = 2). Light orange, thin, paucispiral, nucleus eccentric.</p><p>External features (N = 2). General appearance fairly dark, black pigmentation particularly dense on snout, behind eyes, anterior parts of mantle and upper parts of whorls; eyes black; tentacles less dark with central white stripe; mirrored, sickle-shaped spots on pharynx; mantle almost unpigmented over distal genital glands in both sexes.</p><p>Mantle cavity (N = 2). Gill with 17–18 filaments almost abutting to pericard; osphradium elongate, about 2 / 5 of length of gill, behind middle of gill.</p><p>Digestive system (N = 2, for radula 1). Radula formula R: 4 1 4 / 1-2 1-2, L: 3 1 4, M 1: 15-18, M 2: 20-23; lateral tooth with solid junction between face and flank; stomach without caecum; rectum running along distal genital glands, less closely to prostate than to pallial oviduct.</p><p>Female genitalia (N = 1; Fig. 7 D, E). Ovary a simple sac, starting 1.5 whorls below apex, comprising 0.75 whorls, extending over proximal parts of stomach; renal oviduct undulating before final loop continuing into ventral channel; bursa copulatrix largely behind albumen gland, elongate-ovoid, bursal duct emerging anteroventrally; no receptaculum seminis, sperm storage in distal part of final oviduct loop; albumen gland milky-white, capsule gland opaque white.</p><p>Male genitalia (N = 1). Testis lobate, starting 1 whorl below apex, comprising about 1 whorl, overlapping proximal part of stomach; vesicula seminalis coiling underneath distal quarter of testis; penis coiling over neck, without pigment, distal part broadened before tapering to pointed tip, small hook pointing backwards on right side just behind broadening (Fig. 8 B).</p><p>Remarks.</p><p>The phylogenetic relationships are discussed in the Remarks to S. lasoloensis sp. nov. above.</p></div>	https://treatment.plazi.org/id/EF22A6C280BD52B1A6FFAAB7830C9DFF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Kadow, Laura;von Rintelen, Thomas;Iqram, Muhammad;Bichain, Jean-Michel;Haase, Martin	Kadow, Laura, von Rintelen, Thomas, Iqram, Muhammad, Bichain, Jean-Michel, Haase, Martin (2025): Three new non-lacustrine species of the endemic freshwater gastropod genus Sulawesidrobia (Caenogastropoda, Truncatelloidea, Tateidae) from Sulawesi, Indonesia, with additional records of species from ancient Lake Poso. Zoosystematics and Evolution 101 (4): 1585-1600, DOI: 10.3897/zse.101.161812
4CF2307217DA5CC2BE05ED3E691ED7AD.text	4CF2307217DA5CC2BE05ED3E691ED7AD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sulawesidrobia tilangaensis Kadow & von Rintelen & Iqram & Bichain & Haase 2025	<div><p>Sulawesidrobia tilangaensis sp. nov.</p><p>Etymology.</p><p>This species is named after the type locality, the Tilanga natural Pool.</p><p>Material examined.</p><p>Holotype (Fig. 2 C): Indonesia • Sulawesi, Tana Toraja Regency, Makale Utara, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=119.88731&amp;materialsCitation.latitude=-3.0348887" title="Search Plazi for locations around (long 119.88731/lat -3.0348887)">Tilanga natural Pool</a>, on karstic rocks. 3°02'05.6"S, 119°53'14.3"E. October 2023, Muhammad Iqram leg. MZB Gst. 12189 .</p><p>Paratypes (Fig. 3 E, F): Indonesia • N = 4; same collection data as for holotype; MZB Gst. 12190 . N = 3; same collection data as for holotype; ZMB 118373 .</p><p>Diagnosis.</p><p>This new species is among the smaller and wider conical species of Sulawesidrobia morphologically falling into the variation of S. bonnei . It is characterized by fixed bases at 18 diagnostic positions in the COI alignment not shared with S. bonnei, S. bulat, or S. posoensis, which are reciprocally paraphyletic (for details see Remarks).</p><p>Shell. (N = 5). Conical, short, 1.4 to 1.6 times higher than wide, 3.25 to 3.75 whorls, sutures shallow, whorls moderately convex, semitranslucent, light brown, periostracum without color (Figs 2 C, 3 E, F); protoconch (N = 1) wrinkled with slightly more than 1 whorl (Fig. 4 C), teleoconch with growth lines, otherwise without structure; umbilicus a narrow slit, outer lip straight.</p><p>Remarks.</p><p>Only five adult specimens could be morphologically examined. Therefore, i. e. because of the lack of anatomical data, the differentiation and diagnosis had to be restricted to genetic data. Phylogenetically, S. tilangaensis sp. nov. was nested among the species from Lake Poso with a long branch indicating considerable genetic differentiation (Figs 10, 11). Morphologically, the new species is similar to S. bonnei (see Haase and Bouchet 2006). As S. bonnei cannot be distinguished from S. bulat and S. posoensis based on mitochondrial data due to paraphyly (Zielske et al. 2011, see below), the new species was compared against those three species. Sulawesidrobia tilangaensis sp. nov. had diagnostic bases at 18 positions (type 1 sensu Kühn and Haase 2018) in the COI alignment (Suppl. material 1): position 50 (base C), 157 (T), 181 (G), 200 (A), 265 (G), 286 (G), 301 (G), 340 (T), 370 (G), 397 (C), 454 (G), 481 (G), 484 (G), 511 (G), 532 (C), 544 (G), 577 (G), 578 (C).</p></div>	https://treatment.plazi.org/id/4CF2307217DA5CC2BE05ED3E691ED7AD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Kadow, Laura;von Rintelen, Thomas;Iqram, Muhammad;Bichain, Jean-Michel;Haase, Martin	Kadow, Laura, von Rintelen, Thomas, Iqram, Muhammad, Bichain, Jean-Michel, Haase, Martin (2025): Three new non-lacustrine species of the endemic freshwater gastropod genus Sulawesidrobia (Caenogastropoda, Truncatelloidea, Tateidae) from Sulawesi, Indonesia, with additional records of species from ancient Lake Poso. Zoosystematics and Evolution 101 (4): 1585-1600, DOI: 10.3897/zse.101.161812
