taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
88ED883E68FD58EB9A682B5DC381D17C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201330	https://doi.org/10.3897/asp.82.e136751.figure3	Figure 3. Habitus of some Skleroprotopus species in lateral view. A Skleroprotopus penglai sp. nov.; B Skleroprotopus longiflagellatus sp. nov.; C Skleroprotopus tiankeng sp. nov.; D Skleroprotopus ampullaceus sp. nov.; E Skleroprotopus conicus sp. nov.; F Skleroprotopus yutiantianae sp. nov.; G Skleroprotopus multistriatus sp. nov.; H Skleroprotopus change sp. nov. Scale bars: 5.0 mm.	Figure 3. Habitus of some Skleroprotopus species in lateral view. A Skleroprotopus penglai sp. nov.; B Skleroprotopus longiflagellatus sp. nov.; C Skleroprotopus tiankeng sp. nov.; D Skleroprotopus ampullaceus sp. nov.; E Skleroprotopus conicus sp. nov.; F Skleroprotopus yutiantianae sp. nov.; G Skleroprotopus multistriatus sp. nov.; H Skleroprotopus change sp. nov. Scale bars: 5.0 mm.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
88ED883E68FD58EB9A682B5DC381D17C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201339	https://doi.org/10.3897/asp.82.e136751.figure12	Figure 12. Skleroprotopus ampullaceus sp. nov. . A, G female paratype; B – F, H – J male paratype. A, B gnathochilarium; C right antenna, lateral view; D legs 1, anterior view; E legs 7, anterior view; F legs 2 with penis, posterior view; G right vulva, mesal view; H anterior gonopods, anterior view; I anterior gonopods, posterior view; J posterior gonopods, anterior view. Abbreviations: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; f, flagella; h, hump; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm.	Figure 12. Skleroprotopus ampullaceus sp. nov. . A, G female paratype; B – F, H – J male paratype. A, B gnathochilarium; C right antenna, lateral view; D legs 1, anterior view; E legs 7, anterior view; F legs 2 with penis, posterior view; G right vulva, mesal view; H anterior gonopods, anterior view; I anterior gonopods, posterior view; J posterior gonopods, anterior view. Abbreviations: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; f, flagella; h, hump; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
677230A3897E50DCAFF58AA30B9A00A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201329	https://doi.org/10.3897/asp.82.e136751.figure2	Figure 2. Pictures of live animals, photos by Prof. Mingyi Tian from SCAU. A Skleroprotopus change sp. nov.; B Skleroprotopus megistus sp. nov.; C Skleroprotopus longissimus sp. nov.; D Skleroprotopus genjudi sp. nov.; E Skleroprotopus tiankeng sp. nov.; F Skleroprotopus laiyuanensis sp. nov.; G Skleroprotopus incisodentatus sp. nov.	Figure 2. Pictures of live animals, photos by Prof. Mingyi Tian from SCAU. A Skleroprotopus change sp. nov.; B Skleroprotopus megistus sp. nov.; C Skleroprotopus longissimus sp. nov.; D Skleroprotopus genjudi sp. nov.; E Skleroprotopus tiankeng sp. nov.; F Skleroprotopus laiyuanensis sp. nov.; G Skleroprotopus incisodentatus sp. nov.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
677230A3897E50DCAFF58AA30B9A00A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201330	https://doi.org/10.3897/asp.82.e136751.figure3	Figure 3. Habitus of some Skleroprotopus species in lateral view. A Skleroprotopus penglai sp. nov.; B Skleroprotopus longiflagellatus sp. nov.; C Skleroprotopus tiankeng sp. nov.; D Skleroprotopus ampullaceus sp. nov.; E Skleroprotopus conicus sp. nov.; F Skleroprotopus yutiantianae sp. nov.; G Skleroprotopus multistriatus sp. nov.; H Skleroprotopus change sp. nov. Scale bars: 5.0 mm.	Figure 3. Habitus of some Skleroprotopus species in lateral view. A Skleroprotopus penglai sp. nov.; B Skleroprotopus longiflagellatus sp. nov.; C Skleroprotopus tiankeng sp. nov.; D Skleroprotopus ampullaceus sp. nov.; E Skleroprotopus conicus sp. nov.; F Skleroprotopus yutiantianae sp. nov.; G Skleroprotopus multistriatus sp. nov.; H Skleroprotopus change sp. nov. Scale bars: 5.0 mm.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
677230A3897E50DCAFF58AA30B9A00A6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201335	https://doi.org/10.3897/asp.82.e136751.figure8	Figure 8. Skleroprotopus change sp. nov. . A, D – I male paratype; B, C female paratype. A gnathochilarium and part of right antenna; B gnathochilarium; C right vulva, lateral view; D legs 2 with penis, posterior view; E legs 1, anterior view; F legs 7, anterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I posterior gonopods, anterior view. Abbreviations: cp, coxal process; cx, coxa; f, flagellum; h, hump; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm.	Figure 8. Skleroprotopus change sp. nov. . A, D – I male paratype; B, C female paratype. A gnathochilarium and part of right antenna; B gnathochilarium; C right vulva, lateral view; D legs 2 with penis, posterior view; E legs 1, anterior view; F legs 7, anterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I posterior gonopods, anterior view. Abbreviations: cp, coxal process; cx, coxa; f, flagellum; h, hump; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
0FCA60387E815676A8CD7C03379F6061.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201330	https://doi.org/10.3897/asp.82.e136751.figure3	Figure 3. Habitus of some Skleroprotopus species in lateral view. A Skleroprotopus penglai sp. nov.; B Skleroprotopus longiflagellatus sp. nov.; C Skleroprotopus tiankeng sp. nov.; D Skleroprotopus ampullaceus sp. nov.; E Skleroprotopus conicus sp. nov.; F Skleroprotopus yutiantianae sp. nov.; G Skleroprotopus multistriatus sp. nov.; H Skleroprotopus change sp. nov. Scale bars: 5.0 mm.	Figure 3. Habitus of some Skleroprotopus species in lateral view. A Skleroprotopus penglai sp. nov.; B Skleroprotopus longiflagellatus sp. nov.; C Skleroprotopus tiankeng sp. nov.; D Skleroprotopus ampullaceus sp. nov.; E Skleroprotopus conicus sp. nov.; F Skleroprotopus yutiantianae sp. nov.; G Skleroprotopus multistriatus sp. nov.; H Skleroprotopus change sp. nov. Scale bars: 5.0 mm.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
0FCA60387E815676A8CD7C03379F6061.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201342	https://doi.org/10.3897/asp.82.e136751.figure15	Figure 15. Skleroprotopus conicus sp. nov. . paratypes from Cave Xianpu Dong. A, D-I male paratype; B, C female paratype. A, B gnathochilarium; C right vulva, mesal view; D legs 1, anterior view; E legs 2 with penis, posterior view; F legs 7, anterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I posterior gonopods, anterior view. Abbreviations: b, bulge; cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm.	Figure 15. Skleroprotopus conicus sp. nov. . paratypes from Cave Xianpu Dong. A, D-I male paratype; B, C female paratype. A, B gnathochilarium; C right vulva, mesal view; D legs 1, anterior view; E legs 2 with penis, posterior view; F legs 7, anterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I posterior gonopods, anterior view. Abbreviations: b, bulge; cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
556B8BDE855659439061281CD7CCB1FF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201329	https://doi.org/10.3897/asp.82.e136751.figure2	Figure 2. Pictures of live animals, photos by Prof. Mingyi Tian from SCAU. A Skleroprotopus change sp. nov.; B Skleroprotopus megistus sp. nov.; C Skleroprotopus longissimus sp. nov.; D Skleroprotopus genjudi sp. nov.; E Skleroprotopus tiankeng sp. nov.; F Skleroprotopus laiyuanensis sp. nov.; G Skleroprotopus incisodentatus sp. nov.	Figure 2. Pictures of live animals, photos by Prof. Mingyi Tian from SCAU. A Skleroprotopus change sp. nov.; B Skleroprotopus megistus sp. nov.; C Skleroprotopus longissimus sp. nov.; D Skleroprotopus genjudi sp. nov.; E Skleroprotopus tiankeng sp. nov.; F Skleroprotopus laiyuanensis sp. nov.; G Skleroprotopus incisodentatus sp. nov.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
556B8BDE855659439061281CD7CCB1FF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201338	https://doi.org/10.3897/asp.82.e136751.figure11	Figure 11. Skleroprotopus genjudi sp. nov. . paratypes from Cave Luohanyan. A, D – I male paratype; B, C female paratype. A, B gnathochilarium; C right vulva, mesal view; D legs 1, anterior view; E legs 2 with penis, posterior view; F legs 7, anterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I posterior gonopods, anterior view. Abbreviations: cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm.	Figure 11. Skleroprotopus genjudi sp. nov. . paratypes from Cave Luohanyan. A, D – I male paratype; B, C female paratype. A, B gnathochilarium; C right vulva, mesal view; D legs 1, anterior view; E legs 2 with penis, posterior view; F legs 7, anterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I posterior gonopods, anterior view. Abbreviations: cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
4AE50CFCEAE75F3BA9F861D0994EF64F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201329	https://doi.org/10.3897/asp.82.e136751.figure2	Figure 2. Pictures of live animals, photos by Prof. Mingyi Tian from SCAU. A Skleroprotopus change sp. nov.; B Skleroprotopus megistus sp. nov.; C Skleroprotopus longissimus sp. nov.; D Skleroprotopus genjudi sp. nov.; E Skleroprotopus tiankeng sp. nov.; F Skleroprotopus laiyuanensis sp. nov.; G Skleroprotopus incisodentatus sp. nov.	Figure 2. Pictures of live animals, photos by Prof. Mingyi Tian from SCAU. A Skleroprotopus change sp. nov.; B Skleroprotopus megistus sp. nov.; C Skleroprotopus longissimus sp. nov.; D Skleroprotopus genjudi sp. nov.; E Skleroprotopus tiankeng sp. nov.; F Skleroprotopus laiyuanensis sp. nov.; G Skleroprotopus incisodentatus sp. nov.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
4AE50CFCEAE75F3BA9F861D0994EF64F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201340	https://doi.org/10.3897/asp.82.e136751.figure13	Figure 13. Skleroprotopus incisodentatus sp. nov. . paratypes from Cave Lingjiu Dong. A, G female paratype; B – I male paratype. A, B gnathochilarium; C legs 2 with penis, posterior view; D legs 1, anterior view; E anterior gonopods, anterior view; F legs 7, anterior view; G vulvae, lateral view; H posterior gonopods, anterior view; I anterior gonopods, posterior view. Abbreviations: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; f, flagellum; h, hump; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm.	Figure 13. Skleroprotopus incisodentatus sp. nov. . paratypes from Cave Lingjiu Dong. A, G female paratype; B – I male paratype. A, B gnathochilarium; C legs 2 with penis, posterior view; D legs 1, anterior view; E anterior gonopods, anterior view; F legs 7, anterior view; G vulvae, lateral view; H posterior gonopods, anterior view; I anterior gonopods, posterior view. Abbreviations: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; f, flagellum; h, hump; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
A53A99BD23CA510193C426B181E3C767.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201329	https://doi.org/10.3897/asp.82.e136751.figure2	Figure 2. Pictures of live animals, photos by Prof. Mingyi Tian from SCAU. A Skleroprotopus change sp. nov.; B Skleroprotopus megistus sp. nov.; C Skleroprotopus longissimus sp. nov.; D Skleroprotopus genjudi sp. nov.; E Skleroprotopus tiankeng sp. nov.; F Skleroprotopus laiyuanensis sp. nov.; G Skleroprotopus incisodentatus sp. nov.	Figure 2. Pictures of live animals, photos by Prof. Mingyi Tian from SCAU. A Skleroprotopus change sp. nov.; B Skleroprotopus megistus sp. nov.; C Skleroprotopus longissimus sp. nov.; D Skleroprotopus genjudi sp. nov.; E Skleroprotopus tiankeng sp. nov.; F Skleroprotopus laiyuanensis sp. nov.; G Skleroprotopus incisodentatus sp. nov.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
A53A99BD23CA510193C426B181E3C767.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201344	https://doi.org/10.3897/asp.82.e136751.figure16	Figure 16. Skleroprotopus laiyuanensis sp. nov. . paratypes from Cave Chuqi Dong. A, G female paratype; B – F, H – J male paratype. A, B gnathochilarium; C right antenna, lateral view; D legs 1, anterior view; E legs 2 with penis, posterior view; F legs 7, anterior view; G left vulva, sublateral view; H anterior gonopods, anterior view; I anterior gonopods, posterior view; J posterior gonopods, anterior view. Abbreviations: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm.	Figure 16. Skleroprotopus laiyuanensis sp. nov. . paratypes from Cave Chuqi Dong. A, G female paratype; B – F, H – J male paratype. A, B gnathochilarium; C right antenna, lateral view; D legs 1, anterior view; E legs 2 with penis, posterior view; F legs 7, anterior view; G left vulva, sublateral view; H anterior gonopods, anterior view; I anterior gonopods, posterior view; J posterior gonopods, anterior view. Abbreviations: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
A53A99BD23CA510193C426B181E3C767.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201345	https://doi.org/10.3897/asp.82.e136751.figure17	Figure 17. A distribution map of the genus Skleroprotopus species in China. 1 Skleroprotopus laticoxalis; 2 S. confucius; 3 S. securifer; 4 S. membranipedalis; 5 S. serratus; 6 S. yutiantianae sp. nov.; 7 S. tiankeng sp. nov.; 8 S. megistus sp. nov.; 9 S. longiflagellatus sp. nov.; 10 S. change sp. nov.; 11 S. penglai sp. nov.; 12 S. multistriatus sp. nov.; 13 S. ampullaceus sp. nov.; 14 S. incisodentatus sp. nov.; 15 S. conicus sp. nov.; 16 S. longissimus sp. nov.; 17 S. genjudi sp. nov.; 18 S. laiyuanensis sp. nov. Abbreviations: BJ, Beijing.	Figure 17. A distribution map of the genus Skleroprotopus species in China. 1 Skleroprotopus laticoxalis; 2 S. confucius; 3 S. securifer; 4 S. membranipedalis; 5 S. serratus; 6 S. yutiantianae sp. nov.; 7 S. tiankeng sp. nov.; 8 S. megistus sp. nov.; 9 S. longiflagellatus sp. nov.; 10 S. change sp. nov.; 11 S. penglai sp. nov.; 12 S. multistriatus sp. nov.; 13 S. ampullaceus sp. nov.; 14 S. incisodentatus sp. nov.; 15 S. conicus sp. nov.; 16 S. longissimus sp. nov.; 17 S. genjudi sp. nov.; 18 S. laiyuanensis sp. nov. Abbreviations: BJ, Beijing.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
02ABD0308B975F3280C21BE8831D8F53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201330	https://doi.org/10.3897/asp.82.e136751.figure3	Figure 3. Habitus of some Skleroprotopus species in lateral view. A Skleroprotopus penglai sp. nov.; B Skleroprotopus longiflagellatus sp. nov.; C Skleroprotopus tiankeng sp. nov.; D Skleroprotopus ampullaceus sp. nov.; E Skleroprotopus conicus sp. nov.; F Skleroprotopus yutiantianae sp. nov.; G Skleroprotopus multistriatus sp. nov.; H Skleroprotopus change sp. nov. Scale bars: 5.0 mm.	Figure 3. Habitus of some Skleroprotopus species in lateral view. A Skleroprotopus penglai sp. nov.; B Skleroprotopus longiflagellatus sp. nov.; C Skleroprotopus tiankeng sp. nov.; D Skleroprotopus ampullaceus sp. nov.; E Skleroprotopus conicus sp. nov.; F Skleroprotopus yutiantianae sp. nov.; G Skleroprotopus multistriatus sp. nov.; H Skleroprotopus change sp. nov. Scale bars: 5.0 mm.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
02ABD0308B975F3280C21BE8831D8F53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201334	https://doi.org/10.3897/asp.82.e136751.figure7	Figure 7. Skleroprotopus longiflagellatus sp. nov. . A, D – I male paratype; B, C female paratype. A, B gnathochilarium; C left vulva, lateral view; D legs 1, anterior view; E legs 2 with penis, posterior view; F legs 7, anterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I posterior gonopods, anterior view. Abbreviations: cp, coxal process; cx, coxa; f, flagella; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm.	Figure 7. Skleroprotopus longiflagellatus sp. nov. . A, D – I male paratype; B, C female paratype. A, B gnathochilarium; C left vulva, lateral view; D legs 1, anterior view; E legs 2 with penis, posterior view; F legs 7, anterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I posterior gonopods, anterior view. Abbreviations: cp, coxal process; cx, coxa; f, flagella; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
375FCDC36A6A5496BE22B9AEAE52D4C7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201329	https://doi.org/10.3897/asp.82.e136751.figure2	Figure 2. Pictures of live animals, photos by Prof. Mingyi Tian from SCAU. A Skleroprotopus change sp. nov.; B Skleroprotopus megistus sp. nov.; C Skleroprotopus longissimus sp. nov.; D Skleroprotopus genjudi sp. nov.; E Skleroprotopus tiankeng sp. nov.; F Skleroprotopus laiyuanensis sp. nov.; G Skleroprotopus incisodentatus sp. nov.	Figure 2. Pictures of live animals, photos by Prof. Mingyi Tian from SCAU. A Skleroprotopus change sp. nov.; B Skleroprotopus megistus sp. nov.; C Skleroprotopus longissimus sp. nov.; D Skleroprotopus genjudi sp. nov.; E Skleroprotopus tiankeng sp. nov.; F Skleroprotopus laiyuanensis sp. nov.; G Skleroprotopus incisodentatus sp. nov.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
375FCDC36A6A5496BE22B9AEAE52D4C7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201337	https://doi.org/10.3897/asp.82.e136751.figure10	Figure 10. Skleroprotopus longissimus sp. nov. . paratypes from Cave Bidongfeiyan Dong. A, D – I male paratype; B, C female paratype. A, B gnathochilarium; C right vulva, mesal view; D legs 1, anterior view; E legs 2 with penis, posterior view; F legs 7, anterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I posterior gonopods, anterior view. Abbreviations: cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm.	Figure 10. Skleroprotopus longissimus sp. nov. . paratypes from Cave Bidongfeiyan Dong. A, D – I male paratype; B, C female paratype. A, B gnathochilarium; C right vulva, mesal view; D legs 1, anterior view; E legs 2 with penis, posterior view; F legs 7, anterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I posterior gonopods, anterior view. Abbreviations: cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
9E190A1864AF5B00A20D3C9862598257.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201329	https://doi.org/10.3897/asp.82.e136751.figure2	Figure 2. Pictures of live animals, photos by Prof. Mingyi Tian from SCAU. A Skleroprotopus change sp. nov.; B Skleroprotopus megistus sp. nov.; C Skleroprotopus longissimus sp. nov.; D Skleroprotopus genjudi sp. nov.; E Skleroprotopus tiankeng sp. nov.; F Skleroprotopus laiyuanensis sp. nov.; G Skleroprotopus incisodentatus sp. nov.	Figure 2. Pictures of live animals, photos by Prof. Mingyi Tian from SCAU. A Skleroprotopus change sp. nov.; B Skleroprotopus megistus sp. nov.; C Skleroprotopus longissimus sp. nov.; D Skleroprotopus genjudi sp. nov.; E Skleroprotopus tiankeng sp. nov.; F Skleroprotopus laiyuanensis sp. nov.; G Skleroprotopus incisodentatus sp. nov.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
9E190A1864AF5B00A20D3C9862598257.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201333	https://doi.org/10.3897/asp.82.e136751.figure6	Figure 6. Skleroprotopus megistus sp. nov. . A, D – J male paratype; B, C female paratype. A, B gnathochilarium; C left vulva, lateral view; D legs 7, anterior view; E legs 1, anterior view; F legs 2 with penis, posterior view; G posterior gonopods, anterior view; H anterior gonopods, anterior view; I anterior gonopods, posterior view; J posterior gonopods, posterior view. Abbreviations: cp, coxal process; cx, coxa; f, flagella; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 - t 4, telopoditomeres 1-4; te, telopodite. Scale bars: 0.2 mm.	Figure 6. Skleroprotopus megistus sp. nov. . A, D – J male paratype; B, C female paratype. A, B gnathochilarium; C left vulva, lateral view; D legs 7, anterior view; E legs 1, anterior view; F legs 2 with penis, posterior view; G posterior gonopods, anterior view; H anterior gonopods, anterior view; I anterior gonopods, posterior view; J posterior gonopods, posterior view. Abbreviations: cp, coxal process; cx, coxa; f, flagella; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 - t 4, telopoditomeres 1-4; te, telopodite. Scale bars: 0.2 mm.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
E79A521BC87F54558A813EAC186A270A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201330	https://doi.org/10.3897/asp.82.e136751.figure3	Figure 3. Habitus of some Skleroprotopus species in lateral view. A Skleroprotopus penglai sp. nov.; B Skleroprotopus longiflagellatus sp. nov.; C Skleroprotopus tiankeng sp. nov.; D Skleroprotopus ampullaceus sp. nov.; E Skleroprotopus conicus sp. nov.; F Skleroprotopus yutiantianae sp. nov.; G Skleroprotopus multistriatus sp. nov.; H Skleroprotopus change sp. nov. Scale bars: 5.0 mm.	Figure 3. Habitus of some Skleroprotopus species in lateral view. A Skleroprotopus penglai sp. nov.; B Skleroprotopus longiflagellatus sp. nov.; C Skleroprotopus tiankeng sp. nov.; D Skleroprotopus ampullaceus sp. nov.; E Skleroprotopus conicus sp. nov.; F Skleroprotopus yutiantianae sp. nov.; G Skleroprotopus multistriatus sp. nov.; H Skleroprotopus change sp. nov. Scale bars: 5.0 mm.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
E79A521BC87F54558A813EAC186A270A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201341	https://doi.org/10.3897/asp.82.e136751.figure14	Figure 14. Skleroprotopus multistriatus sp. nov. . A, J female paratype; B – I male paratype. A, B gnathochilarium; C leg 2 with penis, posterior view; D legs 1, anterior view; E legs 7, anterior view; F right antenna, lateral view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I posterior gonopods, anterior view; J right vulva, mesal view. Abbreviations: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; f, flagellum; h, hump; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm.	Figure 14. Skleroprotopus multistriatus sp. nov. . A, J female paratype; B – I male paratype. A, B gnathochilarium; C leg 2 with penis, posterior view; D legs 1, anterior view; E legs 7, anterior view; F right antenna, lateral view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I posterior gonopods, anterior view; J right vulva, mesal view. Abbreviations: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; f, flagellum; h, hump; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
D538F573E57D50CC8A40344706BB2A51.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201330	https://doi.org/10.3897/asp.82.e136751.figure3	Figure 3. Habitus of some Skleroprotopus species in lateral view. A Skleroprotopus penglai sp. nov.; B Skleroprotopus longiflagellatus sp. nov.; C Skleroprotopus tiankeng sp. nov.; D Skleroprotopus ampullaceus sp. nov.; E Skleroprotopus conicus sp. nov.; F Skleroprotopus yutiantianae sp. nov.; G Skleroprotopus multistriatus sp. nov.; H Skleroprotopus change sp. nov. Scale bars: 5.0 mm.	Figure 3. Habitus of some Skleroprotopus species in lateral view. A Skleroprotopus penglai sp. nov.; B Skleroprotopus longiflagellatus sp. nov.; C Skleroprotopus tiankeng sp. nov.; D Skleroprotopus ampullaceus sp. nov.; E Skleroprotopus conicus sp. nov.; F Skleroprotopus yutiantianae sp. nov.; G Skleroprotopus multistriatus sp. nov.; H Skleroprotopus change sp. nov. Scale bars: 5.0 mm.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
D538F573E57D50CC8A40344706BB2A51.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201336	https://doi.org/10.3897/asp.82.e136751.figure9	Figure 9. Skleroprotopus penglai sp. nov. . A, I female paratype; B – H male paratype. A, B gnathochilarium; C legs 7, anterior view; D legs 1, anterior view; E posterior gonopods, anterior view; F legs 2 with penis, posterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I left vulva, mesal view. Abbreviations: cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm.	Figure 9. Skleroprotopus penglai sp. nov. . A, I female paratype; B – H male paratype. A, B gnathochilarium; C legs 7, anterior view; D legs 1, anterior view; E posterior gonopods, anterior view; F legs 2 with penis, posterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I left vulva, mesal view. Abbreviations: cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
1E81187968E0580FA7AE94DD1EAC22E7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201329	https://doi.org/10.3897/asp.82.e136751.figure2	Figure 2. Pictures of live animals, photos by Prof. Mingyi Tian from SCAU. A Skleroprotopus change sp. nov.; B Skleroprotopus megistus sp. nov.; C Skleroprotopus longissimus sp. nov.; D Skleroprotopus genjudi sp. nov.; E Skleroprotopus tiankeng sp. nov.; F Skleroprotopus laiyuanensis sp. nov.; G Skleroprotopus incisodentatus sp. nov.	Figure 2. Pictures of live animals, photos by Prof. Mingyi Tian from SCAU. A Skleroprotopus change sp. nov.; B Skleroprotopus megistus sp. nov.; C Skleroprotopus longissimus sp. nov.; D Skleroprotopus genjudi sp. nov.; E Skleroprotopus tiankeng sp. nov.; F Skleroprotopus laiyuanensis sp. nov.; G Skleroprotopus incisodentatus sp. nov.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
1E81187968E0580FA7AE94DD1EAC22E7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201330	https://doi.org/10.3897/asp.82.e136751.figure3	Figure 3. Habitus of some Skleroprotopus species in lateral view. A Skleroprotopus penglai sp. nov.; B Skleroprotopus longiflagellatus sp. nov.; C Skleroprotopus tiankeng sp. nov.; D Skleroprotopus ampullaceus sp. nov.; E Skleroprotopus conicus sp. nov.; F Skleroprotopus yutiantianae sp. nov.; G Skleroprotopus multistriatus sp. nov.; H Skleroprotopus change sp. nov. Scale bars: 5.0 mm.	Figure 3. Habitus of some Skleroprotopus species in lateral view. A Skleroprotopus penglai sp. nov.; B Skleroprotopus longiflagellatus sp. nov.; C Skleroprotopus tiankeng sp. nov.; D Skleroprotopus ampullaceus sp. nov.; E Skleroprotopus conicus sp. nov.; F Skleroprotopus yutiantianae sp. nov.; G Skleroprotopus multistriatus sp. nov.; H Skleroprotopus change sp. nov. Scale bars: 5.0 mm.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
1E81187968E0580FA7AE94DD1EAC22E7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201332	https://doi.org/10.3897/asp.82.e136751.figure5	Figure 5. Skleroprotopus tiankeng sp. nov. . A, J female paratype, B – I male paratype. A, B gnathochilarium; C right antenna, lateral view; D legs 1, anterior view; E legs 7, anterior view; F posterior gonopods, posterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I legs 2 with penis, posterior view; J right vulva, lateral view. Abbreviations: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm.	Figure 5. Skleroprotopus tiankeng sp. nov. . A, J female paratype, B – I male paratype. A, B gnathochilarium; C right antenna, lateral view; D legs 1, anterior view; E legs 7, anterior view; F posterior gonopods, posterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I legs 2 with penis, posterior view; J right vulva, lateral view. Abbreviations: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
4DB77B61D622521E8BABF4497E245AC4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201330	https://doi.org/10.3897/asp.82.e136751.figure3	Figure 3. Habitus of some Skleroprotopus species in lateral view. A Skleroprotopus penglai sp. nov.; B Skleroprotopus longiflagellatus sp. nov.; C Skleroprotopus tiankeng sp. nov.; D Skleroprotopus ampullaceus sp. nov.; E Skleroprotopus conicus sp. nov.; F Skleroprotopus yutiantianae sp. nov.; G Skleroprotopus multistriatus sp. nov.; H Skleroprotopus change sp. nov. Scale bars: 5.0 mm.	Figure 3. Habitus of some Skleroprotopus species in lateral view. A Skleroprotopus penglai sp. nov.; B Skleroprotopus longiflagellatus sp. nov.; C Skleroprotopus tiankeng sp. nov.; D Skleroprotopus ampullaceus sp. nov.; E Skleroprotopus conicus sp. nov.; F Skleroprotopus yutiantianae sp. nov.; G Skleroprotopus multistriatus sp. nov.; H Skleroprotopus change sp. nov. Scale bars: 5.0 mm.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
4DB77B61D622521E8BABF4497E245AC4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1201331	https://doi.org/10.3897/asp.82.e136751.figure4	Figure 4. Skleroprotopus yutiantianae sp. nov., holotype. A gnathochilarium; B head and collum, dorsal view; C left antenna, lateral view; D legs 1, anterior view; E legs 7, anterior view; F legs 2 with penis, posterior view; G posterior gonopods, anterior view; H anterior gonopods, anterior view; I anterior gonopods, posterior view. Abbreviations: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; dl, dividing line; f, flagellum; ll, lamella lingualis; ml, membranous lobe; pn, penis; pr, promentum; t 1 – t 5, telopoditomeres 1–5; te, telopodite. Scale bars: 0.2 mm.	Figure 4. Skleroprotopus yutiantianae sp. nov., holotype. A gnathochilarium; B head and collum, dorsal view; C left antenna, lateral view; D legs 1, anterior view; E legs 7, anterior view; F legs 2 with penis, posterior view; G posterior gonopods, anterior view; H anterior gonopods, anterior view; I anterior gonopods, posterior view. Abbreviations: a 5, a 6, antennomeres 5, 6; cp, coxal process; cx, coxa; dl, dividing line; f, flagellum; ll, lamella lingualis; ml, membranous lobe; pn, penis; pr, promentum; t 1 – t 5, telopoditomeres 1–5; te, telopodite. Scale bars: 0.2 mm.	2024-12-18	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin		Zenodo	biologists	Chen, Rong;Zhao, Yi;Golovatch, Sergei;Liu, Wei-Xin			
