taxonID	type	description	language	source
88ED883E68FD58EB9A682B5DC381D17C.taxon	description	Figures 3 D, 12	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
88ED883E68FD58EB9A682B5DC381D17C.taxon	diagnosis	Diagnosis. Differs from congeners mostly based on the following combination of characters: (1) male leg 1 relatively slender, telopoditomere 4 linguiform, with a small mesal hump (Fig. 12 D); (2) penis much shorter than coxae 2 (Fig. 12 F); (3) male leg 7 with a coxal process of medium size, densely setose at mesal margin, height subequal to telopodite, which is made up of 2 or 3 segements and a claw (Fig. 12 E); (4) anterior gonopod (Fig. 12 H, I) with a rather long coxal process carrying a narrow, lower, membranous lobe, distally with a vague indentation. — In addition, this new species differs from all other species analyzed in a> 9.1 % uncorrected p - distance of the COI barcoding gene.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
88ED883E68FD58EB9A682B5DC381D17C.taxon	etymology	Etymology. Latin “ ampullaceus ” is meaning bottle-shaped. The specific epithet refers to the the promentum of the gnathochilarium being bottle-shaped in males; adjective.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
88ED883E68FD58EB9A682B5DC381D17C.taxon	description	Description. Length of both sexes ca 30.0 – 37.0 mm, 1.8 – 2.0 mm in diameter, body with 48 – 58 podous + 1 – 2 apodous rings + telson. Coloration in alcohol (Fig. 3 D) uniformly yellow to dark brownish. Antennae and legs yellow brownish. Eye patches blackish, subtriangular, arranged in 4 – 6 irregular rows, altogether about 20 – 38 ommatidia per eye patch. Head capsule smooth and hairless, eyes slightly bulged, epicranial suture well-defined. Labral margin with 2 (3) + 2 (3) supra-labral and 9 + 9 labral setae. Antennae of medium length, reaching behind to middle of ring 2 when stretched dorsally. In length, antennomeres 3> 4> 2> 5> 6> 1> 7 (Fig. 12 C). Antennomeres 5 and 6 each with a distal corolla of sensilla basiconica. Mandibular stipes well-rounded, with two small lobes in male, rounded in female. Gnathochilarium with at least 5 setae on each lamella lingualis (ll); promentum (pr) lance-shaped at base, swollen anteriorly, bottle-shaped in males (Fig. 12 B), vs a narrow, oblong, rhomboid pr in females (Fig. 12 A). Collum with 6 – 14 lower striae laterally, but lowest 4 striae not reaching the anterior margin. Prozona with 2 – 4 irregular subtransverse striae and metazona with 13 – 19 longitudinal striae laterally (Fig. 3 D). Suture dividing pro- and metazona very narrow, a regular comb. Ozopores starting with ring 6, lying mid-laterally on sides of metatergites. Epiproct slightly protruding caudad, with 4 + 4 setae at posterior margin. Paraprocts convex, with 1 + 1 setae. Hypoproct eye-shaped, with 1 + 1 setae. Legs short, but slender, about 0.9 × as long as midbody height. Male leg-pair 1 (Fig. 12 D) hypertrophied, relatively slender, 5 - segmented, strongly curved anteriad; coxa (cx) and telopoditomeres 1 and 2 (t 1, t 2) with some long setae. Telopoditomere 3 (t 3) longest, 1.5 × as long as telopoditomere 4 (t 4), t 4 linguiform, with a small mesal hump (h) subapically; t 3 and t 4 with long setae medially. Male leg-pair 2 (Fig. 12 F) reduced in size; penis (pn) fat, much shorter than coxae (cx), distal part with two long setae. Male leg-pair 7 (Fig. 12 E) reduced, each leg with a 2 or 3 - segmented asymmetrical telopodite (te) and a claw, telopoditomere 1 (t 1) largest and setose subapically; coxal process (cp) medium-sized, about as high as te, densely setose at mesal margin. Anterior gonopods (Fig. 12 H, I) with a very long and relatively narrow coxal process (cp) bearing a long row of short villi laterally, posteriorly with a narrow, lower, membranous lobe (ml), outer margin of ml smooth, with a vague indentation distally. Flagellum (f) very long and slender, distally villose and branched. Telopodite (te) about half as long as cp, with long setae both apically and mesally, laterobasally with a minute remnant (r) of a second podomere. Posterior gonopods (Fig. 12 J) erect, with bristles medially, distally branched, anterior branch smooth, with a pointed tip, posterior one densely setose. Vulva (Fig. 12 G) as usual, with two parallel rows of short setae on both anterior and posterior surfaces, operculum (op) with several long setae distally.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
677230A3897E50DCAFF58AA30B9A00A6.taxon	description	Figures 2 A, 3 H, 8	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
677230A3897E50DCAFF58AA30B9A00A6.taxon	diagnosis	Diagnosis. Differs from congeners mostly based on the following combination of characters: (1) telopoditomere 4 of male leg 1 rounded, with a small mesal hump (Fig. 8 E); (2) penis slightly longer than male coxae 2 (Fig. 8 D); (3) male leg 7 with a very small coxal process and a 4 - segmented telopodite (Fig. 8 F); (4) anterior gonopod (Fig. 8 G, H) with a very long coxal process carrying a somewhat broadened and dentated membranous lobe. — In addition, this new species differs from all other Skleroprotopus species analyzed in uncorrected p - distances ranging from between 6.6 % (compared to S. longiflagellatus sp. nov.) and 15.6 % (compared to S. longissimus sp. nov.).	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
677230A3897E50DCAFF58AA30B9A00A6.taxon	etymology	Etymology. The specific epithet is primarily derived from its type locality, which is named after Chang’e. She is the fairy of the moon palace in the ancient Chinese mythology. “ Change ” in Chinese “ 嫦娥 ”, a noun in apposition.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
677230A3897E50DCAFF58AA30B9A00A6.taxon	description	Description. Length of both sexes ca 20.5 – 28 mm, width 1.7 – 1.8 mm, midbody height 1.5 – 1.8 mm, body with 48 – 56 podous + 1 – 2 apodous rings + telson. Natural coloration marbled reddish- to purple brownish (Fig. 2 A). Anterior 2 / 3 body in alcohol yellowish brown, posterior 1 / 3 body yellowish. Antennae and legs pale yellowish (Fig. 3 H). Eye patches blackish, subtriangular, arranged in 4 – 6 irregular rows, altogether about 15 – 30 ommatidia per eye patch. Head capsule smooth and hairless, eyes slightly bulged, epicranial suture well-defined. Labral margin with 2 (3) + 2 (3) supra-labral and 11 + 11 labral setae. Antennae of medium length, reaching behind to middle of ring 2 when stretched dorsally. In length, antennomeres 3 ≈ 2> 6 ≈ 5> 4> 1> 7. Antennomeres 5 and 6 each with a distal corolla of sensilla basiconica. Mandibular stipes well-rounded in males, more strongly rounded in females. Gnathochilarium with at least 5 setae on each lamella lingualis (ll); promentum (pr) lanceolate, slightly swollen anteriorly in males (Fig. 8 A), vs a narrow drop-shaped pr in females (Fig. 8 B). Collum with 6 – 8 lower striae laterally, but lowest 2 striae not reaching the anterior margin (Fig. 3 H). Prozona with 4 irregular subtransverse striae and metazona with 13 – 17 longitudinal striae laterally. Suture dividing pro- and metazona very narrow, a regular comb. Ozopores starting with ring 6, lying mid-laterally on sides of metatergites (Fig. 3 H). Epiproct slightly protruding caudad, laterally with 2 + 2 setae. Paraprocts convex, smooth. Hypoproct eye-shaped, with 1 + 1 setae. Legs short, about 0.6 × as long as midbody height, claw weakly curved, without modifications. Male leg-pair 1 (Fig. 8 E) hypertrophied, 5 - segmented, strongly curved anteriad; coxa (cx) and telopoditomeres 1 and 2 (t 1, t 2) with some long setae. Telopoditomere 3 (t 3) longest, about as long as all other telopoditomeres combined. Telopoditomere 4 (t 4) irregularly shaped, with a small mesal hump (h) subapically; t 3 and t 4 with few scattered setae and short bristles on mesal face. Male leg-pair 2 (Fig. 8 D) reduced in size and slender; penis (pn) unpaired, slightly longer than coxae (cx), cylindrical, distal part undulating, with five long setae. Male leg-pair 7 (Fig. 8 F) strongly reduced, each leg with a 4 - segmented asymmetrical telopodite (not including claw); coxal process (cp) small, densely setose. Anterior gonopods (Fig. 8 G, H) with a very long coxal process (cp) bearing two long rows of short villi in anterior view, posteriorly carrying with a membranous lobe (ml), outer margin of ml somewhat broadened, with small and irregular denticulations. Flagellum (f) very long and slightly thickened, distally villose. Telopodite (te) about half as long as cp, with long setae both distally and mesally, laterobasally with a minute remnant (r) of a second podomere. Posterior gonopods (Fig. 8 I) erect, branched at tip, anterior branch smooth, with a pointed tip, posterior one densely setose. Vulva (Fig. 8 C) mostly wrapped inside a membrane, characteristic in shape, strongly elongated; operculum (op) longer than bursa, with two parallel rows of short setae on anterior face and 2 long setae distally.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
0FCA60387E815676A8CD7C03379F6061.taxon	description	Figures 3 E, 15	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
0FCA60387E815676A8CD7C03379F6061.taxon	diagnosis	Diagnosis. Differs from congeners mostly based on the following combination of characters: (1) telopoditomere 3 of male leg 1 longest, with a faint bulge in the middle (Fig. 15 D); (2) penis very flat, much shorter than coxae 2 (Fig. 15 E); (3) male leg 7 with a very small, cone-shaped coxal process and a basically normal telopodite (Fig. 15 F); (4) anterior gonopod (Fig. 15 G, H) with a very long coxal process carrying a narrow membranous lobe, outer margin distinctly serrate distally. — In addition, this new species differs from all other species analyzed in a> 10.0 % uncorrected p - distance of the COI barcoding gene.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
0FCA60387E815676A8CD7C03379F6061.taxon	etymology	Etymology. The specific epithet refers to the male leg 7 with a rather small and cone-shaped coxal process; adjective.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
0FCA60387E815676A8CD7C03379F6061.taxon	description	Description. Length of both sexes ca 27.0 – 47.5 mm, width 1.8 – 2.0 mm in diameter, body with 55 – 68 podous + 1 apodous ring + telson. Coloration in alcohol uniformly brownish (Fig. 3 E), prozona slightly red or dark brownish. Antennae and legs yellowish to light brown. Eye patches blackish, subtriangular, arranged in 6 – 14 irregular rows, altogether about 32 – 78 ommatidia per eye patch. Head capsule smooth and hairless, vertex bulged, epicranial suture clearly concave. Labral margin with 2 + 2 supra-labral and 7 + 7 labral setae. Antennae long and slender, reaching behind to middle of ring 4 when stretched dorsally. In length, antennomeres 3> 4> 5> 2 ≈ 6> 7> 1. Antennomeres 5 and 6 each with a distal corolla of sensilla basiconica. Mandibular stipes well-rounded, with two small lobes in males, rounded in females. Gnathochilarium with at least 3 setae on each lamella lingualis (ll); promentum (pr) short and lanceolate, swollen anteriorly, tube-shaped in males (Fig. 15 A), vs an oblong rhomboid pr in females (Fig. 15 B). Collum (Fig. 3 E) with 5 – 7 lower striae laterally, but lowest 2 striae not reaching the anterior margin. Prozona with at least 10 irregularly scattered striae and metazona with 13 – 18 longitudinal striae laterally (Fig. 3 E). Suture dividing pro- and metazona very narrow, a regular comb. Ozopores starting with ring 6, lying anteriorly on lateral sides of metatergites. Epiproct slightly protruding caudad, posterior margin with 2 + 2 setae. Paraprocts convex, medially with 1 + 1 setae. Hypoproct eye-shaped, with 1 + 1 setae. Legs medium-sized, about as long as midbody height. Male leg-pair 1 (Fig. 15 D) hypertrophied, 5 - segmented, strongly curved anteriad; coxa (cx) and telopoditomeres 1 and 2 (t 1, t 2) with some long setae. Telopoditomere 3 (t 3) longest, with a faint bulge (b) in the middle. Telopoditomere 4 (t 4) irregularly shaped, with few setae medially. Male leg-pair 2 (Fig. 15 E) reduced in size and stout; penis (pn) very flat, much shorter than coxae (cx), distal part with 6 long setae. Male leg-pair 7 (Fig. 15 F) with a 4 - or 5 - segmented telopodite (te) sometimes asymmetrical (not including claw); coxal process (cp) rather small, cone-shaped, with few long setae apically. Anterior gonopods (Fig. 15 G, H) with a very long coxal process (cp) bearing a row of short villi in anterior view, posteriorly carrying with a narrow membranous lobe (ml), distal outer margin of ml distinctly serrate. Flagellum (f) very long and slender, with spikes at margin. Telopodite (te) about 3 / 4 as long as cp, with long setae both apically and mesally, laterobasally with a minute remnant (r) of a second podomere. Posterior gonopods (Fig. 15 I) erect, branched distally, anterior branch smooth, with a pointed tip, posterior one densely setose. Vulva (Fig. 15 C) as usual, with two parallel rows of short setae on anterior and posterior surfaces; operculum (op) with 2 long distal setae.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
556B8BDE855659439061281CD7CCB1FF.taxon	description	Figures 2 D, 11	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
556B8BDE855659439061281CD7CCB1FF.taxon	diagnosis	Diagnosis. Differs from congeners mostly based on the following combination of characters: (1) penis much shorter than coxae 2 (Fig. 11 E); (2) male leg 7 with an extremely long coxal process, strongly bent anteriad, telopodite rather small, 2 or 3 - segmented (Fig. 11 F); (3) anterior gonopod (Fig. 11 G, H) with a coxal process of medium length and carrying a much lower, irregularly dentate, membranous lobe, and a very long and slender flagellum, distally branched. — In addition, this new species differs from all other species analyzed in a> 8.2 % uncorrected p - distance of the COI barcoding gene.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
556B8BDE855659439061281CD7CCB1FF.taxon	etymology	Etymology. The species name is primarily derived from its collection site, which is often used as a place for our graduate students to conduct internships on cave biodiversity. “ Genjudi ” here refers to the main operational base, in Chinese is “ 根据地 ”; a noun in apposition.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
556B8BDE855659439061281CD7CCB1FF.taxon	description	Description. Length of both sexes ca 30.0 – 42.0 mm, 1.7 – 2.1 mm in diameter, body with 51 – 63 podous + 1 – 2 apodous rings + telson. Natural coloration (Fig. 2 D) uniformly grayish yellow. Antennae and legs light yellow. Eye patches grey to blackish, subtriangular, arranged in 4 – 7 irregular rows, altogether about 30 – 40 ommatidia per eye patch. Head capsule smooth and hairless, eyes slightly bulged, epicranial suture well-defined. Labral margin with 1 (2) + 1 (2) supra-labral and 11 + 11 labral setae. Antennae long and slender, reaching behind to middle of ring 4 when stretched dorsally. In length, antennomeres 3> 2> 4> 5> 6> 1> 7. Antennomeres 5 and 6 each with distal corolla of sensilla basiconica. Antennomere 7 with 4 sensory cones. Mandibular stipes enlarged, with two small lobes in males, rounded in females. Gnathochilarium with at least 7 setae on each lamella lingualis (ll); promentum (pr) lance-shaped at base, swollen anteriorly, short and rod-like in males (Fig. 11 A), vs a drop-shaped pr in females (Fig. 11 B). Collum with 4 – 6 lower striae laterally, but lowest 2 striae not reaching the anterior margin. Prozona with 2 – 6 irregular subtransverse striae and metazona with 16 – 24 longitudinal striae laterally. Suture dividing pro- and metazona very narrow, a regular comb. Ozopores starting with ring 6, lying mid-laterally on sides of metatergites. Epiproct slightly protruding caudad, posterior margin with 3 + 3 setae. Paraprocts convex, with 1 + 1 setae. Hypoproct eye-shaped, with 1 + 1 setae. Legs long and slender, about 1.2 × as long as midbody height. Male leg-pair 1 (Fig. 11 D) hypertrophied, 5 - segmented, strongly curved anteriad; coxa (cx) and telopoditomeres 1 and 2 (t 1, t 2) with several long setae. Telopoditomere 3 (t 3) longest, about as long as all other telopoditomeres combined. Telopoditomere 4 (t 4) irregularly shaped, with two vague emarginations, one lateral, the other dorsal. Male leg-pair 2 (Fig. 11 E) reduced in size and slender; penis (pn) shorter than coxae (cx), distal part with five long setae. Male leg-pair 7 (Fig. 11 F) strongly modified, coxal process (cp) extremely long, flattened dorsoventrally, only slightly attenuated distad, strongly bent anteriad, densely setose nearly throughout at both mesal and lateral margins; telopodite (te) remains poorly visible laterally near base of cp, vestigial, consisting of 2 or 3 very small telopoditomeres. Anterior gonopods (Fig. 11 G, H) with a coxal process (cp) of medium length, laterally bearing a long row of short villi, posteriorly carrying with a lower membranous lobe (ml), outer margin of ml irregularly dentate, somewhat broadened and reflexed in proximal half. Flagellum (f) very long and slender, distally serrate and branched. Telopodite (te) about 2 / 3 as long as cp, with long setae both apically and mesally, laterobasally with a minute remnant (r) of a second podomere. Posterior gonopods (Fig. 11 I) erect, with bristles medially, distally branched, anterior branch smooth and slender, with a pointed tip, posterior one densely setose. Vulva (Fig. 11 C) as usual, with two parallel rows of short setae on both anterior and posterior surfaces, operculum (op) with several long setae distally.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
4AE50CFCEAE75F3BA9F861D0994EF64F.taxon	description	Figures 2 G, 13	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
4AE50CFCEAE75F3BA9F861D0994EF64F.taxon	diagnosis	Diagnosis. Differs from congeners mostly based on the following combination of characters: (1) telopoditomere 4 of male leg 1 with a small mesal hump (Fig. 13 D); (2) penis slightly higher than coxae 2 (Fig. 13 C); (3) male leg 7 with a very small coxal process, much lower than telopodite, which is made up of 4 segments and a claw (Fig. 13 F); (4) anterior gonopod (Fig. 13 E, I) with a very long coxal process carrying a narrow, lower, membranous lobe, distally with a vague indentation. — In addition, this new species differs from all other species analyzed in a> 8.0 % uncorrected p - distance of the COI barcoding gene.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
4AE50CFCEAE75F3BA9F861D0994EF64F.taxon	etymology	Etymology. The specific epithet refers to the distally vague indentation of a membranous lobe of anterior gonopod; adjective.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
4AE50CFCEAE75F3BA9F861D0994EF64F.taxon	description	Description. Length of both sexes ca 26.0 – 44.0 mm, 1.7 – 2.0 mm in diameter, body with 48 – 68 podous + 1 – 2 apodous rings + telson. Natural coloration (Fig. 2 G) red brownish to purple brownish; generally yellow brownish in alcohol, prozona dark brownish. Antennae and legs yellow. Eye patches blackish, subtriangular, arranged in 6 – 8 irregular rows, altogether about 24 – 48 ommatidia per eye patch. Head capsule smooth and hairless, eyes slightly bulged, epicranial suture well-defined. Labral margin with 2 + 2 supra-labral and 8 + 8 labral setae (Fig. 13 B). Antennae long and slender, reaching behind to middle of ring 4 when stretched dorsally. In length, antennomeres 3> 2> 4 ≈ 5> 4> 6> 7> 1. Antennomeres 5 and 6 each with a distal corolla of sensilla basiconica. (Fig. 13 B). Mandibular stipes well-rounded, with two small lobes in males, regularly rounded in females. Gnathochilarium with at least 5 setae on each lamella lingualis (ll); promentum (pr) short and lanceolate, anteriorly swollen in males (Fig. 13 B), vs a narrow rhombic pr in females (Fig. 13 A). Collum with 11 – 14 lower striae laterally, but lowest 2 striae not reaching the anterior margin. Prozona with 2 – 3 irregular subtransverse striae and metazona with 16 – 25 longitudinal striae laterally. Suture dividing pro- and metazona very narrow, a regular comb. Ozopores starting with ring 6, lying mid-laterally on sides of metatergites. Epiproct slightly protruding caudad, with 3 + 3 setae laterally. Paraprocts convex, with 1 + 1 setae. Hypoproct eye-shaped, with 1 + 1 setae. Legs medium-sized, about as long as midbody height. Male leg-pair 1 (Fig. 13 D) hypertrophied, 5 - segmented, strongly curved anteriad; coxa (cx) and telopoditomeres 1 and 2 (t 1, t 2) with some scattered long setae. Telopoditomere 3 (t 3) longest, about 1.5 × as long as telopoditomere 4 (t 4), t 4 irregularly shaped, subapically with a small mesal hump (h). Male leg-pair 2 (Fig. 13 C) reduced in size; penis (pn) relatively thick, only slightly higher than coxae (cx), distal part with 3 long setae. Male leg-pair 7 (Fig. 13 F) reduced, each leg with a 4 - segmented telopodite (te) and a claw, telopoditomere 2 (t 2) with a long seta subapically; coxal process (cp) small, median margin setose. Anterior gonopods (Fig. 13 E, I) with a very long coxal process (cp), bearing several short villi laterally, posteriorly carrying with a narrow, lower, membranous lobe (ml), outer margin of ml smooth, with a vague indentation distally. Flagellum (f) very long and slender, distally branched and villose. Telopodite (te) about 3 / 4 as long as cp, with long setae both apically and mesally, laterobasally with a minute remnant (r) of a second podomere. Posterior gonopods (Fig. 13 H) branched distally, anterior branch smooth, with a pointed tip, posterior one densely setose. Vulvae (Fig. 13 G) as usual, with two parallel rows of short setae on anterior and posterior surfaces, operculum (op) with several long setae distally.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
A53A99BD23CA510193C426B181E3C767.taxon	description	Figures 2 F, 16	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
A53A99BD23CA510193C426B181E3C767.taxon	diagnosis	Diagnosis. Differs from congeners mostly based on the following combination of characters: (1) telopoditomere 3 of male leg-pair 1 extremely long, longer than all other telopoditomeres combined (Fig. 16 D); (2) penis a little longer than coxae 2 (Fig. 16 E); (3) male leg-pair 7 with very large coxal processes, about 4 × as long as the telopodite (Fig. 16 F); (4) anterior gonopod (Fig. 16 H, I) with a very long coxal process carrying a narrow, lower, membranous lobe, outer margin distinctly microdentate; and an extremely slender flagellum. — In addition, this new species differs from all other species analyzed in a> 10.8 % uncorrected p - distance of the COI barcoding gene.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
A53A99BD23CA510193C426B181E3C767.taxon	etymology	Etymology. The species is named after its tye locality within the Laiyuan Karst Cave, which is the largest cave group known in Fujian Province, southeastern China; adjective.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
A53A99BD23CA510193C426B181E3C767.taxon	description	Description. Length of males ca 18.5 – 35.0 mm, 1.3 – 1.8 mm in diameter, body with 49 – 58 podous + 1 apodous rings + telson. Length of females ca 27.0 – 38.0 mm, 1.8 mm in diameter, body with 53 – 58 podous + 1 apodous rings + telson. Natural coloration (Fig. 2 F) yellowish, in alcohol marbled brownish or red brownish. Antennae and legs yellowish. Eye patches blackish, subtriangular, arranged in 4 – 6 irregular rows, altogether about 22 – 43 ommatidia per eye patch. Head capsule smooth and hairless, eyes slightly bulged, epicranial suture well-defined. Labral margin with 2 (3) + 2 (3) supra-labral and 12 + 12 labral setae. Antennae very long and slender, reaching behind to middle of ring 5 when stretched dorsally. Antennomeres: 3> 4> 2 ≈ 5> 6> 1> 7. Antennomeres 5 and 6 each with a distal corolla of sensilla basiconica (Fig. 16 C). Mandibular stipes well-rounded, with two small lobes in males, regularly rounded in females. Gnathochilarium with at least 6 setae on each lamella lingualis (ll); promentum (pr) heart-shaped at base, the remainder swollen and fusiform in males (Fig. 16 B), vs a narrow, oblong and rhomboid pr in females (Fig. 16 A). Collum with marbled brownish spots and 11 lower striae laterally, but lowest 2 striae not reaching the anterior margin. Prozona marbled, with 3 – 4 subtransverse striae and metazona with 20 – 23 longitudinal striae laterally. Suture dividing pro- and metazona very narrow, a regular comb. Ozopores starting with ring 6, lying mid-laterally on sides of metatergites. Epiproct slightly protruding caudad, with 7 + 7 setae at posterior margin. Paraprocts convex, medially with 3 + 3 setae. Hypoproct eye-shaped, with 1 + 1 setae. Legs medium-sized, about as long as midbody height. Male leg-pair 1 (Fig. 16 D) hypertrophied, 5 - segmented strongly curved anteriad; coxa (cx) and telopoditomeres 1 and 2 (t 1, t 2) with several long setae. Telopoditomere 3 (t 3) longest, longer than all other telopoditomeres combined. Telopoditomere 4 (t 4) irregularly shaped, medially with a few long setae. Male leg-pair 2 (Fig. 16 E) reduced in size and rather slender; penis (pn) slightly longer than coxae (cx), distal part with three long setae. Male leg-pair 7 (Fig. 16 F) strongly modified, with a very small 2 - segmented telopodite (te), telopoditomere 2 (t 2) larger, with a few long setae subapically; coxal process (cp) about 4.0 × as long as te, densely setose. Anterior gonopods (Fig. 16 H, I) with a very long coxal process (cp) bearing a short row of short villi in anterior view, posteriorly with a narrow and low membranous lobe (ml), outer margin of ml distinctly microdentate. Flagellum (f) very long and extremely slender, distally villose. Telopodite (te) about 2 / 3 as long as cp, densely setose apically and mesally. Posterior gonopods (Fig. 16 J) erect and slenderer, distally densely setose; anteriorly with a rather small pointed tip. Vulva (Fig. 16 G) as usual, with two parallel rows of short setae on both anterior and posterior surfaces. Operculum (op) subapically with three pairs of long setae, and two long distal setae on bursa.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
02ABD0308B975F3280C21BE8831D8F53.taxon	description	Figures 3 B, 7	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
02ABD0308B975F3280C21BE8831D8F53.taxon	diagnosis	Diagnosis. Differs from congeners mostly based on the following combination of characters: (1) penis much shorter than male coxae 2 (Fig. 7 E); (2) male leg 7 strongly reduced, with a rather small coxal process and a very small telopodite (Fig. 7 F); (3) anterior gonopod (Fig. 7 G, H) with a very long coxal process carrying a much lower and narrow membranous lobe, outer margin with an obvious groove and (4) an extremely long and slender flagellum, higher than coxal process. — In addition, this new species differs from all other Skleroprotopus species analyzed in uncorrected p - distances ranging between 6.6 % (compared to S. change sp. nov.) and 16.2 % (compared to S. longissimus sp. nov.).	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
02ABD0308B975F3280C21BE8831D8F53.taxon	etymology	Etymology. The specific epithet refers to the anterior gonopod flagellum being particularly long; adjective.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
02ABD0308B975F3280C21BE8831D8F53.taxon	description	Description. Length of males ca 24.0 – 28.5 mm, 1.7 – 1.8 mm in diameter, body with 47 – 59 podous + 1 – 2 apodous rings + telson. Length of females ca 30.0 – 37.0 mm, height 1.8 – 2.0 mm in diameter, body with 57 – 66 podous + 1 apodous rings + telson. Coloration in alcohol (Fig. 3 B) uniformly dark brown, prozona dark brownish, metazona yellow. Antennae and legs brownish. Eye patches blackish, subtriangular, arranged in 5 – 7 irregular rows, altogether about 25 – 55 ommatidia per eye patchr. Head capsule smooth and hairless, eyes slightly bulged, epicranial suture well-defined. Labral margin with about 26 setae in the first row and 4 setae in the second row. Antennae of medium length, reaching behind to anterior of ring 3 when stretched dorsally. In length, antennomeres 3> 2> 4> 5> 6> 1> 7. Antennomeres 5 and 6 each with a distal corolla of sensilla basiconica. Mandibular stipes well-rounded, more strongly rounded in females. Gnathochilarium with at least 4 setae on each lamella lingualis (ll); promentum (pr) lanceolate, swollen anteriorly in males (Fig. 7 A), vs a drop-shaped pr in females (Fig. 7 B). Collum (Fig. 3 B) with 5 – 7 lower striae laterally, but lowest 2 striae not reaching the anterior margin. Prozona with 2 – 5 irregular subtransverse striae and metazona with 14 – 18 longitudinal striae laterally. Suture dividing pro- and metazona very narrow, a regular comb. Ozopores starting with ring 6, lying mid-laterally on sides of metatergites (Fig. 3 B). Epiproct (Fig. 3 B) slightly protruding caudad, with 5 + 5 setae at posterior margin. Paraprocts convex, with 3 + 3 setae. Hypoproct eye-shaped, with 1 + 1 setae. Legs short, about 0.8 × as long as midbody height. Male leg-pair 1 (Fig. 7 D) hypertrophied, 5 - segmented, strongly curved anteriad; coxa (cx) and telopoditomeres 1 and 2 (t 1, t 2) with some long setae. Telopoditomere 3 (t 3) longest, about as long as all other telopoditomeres combined. Telopoditomere 4 (t 4) irregularly shaped, mid-point slightly concave and with several long setae caudally. Male leg-pair 2 (Fig. 7 E) reduced in size and slender; penis (pn) much shorter than coxae (cx), distal part with three long setae. Male leg-pair 7 (Fig. 7 F) strongly reduced, each leg with 1 - segmented telopodite, setose apically; coxal process (cp) rather small, higher than telopodite, medial margin and distally densely setose. Anterior gonopods (Fig. 7 G, H) with a very long and narrow coxal process (cp) bearing a long row of short villi laterally, posteriorly carrying with a narrow, much lower membranous lobe (ml), outer margin of ml somewhat broadened, but smooth, with an obvious groove distally. Flagellum (f) extremely long and slender, slightly higher than cp, one side of distal part villose. Telopodite (te) about 3 / 5 as long as cp, with long setae both distally and mesally, laterobasally with a minute remnant (r) of a second podomere. Posterior gonopods (Fig. 7 I) erect and slightly stout, clearly branched at tip, anterior branch smooth, with a pointed tip; posterior one densely setose. Vulva (Fig. 7 C) as usual, with two parallel rows of short setae on anterior and posterior surfaces, operculum (op) with long setae distally.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
375FCDC36A6A5496BE22B9AEAE52D4C7.taxon	description	Figures 2 C, 10	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
375FCDC36A6A5496BE22B9AEAE52D4C7.taxon	diagnosis	Diagnosis. Differs from congeners mostly based on the following combination of characters: (1) penis much shorter than coxae 2 (Fig. 10 E); (2) male leg-pair 7 with a rather long and slender coxal process (Fig. 10 F); (3) anterior gonopod (Fig. 10 G, H) with a very long coxal process carrying an irregularly dentate and membranous lobe, and a very long and slender flagellum. — In addition, this new species differs from all other species analyzed in a> 12.4 % uncorrected p - distance of the COI barcoding gene.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
375FCDC36A6A5496BE22B9AEAE52D4C7.taxon	etymology	Etymology. Latin “ longissimus ” is meaning “ the longest ”. The specific epithet refers to the coxal process of male leg-pair 7 being particularly long and relatively slender; ajective.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
375FCDC36A6A5496BE22B9AEAE52D4C7.taxon	description	Description. Length of both sexes ca 25.0 – 40.0 mm, 1.6 – 1.9 mm in diameter, body with 47 – 54 podous + 1 apodous rings + telson. Natural coloration (Fig. 2 C) red brownish, in alcohol uniformly brown yellowish, prozona dark brownish, metazona yellowish. Antennae and legs light yellowish. Eye patches blackish, subtriangular, arranged in 4 – 7 irregular rows, altogether about 20 – 37 ommatidia per eye patch. Head capsule smooth and hairless, eyes slightly bulged, epicranial suture well-defined. Labral margin with 2 (3) + 2 (3) supra-labral and 9 + 9 labral setae. Antennae of medium length, reaching behind to anterior of ring 3 when stretched dorsally. In length, antennomeres 3> 2> 4 ≈ 5> 6> 1> 7. Antennomeres 5 and 6 each with a distal corolla of sensilla basiconica. Mandibular stipes well-rounded, with two small lobes in males, more rounded in females. Gnathochilarium with at least 4 setae on each lamella lingualis (ll); promentum (pr) relatively narrow, lance-shaped at base, swollen anteriorly and cup-shaped in males (Fig. 10 A), vs a slightly broadened and oblong rhomboid pr in females (Fig. 10 B). Collum with 5 – 7 lower striae laterally, but lowest 2 striae not reaching the anterior margin. Prozona with 4 – 6 irregular subtransverse striae and metazona with 11 – 25 longitudinal striae laterally. Suture dividing pro- and metazona very narrow, a regular comb. Ozopores starting with ring 6, lying mid-laterally on sides of metatergites. Epiproct slightly protruding caudad, with 4 + 4 setae at posterior margin. Paraprocts convex, with 2 + 2 setae. Hypoproct eye-shaped, with 1 + 1 setae. Legs long and slender, about 1.1 × as long as midbody height. Male leg-pair 1 (Fig. 10 D) hypertrophied, 5 - segmented, strongly curved anteriad; coxa (cx) and telopoditomeres 1 and 2 (t 1, t 2) with some long setae. Telopoditomere 3 (t 3) longest, about as long as all other telopoditomeres combined; Telopoditomere 4 (t 4) irregularly shaped, with two vague emarginations, one lateral, the other dorsal. Male leg-pair 2 (Fig. 10 E) reduced in size and very slender; penis (pn) shorter than coxae (cx), distal part with three long setae. Male leg-pair 7 (Fig. 10 F) strongly modified, coxal process (cp) extremely long and slender, crowned with a small hook facing anteriorly, strongly bent frontally, mostly subparallel-sided, considerably more slender, and free from setae along distal 2 / 3; telopodite (te) 1 - segmented, rather small, carrying a long apical seta. Anterior gonopods (Fig. 10 G, H) with a medium length coxal process (cp), posteriorly carrying with an irregularly dentate membranous lobe (ml), outer margin of ml somewhat broadened and reflexed in proximal half, margin of reflexed part irregular. Flagellum (f) very long and slender, distally serrate and branched. Telopodite (te) about 3 / 4 as long as cp, with long setae both apically and mesally, laterobasally with a minute remnant (r) of a second podomere. Posterior gonopods (Fig. 10 I) erect, branched at tip, anterior branch smooth, with a pointed tip, posterior one densely setose. Vulva (Fig. 10 C) as usual, with two parallel rows of short setae on both anterior and posterior surfaces, operculum (op) with several long setae distally.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
9E190A1864AF5B00A20D3C9862598257.taxon	description	Figures 2 B, 6	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
9E190A1864AF5B00A20D3C9862598257.taxon	diagnosis	Diagnosis. Differs from congeners mostly based on the following combination of characters: (1) telopoditomere 3 of male leg 1 particularly broad and bulging in the middle (Fig. 6 E); (2) penis subequal in height to coxae 2 (Fig. 6 F); (3) coxal process of male leg 7 very small, lower than the telopodite (Fig. 6 D); (4) anterior gonopod with a long coxal process carrying a highly developed, axe-shaped, membranous lobe (Fig. 6 I). — In addition, this new species differs from all other species analyzed in a> 10.3 % uncorrected p - distance of the COI barcoding gene.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
9E190A1864AF5B00A20D3C9862598257.taxon	etymology	Etymology. Latinised Greek “ megistus ” is meaning “ large ”. The specific epithet refers to the male leg-pair 1 being particularly large; adjective.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
9E190A1864AF5B00A20D3C9862598257.taxon	description	Description. Length of both sexes ca 31.5 – 40.0 mm, 1.8 – 2.2 mm in diameter, body with 53 – 65 podous + 1 apodous rings + telson. Natural coloration generally pale reddish brown (Fig. 2 B), in alcohol marbled yellowish brown. Antennae and legs light yellowish. Eye patches blackish, triangular, arranged in 4 – 6 irregular rows, altogether about 22 – 52 ommatidia per eye patch. Head capsule smooth and hairless, eyes slightly bulged, epicranial suture well-defined. Labral margin with 4 + 4 supra-labral and 12 + 12 labral setae. Antennae very long and slender, reaching behind to middle of ring 5 when stretched dorsally. In length, antennomeres 3> 2> 4 ≈ 5> 6> 1> 7. Antennomeres 5, 6 and 7 each with a distal corolla of sensilla basiconica. Mandibular stipes well-rounded in males, more rounded in females. Gnathochilarium with at least 5 setae on each lamella lingualis (ll); promentum (pr) lance-shaped at base, swollen anteriorly, olive-shaped in males (Fig. 6 A), vs a narrow rhomboid pr in females (Fig. 6 B). Collum with marbled brownish spots and 8 – 10 lower striae laterally, but lowest 3 striae not reaching the anterior margin. Prozona with 3 – 4 subtransverse striae and metazona with 13 – 22 longitudinal striae laterally. Suture dividing pro- and metazona very narrow, a regular comb. Ozopores starting with ring 6, lying mid-laterally on sides of metatergites (Fig. 2 B). Epiproct slightly protruding caudad, posterior margin with 3 + 3 setae. Paraprocts convex, medially with 1 + 1 setae. Hypoproct eye-shaped, with 1 + 1 setae. Legs medium-sized, about as long as midbody height. Male leg-pair 1 (Fig. 6 E) hypertrophied, 5 - segmented, tip slightly curved anteriad. Coxa (cx) and telopoditomeres 1 and 2 (t 1, t 2) with some long setae. Telopoditomere 3 (t 3) widest and largest, bulging in the middle. Telopoditomere 4 (t 4) subtriangular; t 3 and t 4 devoid of long setae. Male leg-pair 2 (Fig. 6 F) reduced in size; penis (pn) subequal in height to coxae (cx), distal part with three long setae. Male leg-pair 7 (Fig. 6 D) reduced, with 2 - segmented telopodites (te), t 1 large and setose subapically; coxal process (cp) very small, lower than telopodite (te), with several long setae distally. Anterior gonopods (Fig. 6 H, I) with a very long coxal process (cp) bearing an oval field of short villi in anterior view, posteriorly carrying with a highly developed, axe-shaped, membranous lobe (ml), slightly lower than cp, outer margin of ml with a large, sharp and triangular protrusion, clearly wider than cp. Flagellum (f) very long and slender, villose in distal part. Telopodite (te) about half as long as cp, with long setae apically, laterobasally with a minute remnant (r) of a second podomere. Posterior gonopods (Fig. 6 G, J) stout, anterior branch slender and with a small pointed tip, posterior one semi-annulate and densely setose distally. Vulva (Fig. 6 C) as usual, with two parallel rows of short setae on anterior and posterior surfaces, and two long distal setae on operculum (op).	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
E79A521BC87F54558A813EAC186A270A.taxon	description	Figures 3 G, 14	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
E79A521BC87F54558A813EAC186A270A.taxon	diagnosis	Diagnosis. Differs from congeners mostly based on the following combination of characters: (1) collum with 10 – 16 lower striae and 18 – 27 longitudinal striae laterally on metazona (Fig. 3 G). (2) telopoditomere 4 of male leg-pair 1 with two small mesal humps (Fig. 14 D); (3) penis stout, distally sharpened, much shorter than coxae 2 (Fig. 14 C); (4) male leg 7 with a very small coxal process, only slightly higher than 2 - segmented telopodite (Fig. 14 E); (5) anterior gonopod (Fig. 14 G, H) with a very long and slender coxal process carrying a narrow, lower, membranous lobe, distally with a vague indentation. — In addition, this new species differs from all other Skleroprotopus species analyzed in uncorrected p - distances ranging from between 8.0 – 8.3 % (compared to S. incisodentatus sp. nov.) and 14.6 % (compared to S. longissimus sp. nov.).	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
E79A521BC87F54558A813EAC186A270A.taxon	etymology	Etymology. The species is named after the more numerous striae on the collum and metazona; adjective.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
E79A521BC87F54558A813EAC186A270A.taxon	description	Description. Length of males ca 29.5 – 34.5 mm, 1.8 – 2.0 mm in diameter, body with 53 – 59 podous + 1 apodous rings + telson. Length of females ca 34.0 – 37.0 mm, 1.9 – 2.1 mm in diameter, body with 57 – 62 podous + 1 – 2 apodous rings + telson. Coloration in alcohol (Fig. 3 G) generally marbled dark brownish, antennae and legs light brownish. Eye patches blackish, subtriangular, arranged in 4 – 6 irregular rows, altogether about 26 – 36 ommatidia per eye patch. Head capsule smooth and hairless, eyes slightly bulged, epicranial suture well-defined. Labral margin with 2 + 2 supra-labral and 9 + 9 labral setae. Antennae long and slender, reaching behind to middle of ring 4 when stretched dorsally. In length, antennomeres 3> 2> 5> 4> 6> 7> 1 (Fig. 14 F). Antennomeres 5 and 6 (Fig. 14 F) each with a distal corolla of sensilla basiconica. Mandibular stipes well-rounded in male, even more strongly rounded in female. Gnathochilarium with at least 5 setae on each lamella lingualis (ll); promentum (pr) lute-shaped, anteriorly swollen and very narrow in males (Fig. 14 B), vs a narrow, oblong and rhomboid pr in females (Fig. 14 A). Collum with 10 – 16 lower striae laterally, but lowest 4 striae not reaching the anterior margin. Prozona with 2 – 6 irregular subtransverse striae and metazona with 18 – 27 longitudinal striae laterally (Fig. 3 G). Suture dividing pro- and metazona very narrow, a regular comb. Ozopores starting with ring 6, lying mid-laterally on sides of metatergites. Epiproct slightly protruding caudad, posterior margin with 4 + 4 setae. Paraprocts convex, with 1 + 1 setae. Hypoproct eye-shaped, with 1 + 1 setae. Legs medium-sized, about as long as midbody height. Male leg-pair 1 (Fig. 14 D) hypertrophied, 5 - segmented, strongly curved anteriad; Coxa (cx) and telopoditomeres 1 and 2 (t 1, t 2) with some scattered long setae. Telopoditomere 3 (t 3) longest, about 1.5 × as long as telopoditomere 4 (t 4), t 4 irregularly shaped, caudally densely setose, subapically with two small mesal humps (h). Male leg-pair 2 (Fig. 14 C) reduced in size and somewhat stout; coxae (cx) elongated, penis (pn) very stout, much shorter than cx, distal part sharpened, seta absent. Male leg-pair 7 (Fig. 14 E) strongly reduced, each leg with a 2 - segmented asymmetrical telopodite (te), telopoditomere 1 (t 1) with several long setae; coxal process (cp) very small, slightly higher than te, medial margin and distally densely setose. Anterior gonopods (Fig. 14 G, H) with a very long and narrow coxal process (cp) bearing a row of short villi in anterior view, posteriorly carrying with a low, narrow, membranous lobe (ml), outer margin of ml smooth, with a vague indentation distally. Flagellum (f) very long and slender, distally branched and villose. Telopodite (te) about 2 / 3 as long as cp, with long setae at mesal margin and apically, laterobasally with a minute remnant (r) a second podomere. Posterior gonopods (Fig. 14 I) erect, branched distally, anterior branch smooth, with a very small and pointed tip, posterior one densely setose. Vulva (Fig. 14 J) as usual, with two parallel rows of short setae on both anterior and posterior surfaces; operculum (op) with 3 long distal setae.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
D538F573E57D50CC8A40344706BB2A51.taxon	description	Figures 3 A, 9	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
D538F573E57D50CC8A40344706BB2A51.taxon	etymology	Etymology. The name is derived from the overseas immortal mountains in Chinese mythology to emphasize the local cave harboring this species. “ Penglai ” in Chinese “ 蓬莱 ”, a noun in apposition.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
D538F573E57D50CC8A40344706BB2A51.taxon	diagnosis	Diagnosis. Differs from congeners mostly based on the following combination of characters: (1) penis much shorter than coxae 2 (Fig. 9 F); (2) male leg 7 with a very long coxal process and a rather small telopodite (Fig. 9 C); (3) anterior gonopod (Fig. 9 G, H) with a very long coxal process carrying a much lower, distally grooved membranous lobe, and a very long and thick flagellum. — In addition, this new species differs from all other species analyzed in a> 10.1 % uncorrected p - distance of the COI barcoding gene.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
D538F573E57D50CC8A40344706BB2A51.taxon	description	Description. Length of both sexes ca 20.0 – 36.5 mm, 1.5 – 1.9 mm in diameter, body with 50 – 62 podous + 1 – 3 apodous rings + telson. Coloration in alcohol (Fig. 3 A) uniformly brown yellowish, prozona dark brownish, metazona yellow. Antennae and legs light yellowish. Eye patches blackish, subtriangular, arranged in 5 – 7 irregular rows, altogether about 22 – 39 ommatidia per eye patch. Head capsule smooth and hairless, eyes slightly bulged, epicranial suture well-defined. Labral margin with 2 (3) + 2 (3) supra-labral and 7 + 7 labral setae. Antennae of medium length, reaching behind to middle of ring 2 when stretched dorsally. In length, antennomeres 3> 2> 4> 5> 6> 7> 1. Antennomeres 5 and 6 each with a distal corolla of sensilla basiconica. Mandibular stipes enlarged, with two faint lobes in males (Fig. 3 A), rounded in females. Gnathochilarium with at least 6 setae on each lamella lingualis (ll); promentum (pr) lance-shaped at base, swollen anteriorly and olive-shaped in males (Fig. 9 B), vs a conical pr in females (Fig. 9 A). Collum (Fig. 3 A) with 9 – 11 lower striae laterally, but lowest 2 striae not reaching the anterior margin. Dorsum with 5 + 5 short setae at rear edge of metatergites (Fig. 3 A). Prozona with 2 – 4 irregular subtransverse striae and metazona with 15 – 23 longitudinal striae laterally. Suture dividing pro- and metazona very narrow, a regular comb. Ozopores starting with ring 6, lying mid-laterally on sides of metatergites (Fig. 3 A). Epiproct slightly protruding caudad, with 4 + 4 setae along posterior margin. Paraprocts convex, with 3 + 3 setae (Fig. 3 A). Hypoproct eye-shaped, with 1 + 1 setae. Legs long and slender, about 1.2 × as long as midbody height. Male leg-pair 1 (Fig. 9 D) hypertrophied, 5 - segmented, strongly curved anteriad; coxa (cx) and telopoditomeres 1 and 2 (t 1, t 2) with some long setae. Telopoditomere 3 (t 3) longest. Telopoditomere 4 (t 4) irregularly shaped, mid-point slightly concave. Male leg-pair 2 (Fig. 9 F) reduced in size and very slender; penis (pn) much shorter than coxae (cx), distal part with three long setae. Male leg-pair 7 (Fig. 9 C) strongly modified, with 2 - segmented, rather small telopodites (te), telopoditomere 1 (t 1) larger, with a few long setae subapically; coxal processes (cp) V-shaped, very long, densely setose. Anterior gonopods (Fig. 9 G, H) with a very long coxal process (cp) bearing a long row of short villi laterally, posteriorly carrying with a narrow, much lower, membranous lobe (ml), outer margin of ml somewhat broadened, but smooth, with an obvious groove distally. Flagellum (f) very long and thick, slightly shorter than cp, distally villose. Telopodite (te) about half as long as cp, with long setae both apically and mesally, laterobasally with a minute remnant (r) of a second podomere. Posterior gonopods (Fig. 9 E) erect, branched at tip, anterior branch smooth, with a pointed tip, posterior one densely setose. Vulva (Fig. 9 I) as usual, with two parallel rows of short setae on both anterior and posterior surfaces, operculum (op) with long setae distally.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
1E81187968E0580FA7AE94DD1EAC22E7.taxon	description	Figures 2 E, 3 C, 5	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
1E81187968E0580FA7AE94DD1EAC22E7.taxon	diagnosis	Diagnosis. Differs from congeners mostly based on the following combination of characters: (1) penis much longer than coxae 2 (Fig. 5 I); (2) male leg 7 with a very small coxal process, lower than the telopodite (Fig. 5 E); (3) anterior gonopod (Fig. 5 G, H) with a large and foot-shaped like coxal process, carrying a broadened membranous lobe, outer margin with a triangular protrusion, and a relatively thick flagellum. — In addition, this new species differs from all other species analyzed in a> 10.3 % uncorrected p - distance of the COI barcoding gene.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
1E81187968E0580FA7AE94DD1EAC22E7.taxon	etymology	Etymology. The species is named after its type locality that is the Hanzhong sinkhole group, a geological wonder composed of multiple heavenly pits. “ Tiankeng ” in Chinese “ 天坑 ”, a noun in apposition.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
1E81187968E0580FA7AE94DD1EAC22E7.taxon	description	Description. Length of both sexes ca 29.0 – 38.9 mm, width 1.9 – 2.2 mm, midbody height 1.8 – 2.1 mm, body with 49 – 63 podous + 1 apodous rings + telson. Natural coloration generally yellow to brownish. Antennae and legs light yellowish (Fig. 2 E). Eye patches transparent, subtriangular, arranged in 4 – 6 irregular rows, altogether about 18 – 27 (males) or 15 – 40 (females) ommatidia per eye patch. Head capsule smooth and hairless, eyes slightly bulged, epicranial suture well-defined. Labral margin with 2 + 2 supra-labral and 8 + 8 labral setae. Antennae long, reaching behind to anterior of ring 4 when stretched dorsally (Fig. 3 C). In length, antennomeres 3> 5> 2> 4 ≈ 6> 1 ≈ 7 (Fig. 5 C). Antennomeres 5, 6 and 7 each with a distal corolla of sensilla basiconica. Mandibular stipes enlarged, with two small lobes in males, well-rounded in females. Gnathochilarium with at least 3 – 8 setae on each lamella lingualis (ll) in males; promentum (pr) columnar in males, anteriorly swollen and tubular in shape (Fig. 5 B), vs a broad conical pr in females (Fig. 5 A). Collum with marbled brownish spots and 8 – 10 lower striae laterally, but lowest 2 striae not reaching the anterior margin. Prozona with 3 irregular subtransverse striae and metazona with 13 – 24 longitudinal striae laterally (Fig. 3 C). Suture dividing pro- and metazona rather narrow, a nearly regular comb. Ozopores starting with ring 6, lying mid-laterally on sides of metatergites. Epiproct slightly protruding caudad, posterior margin with 6 + 6 setae. Paraprocts convex, with 1 + 1 setae. Hypoproct eye-shaped, with 1 + 1 setae. Legs long and slender, about 1.4 × as long as midbody height. Male leg-pair 1 (Fig. 5 D) hypertrophied, 5 - segmented, strongly curved anteriad. Coxa (cx) and telopoditomeres 1 and 2 (t 1, t 2) each with few long setae, others mostly short. Telopoditomere 3 (t 3) longest, about equivalent in length to the sum of the cx and t 1 – 2; Telopoditomere 4 (t 4) irregularly shaped, with several long setae subapically. Male leg-pair 2 (Fig. 5 I) reduced in size; penis (pn) pyramid-shaped, much longer than coxae (cx), distal part with two long setae (Fig. 5 I). Male leg-pair 7 (Fig. 5 E) reduced, with 3 - segmented telopodites, telopoditomere 2 (t 2) largest and setose subapically; coxal process (cp) small, much lower than telopodite, densely setose at mesal margin. Anterior gonopods (Fig. 5 G, H) with a large, foot-shaped like coxal process (cp) bearing with a long row of short villi in anterior view, posteriorly carrying with a broadened, and high membranous lobe (ml), slightly shorter than cp, middle outer margin of ml with a triangular protrusion. Flagellum (f) very long and relatively thick, distally spinulose-villose. Telopodite (te) about half as long as cp, with long setae apically, laterobasally with a minute remnant (r) of a second podomere. Posterior gonopods (Fig. 5 F) erect and stout, deeply branched at tip, anterior branch smooth, with a pointed tip, higher than a densely setose posterior one. Vulva (Fig. 5 J) mostly wrapped inside a membrane, characteristic in shape, strongly elongated; operculum (op) longer than bursa, with two parallel rows of short setae on anterior and posterior surfaces, and with at least 10 long distal setae.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
4DB77B61D622521E8BABF4497E245AC4.taxon	description	Figures 3 F, 4	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
4DB77B61D622521E8BABF4497E245AC4.taxon	diagnosis	Diagnosis. Differs from congeners mostly based on the following combination of characters: (1) leg 1, 6 - segmented (Fig. 4 D); (2) penis much shorter than coxae 2 (Fig. 4 F); (3) coxal process of leg 7 (Fig. 4 E) tongue-shaped, about 2 × as long as a club-shaped telopodite; (4) anterior gonopod with a very long and narrow coxal process, carrying an irregularly dentate and membranous lobe (Fig. 4 I). — In addition, this new species differs from all other species analyzed in a> 12.7 % uncorrected p - distance of the COI barcoding gene.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
4DB77B61D622521E8BABF4497E245AC4.taxon	etymology	Etymology. The specific epithet is honored to Dr. Yu Tiantian (余甜甜) (Kunming Institute of Zoology, Chinese Academy of Sciences), who is the collector; noun in the genitive case.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
4DB77B61D622521E8BABF4497E245AC4.taxon	description	Description. Length of holotype ca 25.0 mm, 1.6 mm in diameter, body with 53 podous + 1 apodous rings + telson. Coloration in alcohol marbled dark brownish, slightly lighter in anterior 1 / 3 body (Fig. 3 F). Antennae semi-transparent, gradually infuscate from base to end, antennomeres 5 and 6 darkest, frosted violet (Fig. 4 C). Eye patches blackish, triangular, arranged in 6 irregular rows, about 35 (left) or 43 (right) ommatidia per eye patch (Fig. 4 B). Legs pale yellow. Head capsule (Fig. 4 B) smooth and hairless, vertex bulged, epicranial suture obviously concave. Labral margin with 2 + 2 supra-labral and 6 (7) + 6 (7) labral setae (Fig. 4 A). Antennae (Fig. 4 C) medium in length, reaching behind to middle of ring 2 when stretched dorsally. In length, antennomeres 3> 2> 4> 6 ≈ 5> 1> 7. Antennomeres 5 and 6 each with a distal corolla of sensilla basiconica. Antennomere 7 invaginated terminally, sensory cones inconspicuous. Mandibular stipes enlarged, with two nearly equal lobes (Fig. 3 F). Gnathochilarium with at least 4 setae on each lamella lingualis (ll); promentum (pr) long and diamond-shaped, swollen anteriorly (Fig. 4 A). Collum (Fig. 3 F) with 6 lower striae laterally, but lowest stria not reaching the anterior margin. Prozona delicately alveolate, metazona laterally with 13 – 18 longitudinal striae (Fig. 3 F). Suture dividing pro- and metazona narrow, a regular comb. Ozopores starting with ring 6, lying anteriorly on lateral sides of metatergites. Epiproct slightly protruding caudad, with 2 + 2 setae along posterior margin. Paraprocts convex, medially with 2 + 2 setae. Hypoproct eye-shaped, with 1 + 1 setae. Legs (Fig. 3 F) short, about 0.7 × as long as midbody height, claw weakly curved, without modifications. Male leg-pair 1 (Fig. 4 D) hypertrophied, strongly curved anteriad, 6 - segmented, but dividing line (dl) between telopoditomeres 4 and 5 (t 4, t 5) incomplete. Both t 4 and t 5 combined longest, irregularly shaped, with a few scattered setae and short bristles medially. Coxa (cx) and telopoditomeres 1 and 2 (t 1, t 2) with some long setae. Telopoditomere 3 (t 3) broadest. Male leg-pair 2 (Fig. 4 F) reduced in size and slender; penis (pn) unpaired, shorter than coxae (cx), cylindrical, distal part with two long setae. Male leg-pair 7 (Fig. 4 E) strongly modified, with 2 - segmented telopodites, club-shaped, densely setose apically, telopoditomere 2 rather small. Coxal process (cp) strongly elongated, tongue-shaped, about 2 × as long as telopodite (te), clothed with dense long bristles. Anterior gonopods (Fig. 4 H, I) with a very long and rather slender coxal process (cp) bearing a long row of short villi in anterior view, posteriorly carrying with an irregularly dentate membranous lobe (ml), outer margin of ml distinctly broadened and reflexed, subtriangular, distally sharp. Flagellum (f) long and slender, a little barbed. Telopodite (te) about 2 / 3 as long as cp, with long setae apically. Posterior gonopods (Fig. 4 G) erect and thin, anterior branch slender, tapering regularly towards tip, higher than posterior branch, and both densely setose distally. Female unknown.	en	Chen, Rong, Zhao, Yi, Golovatch, Sergei, Liu, Wei-Xin (2024): Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae). Arthropod Systematics & Phylogeny 82: 659-691, DOI: 10.3897/asp.82.e136751
