taxonID	type	description	language	source
A91587910B45D21EFD4622C020F2156C.taxon	type_taxon	Type Species. Brachinus quadrimaculatus Dejean, 1831 (designated by Andrewes 1939)	en	Akhil, S. V., Thomas, Sabu K. (2019): New Species of Bombardier Beetles of the Genus Styphlomerus Chaudoir, 1875 (Coleoptera: Carabidae) from Southern India, with a Key to the Indomalayan and Palearctic Species. The Coleopterists Bulletin 73 (2): 465-471, DOI: 10.1649/0010-065X-73.2.465, URL: http://dx.doi.org/10.1649/0010-065x-73.2.465
A91587910B45D21EFD4622C020F2156C.taxon	description	Description. Length 4.5 – 12.0 mm. Mouthparts: Ligula smooth, translucent, truncate, generally bisetose (otherwise plurisetose with 2 long setae over apex and many short setae centrally); paraglossae membranous, wide, glabrous, truncate, subequal to length of ligula. Palpi thick, densely pubescent, maxillary palpi with penultimate article short, conical, terminal segment long, slightly swollen, oval; labial palpi with terminal segment subsecuriform, obliquely truncate. Mentum deeply emarginate, median tooth triangular, acute; lobes divergent, wide; epilobes with distinct suture, triangular, apex rounded. Labrum glabrous, flat, transverse, narrow, with 6 evenly spaced setae. Antennae: Thick, covered with dense setae. Mandibles thick, not crossing each other, scrobe unisetose. Elytra: Generally rectangular, sides subparallel, humerus square. Hind wings: Present. Legs: Femora densely pubescent; tarsi with basal article scarcely attenuate. Male protarsomeres 1 – 3 asymmetrical (symmetrical in females), adhesive pads beneath forming 2 parallel rows of spatulate setae.	en	Akhil, S. V., Thomas, Sabu K. (2019): New Species of Bombardier Beetles of the Genus Styphlomerus Chaudoir, 1875 (Coleoptera: Carabidae) from Southern India, with a Key to the Indomalayan and Palearctic Species. The Coleopterists Bulletin 73 (2): 465-471, DOI: 10.1649/0010-065X-73.2.465, URL: http://dx.doi.org/10.1649/0010-065x-73.2.465
A91587910B46D218FD0826B9227A10B1.taxon	description	Specimens Examined: Two females, India: Tamil Nadu state: Virudhunagar district: Krishnan Kovil, light attracted, 15. X. 2017, coll. S. V. Akhil.	en	Akhil, S. V., Thomas, Sabu K. (2019): New Species of Bombardier Beetles of the Genus Styphlomerus Chaudoir, 1875 (Coleoptera: Carabidae) from Southern India, with a Key to the Indomalayan and Palearctic Species. The Coleopterists Bulletin 73 (2): 465-471, DOI: 10.1649/0010-065X-73.2.465, URL: http://dx.doi.org/10.1649/0010-065x-73.2.465
A91587910B46D218FD0826B9227A10B1.taxon	materials_examined	Type Material. Holotype labeled “ Holotype, Female, India: Tamil Nadu state: Virudhunagar district: Krishnan Kovil, light attracted, 15. X. 2017, coll. S. V. Akhil ”. Paratype labeled “ Paratype, Female, India: Tamil Nadu state: Virudhunagar district: Krishnan Kovil, light attracted, 15. X. 2017, coll. S. V. Akhil ”. Deposited in ZSIC.	en	Akhil, S. V., Thomas, Sabu K. (2019): New Species of Bombardier Beetles of the Genus Styphlomerus Chaudoir, 1875 (Coleoptera: Carabidae) from Southern India, with a Key to the Indomalayan and Palearctic Species. The Coleopterists Bulletin 73 (2): 465-471, DOI: 10.1649/0010-065X-73.2.465, URL: http://dx.doi.org/10.1649/0010-065x-73.2.465
A91587910B46D218FD0826B9227A10B1.taxon	description	Description. TLA = 6.3 – 6.6 mm. Color: Head, pronotum, scape, palpi, scutellum, region around scutellum, and legs reddish yellow, scape, palpi, and legs a lighter hue of reddish yellow; elytra bluish black; head to metasternum pale yellow; mesepimeron yellow with brown hue. Head: Stout, widest behind eyes, as wide as pronotum, as long as broad, vertex and neck rough with dense punctation and wrinkles, frons smooth with fine punctures and faintly wrinkled; labrum subrectangular, apex slightly convex, with 6 setae on apical margin; clypeus subrectangular, glabrous, base wider than apex; frontoclypeal margin marked by a faint brownish black line; frontal fovea shallow; frons impunctate; lateral lobes of mentum broad, with a tooth-like projection laterally, glabrous, impunctate, median tooth short, stout, blunt; submentum very narrow; antennae short, thick, antennomeres cylindrical, apical antennomere pointed, elongate-oval; apical palpomeres subcylindrical, apex tapering, penultimate palpomere shaped as inverted cone, 2 nd palpomere curved, palpomeres sparsely pubescent. Pronotum: Wider than long, flat, rough with wrinkles and punctation throughout disc, disc matt, sparsely pubescent; anterior margin straight, lateral angles of anterior margin projecting forward, posterior margin concave, sinuate; disk with anterior half broad, convex, posterior half narrowed but widening towards base, hind angles pointed, obtuse, slightly projecting laterally; median furrow shallow, not reaching anterior and posterior margins; lateral ridge narrow, margin brownish black. Scutellum: Broad, pointed. Elytra: Subparallel, dorsal surface matt, rough, wrinkled, impunctate, sparsely pubescent; base narrowed, humerus pronounced, projecting anteriorly; widest just before apex, apex rounded; sutural line reddish yellow, widest just below scutellum, narrowed after that, continuing very faintly to apex; striations weak, with sparse pubescence on both sides of striae, intervals rounded. Hind wings: Welldeveloped. Venter: Genae broad, with sparse pubescence under eyes, remainder glabrous, smooth; gula divergent posteriorly, distinctly marked by brownish black gular suture; prothorax with sparse pubescence in anterolateral region of prosternum, midregion glabrous; prosternal process curved downwards, glabrous; apex of abdomen pointed. Measurements. Holotype (female): TLA = 6.36 mm, TLB = 5.7 mm, PL = 0.97 mm, PW = 1.2 mm, EL = 3.68 mm, TW = 2.62 mm. Paratype (female): TLA = 6.62 mm, TLB = 5.64 mm, PL = 0.98 mm, PW = 1.2 mm, EL = 3.6 mm, TW = 2.52 mm. Geographical Distribution. INDIA: Tamil Nadu State: Virudhunagar district: Krishnan Kovil. Collection Method. Collected under fluorescent light.	en	Akhil, S. V., Thomas, Sabu K. (2019): New Species of Bombardier Beetles of the Genus Styphlomerus Chaudoir, 1875 (Coleoptera: Carabidae) from Southern India, with a Key to the Indomalayan and Palearctic Species. The Coleopterists Bulletin 73 (2): 465-471, DOI: 10.1649/0010-065X-73.2.465, URL: http://dx.doi.org/10.1649/0010-065x-73.2.465
A91587910B46D218FD0826B9227A10B1.taxon	etymology	Etymology. Named after the locality (Devagiri) of our host institution.	en	Akhil, S. V., Thomas, Sabu K. (2019): New Species of Bombardier Beetles of the Genus Styphlomerus Chaudoir, 1875 (Coleoptera: Carabidae) from Southern India, with a Key to the Indomalayan and Palearctic Species. The Coleopterists Bulletin 73 (2): 465-471, DOI: 10.1649/0010-065X-73.2.465, URL: http://dx.doi.org/10.1649/0010-065x-73.2.465
A91587910B46D218FD0826B9227A10B1.taxon	discussion	Remarks. This species is similar to Styphlomerus striatus Akhil and Thomas, new species but differs by having smaller form, wider head, bluish black elytra, elytral humerus projecting forward, narrowed elytral base, and narrow, reddish yellow sutural line.	en	Akhil, S. V., Thomas, Sabu K. (2019): New Species of Bombardier Beetles of the Genus Styphlomerus Chaudoir, 1875 (Coleoptera: Carabidae) from Southern India, with a Key to the Indomalayan and Palearctic Species. The Coleopterists Bulletin 73 (2): 465-471, DOI: 10.1649/0010-065X-73.2.465, URL: http://dx.doi.org/10.1649/0010-065x-73.2.465
A91587910B40D219FD74222924471439.taxon	description	Specimens Examined. India: Kerala state: Tholpetty, 1 male, light attracted, 06. V. 2015, coll. S. V. Akhil; India: Tamil Nadu State: Ettimadai, 1 male, 4 female, light attracted, 08. IX. 2017, coll. S. V. Akhil; India: Tamil Nadu State: Rajapalayam, 1 male, hand picking, 15. X. 2017, coll. S. V. Akhil.	en	Akhil, S. V., Thomas, Sabu K. (2019): New Species of Bombardier Beetles of the Genus Styphlomerus Chaudoir, 1875 (Coleoptera: Carabidae) from Southern India, with a Key to the Indomalayan and Palearctic Species. The Coleopterists Bulletin 73 (2): 465-471, DOI: 10.1649/0010-065X-73.2.465, URL: http://dx.doi.org/10.1649/0010-065x-73.2.465
A91587910B40D219FD74222924471439.taxon	materials_examined	Type Material. Holotype labeled “ Holotype, Male, India: Tamil Nadu state: Ettimadai, Light attracted, 08. IX. 2017, coll. S. V. Akhil ”. Paratypes: Four females labeled “ Paratype, Female, India: Tamil Nadu state: Ettimadai, Light attracted, 08. IX. 2017, coll. S. V. Akhil ” / One male labeled “ Paratype, Male, India: Kerala state: Tholpetty, Light attracted, 06. V. 2015, coll. S. V. Akhil ” / One male labeled “ Paratype, Male, India: Tamil Nadu state: Rajapalayam, Hand picking, 15. X. 2017, coll. S. V. Akhil. Deposited in ZSIC.	en	Akhil, S. V., Thomas, Sabu K. (2019): New Species of Bombardier Beetles of the Genus Styphlomerus Chaudoir, 1875 (Coleoptera: Carabidae) from Southern India, with a Key to the Indomalayan and Palearctic Species. The Coleopterists Bulletin 73 (2): 465-471, DOI: 10.1649/0010-065X-73.2.465, URL: http://dx.doi.org/10.1649/0010-065x-73.2.465
A91587910B40D219FD74222924471439.taxon	description	Description. TLA = 7.5 – 7.7 mm. Color: Head, pronotum, labrum, mandible, scutellum, and region around scutellum reddish brown; scape and legs reddish yellow; pedicel darker reddish yellow, antennomeres 3 and 4 brownish black, other antennomeres dark brown; elytra black; genae, gula, prosternum, ventral prothorax, and mesosternum dark reddish yellow; metasternum, mesepisternum, metepisternum, and mesepimeron brownish yellow; abdomen brownish black. Head: Broader than long, widest between eyes, slightly narrower than pronotum, stout, punctate, region around eyes, sides of frons, and clypeus sparsely pubescent; eyes black, protruding; labral apical angles strongly rounded, form subrectangular, with 6 setae on apical margin; clypeus rectangular, wider than long, apical margin straight, with short setae on lateral margins, midregion glabrous; frontoclypeal suture impressed, distinct; frons with anterolateral setose depressions, midregion glabrous; mandibles sharp, broad; mentum broad, margins pronounced, with short, broad, pointed median tooth, lateral lobes triangular with lateral tooth-like projection, a single seta apicolaterally, remainder of mentum glabrous; palpi pubescent, apical palpomeres pointed, conical, penultimate palpomere with apex wide, base narrow; antennae short, not reaching middle of elytra, antennomeres slender. Pronotum: Punctate, pubescent; anterior margin slightly concave, lateral angles of anterior margin projecting slightly forward, posterior margin straight; anterior 3 / 4 of disc convex, posterior fourth narrowed, parallel towards base, hind angles quadrate without lateral projection; median furrow weak, shallow, not reaching anterior and posterior margins; lateral ridge wider, margin dark brown. Scutellum: Pointed, broad. Elytra: Subparallel, matt, faintly punctate, pubescent with golden brown setae; humerus prominent, rounded; surface flat, apex depressed; striations visible, weak, rounded, intervals narrow, setae along striations and interstices. Hind wings: Well-developed. Venter: Genae broad, smooth, with few short setae under eyes, remainder glabrous, genae with fine, superficial wrinkles; gula wide, divergent posteriorly, gular sutures brownish black; prothorax ventrally smooth, sparsely pubescent laterally towards neck region; prosternal process smooth, with 4 short setae apically in 2 rows; apical portion of abdomen pointed. Measurements. Holotype (male): TLA = 7.55 mm, TLB = 6.34 mm, PL = 1.05 mm, PW = 1.38 mm, EL = 3.97 mm, TW = 2.61 mm. Paratype (female): TLA = 7.62 – 8.03 mm, TLB = 6.48 – 6.58 mm, PL = 0.96 – 1.06 mm, PW = 1.28 – 1.37 mm, EL = 4.00 – 1.10 mm, TW = 2.50 – 2.74 mm. Geographical Distribution. INDIA: Tamil Nadu state: Rajapalayam, Ettimadai; Kerala state: Tholpetty. Collection Method. Collected under fluorescent light.	en	Akhil, S. V., Thomas, Sabu K. (2019): New Species of Bombardier Beetles of the Genus Styphlomerus Chaudoir, 1875 (Coleoptera: Carabidae) from Southern India, with a Key to the Indomalayan and Palearctic Species. The Coleopterists Bulletin 73 (2): 465-471, DOI: 10.1649/0010-065X-73.2.465, URL: http://dx.doi.org/10.1649/0010-065x-73.2.465
A91587910B40D219FD74222924471439.taxon	etymology	Etymology. Named for the elytral striations that differentiate it from the closely related S. ruficeps.	en	Akhil, S. V., Thomas, Sabu K. (2019): New Species of Bombardier Beetles of the Genus Styphlomerus Chaudoir, 1875 (Coleoptera: Carabidae) from Southern India, with a Key to the Indomalayan and Palearctic Species. The Coleopterists Bulletin 73 (2): 465-471, DOI: 10.1649/0010-065X-73.2.465, URL: http://dx.doi.org/10.1649/0010-065x-73.2.465
A91587910B40D219FD74222924471439.taxon	discussion	Remarks. Styphlomerus striatus is similar to S. ruficeps but differs by having weak, shallow striations on the elytra, a pointed scutellum, and the anterior margin of the pronotum concave.	en	Akhil, S. V., Thomas, Sabu K. (2019): New Species of Bombardier Beetles of the Genus Styphlomerus Chaudoir, 1875 (Coleoptera: Carabidae) from Southern India, with a Key to the Indomalayan and Palearctic Species. The Coleopterists Bulletin 73 (2): 465-471, DOI: 10.1649/0010-065X-73.2.465, URL: http://dx.doi.org/10.1649/0010-065x-73.2.465
A91587910B42D21AFD7726CD23BE140E.taxon	materials_examined	Type Material. Holotype, female, India: Rajasthan state: Alwar district: Naranimata env., 460 m (Hrdlička), in NMPC.	en	Akhil, S. V., Thomas, Sabu K. (2019): New Species of Bombardier Beetles of the Genus Styphlomerus Chaudoir, 1875 (Coleoptera: Carabidae) from Southern India, with a Key to the Indomalayan and Palearctic Species. The Coleopterists Bulletin 73 (2): 465-471, DOI: 10.1649/0010-065X-73.2.465, URL: http://dx.doi.org/10.1649/0010-065x-73.2.465
