identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
A93787D0184C181CFF00FA94FBEB509C.text	A93787D0184C181CFF00FA94FBEB509C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eriosema grearii Candido & Fort.	<div><p>Eriosema grearii Cândido &amp; Fort. -Perez, spec. nov. (Fig. 1)</p> <p>Morphologically similar to Eriosema heterophyllum and differing from it mainly by free stipules (vs. joined in E. heterophyllum), membranaceous leaflets (vs. papyraceous to cartaceous) with secretory-base trichomes (vs. absence of secretory-base trichomes), flowers distributed along the inflorescences axis (vs. congested at its apex) in inflorescences not opposite the leaves (vs. opposite).</p> <p>Type:— BRAZIL. Mato Grosso do Sul: Municipality of Ponta Porã. Cabeceira do APA, 23°53’S; 55°46’W, 16 October 1980, J. G. Guimarães 1196 (holotype HRB!).</p> <p>Subshrub ca. 0.25 m tall, stems erect, unbranched, striated, velutinous-tomentose. Stipules 5–9 × 2–4 mm, free, elliptic, lanceolate to ovate, apex acute, persistent, externally tomentose. Stipels absent. Leaves unifoliolate, petioles 8–10 mm long. Leaflets 3–7 × 2–4 cm, adaxial and abaxial surfaces pilose, with glands, elliptic to obovate, apex mucronate, base attenuate, margins entire. Racemes on the lower third of the stems, not opposite the leaves, 6–10 flowers, flowers distributed along the inflorescence axis, racemes (including peduncle) 7–15 cm long, villous, pedicels 3–5 mm long, flowers 6–13 mm long. Calyx 5–lobed, villous, margins entire, 5–veined; tube 3–4 mm long, two vexillar lobes 2–4 mm long, three carenal lobes 3–8 mm long; corolla yellow, the standard 9–12× 8–10 mm (excluding claw), orbicular, multi-veined, apex retuse or slightly emarginate, base auriculate, auricles 1 mm long, adaxial surface pubescent, abaxial surface villous; claw 2–4 mm; wing petals 6–10 × 2.5–5 mm (excluding claw), oblong to obovate, apex rounded, glabrous, slightly auriculate; claw 2 mm long; keel petals 6–10 × 3–5 mm (excluding claw), falcate to obovate, apex rounded, glabrous to pilose at the apex in both surface; claw 2–4 mm long; stamens 12–14 mm long, anthers uniform; ovary 5 mm long, villous; style 10 mm long, curved, stigma minute. Fruit not seen.</p> <p>Distribution and Habitat: —The taxon is known from a single collection in the municipality of Ponta Porã, in the Cerrado region of Mato Grosso do Sul, Brazil (Fig. 4). The area is a disturbed grassy cerrado, with red latosol and stony soils, and liable to frequent fires.</p> <p>stereomicroscope.</p> <p>Phenology: —The species was collected with flowers in October.</p> <p>Etymology: —The epithet honors Dr. John Grear, U.S. scientist who published a monograph for the American species of Eriosema and described several new species of the genus.</p> <p>Conservation Status: —Despite an intensive fieldwork and herbaria study, no other specimens of the new species but that in the type location was found, suggesting a narrow geographical distribution with high degree of vulnerability. However, for the assignment of a conservation status our data are insufficient, therefore, in accordance with the IUCN Red List criteria (IUCN 2014), the species must be considered Data Deficient (DD).</p> <p>Taxonomic Comments: — Eriosema grearii resembles E. heterophyllum Benth. but is easily differentiated by its free stipules (vs. joined), membranaceous leaflets (vs. papyraceous to cartaceous), the calyx divided into two parts, one of two lobes of the same size and the other of three carenal unequal lobes, with one much higher than the rest (Fig. 1E) (vs. calyx with all lobes of the same size), lax flowers (Fig. 1A) distributed along the inflorescences axis (vs. flowers concentrated at the apex), inflorescence not opposite the leaves (Fig. 1A) (vs. opposite), and leaflets with secretory-base trichomes (vs. absence of secretory-base trichomes).</p> <p>Secretory Structure Study: — Eriosema grearii exhibits a peculiar type of trichomes on the abaxial leaflet surface, referred to here as secretory-base trichomes and reported for the first time for the Phaseoleae (Fig. 2 A–D), as well as spherical capitate trichomes (Fig. 2 E–F), and vesicular glands (Fig. 2 G), with idioblasts in the mesophyll (Fig. 2 E, G).</p> <p>The vesicular glands have bulky heads and one-celled peduncles embedded in the epidermis depressions (Fig. 2 G). The spherical capitate trichomes have seven-celled heads with inconspicuous (Fig. 2 E) or conspicuous (Fig. 2 F) stalks not inserted in the epidermis depressions.</p> <p>The secretory structures, referred here as secretory-base trichomes and observed for the first time in Phaseoleae (Fig. 2 A–D, H), are multicellular, unbranched, uniseriate trichomes, comprised of an elongated, acuminate apical cell with tick and lignified walls, two intermediate cells with thick unlignified walls, and a large, hexagonal basal cell (Fig. 2 C). The basal cell shows a reduced cytoplasm and one large vacuole filled with a flocculent material, stained pink by toluidine blue (Fig. 2 D).</p> <p>In contrast, Eriosema heterophyllum shows bulky capitate trichomes with large heads and one-celled peduncles embedded in the epidermis depressions (Fig. 3 A, D) besides non-glandular multicellular unbranched uniseriate trichomes with one elongated acuminate apical cell and one basal cell (Fig. 3 A–C, E).</p> </div>	https://treatment.plazi.org/id/A93787D0184C181CFF00FA94FBEB509C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Cândido, Elisa Silva;Vargas, Wanderleia De;Vatanparast, Mohammad;Mansano, Vidal De Freitas;Machado, Silvia Rodrigues;Fortuna-Perez, Ana Paula	Cândido, Elisa Silva, Vargas, Wanderleia De, Vatanparast, Mohammad, Mansano, Vidal De Freitas, Machado, Silvia Rodrigues, Fortuna-Perez, Ana Paula (2016): A New Species of Eriosema (Leguminosae, Papilionoideae, Phaseoleae) from Mato Grosso do Sul, Brazil, with a Secretory Structure Novel to the Genus. Phytotaxa 263 (2): 122-130, DOI: 10.11646/phytotaxa.263.2.4, URL: http://dx.doi.org/10.11646/phytotaxa.263.2.4
A93787D0184B181FFF00FE00FA1752F7.text	A93787D0184B181FFF00FE00FA1752F7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eriosema (De Candolle 1825) Desvaux 1826	<div><p>Key to the species of Eriosema in Mato Grosso do Sul, Brazil</p> <p>1 Leaves unifoliolate............................................................................................................................................................................ 2</p> <p>- Leaves trifoliolate.............................................................................................................................................................................. 6</p> <p>2 Stipules free........................................................................................................................................................................................3</p> <p>- Stipules fully or partially fused..........................................................................................................................................................4</p> <p>3 Racemes 3–5 cm long, with 3–5 flowers, pubescent........................................................................................... E. obovatum Benth.</p> <p>- Racemes 7–15 cm long, with 6–10 flowers, villous............................................................................................................. E. grearii</p> <p>4 Stipules deciduous; leaflets conspicuously variable in size on the same plant............................ E. benthamianum Mart. ex Benth.</p> <p>- Stipules persistent; leaflets usually uniform in size on the same plant..............................................................................................5</p> <p>5 Racemes with 2–7 flowers, usually shorter than mature leaves....................................................... E. simplicifolium (DC.) G.Don</p> <p>- Racemes with 8–20 flowers, usually longer than mature leaves................................................................. E. heterophyllum Benth.</p> <p>6 Flowers 21–30 mm long........................................................................................................................................... E. riedelii Benth.</p> <p>- Flowers up to 18 mm long..................................................................................................................................................................7</p> <p>7 Leaflets usually less than five times as long as wide.........................................................................................................................8</p> <p>- Leaflets usually more than five times as long as wide.................................................................................................................... 10</p> <p>8 Stems glabrescent or cinereous-pubescent..................................................................................................... E. platycarpon Micheli</p> <p>- Stems yellowish, rufous- or ferruginous-pubescent...........................................................................................................................9</p> <p>9 Plants densely rufous-pilose or ferruginous-pubescent............................................................................... E. rufum (Kunth) G.Don</p> <p>- Plants yellow-pubescent...................................................................................................................................... E. campestre Benth.</p> <p>10 Leaflets elliptic to narrowly elliptic, the standard with a retuse apex................................................... E. tacuaremboense Arechav.</p> <p>- Leaflets linear, lanceolate or rarely narrowly elliptic; the standard with an apiculate apex............................................................11</p> <p>11 Stems whitish to cinereous-pubescent.................................................................................................................... E. strictum Benth.</p> <p>- Stems ferruginous or rufous-pubescent............................................................................................................................................12</p> <p>12 Racemes with 4–5 flowers....................................................................................................................... E. crinitum (Kunth) G.Don</p> <p>- Racemes with more than 8 flowers.................................................................................................................. E. longifolium Benth.</p></div> 	https://treatment.plazi.org/id/A93787D0184B181FFF00FE00FA1752F7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Cândido, Elisa Silva;Vargas, Wanderleia De;Vatanparast, Mohammad;Mansano, Vidal De Freitas;Machado, Silvia Rodrigues;Fortuna-Perez, Ana Paula	Cândido, Elisa Silva, Vargas, Wanderleia De, Vatanparast, Mohammad, Mansano, Vidal De Freitas, Machado, Silvia Rodrigues, Fortuna-Perez, Ana Paula (2016): A New Species of Eriosema (Leguminosae, Papilionoideae, Phaseoleae) from Mato Grosso do Sul, Brazil, with a Secretory Structure Novel to the Genus. Phytotaxa 263 (2): 122-130, DOI: 10.11646/phytotaxa.263.2.4, URL: http://dx.doi.org/10.11646/phytotaxa.263.2.4
