taxonID	type	description	language	source
A9379460142A540C49D46491FA1FF80E.taxon	description	(Figs 1 – 13)	en	Mai, Zuqi, Jiang, Zhuoyin, Hu, Jian, Hendrich, Lars, Jia, Fenglong (2022): A new species of Clypeodytes Régimbart, 1894 from China (Coleoptera, Dytiscidae: Bidessini). Zootaxa 5124 (1): 50-60, DOI: 10.11646/zootaxa.5124.1.2
A9379460142A540C49D46491FA1FF80E.taxon	materials_examined	Type locality: Xintanghe Village (25 ° 9 ′ 35.7 ″ N 98 ° 14 ′ 42 ″ E), 1.875 m, Tengchong County, Baoshan Prefecture, Yunnan, China. Type material. HOLOTYPE: Male (SYSU), “ China, Yunnan, Baoshan Prefecture, Tengchong County, Xintanghe Village (ŕä 河村), 25 ° 9 ′ 35.7 ″ N 98 ° 14 ′ 42 ″ E, 1875 m, swamp with extensive vegetation, 12. V. 2021, Zuqi Mai, Zhuoyin Jiang & Zhenming Yang leg. ” PARATYPES: 39 exs (SYSU), same data as the holotype; 1 male and 8 exs (NMPC), “ China, Yunnan province, GUDONG env., 8. VI. 2007, YUNFENG SHAN Mt., 25 ° 22.7 ′ N, 098 ° 25.4 ′ E, 1825 m, J. Hájek & J. Růžička leg. ”; 1 female (ZSM), “ China, Yunnan, Baoshan Prefecture, mountain range 22 km S Tengchong County, 24 ° 49 ′ 29 ″ N 98 ° 29 ′ 27 ″ E, 1759 m, secondary forest, litter dead wood sifted, 2.6.2007, A. Pütz leg. ”	en	Mai, Zuqi, Jiang, Zhuoyin, Hu, Jian, Hendrich, Lars, Jia, Fenglong (2022): A new species of Clypeodytes Régimbart, 1894 from China (Coleoptera, Dytiscidae: Bidessini). Zootaxa 5124 (1): 50-60, DOI: 10.11646/zootaxa.5124.1.2
A9379460142A540C49D46491FA1FF80E.taxon	description	Description of holotype. Total length 1.75 mm; maximum width 1.1 mm. Body short and robust, attenuated posteriad. Body surface with dense coarse punctures. Coloration. Dorsal yellow brown to dark brown, with black patterns on basal pronotum and elytra (Fig. 1). Ventral surface to large extent reddish brown (Fig. 3). Legs, antennae, maxillary palps and labial palps yellowish (Figs 1 – 4). Head. Anterior margin of clypeus with thick bead (Fig. 9). Frons with sparse fine setiferous punctures. Eyes large, posterior margin of eye attached to occipital line. Behind occipital line microreticulate (Fig. 9). Prothorax. Pronotum with setiferous punctures sparser than those on frons, becoming coarser posteriorly; lateral pronotal bead present; basal striae distinct (Fig. 8). Prosternal process lanceolate and slightly tectiform and, apex broadly rounded. Meso- and metathorax. Elytron without basal stria and without sutural line, with prominent longitudinal keel sublaterally (Fig. 10) and a very weak keel more close to epipleuron; surface of elytron with dense coarse punctures, a few punctures with setae apically. Epipleuron in anterior third broad, becoming narrower posteriad; anteriorly with humeral cavity, limited posteriorly by an oblique subhumeral carina (Fig. 11). Hind wings well developed. Anepisternum, metaventrite and metacoxa with sparse setiferous punctures; metaventrite with a line of fine punctures left and right a mid-line (= discrimen; see Fig. 12). Metacoxal lines short, more or less parallel; distance between lines not much smaller than length of lines. Lobes of metacoxal processes not prominent. Legs. Metatibia elongate, distally gradually expanded (Fig. 8). Abdomen. Ventrites 1 – 2 with coarse punctures similar to those on metacoxa. Ventrites 3 – 6 with finer punctation, near apex of last abdominal ventrite denser. Male genitalia (Figs 5 – 7). Median lobe of aedeagus in lateral view as in Fig. 6; apical portion straight and slender, nearly acicular. Paramere (Fig. 7) two-segmented and with hook-shaped apex. Variation. Total length 1.7 – 2.0 mm; maximum width 1.0 – 1.15 mm; several paratypes with paler coloration on pronotum and elytra. If so, each elytron with two black spots separated from the dark pattern on upper side (Figs 1 – 2).	en	Mai, Zuqi, Jiang, Zhuoyin, Hu, Jian, Hendrich, Lars, Jia, Fenglong (2022): A new species of Clypeodytes Régimbart, 1894 from China (Coleoptera, Dytiscidae: Bidessini). Zootaxa 5124 (1): 50-60, DOI: 10.11646/zootaxa.5124.1.2
A9379460142A540C49D46491FA1FF80E.taxon	etymology	Etymology. This specific name of the new species is the Latin adjective limpidus (= clear), referring to the species habitat, a swamp with limpid fresh water.	en	Mai, Zuqi, Jiang, Zhuoyin, Hu, Jian, Hendrich, Lars, Jia, Fenglong (2022): A new species of Clypeodytes Régimbart, 1894 from China (Coleoptera, Dytiscidae: Bidessini). Zootaxa 5124 (1): 50-60, DOI: 10.11646/zootaxa.5124.1.2
A9379460142A540C49D46491FA1FF80E.taxon	distribution	Distribution. Only known from three localities in western Yunnan close to the border to Myanmar. Baoshan is a prefecture-level city in China and the second-largest metropolitan area in western Yunnan province. It is located between the border of Myanmar and the Lanchang river (Mekong river).	en	Mai, Zuqi, Jiang, Zhuoyin, Hu, Jian, Hendrich, Lars, Jia, Fenglong (2022): A new species of Clypeodytes Régimbart, 1894 from China (Coleoptera, Dytiscidae: Bidessini). Zootaxa 5124 (1): 50-60, DOI: 10.11646/zootaxa.5124.1.2
A9379460142A540C49D46491FA1FF80E.taxon	diagnosis	Diagnosis. The new species can be easily recognized from the other known Chinese species C. bufo by its smaller size (C. limpidus sp. nov. total length: 1.7 – 2.0 mm, C. bufo total length: 2.0 – 2.3 mm), the dorsal coloration with black patterns on the basal part of the pronotum and on the elytra, the narrower shape of the pronotum, the more roundish habitus, and the shape of the median lobe (Figs 5 – 7, 15 – 17). Habitat. At the type locality the new species was collected along the margins of a swamp with dense vegetation, situated in a valley (Fig. 13 A). Here C. limpidus sp. nov. co-occurs with several other Dytiscidae: Hydroglyphus sp., Hydaticus major Régimbart, 1899, Hydrovatus sp., Laccophilus sp. and Rhantus sp. Specimens from Yunfeng Shan were found in a deep, densely vegetated swampy pool (Fig. 13 B). At this locality the new species was associated with Agabus amoenus sinuaticollis Régimbart, 1899, Hydaticus bipunctatus Wehncke, 1876, and H. major Régimbart, 1899. The single specimen collected by A. Pütz in 2007 was obtained by sifting leaf litter and dead rotten wood.	en	Mai, Zuqi, Jiang, Zhuoyin, Hu, Jian, Hendrich, Lars, Jia, Fenglong (2022): A new species of Clypeodytes Régimbart, 1894 from China (Coleoptera, Dytiscidae: Bidessini). Zootaxa 5124 (1): 50-60, DOI: 10.11646/zootaxa.5124.1.2
A9379460142D540649D467EDFC8AFBD0.taxon	description	(Figs 14 – 21)	en	Mai, Zuqi, Jiang, Zhuoyin, Hu, Jian, Hendrich, Lars, Jia, Fenglong (2022): A new species of Clypeodytes Régimbart, 1894 from China (Coleoptera, Dytiscidae: Bidessini). Zootaxa 5124 (1): 50-60, DOI: 10.11646/zootaxa.5124.1.2
A9379460142D540649D467EDFC8AFBD0.taxon	materials_examined	Material examined (China and Laos). China, Guangdong: 3 exs (SYSU), “ Xingyi District (信fi), 1960. VI. 7, Zhe-long Pu leg. ”; 2 exs (SYSU), “ Gaozhou District (š 州), Dongzhen (东đ), 1960. VI. 7, Zhe-long Pu leg. ”; Guangxi: 1 ex. (SYSU), “ Shangsi County (上思), 1997. VI. 24, Zhi-he Huang leg. ”; Guizhou: 1 ex. (SYSU): “ Guizhou Dushan (Ė 州â山) ” / “ 1940.10.10 ” // “ Clypeodytes bufo (Sharp) ” / “ Det. Hong Zeng ”; Hainan: 1 ex. (SYSU): “ Guangdong Hainan (广东海南) ” / “ 1957.12.16 ” / “ Cui-ying Li ” // “ Clypeodytes bufo (Sharp) ” / “ Det. Hong Zeng ” / “ 1988. XI ”. Laos: 3 exs (ZSM), “ Laos centr. Kham Mouan Prov. Nakai village env. ca. 70 km NNE Muang Khammouan, 500 – 600 m 17 ° 43 ’ N 105 ° 09 ’ E, M. Strba leg. ”; 1 male (NMPC), “ LAOS, Vientiane prov., LAO PAKO env., 200 m, 55 km NE Vientiane, 1. - 5. IV. 2004, J. Bezděk leg. ”.	en	Mai, Zuqi, Jiang, Zhuoyin, Hu, Jian, Hendrich, Lars, Jia, Fenglong (2022): A new species of Clypeodytes Régimbart, 1894 from China (Coleoptera, Dytiscidae: Bidessini). Zootaxa 5124 (1): 50-60, DOI: 10.11646/zootaxa.5124.1.2
A9379460142D540649D467EDFC8AFBD0.taxon	discussion	Remarks. Clypeodytes bufo is widespread in the Oriental region, occurring as far as southern China in the Palearctic. Since the species was described from Sri Lanka and the authors could not study the types or freshly collected specimens from that island, a detailed re-description of the species is not possible. Based on the literature available to us, however, we have come to the conclusion that the specimens of the second Clypeodytes species from China should be attributed to C. bufo. However, it must be mentioned here that no lateral keel on elytron at all could be found in specimens of C. bufo from Laos, so that these specimens differ significantly morphologically from the Chinese ones. Only further morphological studies, including molecular data, from different countries could finally clarify the systematic position of C. bufo, and whether this taxon in its entire distribution area is only one species or must be treated as a species complex.	en	Mai, Zuqi, Jiang, Zhuoyin, Hu, Jian, Hendrich, Lars, Jia, Fenglong (2022): A new species of Clypeodytes Régimbart, 1894 from China (Coleoptera, Dytiscidae: Bidessini). Zootaxa 5124 (1): 50-60, DOI: 10.11646/zootaxa.5124.1.2
A9379460142D540649D467EDFC8AFBD0.taxon	diagnosis	Diagnosis. The present diagnosis is based on specimens from China and Laos: Total length 2.0 – 2.3 mm. Body robust and comparably elongate, with broad pronotum. Lateral outline distinctly discontinuous between pronotum and elytra. Anterior lateral margins of pronotum conspicuously rounded, nearly parallel-sided laterally. Body dark brown throughout, without any patterns (Fig. 14). Body surface with dense coarse punctures. Anterior margin of clypeus with thick bead; posterior margin of eye attached with occipital line. Basal pronotal striae distinct (Fig. 18). Elytron without sutural line but with basal stria (Fig. 18). Each elytron with a weak keel laterally (Fig. 19). Epipleuron with a weak subhumeral, almost transverse carina (Fig. 20). Metaventrite medially with a pair of coarse serial punctures (Fig. 21). Median lobe of aedeagus in lateral and ventral view as in Figs 15 – 16; paramere twosegmented, with pointed hook-shaped apex (Fig. 17). Clypeodytes bufo can be easily recognized from C. limpidus sp. nov. by its larger size (C. bufo total length = 2.0 – 2.3 mm, C. limpidus sp. nov. total length = 1.7 – 2.0 mm), the throughout reddish brown dorsal coloration, the very broad pronotum, the more elongate habitus, and the shape of the median lobe (Figs 15 – 17, 5 – 7).	en	Mai, Zuqi, Jiang, Zhuoyin, Hu, Jian, Hendrich, Lars, Jia, Fenglong (2022): A new species of Clypeodytes Régimbart, 1894 from China (Coleoptera, Dytiscidae: Bidessini). Zootaxa 5124 (1): 50-60, DOI: 10.11646/zootaxa.5124.1.2
A9379460142D540649D467EDFC8AFBD0.taxon	distribution	Distribution. China (Guangdong, Guangxi, Guizhou, Hainan, Hubei, Hunan), India, Nepal,? Bhutan, Bangladesh, Sri Lanka, Myanmar, Vietnam (Ghosh & Nilsson 2012; Nilsson & Hájek 2021; Shaverdo et al. 2021; Vazirani 1969, 1971) and Laos (first record). Habitat. The collecting circumstances of the Chinese specimens are unknown. The few specimens from Laos were collected at light. In India it was collected in exposed and partly shaded lentic sites of mainly alkaline ponds and smaller lakes (Senthil Kumar et al. 2017, 2018). At Rambha, near the brackish Chilika Lake in Odisha (India), the species was found in fresh water (Ghosh et al. 2016).	en	Mai, Zuqi, Jiang, Zhuoyin, Hu, Jian, Hendrich, Lars, Jia, Fenglong (2022): A new species of Clypeodytes Régimbart, 1894 from China (Coleoptera, Dytiscidae: Bidessini). Zootaxa 5124 (1): 50-60, DOI: 10.11646/zootaxa.5124.1.2
