taxonID	type	description	language	source
38CD0A572CF3516EAE494A60B0331D0B.taxon	materials_examined	Type. Chile. Region I Tarapaca and Region II Antofagasta: " Peruvia meridionalis: Cobija, Iquiqui et Arica ", H. Cuming 953 (lectotype, selected by Plos and Sancho (2013: 338): K [K 000486684, photo!]; isolectotypes E [E 00322766, photo!], GH [GH 00010778!], K [K 000486685, photo!], P [P 02673192, photo!]); remaining syntypes: Gaudichaud s. n. (B [probably destroyed, could not be found], F [F 1012247, photo!]). Ophryosporus anomalus has been cited for Peru (King and Robinson 1972 b; Brako and Zarucchi 1993), likely due to the type collection label (" Peruvia meridionalis ... "). However, the localities mentioned there (" ... Cobija, Iquiqui et Arica ") are now situated in Chile, and all other reports originate from the coastal zone around Tocopilla (Johnston 1932; Jaffuel 1936) and therefore Luebert et al. (2007) considered it a Chilean endemic. We hereby extend the distribution of the species to include the populations from the coastal area between Rio Loa and Iquique, previously referred to as O. floribundus (Munoz-Schick et al. 2001; Pinto and Luebert 2009). See discussion below under the latter species.	en	Merklinger, Felix F., Luebert, Federico (2020): Notes on the genus Ophryosporus (Asteraceae, Eupatorieae) in Chile. PhytoKeys 161: 61-77, DOI: http://dx.doi.org/10.3897/phytokeys.161.53736, URL: http://dx.doi.org/10.3897/phytokeys.161.53736
38CD0A572CF3516EAE494A60B0331D0B.taxon	materials_examined	Specimens examined. Chile. Region I Tarapaca: Prov. Iquique, Alto Punta Gruesa, 20 ° 22 ' S, 70 ° 09 ' W, 14 Dec 1997, R. Pinto s. n. (SGO 142948); Alto Punta Patache, 20 ° 49 ' S, 70 ° 09 ' W, 6 Dec 1997, R. Pinto s. n. (SGO 142949); Alto Punta Patache, 22 Jan 2000, R. Pinto s. n. (SGO [photo]); Alto Punta Lobos, 21 ° 02 ' S, 70 ° 09 ' W, 14 Jan 1998, R. Pinto s. n. (SGO 142950); Alto Chipana, 21 ° 16 ' S, 70 ° 03 ' W, 15 Oct 1997, W. Sielfeld 7 (SGO 143038); Alto Chipana, 21.304528 S, 70.03204 W, 990 m, 21 Oct 2016, F. Luebert, A. Stoll & T. Boehnert 3427 A (BONN, ULS); Alto Chipana, 21.292633 S, 70.042234 W, 840 m, 1 Oct 2019, F. Luebert, F. F. Merklinger & J. Ruhm 4102 (BONN, EIF, K). Region II Antofagasta: Prov. Tocopilla, Tocopilla, 27 Oct 1930, F. Jaffuel 1026 (GH); Tocopilla, Cerro Rosario, 1 Nov 1941, M. R. Espinosa s. n. (SGO 143254); Cobija, s. a., C. Gaudichaud s. n. (F 1012247); Cobija, quebrada Aguada Canas, 4 Apr 1949, W. Biese 3088 (SGO 096693).	en	Merklinger, Felix F., Luebert, Federico (2020): Notes on the genus Ophryosporus (Asteraceae, Eupatorieae) in Chile. PhytoKeys 161: 61-77, DOI: http://dx.doi.org/10.3897/phytokeys.161.53736, URL: http://dx.doi.org/10.3897/phytokeys.161.53736
E63E026F860E5C3AAFE1F4E9F7CAB5B9.taxon	materials_examined	Type. Peru. " Perou cordilliere, 1834, T. P. X. Haenke s. n. (holotype: G-DC [G 00130596, photo!]; isotype: P [P 00742191, photo!]). This taxon is cited for Chile in Munoz-Schick et al. (2001), Pinto and Luebert (2009), Zuloaga et al. (2008) and Rodriguez et al. (2018) based on four specimens. Three of them were collected by R. Pinto in the late 1990 s at three coastal localities in northern Chile, Alto Chipana, Punta Lobos and Punta Gruesa (Pinto, s. n., SGO 142948, SGO 142949 and SGO 142950). Recent field work at the coastal localities has not resulted in any collections that match the type of O. floribundus. Rather we found plants that we identified as O. anomalus, another species that has been reported for this area (Johnston 1932) and has been only sporadically collected since. The type of O. floribundus has opposite, solitary leaves and long internodes of about 3 - 4 cm in length. The type of O. anomalus has crowded leaves that are borne in fascicles and with very short internodes. The leaves of the Chilean coastal specimens assigned to O. floribundus vary in size and shape, some corresponding well to the type of O. anomalus being narrowly oblanceolate and with entire margins and an obtuse apex, others becoming more broadly lanceolate to triangular with dentate margins and an acute apex thus remotely resembling O. floribundus but actually more similar to those of O. triangularis. On younger shoots the leaves appear more or less opposite but generally they are borne in fascicles. The cypselae of the plants on both sheets of the type specimen of O. anomalus (Cuming 953, K) bear quite visible, slightly spreading squamellate pappus setae, while no pappus is recorded for O. floribundus. We therefore conclude that O. floribundus is restricted to Peru from the area of Lima, while the coastal plants in northern Chile should be referred to O. anomalus.	en	Merklinger, Felix F., Luebert, Federico (2020): Notes on the genus Ophryosporus (Asteraceae, Eupatorieae) in Chile. PhytoKeys 161: 61-77, DOI: http://dx.doi.org/10.3897/phytokeys.161.53736, URL: http://dx.doi.org/10.3897/phytokeys.161.53736
E63E026F860E5C3AAFE1F4E9F7CAB5B9.taxon	materials_examined	Specimens examined. Peru. Dept. Amazonas: Purruchuca [sic], s. a., Mathews, A. 1015 (G). Dept. Lima: Prov. Canta, road from Canta to Lima below San Jose turnoff towards Lima, 11.49383 S, 76.65187 W, 2322 m, 1 Mar 2018, M. Weigend & K. A. Pena Ramos 9719 (BONN); road from Canta to Lima, road down from Canta to turn off San Jose, 11.49383 S, 76.65187 W, 2322 m, 1 Mar 2018, M. Weigend & K. A. Pena Ramos 9722 (BONN). Prov. Huarochiri, between Matucana and Tambo, 26 Dec 1901, Weberbauer, A. 115 (G); Matucana, c. 2400 m, 25 May 1940, E. Asplund 11072 (S); Matucana, c. 2600 m, 25 May 1940, E. Asplund 11079 (S).	en	Merklinger, Felix F., Luebert, Federico (2020): Notes on the genus Ophryosporus (Asteraceae, Eupatorieae) in Chile. PhytoKeys 161: 61-77, DOI: http://dx.doi.org/10.3897/phytokeys.161.53736, URL: http://dx.doi.org/10.3897/phytokeys.161.53736
76F68AB395F5500EAEAE4D5FD1DEFA9C.taxon	materials_examined	Type. Peru. Dept. Puno: " Perou, sur les montagnes, aux environs de la ville de d'Azangaro ", W. Lechler 1751 (lectotype, selected by Plos and Sancho (2013: 336): P [P 00742426, photo!]; isolectotypes: GOET [GOET 001506, photo!], K [K 000542525, photo!], P [P 00742425, P 00742428, photo!], W [W 0018472, photo!]). This species is confirmed for Chile and has been collected by the authors near the village of Chusmiza, at the same locality as Zoellner 2997 (L, LP) and Gardner & Knees 6534 (E, SGO). The pappus setae of this species are formed by conspicuous yellowish-white setae of c. 4 mm in length. At this locality, O. heptanthus grows in local sympatry with O. pinifolius. However, the pappus of the latter is formed by minute squamellae and the two taxa are thus readily distinguishable. Plos (2012) cited specimen Zoellner 2997 twice, once for O. heptanthus (LP) and a second time for O. hoppii (CONC). It is possible, that having collected both specimens at the same locality where they occur sympatrically, Zoellner mistook them for a single species and only later at the two herbaria they were identified as belonging to two distinct taxa. We could not have access to the latter material at CONC, but we assume that it corresponds either to O. heptanthus or O. pinifolius.	en	Merklinger, Felix F., Luebert, Federico (2020): Notes on the genus Ophryosporus (Asteraceae, Eupatorieae) in Chile. PhytoKeys 161: 61-77, DOI: http://dx.doi.org/10.3897/phytokeys.161.53736, URL: http://dx.doi.org/10.3897/phytokeys.161.53736
76F68AB395F5500EAEAE4D5FD1DEFA9C.taxon	materials_examined	Specimens examined. Bolivia. La Paz: Prov. Bautista Saavedra, Chajaya, a few km from Charazani, 15 ° 13 ' S, 69 ° 01 ' W, 3500 m, 30 Mar 1985, J. C. Solomon 13294 (U [U 1145280]). Prov. Omasuyos, Viciniis Ochachache [Achacachi], 4000 m, Jan-Apr 1859, G. Mandon 260 (S). Prov. Murillo, 4 km up the Rio Achumani from Calacoto (La Paz), 16 ° 30 ' S, 68 ° 02 ' W, 3600 m, 11 Apr 1986, J. C. Solomon 15271 (U [U 1145281]); 1 km NW of Ovejuyo, 16 ° 32 ' S, 68 ° 03 ' W, 3700 - 3900 m, 2 Apr 1982, J. C. Solomon 7453 (U [U 1145279]). Chile. Region I Tarapaca: Prov. Tamarugal, Chusmiza, 3200 m, 10 Jan 1969, O. Zoellner 2997 (L 125727); quebrada de Chusmiza, 19 ° 41 ' 4.9 " S, 69 ° 11 ' 01.9 " W, 3350 m, 18 Feb 2003, M. F. Gardner & S. G. Knees 6534 (E, SGO); Chusmiza, 19 ° 40 ' 48.2 " S, 69 ° 10 ' 49.5 " W, 3380 m, 27 Apr 2008, M. Munoz & A. Moreira 4940 (SGO 157269); at the entrance of the Andean village Chusmiza, 19.67880 S, 69.17956 W, 3392 m, 2 Oct 2019, F. F. Merklinger, F. Luebert & J. Ruhm 2019 - 20 (BONN, EIF, K). Peru. Dept. Ayacucho: Prov. Lucanas, a few km from Puente Toro Muerte, 14 ° 42 ' 55.1 " S, 74 ° 32 ' 44.7 " W, 3589 m, 21 Mar 2019, M. Weigend 9841 / 19 - 32 (BONN). Dept. Cusco: Prov. Cusco, alrededore Cusco, 17 May 1958, A. L. Cabrera & H. A. Fabris 13536 (S); Rio Blanco, 1500 ft [450 m], 8 - 19 May 1922, J. F. Macbride & W. Featherstone 718 (S). Dept. Puno: Prov. Puno, Checayani, NE of Azangaro, 3980 m, 28 Mar 1957, H. Ellenberg 461 (U [U 1145288]); Huerta N of Puno, 3840 m, 22 Mar 1957, H. Ellenberg 238 A (U [U 1145284]); Huerta N of Puno, 4100 m, 22 Mar 1957, H. Ellenberg 292 (U [U 1145286]); Huerta N of Puno, 3840 m, 22 Mar 1957, H. Ellenberg 238 (U [U 1145285]). Prov. Lampa, Pucara, 3900 m, 22 Aug 1957, H. Ellenberg 2753 A (U [U 1145287]). Dept. Moquegua: Prov. de Mariscal Nieto, Carumas, 3200 m, 21 Feb- 6 Mar 1925, A. Weberbauer 7333 (S); Prov. General Sanchez Cerro, Puquina, outside Puquina towards Arequipa, 16 ° 36 ' 39.4 " S, 70 ° 11 ' 30.8 " W, 3174 m, 29 Mar 2019, M. Weigend 9994 / 19 - 182 (BONN).	en	Merklinger, Felix F., Luebert, Federico (2020): Notes on the genus Ophryosporus (Asteraceae, Eupatorieae) in Chile. PhytoKeys 161: 61-77, DOI: http://dx.doi.org/10.3897/phytokeys.161.53736, URL: http://dx.doi.org/10.3897/phytokeys.161.53736
5A65173D56905FB9A921A240638B0A25.taxon	materials_examined	Type. Peru. Dept. Arequipa: Jul 1925, W. Hopp 28 (holotype: B [probably destroyed], F neg. 14723!; lectotype, designated here: GH [GH 00013302]). Epitype (designated here): Peru. Dept. Arequipa. Lomas of Mollendo, c. 4 km N of Islay, 230 m, 20 Nov 1983, M. O. Dillon & D. Dillon 3926 (USM [74666]; isoepitypes: BONN!, F!, US [3026292, photo!]). This taxon was originally described as a new genus and species, Trychinolepis hoppii (Robinson 1928: 6), because of its irregularly lobed, squamellate pappus, which resembled that of the West Indian genus Phania, even though a habitual resemblance to the genus Ophryosporus was stated by the author (Robinson 1928). Subsequent analyses led to this monotypic genus to be allocated to Ophryosporus, because the pappus remained the only difference with other species of Ophryosporus (King and Robinson 1972 a). Specimens assigned to O. hoppii in Peru have a squamellate pappus, where the setae are fused at the base and are rather conspicuous. Cypselae with this type of pappus are visible on the type specimen and can even be seen on online images. However, due to the loss of the holotype in B and the very fragmentary remnants of the isotype at GH, we have decided to epitypify the specimen collected by M. O. Dillon and D. Dillon in 1983, as it corresponds to the protologue of T. hoppii and has been collected in the same region. This specimen has all necessary characters suitable for identification and is a good reference for potential future work. Specimens which we could positively identify as Ophryosporus hoppii were all collected in Peru. Records of O. hoppii for Chile appear to be miss-identifications and belong to O. pinifolius, e. g. Schlegel 4879 & 5092 (CONC, F). The pappus of O. pinifolius on the contrary, although squamellate, are much more inconspicuous (Fig. 2). Our own observations and extensive sampling in northern Chile failed to positively identify a plant that clearly fits into the species concept of O. hoppii. For example, a specimen from Chusmiza, N of Iquique (O. Zoellner 2997, U) is cited twice in Plos (2012), once for O. hoppii and a second time for O. heptanthus. The locality at Chusmiza was visited by the authors, and two species, O. pinifolius and O. heptanthus, were confirmed. These two taxa grow in local sympatry here, yet are distinguished from one another by their pappus, which in O. pinifolius consists of minute squamellae and which in O. heptanthus is formed by setae, up to 4 mm long. It is possible that the two taxa were collected by Zoellner as one species, only for later herbarium identification to recognize the two different taxa in the two herbarium vouchers. A further herbarium record from Alto Patache in Chile (H. Larrain 98200, CONC) could not be found in CONC.	en	Merklinger, Felix F., Luebert, Federico (2020): Notes on the genus Ophryosporus (Asteraceae, Eupatorieae) in Chile. PhytoKeys 161: 61-77, DOI: http://dx.doi.org/10.3897/phytokeys.161.53736, URL: http://dx.doi.org/10.3897/phytokeys.161.53736
5A65173D56905FB9A921A240638B0A25.taxon	materials_examined	Specimens examined. Peru. Dept. Arequipa: Prov. Islay, Quebrada Guerreros, 456 m, 6 Apr 1998, FLSP 2411 (HUSA, US); Ocona, 5 Feb 1969, J. Soukup 6426 / 6216 (US, USM). Prov. Castilla, Chuquibamba towards Aplao 15 ° 51 ' 52.3 " S, 72 ° 36 ' 56.4 " W, 2231 m, 24 Mar 2019, M. Weigend 9862 / 19 - 52 (BONN, USM).	en	Merklinger, Felix F., Luebert, Federico (2020): Notes on the genus Ophryosporus (Asteraceae, Eupatorieae) in Chile. PhytoKeys 161: 61-77, DOI: http://dx.doi.org/10.3897/phytokeys.161.53736, URL: http://dx.doi.org/10.3897/phytokeys.161.53736
572AD3B86FC25555B7A46E3257DEE485.taxon	materials_examined	Type. Chile. Region II, Antofagasta: Prov. Antofagasta, dept. Taltal, Aguada del Panul, I. M. Johnston 5424 (holotype: GH [GH 00010781, photo!]; isotypes: S [S-R- 3810!, S 10 - 19704!], SGO [SGO 59043!]). This peculiar species is known only from three localities north of the town of Paposo (Johnston 1929). It is distributed very narrowly within the range of O. triangularis and is sympatric with this latter species. Morphologically, O. johnstonii is very similar to O. triangularis.	en	Merklinger, Felix F., Luebert, Federico (2020): Notes on the genus Ophryosporus (Asteraceae, Eupatorieae) in Chile. PhytoKeys 161: 61-77, DOI: http://dx.doi.org/10.3897/phytokeys.161.53736, URL: http://dx.doi.org/10.3897/phytokeys.161.53736
572AD3B86FC25555B7A46E3257DEE485.taxon	materials_examined	Specimens examined. Chile. Region II Antofagasta: Prov. Antofagasta, vicinity of Miguel Diaz, directly N of quebrada Iscuna, c. 55 km N of Paposo, 24 ° 33 ' S, 70 ° 33 ' W, 100 - 300 m, 15 Dec 1987, M. O. Dillon & S. Teillier 5292 (BONN); vicinity of Aguada de Miguel Diaz, 24 ° 35 ' S, 1 - 4 Dec 1925, I. M. Johnston 5310 (SGO 059042); vicinity of Aguada Cardon, 24 ° 45 ' S, 30 Nov 1925, I. M. Johnston 5259 (S); rocky slopes of Aguada Cardon, 24.74173 S, 70.54385 W, 210 m, 15 Oct 2016, F. Luebert, A. Stoll & T. Boehnert 3384 (BONN, ULS); Aguada Cardon, 24.741717 S, 70.542687 W, 210 m, 5 Oct 2017, F. Luebert, T. Boehnert & F. F. Merklinger 3950 (BONN, ULS).	en	Merklinger, Felix F., Luebert, Federico (2020): Notes on the genus Ophryosporus (Asteraceae, Eupatorieae) in Chile. PhytoKeys 161: 61-77, DOI: http://dx.doi.org/10.3897/phytokeys.161.53736, URL: http://dx.doi.org/10.3897/phytokeys.161.53736
5B4FB92DF4CE5E63919C92E700DB8779.taxon	materials_examined	Type. Chile. Region V Valparaiso: T. C. Bridges 52 (lectotype, selected by Plos and Sancho (2013: 336): E [E 00249901, photo!]; isolectotypes: K [K 486667, photo!], W [W 0018468, photo!]); remaining syntypes: Valparaiso, H. Cuming 337, K [K 486668, photo!], E [E 249900, E 249902, photo!]; W [W 0018467, photo!]. Valparaiso, J. Gillies (not seen). This is the southernmost species of Ophryosporus in Chile, distributed from the region Metropolitana de Santiago northward to the region Atacama. This species is not strictly limited to the coast but also occurs further inland, for example in the Cuesta Las Chilcas or Andacollo. Ophryosporus paradoxus is a very distinct species that can be differentiated from O. triangularis by its larger, lanceolate and rather papery leaves with strongly lobed margins, as opposed to the small, triangular, slightly fleshy leaves with revolute margins in O. triangularis. The secondary inflorescences are thyrsoid, emerge terminally and produce florets with white corollas and a pappus of white setae up to c. 3 mm long.	en	Merklinger, Felix F., Luebert, Federico (2020): Notes on the genus Ophryosporus (Asteraceae, Eupatorieae) in Chile. PhytoKeys 161: 61-77, DOI: http://dx.doi.org/10.3897/phytokeys.161.53736, URL: http://dx.doi.org/10.3897/phytokeys.161.53736
5B4FB92DF4CE5E63919C92E700DB8779.taxon	materials_examined	Specimens examined. Chile. Region III Atacama: Prov. Copiapo, Jorquera-valley, 12 Jan 1970, O. Zoellner 4682 (L 3661656). Prov. Huasco, below El Chivato, 28 ° 54 ' S, 70 ° 04 ' W, 1800 m, 4 Jan 1926, I. M. Johnston 5870 (S); Resguardo, 28 ° 58 ' S, 70 ° 10 ' W, 1530 m, 4 Jan 1926, I. M. Johnston 5863 (S); valley San Felix, 1180 m, 16 Dec 1941, E. Pisano V. & R. Bravo F. 1089 (SGO). Region IV Coquimbo: Prov. Elqui, valley of Rio Turbio between Rivadavia and Guanta, 900 m, 18 - 19 Jan 1926, I. M. Johnston 6271 (S); near Guanaqueros, 24 Jul 1973, O. Zoellner 6827 (L 3661655); road from Marquesa to Vinita Baja, 29.954529 S, 70.964978 W, 340 m, 26 Sep 2017, F. Luebert, T. Boehnert & F. F. Merklinger 3822 (BONN, ULS); road from Marquesa to Vinita Baja, 29.873778 S, 70.860812 W, 700 m, 26 Sep 2017, F. Luebert, T. Boehnert & F. F. Merklinger 3825 (BONN, ULS); road to Andacollo, c. 6 km before Andacollo, 30.201205 S, 71.092169 W, 900 m, 24 Sep 2017, F. Luebert, T. Boehnert & F. F. Merklinger 3807 (BONN, ULS). Prov. Limari, Ovalle, Villaseca near Huamalata, 30.568064 S, 71.150966 W, 270 m, 24 Sep 2017, F. Luebert, T. Boehnert & F. F. Merklinger 3799 (BONN, ULS); Fray Jorge, 215 m, 13 Aug 1917, C. & I. Skottsberg 746 (S); Ovalle, Fray Jorge, 200 m, Nov 1925, E. Werdermann 892 (U [U 1145306], S); Ovalle, Fray Jorge, Oct 1947, B. Sparre 3061 (S); hotel Termas de Socos, 30.732502 S, 71.493507 W, 80 m, 23 Sep 2017, F. Luebert, T. Boehnert & F. F. Merklinger 3777 (BONN, ULS); road from Los Loros to Caleta El Toro, 30.741021 S, 71.65348 W, 50 m, 23 Sep 2017, F. Luebert, T. Boehnert & F. F. Merklinger 3790 (BONN, ULS); road from Alcones to Los Loros, 30.78262 S, 71.587161 W, 330 m, 23 Sep 2017, F. Luebert, T. Boehnert & F. F. Merklinger 3781 (BONN, ULS). Prov. Choapa, road from Combarbala to Canela Baja, a few km after Los Pozos, 31.363888 S, 71.260395 W, 500 m, 22 Sep 2017, F. Luebert, T. Boehnert & F. F. Merklinger 3775 (BONN, ULS); road from Illapel to Combarbala, near Illapel, 31.604095 S, 71.125953 W, 450 m, 22 Sep 2017, F. Luebert, T. Boehnert & F. F. Merklinger 3772 (BONN, ULS). Region V Valparaiso: Prov. Petorca, La Ligua, 5 km from Petorca on road from Pedequa, 550 m, 27 Nov 1938, C. R. Worth & J. L. Morrison 16704 (S). Prov. San Felipe de Aconcagua, Cuesta Las Chilcas, 560 m, 19 Jul 2003, F. Luebert & L. Kritzner 1757 (EIF); Cuesta Las Chilcas, 32.851797 S, 70.875068 W, 380 m, 20 Sep 2017, F. Luebert, T. Boehnert & F. F. Merklinger 3741 (BONN, ULS); Cuesta Las Chilcas, 32.851797 S, 70.875068 W, 380 m, 20 Sep 2017, F. Luebert, T. Boehnert & F. F. Merklinger 3741 A (BONN, ULS). Prov. Valparaiso, 14 Jan 1947, E. Wall & B. Sparre 45 (S); rocks near the sea, 17 Nov 1895, O. Buchtien s. n. (S). Prov. Quillota, Parque Nacional La Campana, Cerro La Campana, 32 ° 58.777 " S, 71 ° 7.670 " W, 480 m, 30 Dec 2000, F. Luebert 1398 (EIF). Prov. San Antonio, Nov 1927, O. Buchtien 3450 (L 3661663). Prov. Melipilla, Curacavi, Dec 1967, (L 3661657); La Barriga, Oct 1964, O. Zoellner 1765 (L 3661658).	en	Merklinger, Felix F., Luebert, Federico (2020): Notes on the genus Ophryosporus (Asteraceae, Eupatorieae) in Chile. PhytoKeys 161: 61-77, DOI: http://dx.doi.org/10.3897/phytokeys.161.53736, URL: http://dx.doi.org/10.3897/phytokeys.161.53736
EC8C263C7D2B57A09BCA67C9A8801359.taxon	materials_examined	Type. Chile. Region I Tarapaca: Usmagama, 15 Mar 1885, R. A. Philippi s. n. (lectotype, selected by Plos and Sancho (2013: 339): SGO [SGO 044738!]; isolectotype: K [K 000486664, photo!]). Ophryosporus pinifolius has an inconspicuous pappus that consists of minute squamellae. Its leaves are extremely variable and range from linear-lanceolate with entire margins to irregularly dentate ones. It is one of two Andean species in the genus known to occur in Chile. Based on herbarium records, its distribution is centered in the northern regions of Tarapaca and Arica y Parinacota where it is widespread (Fig. 1).	en	Merklinger, Felix F., Luebert, Federico (2020): Notes on the genus Ophryosporus (Asteraceae, Eupatorieae) in Chile. PhytoKeys 161: 61-77, DOI: http://dx.doi.org/10.3897/phytokeys.161.53736, URL: http://dx.doi.org/10.3897/phytokeys.161.53736
EC8C263C7D2B57A09BCA67C9A8801359.taxon	materials_examined	Specimens examined. Chile. Region XV Arica y Parinacota: Prov. Arica, Timar, between Timar and Tignamar, 18.717336 S, 69.663483 W, 2840 m, 29 Mar 2017, F. F. Merklinger & A. Stoll 2017 - 51 (BONN, ULS); shortly after Timar, 18.736382 S, 69.706777 W, 2447 m, 29 Mar 2017, F. F. Merklinger & A. Stoll 2017 - 54 (BONN, ULS); shortly after Timar, 18.747992 S, 69.699137 W, 2307 m, 29 Mar 2017, F. F. Merklinger & A. Stoll 2017 - 47 (BONN, ULS); Codpa, between Codpa and Timar, 18.762747 S, 69.69828 W, 2393 m, 29 Mar 2017, F. F. Merklinger & A. Stoll 2017 - 46 (BONN, ULS); quebrada de Vitor NW of Palca, 18.827041 S, 69.677724 W, 2085 m, 28 Mar 2017, F. F. Merklinger & A. Stoll s. n. (BONN, ULS); quebrada Chokaya, from Codpa into Camarones valley toward Pachica, 18.86473 S, 69.68034 W, 2235 m, 28 Mar 2017, F. F. Merklinger & A. Stoll 2017 - 30 (BONN, ULS); quebrada Chokaya, from Codpa into Camarones valley toward Pachica, 18.88289 S, 69.664972 W, 2373 m, 28 Mar 2017, F. F. Merklinger & A. Stoll 2017 - 40 (BONN, ULS); between Esquina and Pachica, 18.927444 S, 69.552944 W, 2298 m, 28 Mar 2017, F. F. Merklinger & A. Stoll 2017 - 43 (BONN, ULS); along road through Illapata, 18.94831 S, 69.50272 W, 2300 m, 28 Mar 2017, F. F. Merklinger & A. Stoll 2017 - 41 (BONN, ULS); road A- 135 from Panamericana to Puquios, 18.254661 S, 69.828852 W, 3100 m, 14 Oct 2017, F. Luebert, T. Boehnert & F. F. Merklinger 4008 (BONN, ULS). Prov. Parinacota, along road Putre-Arica, 18.21033 S, 69.56082 W, 3500 m, 1 Apr 2017, F. F. Merklinger & A. Stoll 2017 - 65 (BONN, ULS); Paychama [Pachama], 3600 m, 10 Mar 1927, C. Troll 3244 (B, M); along road Putre-Arica, 18.45314 S, 69.76415 W, 3102 m, 1 Apr 2017, F. F. Merklinger & A. Stoll 2017 - 62 (BONN, ULS); along A- 31 near Belen, 18.48545 S, 69.52782 W, 3500 m, 29 Mar 2017, F. F. Merklinger & A. Stoll 2017 - 56 (BONN, ULS); NE of Saxamar, 18.55108 S, 69.50015 W, 3500 m, 29 Mar 2017, F. F. Merklinger & A. Stoll 2017 - 57 (BONN, ULS); Saxamar, 18.56667 S, 69.48333 W, c. 3000 m, 20 Mar 2015, A. Moreira & F. Luebert 2456 (BONN); W of Tignamar along road, 18.57918 S, 69.52785 W, 3300 m, 29 Mar 2017, F. F. Merklinger & A. Stoll 2017 - 55 (BONN, ULS); Tignamar, 18 ° 37 ' S, 69 ° 28 ' W, 3100 m, 11 Sep 1963, F. Schlegel 4879 (CONC, F [photo; as Piqueria floribunda]); between Timar and Tignamar, shortly before Tignamar, 18.663898 S, 69.560963 W, 3354 m, 29 Mar 2017, F. F. Merklinger & A. Stoll 2017 - 53 (BONN, ULS). Prov. Arica, Palca, 18 ° 50 ' S, 69 ° 40 ' W, 2200 m, 30 Oct 1964, F. Schlegel 5092 (CONC, F [photo, as Piqueria floribunda]). Region I Tarapaca: Prov. Tamarugal, quebrada de Soga, 19.514361 S, 69.381274 W, 2400 m, 16 Mar 2017, F. Luebert, T. Boehnert & F. F. Merklinger, A. Stoll & D. Quandt 3455 (BONN, ULS); quebrada Aroma, 19.514361 S, 69.381274 W, 2300 m, 16 Mar 2017, F. F. Merklinger 2017 - 6 (BONN, ULS); above Jaina, 19.551255 S, 69.242848 W, 2750 m, 21 Mar 2017, F. Luebert & T. Boehnert 3631 (BONN, ULS); at entrance of Andean village Chusmiza, 19.67830 S, 69.17930 W, 3393 m, 2 Oct 2019, F. F. Merklinger, F. Luebert & J. Ruhm 2019 - 21 (BONN, EIF, K); Chusmiza, above town, 19.683025 S, 69.183376 W, 3400 m, 21 Mar 2017, F. Luebert & T. Boehnert 3642 (BONN, ULS); 2 km above the village of Chusmiza in quebrada de Chusmiza at km 76 alongside the new road to Colchane, 19 ° 40 ' 53.7 " S, 69 ° 11 ' 11.0 " W, 3406 m, 18 Feb 2003, M. F. Gardner & S. G. Knees 6512 (SGO 150393); road to Usmagama, turnoff ruta CH- 15, 19.730253 S, 69.218684 W, 2956 m, 26 Mar 2017, F. F. Merklinger & A. Stoll 2017 - 26 (BONN, ULS); road to Usmagama, turnoff ruta CH- 15, 19.730253 S, 69.218684 W, 2956 m, 26 Mar 2017, F. F. Merklinger & A. Stoll 2017 - 19 (BONN, ULS); cuesta Usmagama, km 3.9, 19.730154 S, 69.217046 W, 3050 m, 28 Oct 2016, F. Luebert & T. Boehnert 3452 (BONN, ULS); Usmagama, road from Usmagama to Limacsina, 19.78771 S, 69.207368 W, 2434 m, 26 Mar 2017, F. F. Merklinger & A. Stoll 2017 - 22 (BONN, ULS); quebrada de Parca, 19.985261 S, 69.098117 W, 3261 m, 22 Mar 2017, F. F. Merklinger & A. Stoll 2017 - 12 (BONN, ULS).	en	Merklinger, Felix F., Luebert, Federico (2020): Notes on the genus Ophryosporus (Asteraceae, Eupatorieae) in Chile. PhytoKeys 161: 61-77, DOI: http://dx.doi.org/10.3897/phytokeys.161.53736, URL: http://dx.doi.org/10.3897/phytokeys.161.53736
80E4D9E96B0A56F1A3B39D84A76A3E23.taxon	materials_examined	Type. Chile. Region III Atacama: Copiapo, F. J. F. Meyen s. n. (holotype: B [probably destroyed]; F neg. 14718!).	en	Merklinger, Felix F., Luebert, Federico (2020): Notes on the genus Ophryosporus (Asteraceae, Eupatorieae) in Chile. PhytoKeys 161: 61-77, DOI: http://dx.doi.org/10.3897/phytokeys.161.53736, URL: http://dx.doi.org/10.3897/phytokeys.161.53736
80E4D9E96B0A56F1A3B39D84A76A3E23.taxon	materials_examined	Specimens examined. Chile. Region II Antofagasta: Prov. Tocopilla, quebrada 2 - 3 km N of Tocopilla above old Caleta Duendes, 150 - 200 m, 18 Oct 1988, M. O. Dillon & D. Dillon 5718 (BONN); Tercera quebrada Tocopilla, 22.0558 S, 70.17662 W, 300 m, 18 Oct 2016, F. Luebert, A. Stoll & T. Boehnert 3413 (BONN, ULS); Quebrada La Higuera, S Tocopilla, 22 ° 18 ' 7 " S, 70 ° 12 ' 58 " W, 30 Sep 2005, F. Luebert, N. Garcia & N. Schulz 2569 / 963 (EIF); Quebrada above Caleta Buena, S Mantos de la Luna, 22.43075 S, 70.22186 W, 640 m, 10 Oct 2017, F. Luebert, T. Boehnert & F. F. Merklinger 3989 (BONN, ULS). Prov. Antofagasta, near Cobija, 19 Dec 1971, O. Zoellner 4593 (L 153863); Juan Lopez, 23.51205 S, 70.53365 W, 150 m, 19 Jul 2003, M. Antonissen 7 (EIF); Quebrada La Chimba, 17 Dec 1987, M. O. Dillon & J. T. S. Teillier 5321 (BONN); Quebrada La Chimba, 23 ° 33 ' S, 70 ° 22 ' W, 380 - 480 m, 11 Nov 1988, M. O. Dillon & D. Dillon 5881 (BONN); Quebrada La Chimba, 23.53567 S, 70.35887 W, 460 m, 17 Oct 2016, F. Luebert, A. Stoll & T. Boehnert 3408 (BONN, ULS); Quebrada Cardon, 24.741717 S, 70.542687 W, 210 m, 5 Oct 2017, F. Luebert, T. Boehnert & F. F. Merklinger 3949 (BONN, ULS); Aguada Cardon, 24.74173 S, 70.54385 W, 210 m, 15 Oct 2016, F. Luebert, A. Stoll & T. Boehnert 3389 (BONN, ULS); Quebrada Panul, 24.773468 S, 70.533915 W, 180 m, 4 Oct 2017, F. Luebert, T. Boehnert & F. F. Merklinger 3939 (BONN, ULS); Quebrada Panul, 24.777263 S, 70.531618 W, 190 m, 5 Oct 2017, F. Luebert, T. Boehnert & F. F. Merklinger 3944 (BONN, ULS); Quebrada Panul, 24.777263 S, 70.531618 W, 190 m, 5 Oct 2017, F. Luebert, T. Boehnert & F. F. Merklinger 3944 A (BONN, ULS); c. 7 km N of Paposo, 24 ° 57 ' S, 70 ° 29 ' W, 40 m, 14 Dec 1987, M. O. Dillon & J. T. S. Teillier 5262 (BONN); El Rincon, al N de Paposo, 17 Sep 1941, C. Munoz P. & G. T. Johnson 2902 (SGO 118350); Paposo, base Cerro Carneros, 24 ° 56 ' 24 " S, 70 ° 28 ' 44.6 " W, 160 m, 23 Oct 2009, A. Moreira & F. Luebert 1200 (SGO 158780); Quebrada Portezuelo, 25.0124 S, 70.446467 W, 550 m, 13 Oct 2016, F. Luebert, A. Stoll & T. Boehnert 3357 (BONN, ULS); Quebrada Matancilla, c. 5 km S of Punta Grande, 25 ° 07 ' S, 70 ° 27 ' W, 170 - 350 m, 27 Oct 1988, M. O. Dillon, D. Dillon, V. Asencio & M. Villarroel O. 5750 (BONN); Cachinalcito, 25 ° 10 ', 28 Nov 1925, I. M. Johnston 5173 (S); c. 20 km N of Taltal, quebrada Anchuna, 25.23543 S, 70.42594 W, 183 m, 31 Aug 2018, F. F. Merklinger, A. Kozok & D. Quandt 2018 - 63 (BONN, ULS); c. 20 km N of Taltal, quebrada Anchuna, 25.23543 S, 70.42594 W, 183 m, 31 Aug 2018, F. F. Merklinger, A. Kozok & D. Quandt 2018 - 64 (BONN, ULS); c. 20 km N of Taltal, quebrada Anchuna, 25.23543 S, 70.42594 W, 183 m, 31 Aug 2018, F. F. Merklinger, A. Kozok & D. Quandt 2018 - 65 (BONN, ULS); c. 20 km N of Taltal, quebrada Anchuna, 25.23543 S, 70.42594 W, 183 m, 31 Aug 2018, F. F. Merklinger, A. Kozok & D. Quandt 2018 - 66 (BONN, ULS); Quebrada El Medano, 300 m, 8 Oct 1941, E. Pisano V. & R. Bravo F. 398 (SGO); Quebrada San Ramon, 25.38578 S, 70.43658 W, 120 m, 11 Oct 2016, F. Luebert, A. Stoll & T. Boehnert 3346 (BONN, ULS); Quebrada San Ramon, 17 Sep 1968, O. Zoellner 2851 (U [U 3661650]); Cerro Perales, c. 5 km E of Taltal, 25 ° 25 ' S, 70 ° 25 ' W, 550 m, 21 Nov 1988, M. O. Dillon & D. Dillon 6002 (BONN); Hills SE of Taltal, 25 ° 29 ' S, 25 Nov 1925, I. M. Johnston 5080 (S); Posado Hidalgo, 25 ° 45 ' S, 70 ° 35 ' W, 13 Dec 1925, I. M. Johnston 5661 (S); along road Panamericana toward Caleta Esmeralda, 25.895921 S, 70.581052 W, 500 m, 9 Oct 2016, F. Luebert, A. Stoll & T. Boehnert 3308 (BONN, ULS). Region III Atacama: Prov. Chanaral, 21 km W of Panamericana on northern route to Pan de Azucar NP, 26 ° 08 ' S, 70 ° 37 ' W, 85 m, 30 Sep 1988, M. O. Dillon, D. Dillon & V. Pobleto 5609 (BONN); Falda Verde, hills N of Chanaral, 26.296721 S, 70.631252 W, 75 - 600 m, 1 Oct 2017, F. Luebert, T. Boehnert & F. F. Merklinger 3905 (BONN); Hills back of El Barquito, 26 ° 23 ' S, 28 - 29 Oct 1925, I. M. Johnston 4809 (S). Prov. Copiapo, Sector quebrada El Leon, 26.976625 S, 70.773903 W, 70 m, 8 Oct 2016, F. Luebert, A. Stoll & T. Boehnert 3290 (BONN, ULS); Caldera on small point just north of town, 27 ° 3 ' S, 22 Nov 1925, I. M. Johnston 5067 (S); Copiapo, 400 m, 13 Jul 1938, Ch. H. Andreas 885 (U [U 1145297]); Tierra Amarilla, 700 m, Oct 1924, E. Werdermann 456 (U [U 1145305]); Estancia Castilla, road from Totoral to Panamericana, 27.919994 S, 70.84531 W, 240 m, 28 Sep 2017, F. Luebert, T. Boehnert & F. F. Merklinger 3869 (BONN, ULS). Prov. Huasco, Road Carrizal Bajo to Canto de Agua, c. 2 km from Carrizal Bajo, 28.112004 S, 71.116139 W, 45 m, 7 Oct 2016, F. Luebert, A. Stoll & T. Boehnert 3279 (BONN, ULS); Quebrada Baratillo, 28 ° 21 ' 57 " S, 71 ° 7 ' 21 " W, 150 m, 14 Sep 2003, F. Luebert & L. Kritzner 1805 (EIF); Quebrada Baratillo, 28.363325 S, 71.096321 W, 110 m, 7 Oct 2016, F. Luebert, A. Stoll & T. Boehnert 3269 (BONN, ULS); Hills E of Tres Playitas, 28.400532 S, 71.16755 W, 160 m, 27 Sep 2017, F. Luebert, T. Boehnert & F. F. Merklinger 3854 (BONN, ULS). Region IV Coquimbo: Prov. Elqui, Guayacan, s. a., unknown s. n. (S); Puente Juan Soldado, 29.656542 S, 71.301174 W, 200 m, 27 Sep 2017, F. Luebert, T. Boehnert & F. F. Merklinger 3831 (BONN, ULS); La Serena, 16 Sep 1947, B. Sparre 2595 (S); Coquimbo, Jul-Aug 1958, W. H. Harvey s. n. (S); Coquimbo, 100 m, Nov 1923, E. Werdermann 124 (U [U 1145298]); Coquimbo, rocks behind the Fort, 29.93372 S, 71.33691 W, 25 m, 24 Sep 2017, F. Luebert, T. Boehnert & F. F. Merklinger 3809 (BONN, ULS); Coquimbo, rocks behind the Fort, 29.93372 S, 71.33691 W, 25 m, 24 Sep 2017, F. Luebert, T. Boehnert & F. F. Merklinger 3809 A (BONN, ULS); Herradura, 9 Aug 1917, C. & I. Skottsberg 704 (S). Prov. Limari, Ovalle, Rio Limari, 11 Oct 1947, B. Sparre 2994 (S); N of Caleta El Toro, 30.737239 S, 71.699907 W, 25 m, 23 Sep 2017, F. Luebert, T. Boehnert & F. F. Merklinger 3794 (BONN, ULS); along road from Los Loros to Caleta El Toro, 30.741021 S, 71.65348 W, 50 m, 23 Sep 2017, F. Luebert, T. Boehnert & F. F. Merklinger 3786 (BONN, ULS).	en	Merklinger, Felix F., Luebert, Federico (2020): Notes on the genus Ophryosporus (Asteraceae, Eupatorieae) in Chile. PhytoKeys 161: 61-77, DOI: http://dx.doi.org/10.3897/phytokeys.161.53736, URL: http://dx.doi.org/10.3897/phytokeys.161.53736
