identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
AF1DA83BD457FFE6FF3ADFB844537EBB.text	AF1DA83BD457FFE6FF3ADFB844537EBB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Albericus murritus Kraus & Allison 2009	<div><p>Albericus murritus sp. n.</p><p>urn:lsid:zoobank.org:act: C1F51397-8A1C-404B-AAF5-2B68F46C6A53</p><p>Fig. 1A</p><p>Holotype. BPBM 33657 (field tag FK 13097), collected by F. Kraus, S of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=142.63423&amp;materialsCitation.latitude=-5.656703" title="Search Plazi for locations around (long 142.63423/lat -5.656703)">Tumbutu River</a>, Muller Range, 5.6567028°S, 142.6342342°E, 1700 m, Southern Highlands Province, Papua New Guinea, 2 April 2009.</p><p>Paratypes (n = 31). BPBM 33636–40, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=142.63423&amp;materialsCitation.latitude=-5.643116" title="Search Plazi for locations around (long 142.63423/lat -5.643116)">Kunida</a>, Muller Range, 5.6431159°S, 142.6342342°E, 1700 m, 21–22 March 2009 ; BPBM 33641, E slope <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=142.62334&amp;materialsCitation.latitude=-5.66954" title="Search Plazi for locations around (long 142.62334/lat -5.66954)">Mt. Itukua</a>, Muller Range, 5.66954°S, 142.62334°E, 2177 m, 27 March 2009 ; BPBM 33642– 53, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=142.63963&amp;materialsCitation.latitude=-5.6503625" title="Search Plazi for locations around (long 142.63963/lat -5.6503625)">Tumbutu River</a> below Mt. Paramo, 5.6503623°S, 142.63963°E, 31 March 2009 ; BPBM 33654–56, same data as BPBM 33642–53 except collected 1 April 2009; BPBM 33658, PNGNM 24095–96, same data as holotype; BPBM 33659, PNGNM 24093–94, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=142.63904&amp;materialsCitation.latitude=-5.64545" title="Search Plazi for locations around (long 142.63904/lat -5.64545)">Mt. Paramo</a>, 5.64545°S, 142.63904°E, 1777 m, 2 April 2009 ; BPBM 33660–63, same data as BPBM 33642–53 except collected 2 April 2009 .</p><p>Diagnosis. A small species (adult SV = 14.6–18.3 mm) distinguished by its combination of oblique lores, distinct tympanum, relatively broad snout (IN/SV = 0.081 – 0.096, mean 0.086), relatively wide finger discs (3rdF/SV = 0.059 –0.081), and advertisement call consisting of a single peep uttered in a continuous series.</p><p>Comparisons with other species. The new species differs from all congeners except A. swanhildae Menzies and A. exclamitans Kraus and Allison in having a call</p><p>Figure |. Photos in life of A paratype of Albericus murritus sp. n. (BPBM 33656) B paratype of Cophixalus caverniphilus sp. n. (BPBM 33711) C paratype of Oreophryne anamiatoi sp. n. (BPBM 33764), and D paratype of Oreophryne anamiatoi sp. n. (BPBM 33765).</p><p>consisting of a peep; all other Albericus have calls consisting of a single honk/buzz or a series of clicks. From A. swanhildae the new species differs in its larger size (SV = 13.6–15.4 mm in A. swanhildae), oblique (vs. vertical) lores, wider finger discs (3rdF/ SV = 0.055 –0.061 in A. swanhildae), and in having a single broad dark band across the center of each shank (vs. three narrow dark bars across each shank in A. swanhildae). From A. exclamitans the new species differs in having the tympanum evident (vs. hidden) in males and in having the call consist of an infrequently produced single peep (vs. rapid burst of 3–48 peeps in A. exclamitans). Albericus murritus is also slightly smaller than A. exclamitans (male SV = 14.6–18.3 mm, female SV = 15.0– 18.3 mm in A. murritus vs. 15.3–20.7 and 18.0–22.0 in A. exclamitans) and has a somewhat broader snout (IN/SV = 0.074 –0.085, mean 0.079 in A. exclamitans).</p><p>Description of holotype. Adult male. Head relatively wide (HW/SV = 0.38), with oblique and shallowly concave loreal region; canthus rostralis broadly rounded; nostrils small, crescent-shaped, much closer to tip of snout than to eyes; distance from external naris to eye larger than internarial distance (EN/IN = 1.14, IN/SV = 0.083, EN/SV = 0.095); snout bluntly rounded when viewed from above, truncate when viewed from side; eyes moderately large (EY/SV = 0.13), eyelid approximately 2/3 width of interorbital distance; tympanum indistinct, partially hidden by surrounding skin. Dorsum pustulose on body and limbs; supratympanic fold absent but row of dorsal pustules occupy that area; ventral surfaces coarsely granular. Fingers unwebbed, bearing discs with terminal grooves; relative lengths 3&gt;4&gt;2&gt;1. Finger discs approximately twice widths of penultimate phalanges. Subarticular tubercles not well developed; metacarpal tubercles absent. Toes unwebbed, bearing discs with terminal grooves; relative lengths 4&gt;5&gt;3&gt;2&gt;1. Toe discs smaller than those of fingers (3rdF/4thT = 1.24), approximately 1.5 times width of penultimate phalanges. Subarticular tubercles low; metatarsal tubercles lacking. Hind legs rather short (TL/SV = 0.40); arms rather long (ArmL/SV = 0.54).</p><p>In preservative, dorsal ground color an irregular mix of yellow-tan and brown, with the former predominating dorsolaterally and the latter mid-dorsally; parts of both fields suffused with russet. Irregular black flecks and markings scattered throughout, concentrated above shoulders, on rear of head, and above tympana. Traces of a short, lighter, yellow-brown bar on each scapula; similar-colored interocular bar and vestiges of lumbar ocelli. Sides dark gray flecked with black and light blue-gray. Face yellow-tan flecked with black and russet. Legs yellow-tan with one broad, dark, centrally placed band on each thigh and shank. Rear of thighs dark brown with narrow yellow-tan band proximally. Venter dirty light gray evenly and densely peppered with dark gray; palmar and plantar surfaces same. Front margin of mandible russet. Iris black flecked with silver.</p><p>Variation. Mensural variation for the type series is shown in Table 1. Th ere is no obvious sexual dimorphism in morphometric features, although there is slight evidence that females may average larger in body size. However, this difference is slight considering the normal pattern of larger female size in most Papuan frogs.</p><p>Most specimens appear dark brown in preservative (darker than the holotype), with a few contrasting light-brown streaks or lines scattered on dorsum. These typically involve a single short line on each scapula, an interocular bar, and often traces of lumbar ocelli. Lighter specimens are similar but show more clearly the variable mottling seen in the holotype. Density of dark ventral stippling varies from sparse to dense, making the overall appearance of the venter vary from light gray to black. Both ventral extremes appear in frogs with both light and dark dorsa, but venters of frogs that are lighter dorsally average somewhat paler than those of dark frogs.</p><p>Color in life. BPBM 33636: “Dorsum dark tan with irregular black blotches and tiny white or tan dots on some warts; fairly warty. Orange-brown interocular bar, suprascapular marks, and on arms and heels. Venter charcoal gray with tiny light-gray punctations. Light-tan patch from eye to rictus. Iris brown.” BPBM 33637 was yellow-brown with cream and black markings, venter densely punctated with light gray, and iris tan.</p><p>Call. This species begins calling at dusk and calls in highest numbers during the first few hours of darkness. Th e call consists of a single “peep” note uttered in a continuous series with occasional breaks (Fig 2). We recorded calls of two individuals, BPBM 33641 and BPBM 33657 (Table 2). The notes had a mean duration of 0.151 s (range 0.093 –0.213). The inter-note interval for BPBM 33641 ranged from 4.1– 9.0 s (n = 20), except for two instances of 31.6 and 35.9 s. Many species of Albericus produce calls in groups separated by periods of silence (A. Allison, pers. obs.), and our brief recording suggests that this is occurring in BPBM 33641.</p></div>	https://treatment.plazi.org/id/AF1DA83BD457FFE6FF3ADFB844537EBB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Kraus, Fred;Allison, Allen	Kraus, Fred, Allison, Allen (2009): New microhylid Frogs from the Muller Range, Papua New Guinea. ZooKeys 26 (26): 53-76, DOI: 10.3897/zookeys.26.258
AF1DA83BD453FFE9FF3AD80244657B06.text	AF1DA83BD453FFE9FF3AD80244657B06.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cophixalus caverniphilus Kraus & Allison 2009	<div><p>Cophixalus caverniphilus sp. n.</p><p>urn:lsid:zoobank.org:act: 90795C42-34A3-4D52-B299-4025E0389CA3</p><p>Fig. 1B</p><p>Holotype. BPBM 33748 (field tag FK 13180), collected by F. Kraus and D. Gibson, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=142.6394&amp;materialsCitation.latitude=-5.64728" title="Search Plazi for locations around (long 142.6394/lat -5.64728)">Mt. Paramo</a>, Muller Range, 5.64728°S, 142.63941°E, 1718 m, Southern Highlands Province, Papua New Guinea, 5 April 2009.</p><p>Paratypes (n = 48). BPBM 33707, E slope <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=142.62334&amp;materialsCitation.latitude=-5.66954" title="Search Plazi for locations around (long 142.62334/lat -5.66954)">Mt. Itukua</a>, Muller Range, 5.66954°S, 142.62334°E, 2177 m, 20 March 2009 ; BPBM 33708–40, PNGNM 24101–07, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=142.63663&amp;materialsCitation.latitude=-5.64791" title="Search Plazi for locations around (long 142.63663/lat -5.64791)">Mt. Paramo</a>, Muller Range, 5.64791°S, 142.63663°E, 1780 m, 22 March 2009 ; BPBM 33741–43, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=142.63&amp;materialsCitation.latitude=-5.64" title="Search Plazi for locations around (long 142.63/lat -5.64)">Kunida</a>, 5.64°S, 142.63°E, 1700–1800 m, 23 March 2009 ; BPBM 33744– 46, same data as BPBM 33707, except collected 30 March 2009; BPBM 33747, same data as holotype except collected 4 April 2009 .</p><p>Diagnosis. A moderately large species of Cophixalus (adult SV = 25.5–36.7 mm) distinguished by its combination of expanded finger discs larger than toe discs (3rdF/ SV = 0.069 –0.107, 3rdF/4thT = 1.27–1.70); long legs (TL/SV = 0.50–0.59); third toe distinctly longer than fifth; distinct tympanum; curved scapular ridges; pustulose skin; short, dark postocular stripe; dark W-shaped mark above the shoulders; lacking a dark</p><p>)</p><p>s</p><p>(</p><p>Notes 60</p><p>Between 20</p><p>Interval ļ0</p><p>0 0 5 ļ0 ļ5 20 25 30 35</p><p>Call No.</p><p>face and yellow spotting on the rear of thighs; and having a call consisting of a rapid series of 6–9 musical peeps.</p><p>Comparisons with other species. The new species differs from all congeners except C. balbus Günther, C. biroi (Méhelÿ), C. cheesmanae Parker, C. cryptotympanum, C. nubicola Zweifel, C. riparius Zweifel, and C. verrucosus (Boulenger) in its combination of SV&gt; 25 mm, finger discs larger than toe discs, and third toe longer than fifth. It differs from C. balbus and C. biroi in having (vs. lacking) raised scapular ridges, lacking (vs. having) a triangle of pale coloration on the top of the snout, having a call consisting of only 6–9 peeps (vs. long trains of peeps), and from C. biroi in having (vs. lacking) a dark W-shaped mark above the shoulders; from C. cheesmanae in having pustulose (vs. smooth) skin and in lacking the dorsolateral ridge and uniformly dark face of that species; from C. cryptotympanum in having an exposed (vs. obscure or hidden) tympanum, pustulose (vs. smooth) skin, a dark (vs. light) postocular stripe, and a dark W-shaped mark above the shoulders (absent in C. cryptotympanum); from C. nubicola in its longer legs (TL/SV = 0.35–0.38 in C. nubicola), thighs same color as (vs. conspicuously paler than) dorsum, and dorsum yellow or greenish-yellow with a dark W-shaped mark above the shoulders (vs. reddish-brown with light spots in C. nubicola); from C. riparius in its smaller size (SV = 40–50mm in C. riparius) and dorsum yellow or greenish-yellow with a dark W-shaped mark above the shoulders (vs. brown with two elongated and irregular dark dorsolateral patches or network in C. riparius); and from C. verrucosus in having larger finger discs (3rdF/SV = 0.052 –0.073, 3rdF/4thT = 0.96–1.26), a dark W-shaped mark above the shoulders (absent in C. verrucosus), and rear of thighs yellow dusted with brown (vs. spotted with yellow on dark brown in C. verrucosus).</p><p>Description of holotype. Adult male with small lateral incision on right side. Head moderately wide (HW/SV = 0.39), with steep, almost vertical, loreal region and slightly inflated lip; canthus rostralis rounded, straight when viewed from above; nostrils directed laterally, much closer to tip of snout than to eyes; internarial distance larger than distance from naris to eye (EN/IN = 0.83, IN/SV = 0.106, EN/ SV = 0.088); snout rounded and somewhat projecting when viewed from the side, broadly angulate when viewed from above; eyes moderately large (EY/SV = 0.12); eyelid approximately 2/3 width of interorbital distance; tympanum indistinct and small (TY/SV = 0.049), with a slightly raised annulus anteriorly but covered by ridge of skin dorsally and posteriorly. Skin pustulose dorsally and laterally, with prominent pair of scapular ridges, granulose ventrally. Supratympanic fold present, distinct ventrally but merging with pustulose skin dorsally. Fingers unwebbed, bearing discs with terminal grooves; relative lengths 3&gt;4&gt;2&gt;1; first finger and disc well-developed. Finger discs approximately 3 times widths of penultimate phalanges, except for the first finger disc, which is approximately 2 times width of penultimate phalanx. Subarticular tubercles low; inner metacarpal tubercle weakly developed, outer absent. Toes unwebbed, bearing discs with terminal grooves; relative lengths 4&gt;3&gt;5&gt;2&gt;1. Toe discs smaller than those of fingers (3 rdF /4thT = 1.33); approximately twice widths of penultimate phalanges. Subarticular tubercles weakly developed; inner metatarsal tubercle narrow, elongate, and low; outer lacking. Hind legs moderately long (TL/ SV = 0.54).</p><p>In preservative, dorsum brown with broken, dark-brown scapular W, and indistinct dark-brown markings scattered throughout. Dark-brown interocular bar, postocular stripe, and loreal stripe. Face light brown with small dark-brown flecks and smudges. Rear of thighs pale straw suffused with brown stippling and having dark-brown blotching distally. Ventral surfaces pale straw yellow heavily stippled with black, this more evenly distributed on chin and throat, and more broken on abdomen and under legs. Few white flecks along margin of mandible and sparsely scattered on throat and chest. Palmar and plantar surfaces brown, latter darker. Iris very dark brown, densely flecked with brass.</p><p>Variation. Mensural variation for the type series is shown in Table 3. Sexual dimorphism is slight except that females are of larger size than males. Th ere is relatively little variation in dorsal color pattern. Most specimens are similar to the holotype, but four or five have a slightly lighter ground color, and four are darker; all have the scapular W. Similarly, ventral color pattern varies little. Most specimens are like the holotype, but a few (7 or 8) have darker stippling, which gives a higher contrast to the ventral pattern.</p><p>Color in life. BPBM 33708: “Mustard yellow with irregular dark-brown markings; dark-brown suprascapular W and dorsolateral lines which are ridges. Iris bronze punctated with brown. Venter yellow with blue-white punctations.” BPBM 33728 was mustard yellow with black spots dorsally. Th e yellow color was prominent in all specimens collected.</p><p>Call. We heard this species call only in the morning between 0500–0930 h. We obtained good recordings of 16 calls produced by two individuals, BPBM 33747 and 33748 (Table 4). Both calls were similar, so we combined data for analysis. The call consists of 5–9 (mode = 7) regularly spaced notes delivered at a mean rate of 6.63 notes/s (range 5.65–7.57) with the acoustic impression of bell-like peeping. The mean duration of calls was 1.77 s (range 1.50–2.18). The first note, with one exception, was the longest note in the call, averaging 0.2933 s (range 0.0920 –0.3280) (Fig. 5A). If the exception is excluded, the mean duration of the first note increases to 0.3067 (range 0.2818 –0.3280). This note also generally had a lower amplitude than that of subsequent notes.</p><p>The second note tended to be the second longest in the call, averaging 0.1556 s (range 0.0790 –0.1990). Subsequent notes were generally similar, averaging 0.1324 s in duration (range 0.0602 –0.1960), but in nine of the 16 calls the last note was the shortest in the call, with a duration ranging from 0.0602 – 0.1100 s.</p><p>There was a slight tendency for pulsing of the first note (Fig. 6 A–B). This is more apparent in the second and subsequent notes, particularly in BPBM 33748 (Fig. 6B), in which the first note generally included upwards of 8–9 irregular, often partial pulses. Subsequent notes, particularly in BPBM 33748, generally had at least two components: a short, high-amplitude pulse of ̴0.03–04 s followed by a longer, lower-amplitude pulse that was generally 0.07 s or more in duration.</p><p>0.5 ļ.0 ļ.5 2.0 Time (s)</p></div>	https://treatment.plazi.org/id/AF1DA83BD453FFE9FF3AD80244657B06	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Kraus, Fred;Allison, Allen	Kraus, Fred, Allison, Allen (2009): New microhylid Frogs from the Muller Range, Papua New Guinea. ZooKeys 26 (26): 53-76, DOI: 10.3897/zookeys.26.258
AF1DA83BD458FFF0FF3AD91845017A6F.text	AF1DA83BD458FFF0FF3AD91845017A6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oreophryne anamiatoi Kraus & Allison 2009	<div><p>Oreophryne anamiatoi sp. n.</p><p>urn:lsid:zoobank.org:act: 29D10CFC-A8FB-431B-A61C-289B7619CF78</p><p>Fig. 1C, D</p><p>Holotype. BPBM 33768 (field tag FK 12882), collected by F. Kraus and J. Anamiato, E slope <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=142.62334&amp;materialsCitation.latitude=-5.66954" title="Search Plazi for locations around (long 142.62334/lat -5.66954)">Mt. Itukua</a>, Muller Range, 5.66954° S, 142.62334° E, 2177 m, Southern Highlands Province, Papua New Guinea, 27 March 2009.</p><p>Paratypes (n = 20). BPBM 33763, same data as holotype except collected 17 March; BPBM 33764, same data as holotype except collected 22 March; PNGNM 24097, same data as holotype except collected 25 March; BPBM 33765–66, same data as holotype except collected 26 March; BPBM 33767, same data as holotype; BPBM 33769–71, same data as holotype except collected 28 March; PNGNM 24098, same data as holotype except collected 29 March; BPBM 33772, same data as holotype except collected 30 March; PNGNM 24099, same data as holotype except collected 31 March; BPBM 33773–79, PNGNM 24100, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=142.63618&amp;materialsCitation.latitude=-5.64509" title="Search Plazi for locations around (long 142.63618/lat -5.64509)">Mt. Paramo</a>, Muller Range, 5.64509° S, 142.63618° E, 1874 m, 3 April 2009 .</p><p>Diagnosis. A medium-sized species of Oreophryne (adult SV = 23.4–29.7 mm) distinguished by its combination of a cartilaginous connection of the procoracoid to the scapula, no webbing between the toes, fifth toe longer than the third, relatively short snout (EN/SV = 0.073 –0.086), dark face, venter with a dense array of darkbrown flecks, and call consisting of an extended multi-note chuckle or cackle.</p><p>Comparisons with other species. The new species differs from all Papuan congeners except O. alticola Zweifel, Cogger, and Richards, O. asplenicola Günther, O. brevicrus Zweifel, O. clamata Günther, O. crucifera (van Kampen), O. flava Parker, O. habbemensis Zweifel, Cogger, and Richards, O. idenburgensis Zweifel, O. kampeni Parker, O. notata, O. pseudasplenicola Günther, and O. waira Günther in having (vs. lacking) a procoracoid that reaches the scapula. It differs from O. crucifera, O. idenburgensis, O. kampeni, and O. waira in lacking (vs. having) webbing between the toes; and it differs from O. alticola, O. brevicrus, O. clamata, and O. habbemensis in having the fifth toe longer than (vs. shorter than or subequal to) the third. Oreophryne anamiatoi differs from the remaining species O. asplenicola, O. flava, O. notata, and O. pseudasplenicola in its larger size (23.4–29.7 mm vs. maximum of 21 mm in those four species) in having a uniformly dark face, and in its call (call a series of peeps in the other species, but call unknown in O. flava); it further differs from O. asplenicola and O. pseudasplenicola in its shorter snout (EN/SV&gt; 0.086 in those species), from O. notata and O. pseudasplenicola in lacking (vs. having) an inverted white U on the face, and from O. flava in having an abdomen with dark-brown flecking (vs. immaculate yellow-white), and lacking (vs. having) a dark W-shaped mark between the shoulders.</p><p>Description of holotype. Adult female with incision on right side. Head wide (HW/SV = 0.40), with steep, slightly concave loreal region. Canthus rostralis round- ed, concave when viewed from above. Nostrils directed laterally, closer to tip of snout than to eyes. Internarial distance broader than distance from naris to eye (EN/IN = 0.80, IN/SV = 0.101, EN/SV = 0.081). Snout truncate when viewed from the side, shallowly angulate when viewed from above. Eyes moderately large (EY/SV = 0.11); eyelid approximately two-thirds width of interorbital distance. Tympanum distinct but small (TY/SV = 0.047). Dorsal skin granular with series of weakly raised parallel ridges and scattered small pustules; ventral surfaces coarsely granular. Supratympanic fold narrow. Fingers unwebbed, bearing discs with terminal grooves; relative lengths 3&gt;4&gt;2&gt;1. Finger discs approximately 3 times widths of penultimate phalanges, except for first finger, which is approximately twice width of penultimate phalanx. Subarticular tubercles well developed; inner metacarpal tubercle oval and low; outer rounded and obscure. Toes unwebbed, bearing discs with terminal grooves; relative lengths 4&gt;5&gt;3&gt;2&gt;1. Toe discs smaller than those of fingers (3 rdF /4thT = 1.37), approximately twice width of penultimate phalanges. Subarticular tubercles low but distinct; inner metatarsal tubercle large, oval; outer absent. Hind legs of moderate length (TL/SV = 0.46).</p><p>In preservative, dorsum medium brown with small black flecks scattered throughout, these concentrated dorsolaterally, mid-dorsally, and on top of head. Entire face to posterior of jaw angle uniform dark brown. Dark-brown postocular stripe extends along ventral side of supratympanic ridge. Rear and front of thighs uniform medium brown. Tops of wrist, hand, and first three fingers boldly marked with dark brown. Sides darker brown with very pale straw flecks. Venter very pale straw with bold, large, dark-brown flecks, these dense on chin, throat, and under legs, sparse on abdomen. Palmar and plantar surfaces dark brown spotted with very pale straw. Iris dark brown minutely flecked with silver.</p><p>Variation. Mensural variation for the type series is shown in Table 5. Sexual dimorphism is slight except that females clearly attain larger size than males and may have slightly narrower snouts (reflected in EN/IN values).</p><p>Dorsal ground color varies from light brown to dark brown, and pattern varies from virtually absent to moderately well stippled and streaked with black. Black stippling may be uniformly distributed, or concentrated laterally or dorsolaterally. Two specimens have a narrow tan vertebral stripe margined with black stippling. The dark face and postocular bar are present in all. Venters of all specimens are boldly spotted with dark brown on white, but two specimens have the abdomen clear and a few others have it less spotted than the chin and throat.</p><p>Color in life. BPBM 33764 (Fig. 1C): “Dorsum medium brown with a slight russet cast. Face dark brown; short dark-brown postocular dash; and few dark-brown lateral flecks. Rear of thighs brown, slightly darker than dorsum. Venter pale yellow heavily flecked with dark gray. Iris bronze with narrow red rim around pupil. Upper arms and tarsi burnt orange.” BPBM 33765 was uniform brown dorsally with a dark face mask and postocular stripe (Fig. 1D); rear and front of thighs, and groin, uniform brown, slightly darker than dorsum; venter dirty cream flecked with gray; iris dark brown. BPBM 33766 had black flecks on sides and venter brighter yellow but with fewer gray flecks and with white chromatophores; BPBM 33767 had the dorsum and rear of thighs burnt orange and venter pale yellow; BPBM 33769 and 33771 had a tan vertebral line. BPBM 33774 had a metallic green sheen on eyelids and top of snout; BPBM 33776 same but less extensive. PNGNM 24097 was light brown with straw yellow on sides, with dark-brown spots dorsally and laterally, denser on sides; face to rictus and short postocular stripe dark brown; rear of thighs brown like dorsum but unspotted; venter pale yellow spotted with dark-gray flecks; iris brown.</p><p>Call. Animals called during the first few hours of darkness. We recorded ten calls from two individuals (Table 6). Th e calls from both were similar, so we combined data from both for analysis.</p><p>The call consists of 17–22 pulsed notes (Figs. 8A, 9A). Note duration was similar over the course of the call and averaged 0.0569 s (range 0.0266 –0.0681). Internote intervals were also similar throughout the call and were similar in duration to the notes, averaging 0.0563 s (range 0.0390 –0.0942). Mean call duration was 2.10s (range 1.77–2.33), and notes were delivered at a mean repetition rate of 18.5 notes/s (range 17.3–20.3). Number of pulses/note ranged from 3–11 (Fig. 9A), with a strong tendency to decrease over the course of the call (Fig 10). Modal number of pulses in the first three notes in each call ranged from 10–11, producing a mean rate for those notes of 200.9 pulses/s, while modal number of pulses in the last three notes ranged from 7–9, producing a mean rate for those notes of 146.4 pulses/s (Table 7). Notes are finely tuned, with a mean dominant frequency of 2490</p><p>0.5 ļ.0 ļ.5 2.0 2.5</p><p>Time (s)</p><p>0.05 0.ļ0 0.ļ5 0.20 0.25 0.30</p><p>Time (s)</p><p>Hz (range 2460–2540) (Fig. 8 B–C, 9B–C) and no frequency modulation over the duration of the note.</p><p>Etymology. The species is named for Jim Anamiato of the Papua New Guinea National Museum for his considerable assistance on several of our expeditions, including the one during which this frog was discovered.</p><p>Range. Known only from the vicinity of the type locality on the northeastern slopes of the Muller Range, Southern Highlands Province, Papua New Guinea (Fig. 4).</p><p>Ecological notes. Animals inhabited both primary and secondary rainforest at elevations ranging from 1870–2180 m. Most animals were found in the immediate vicinity of stream banks, but a few were found several meters away in adjacent forest. Males called from dense moss mats on standing or fallen trees from 1–5 m above the ground,</p><p>2 4 6 8 ļ0</p></div>	https://treatment.plazi.org/id/AF1DA83BD458FFF0FF3AD91845017A6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Kraus, Fred;Allison, Allen	Kraus, Fred, Allison, Allen (2009): New microhylid Frogs from the Muller Range, Papua New Guinea. ZooKeys 26 (26): 53-76, DOI: 10.3897/zookeys.26.258
