taxonID	type	description	language	source
59BEEC8D7880AB388853088A475E751C.taxon	description	Description. Body size and form: Beetle small: TL-H 3.1 mm, TL 3.4 mm, MW 1.9 mm, with broader, oval habitus. Coloration: Brownish, with head and pronotum paler. Head reddish brown, darker posterior to eyes. Pronotum reddish brown on sides, dark brown on disc. Elytra brown. Head appendages yellowish red, legs reddish, distally darker, especially metathoracic legs (Fig. 14). Surface sculpture: Submatt dorsally. As in E. sumokedi sp. n., except for more strongly impressed microreticulation. Structures: Pronotum with lateral bead. Its lateral sides with longitudinal impressions. Base of prosternum and neck of prosternal process with distinct ridge, rounded anteriorly. Blade of prosternal process lanceolate, relatively broad, slightly convex, with distinct lateral bead and few setae. Abdominal ventrite 6 slightly truncate. Male: Antennae simple (Fig. 14). Protarsomere 4 with medium-sized, thick, curved anterolateral hook-like seta. Protarsomere 5 long and narrow, with anterior row of 17 and posterior row of 6 relatively short, thin setae (Fig. 38 D). Median lobe in lateral view slightly curved, with thickened, angulate apex; in ventral view, subparallel, very slightly narrowed distally, and with broadly truncate apex. Paramere slightly concave on dorsal side and with dense setae on subdistal part; proximal setae sparser (Fig. 38 A-C). Abdominal ventrite 6 with 4 - 5 lateral striae on each side. Female: Unknown.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
59BEEC8D7880AB388853088A475E751C.taxon	distribution	Distribution. Papua New Guinea: Madang Province. The species is known only from the type locality (Fig. 50).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
59BEEC8D7880AB388853088A475E751C.taxon	etymology	Etymology. The species is named after Adelbert Mountains. The species name is an adjective in the nominative singular.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
C6F678DB4C89060B9E9650806C282E4B.taxon	description	Description. Body size and form: Beetle medium-sized: TL-H 4.3 - 4.7 mm, TL 4.8 - 5.2 mm, MW 2.2 - 2.5 mm (holotype: TL-H 4.4 mm, TL 4.8 mm, MW 2.3 mm), with oblong habitus. Coloration: Brown to piceous, with head and pronotum paler. Head reddish brown to piceous, with small darker areas posterior to eyes. Pronotum dark brown to piceous, paler on sides and darker on disc. Elytra dark brown to piceous, with vague narrow reddish to brownish sutural lines. Head appendages and legs proximally reddish brown, legs distally darker, brownish, especially metathoracic legs (Fig. 21). Surface sculpture: Matt dorsally. Head with dense, coarse punctation (no spaces between punctures or spaces 2 times size of punctures), finer and sparser anteriorly; diameter of punctures equal to diameter of cells of microreticulation. Pronotum and elytra with dense, coarse punctation, sparser and finer than on head. Pronotum and elytra with strongly impressed microreticulation. Head with microreticulation stronger. Metaventrite and metacoxae distinctly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles, abdominal ventrites with distinct microreticulation and strioles. Metaventrite medially, metacoxal plates, and abdominal ventrites with fine, sparse punctation. Structures: Pronotum with distinct lateral bead. Its lateral sides with distinct longitudinal impressions. Base of prosternum and neck of prosternal process with distinct ridge, rounded anteriorly. Blade of prosternal process lanceolate, relatively narrow, slightly convex, and smooth, with distinct lateral bead and few lateral setae. Abdominal ventrite 6 slightly truncate. Male: Antennae simple (Fig. 21). Protarsomere 4 with anterior angle slightly expanded, with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 slightly concave ventrally, with anterior band of ca 70 and posterior band of ca 30 relatively long setae (Fig. 45 D). Median lobe in lateral view short, slightly curved, and evenly tapering to dully pointed apex, apex not bent downwards; in ventral view, almost subparallel and distally slightly narrowed to apex, apex roundly truncate. Paramere slightly concave on dorsal side, with long, dense subdistal setae, proximal ones finer (Fig. 45 A-C). Abdominal ventrite 6 with 13 - 15 lateral striae on each side. Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
C6F678DB4C89060B9E9650806C282E4B.taxon	distribution	Distribution. Papua New Guinea: Southern Highlands Province. The species is known only from the type locality (Fig. 50).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
C6F678DB4C89060B9E9650806C282E4B.taxon	etymology	Etymology. The species is named after Mt Ambua. The name is a noun in the nominative singular standing in apposition.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
E33CD35ED69C227FD8FEE1BF7C00BC9A.taxon	description	Description. Body size and form: Beetle small: TL-H 3.05 - 3.25 mm, TL 3.35 - 3.55 mm, MW 1.8 - 1.9 mm (holotype: TL-H 3.05 mm, TL 3.35 mm, MW 1.8 mm), with broader, oval habitus. Coloration: Brownish, with head and pronotum paler. Head reddish brown to brownish, sometimes paler anteriorly. Pronotum reddish brown on sides, brown to dark brown on disc. Elytra brown to dark brown, sometimes with narrow reddish sutural lines. Head appendages yellowish red, legs reddish, distally darker, especially metathoracic legs (Fig. 13). Surface sculpture: Shiny but with dense, distinct punctation dorsally. As in E. sumokedi sp. n. Structures: Pronotum with lateral bead. Its lateral sides with shallow longitudinal impressions. Base of prosternum and neck of prosternal process with distinct ridge, narrowly rounded anteriorly. Blade of prosternal process lanceolate, relatively broad, slightly convex, with distinct lateral bead and few setae. Abdominal ventrite 6 slightly truncate. Male: Antennae simple (Fig. 13). Protarsomere 4 with medium-sized, thick, curved anterolateral hook-like seta. Protarsomere 5 long and narrow, with anterior row of 21 and posterior row of 4 relatively short setae (Fig. 37 D). Median lobe in lateral view slightly curved, with slightly thickened, angulate, and curved downwards apex; in ventral view, subparallel, with broad and slightly concave apex. Paramere slightly concave on dorsal side and with dense setae on subdistal part; proximal setae sparser (Fig. 37 A-C). Abdominal ventrite 6 with 3 or 4 lateral striae on each side. Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
E33CD35ED69C227FD8FEE1BF7C00BC9A.taxon	distribution	Distribution. Papua New Guinea: Sandaun Province. The species is known only from the type locality (Fig. 50).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
E33CD35ED69C227FD8FEE1BF7C00BC9A.taxon	etymology	Etymology. The species is named after the Bewani Mountains. The name is a noun in the nominative singular standing in apposition.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
FB04BF19480843E14E474AF80F3D48DA.taxon	materials_examined	Additional material. 1 female " IRIAN JAYA: Paniai Prov. road Nabire-Ilaga, km 65 29.8.1996, 250 m leg. M. Balke (96 # 6) " (NHMW). 4 males, 2 females " West New Guinea / Paniai Prov / JR 22 track Nabire-Ilaga km 62 250 m, 24.7.1991, forest pools leg. Balke & Hendrich " (CLH). 1 male " W. - Neuguinea / Paniai Prov. Strasse Nabire-Ilaga km 5 700 m, 22. - 2.9.1990 / IR 11 leg: Balke & Hendrich " (CLH).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
FB04BF19480843E14E474AF80F3D48DA.taxon	diagnosis	Diagnosis. For complete description, see Balke (1998: 328). Beetle medium-sized: TL-H 3.6 - 4.05 mm; oblong-oval; reddish brown to dark brown, sometimes with reddish to reddish brown pronotal sides and head anteriorly; submatt, with fine but rather dense punctation and strongly impressed microreticulation; pronotum without lateral bead; male antennae simple (Fig. 2); male protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta; male protarsomere 5 long and narrow, with anterior band of more than 60 and posterior row of 12 relatively long, thin setae (Fig. 26 D); median lobe in lateral view slightly curved and apically rounded, in ventral view, almost subparallel and not narrowed before truncate or slightly concave apex; paramere slightly concave on dorsal side and with long, dense, thin setae, situated along dorsal margin: subdistal setae strong and dense, setae in middle part shorter and sparser, proximal setae long but sparser than subdistal ones (Fig. 26 A-C). Female without evident differences in external morphology from males, except for non-modified pro- and mesotarsi and abdominal ventrite 6 without striae.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
FB04BF19480843E14E474AF80F3D48DA.taxon	distribution	Distribution. Papua: Nabire Regency. The species is known only from the area close to the type locality (Fig. 50).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
05F7A20804B9F09FD9808916361DBA77.taxon	description	Description. Body size and form: Beetle small: TL-H 3.0 - 3.25 mm, TL 3.25 - 3.55 mm, MW 1.65 - 1.8 mm (holotype: TL-H 3.1 mm, TL 3.4 mm, MW 1.7 mm), with oblong-oval habitus. Coloration: Reddish. Dorsal surface almost uniformly yellowish red to reddish brown, with paler anterior part of head and pronotum laterally; head appendages and legs yellowish red (Fig. 12). All type specimens are teneral, therefore, coloration may be darker. Surface sculpture: Submatt dorsally. As in E. pseudopusilla sp. n. but microreticulation more weakly impressed, dorsal surface shinier. Structures: Pronotum with lateral bead. Its lateral sides with distinct longitudinal impressions. Base of prosternum and neck of prosternal process with distinct ridge, rounded anteriorly. Blade of prosternal process lanceolate, relatively broad, slightly convex, with distinct lateral bead and few setae. Abdominal ventrite 6 slightly truncate. Male: Antennae simple (Fig. 12). Protarsomere 4 with medium-sized, thick, curved anterolateral hook-like seta. Protarsomere 5 long and narrow, with anterior band of 30 and posterior row of 7 relatively long setae (Fig. 36 D). Median lobe in lateral view slightly curved, with slightly thickened, straight apex; in ventral view, subparallel, very slightly narrowed distally, with broad, truncate apex. Paramere slightly concave on dorsal side, with long, dense subdistal setae, median and proximal ones finer and sparser (Fig. 36 A-C). Abdominal ventrite 6 with 4 - 6 lateral striae on each side. Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
05F7A20804B9F09FD9808916361DBA77.taxon	distribution	Distribution. Papua: Jayapura Regency. The species is known only from the Cyclops Mountains (Fig. 50).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
05F7A20804B9F09FD9808916361DBA77.taxon	etymology	Etymology. The species is named after the Cyclops Mountains. The name is a noun in the nominative singular standing in apposition.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
6A26D3C073EB86450E76BD39C3B6E4B5.taxon	materials_examined	Additional material. Western Highlands: 3 females " Papua New Guinea: Western Highlands, Mt. Hagen town area, 1600 m, 7. xii. 1994 05.49.745 S 144.22.357 E Balke & Kinibel (PNG 131) " (ZSM). 1 male, 1 female " Papua New Guinea: Western Highlands, Kurumul, 6 Km SW Kudjip, small stream, 1584 m, 13. vi. 1994, 05.53.426 S 144.36.600 E, John (PNG 78) ", the female with an additional green label " M. Balke 1342 " (ZSM). Eastern Highlands: 5 males " Papua New Guinea: Eastern Highlands, Akameku-Brahmin, Bismarck Range, 1900 m, 23. xi. 1994, 05.54.284 S 145.22.271 E, Balke & Kinibel (PNG 108) ", one of them with an additional green label " M. Balke 1399 " (NHMW, ZSM). 3 males, 2 females " Papua New Guinea: Eastern Highlands, Akameku-Brahmin, Bismarck Range, 1500 m, 24. xi. 1994, 05.51.964 S 145.23.604 E, Balke & Kinibel (PNG 111) " (NHMW, ZSM). Simbu: 3 males, 1 female " Collection Naturhistorisches Museum Basel ", " Papua New Guinea Simbu prov. L. Ciek lgt. ", Kundiawa, Mu vill. 145 ° 02 ' E 4 ° 42 ' S III. 2001, 1900 m " (NHMB). 1 male " Ibisca Niugini, PNG 28 - 30. x. 2012 Mount Wilhelm 1700 m ", " - 5,759269238 145,235611 FIT-MW 1700 - K- 2 / 8 - d 01 / Plot 11 / P 1959 Vial 05833 " (IECA). 1 male " - 5,759269238 145,235611 FIT-MW 1700 - J- 1 / 8 - d 01 / Plot 10 / P 1950 Vial 02485 ", " Ibisca Niugini, PNG 25 - 27. x. 2012 Mount Wilhelm 1700 m " (ZSM). 2 females " Ibisca Niugini, PNG 27 - 29. x. 2012 Mount Wilhelm 1700 m - 5,759269238 145,235611 MW 1700 / P 1943 Vial 04017 " (ZSM). 1 female " Ibisca Niugini, PNG 31. x- 2. xi. 2012 Mount Wilhelm 1700 m - 5,759269238 145,235611 MW 1700 / P 1953 Vial 07522 " (ZSM). 1 female " Ibisca Niugini, PNG 2 - 4. xi. 2012 Mount Wilhelm 1700 m - 5,759269238 145,235611 MW 1700 / P 1914 Vial 07577 " (ZSM). 1 female " Ibisca Niugini, PNG 25 - 27. x. 2012 Mount Wilhelm 1700 m - 5,759269238 145,235611 MW 1700 / P 1878 Vial 02411 " (ZSM). 1 female " Ibisca Niugini, PNG 5 - 7. xi. 2012 Mount Wilhelm 1700 m ", " - 5,759269238 145,235611 FIT-MW 1700 - O- 6 / 8 - d 12 / Plot 15 / P 1995 Vial 05666 " (ZSM). 1 female " Ibisca Niugini, PNG 27 - 29. x. 2012 Mount Wilhelm 1700 m - 5,759269238 145,235611 ", " FIT-MW 1700 - H- 2 / 8 - d 03 / Plot 8 / P 1935 Vial P 1935 - CODYTI " (ZSM). 1 female " Ibisca Niugini, PNG 27 - 29. x. 2012 Mount Wilhelm 1700 m - 5,759269238 145,235611 ", " FIT-MW 1700 - I- 2 / 8 - d 03 / Plot 9 / P 1943 Vial P 1943 - CODYTI " (ZSM). 1 female " Ibisca Niugini, PNG 28 - 30. x. 2012 Mount Wilhelm 1700 m ", " - 5,759269238 145,235611 FIT-MW 1700 - T- 2 / 8 - d 04 / Plot 20 / P 2031 Vial 05553 " (ZSM). 1 female " Ibisca Niugini, PNG 27 - 29. x. 2012 Mount Wilhelm 1700 m ", " - 5,760916233 145,2353363 MW 1700 / P 1895 Vial P 1895 - CODYTI " (ZSM). 1 female " Ibisca Niugini, PNG 4 - 6. xi. 2012 Mount Wilhelm 1700 m ", " - 5,760916233 145,2353363 FIT-MW 1700 - C- 6 / 8 - d 11 / Plot 3 / P 1899 Vial 04070 " (ZSM). 1 female " Ibisca Niugini, PNG 25 - 27. x. 2012 Mount Wilhelm 1700 m - 5,759910107 145,234726 MW 1700 / P 1886 Vial 07540 " (ZSM). 1 female " Ibisca Niugini, PNG 28 - 30. x. 2012 Mount Wilhelm 2200 m ", " - 5,75897789 145,1860657 FIT-MW 2200 - F- 7 / 8 - d 13 / Plot 6 / P 2314 Vial 07373 - CODYTI " (ZSM). 1 female " Ibisca Niugini, PNG 27 - 29. x. 2012 Mount Wilhelm 2200 m ", " - 5,75897789 145,1860657 FIT-MW 2200 - K- 6 / 8 - d 12 / Plot 11 / P 2353 Vial 11687 " (ZSM). 1 female " Ibisca Niugini, PNG 28 - 30. x. 2012 Mount Wilhelm 2200 m - 5,760178089 145,186264 MW 2200 / P 2282 Vial 07152 " (ZSM). 2 females " Ibisca Niugini, PNG 29 - 31. x. 2012 Mount Wilhelm 2200 m - 5,760178089 145,186264 FIT-MW 2200 - N- 7 / 8 - d 14 / Plot 14 / P 2378 Vial 15681 " (ZSM). 2 females " Ibisca Niugini, PNG 27 - 29. x. 2012 Mount Wilhelm 2200 m FIT-MW 2200 - O- 6 / 8 - d 12 / Plot 15 / P 2385 Vial ... - CODYTI " (ZSM). 1 female " 04103 Ibisca Niugini 2012 " (ZSM). 1 female " 04103 " (ZSM).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
6A26D3C073EB86450E76BD39C3B6E4B5.taxon	diagnosis	Diagnosis. For complete description, see Balke (1999: 274). Beetle medium-sized: TL-H 3.5 - 4.3 mm, oblong-oval; reddish brown to brown, with paler pronotum; submatt, with dense, rather coarse punctation and rather strongly impressed microreticulation; pronotum with distinct lateral bead; male antennae simple (Fig. 15); male protarsomere 4 with anterior angle slightly expanded, with large, thick, strongly curved anterolateral hook-like seta; male protarsomere 5 slightly concave ventrally, with anterior band of ca 100 and posterior band ca 40 of relatively long setae (Fig. 39 D); median lobe in lateral view evenly broad, with rounded, not curved downwards apex, in ventral view, evenly tapering, with broadly pointed apex; paramere slightly concave on dorsal side and with long, dense, thin setae, situated along dorsal margin; subdistal setae strong and dense, setae in middle part slightly shorter and sparser, proximal setae long but sparser than subdistal ones (Fig. 39 A-C).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
6A26D3C073EB86450E76BD39C3B6E4B5.taxon	distribution	Distribution. Papua New Guinea: East Sepik, Simbu, Western Highlands, and Eastern Highlands Provinces (Fig. 50).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
D7657B421ACAF4D00F17D9CE636622BE.taxon	diagnosis	Diagnosis. For complete description, see Balke (1998: 330). Beetle medium-sized: TL-H 3.7 - 4.4 mm; oblong-oval; brown to dark brown, with reddish brown pronotal sides, head, and sometimes also sides of elytra, in some specimens, disc of pronotum and elytron almost piceous; submatt, with fine but rather dense punctation and strongly impressed microreticulation; pronotum without lateral bead; male antennae simple (Fig. 3); male protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta; male protarsomere 5 long and narrow, with anterior band of more than 60 and posterior row of 6 relatively long, thin setae (Fig. 27 D); median lobe in lateral view slightly curved, with apex curved downwards, with visible angle on dorsal side, in ventral view, distally distinctly narrowed to truncate apex; paramere slightly concave on dorsal side and with long, dense, thin setae, situated along dorsal margin: subdistal setae strong and dense, setae in middle part shorter and sparser, proximal setae long, only slightly sparser than subdistal ones (Fig. 27 A-C).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
D7657B421ACAF4D00F17D9CE636622BE.taxon	distribution	Distribution. Papua: Pegunungan Bintang. The species is known only from the type locality (Fig. 50).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
7A837C49741C199DF4838F3819531988.taxon	diagnosis	Diagnosis. For complete description, see Balke (1998: 331). Beetle medium-sized: TL-H 4.35 - 4.9 mm; broader, oblong-oval; dark brown, with reddish brown pronotal sides and head anteriorly, in some specimens, disc of pronotum and elytron almost piceous; submatt, with very fine, on elytra often almost invisible punctation and strongly impressed microreticulation; pronotum with lateral bead; male antennae simple (Fig. 16); male protarsomere 4 with anterior angle very slightly expanded, with large, thick, strongly curved anterolateral hook-like seta; male protarsomere 5 long and narrow, slightly concave ventrally, with anterior band of ca 70 and posterior band of ca 30 relatively long setae (Fig. 40 D); median lobe in lateral view evenly broad, with rounded, not curved downwards, only slightly thickened apex, in ventral view, with subparallel sides and roundly truncate apex; paramere slightly concave on dorsal side and with long, dense, thin setae, situated along dorsal margin distinctly divided to dense and strong subdistal setae and sparser proximal ones, setae in middle short and fine (Fig. 40 A-C).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
7A837C49741C199DF4838F3819531988.taxon	distribution	Distribution. Papua New Guinea: Morobe Province. The species is known only from the type locality, Wagau in Herzog Range (Fig. 50).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
5764CC89BD27F3E8DC91B2D15F98A621.taxon	description	Description. Body size and form: Beetle size variable but generally beetle medium-sized: TL-H 3.45 - 4.3 mm, TL 3.75 - 4.8 mm, MW 1.8 - 2.25 mm (holotype: TL-H 4.15 mm, TL 3.75 mm, MW 2 mm), with oblong-oval habitus, slightly more attenuated posteriorly. Coloration: Brown to piceous, with head and pronotum paler. Head reddish brown to piceous, darker posteriorly. pronotum reddish brown to piceous, broadly paler on lateral sides and sometimes also narrowly anteriorly and posteriorly. Elytra uniformly brown to piceous. Head appendages and legs yellowish red to reddish brown, legs distally darker, especially metathoracic legs (Fig. 4). Teneral specimens paler. Surface sculpture: Submatt dorsally. Head with rather dense punctation (spaces between punctures 1 - 2 times size of punctures), evidently finer and sparser anteriorly; diameter of punctures smaller than diameter of cells of microreticulation. Pronotum and elytra with fine but rather dense punctation, sparser and finer than on head. Pronotum and elytra with strongly impressed microreticulation. Head with microreticulation stronger. Metaventrite and metacoxae distinctly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal ventrites with distinct microreticulation, strioles, and very fine sparse punctation. Structures: Pronotum without lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, rounded anteriorly. Blade of prosternal process lanceolate, relatively narrow, slightly convex, with distinct lateral bead and few setae. Abdominal ventrite 6 broadly rounded or slightly truncate. Male: Antennae simple (Fig. 4). Protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 long and narrow, with anterior band of more than 80 and posterior row of 11 relatively long, thin setae (Fig. 28 D). Median lobe in lateral view slightly curved, its apex rounded and not or only very slightly curved downwards; in ventral view, distally distinctly narrowed to truncate apex. Paramere slightly concave on dorsal side and with long, dense, thin setae situated along dorsal margin: subdistal setae denser, proximal setae sparser, setae in middle shorter, thinner (Fig. 28 A-C). Abdominal ventrite 6 with 7 - 11 lateral striae on each side. Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
5764CC89BD27F3E8DC91B2D15F98A621.taxon	distribution	Distribution. Papua New Guinea: Sandaun Province (Fig. 50).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
5764CC89BD27F3E8DC91B2D15F98A621.taxon	etymology	Etymology. The species is named for Sentiko Ibalim, one of the great young PNG entomologists, who collected most of these beetles. The species name is a noun in the genitive case.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
3967073F196D007E333101A799C3431E.taxon	description	Description. Body size and form: Beetle small: TL-H 3.15 - 3.65 mm, TL 3.45 - 4.0 mm, MW 1.65 - 1.9 mm (holotype: TL-H 3.65 mm, TL 4 mm, MW 1.85 mm), with oblong habitus. Coloration: Reddish to reddish brown, with head and pronotum paler. Head yellowish red to reddish brown, with small darker areas posterior to eyes. Pronotum yellowish red to reddish brown, with small brown to dark brown area on disc. Elytra reddish brown to brown, with narrow reddish sutural lines. Head appendages yellowish red, legs reddish, distally darker, especially metathoracic legs (Fig. 6). Teneral specimens paler. Surface sculpture: Submatt dorsally. Head with rather dense punctation (spaces between punctures 1 - 2 times size of punctures), evidently finer and sparser anteriorly; diameter of punctures smaller than diameter of cells of microreticulation. Pronotum and elytra with dense, distinct but fine punctation, sparser and finer than on head. Pronotum and elytra with strongly impressed microreticulation. Head with microreticulation stronger. Metaventrite and metacoxae distinctly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal ventrites with distinct microreticulation, strioles, and very fine sparse punctation. Structures: Pronotum without lateral bead, in few specimens with its traces in posterior part. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively narrow, slightly convex, with distinct lateral bead and few setae. Abdominal ventrite 6 rounded. Male: Antennae simple (Fig. 6). Protarsomere 4 with anterolateral seta thin, long, smaller than more laterally situated large setae, slightly curved downwards. Protarsomere 5 long and narrow, with anterior band of more than 40 and posterior row of 7 relatively long, thin setae (Fig. 30 D). Median lobe in lateral view almost straight, its apex rounded and not curved downwards; in ventral view, distally distinctly narrowed before rounded, narrow apex. Paramere slightly concave on dorsal side and with dorsal setae distinctly divided to long, dense subdistal setae and sparser proximal ones (Fig. 30 A-C). Abdominal ventrite 6 with 3 - 5 lateral striae on each side. Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
3967073F196D007E333101A799C3431E.taxon	distribution	Distribution and variability. Papua New Guinea: Madang and Eastern Highlands (Fig. 50). The species is known mainly from Keki area in Adelbert Mountains; only one beetle was collected in Bena, Eastern Highlands. It shows no morphological difference from the specimens of Keki populations, except for a small difference in the median lobe shape, which could be an expression of species variability. The species might have the same pattern of distribution as Exocelina brahminensis Shaverdo et al., 2012, which has a wide distribution in the Momase Region and is known from Adelbert Mountains and Bena.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
3967073F196D007E333101A799C3431E.taxon	etymology	Etymology. The species is named after Keki Village. The name is a noun in the nominative singular standing in apposition.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
CDD3DC151D4DE56133D6837B59766236.taxon	description	Description. Body size and form: Beetle large: TL-H 5.4 mm, TL 6.0 mm, MW 2.9 mm, with broader, oblong-oval habitus. Coloration: Piceous, with paler pronotum. Head piceous, narrowly brownish anteriorly and with two vague brownish spots between eyes. Pronotum dark brown, piceous on disc. Elytra piceous, with vague narrow brownish sutural lines. Head appendages and legs proximally reddish brown, legs distally darker, brownish, especially metathoracic legs (Fig. 25). Surface sculpture: Submatt dorsally. Head with dense, coarse punctation (no spaces between punctures or spaces of equal size of punctures), finer and sparser anteriorly; diameter of punctures equal to diameter of cells of microreticulation. Pronotum with relatively dense but fine punctation, sparser and finer than on head. Elytra with finer punctation than on pronotum. Pronotum and elytra with strongly impressed microreticulation. Head with microreticulation stronger. Metaventrite and metacoxae distinctly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles, abdominal ventrites with distinct microreticulation and strioles. Metaventrite medially, metacoxal plates, and abdominal ventrites with fine, sparse punctation. Structures: Pronotum with distinct lateral bead. Its lateral sides with distinct longitudinal impressions. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively short, broad, slightly convex and smooth in the middle, with distinct lateral bead and few lateral setae, lateral sides flattened. Abdominal ventrite 6 rounded. Male: Antennae simple (Fig. 25). Protarsomere 4 with anterior angle slightly expanded, with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 slightly concave ventrally, with anterior band of ca 100 and posterior band of ca 40 relatively long setae (Fig. 49 D). Median lobe in lateral view long, slightly curved, with small, very slightly bent downwards, thickened apex; in ventral view, evenly tapering to broadly pointed apex. Paramere slightly concave on dorsal side and with weak dorsal setation, setae on subdistal part stronger, denser, more evident than proximal setae (Fig. 49 A-C). Abdominal ventrite 6 with 17 - 18 lateral striae on each side. Female: Unknown.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
CDD3DC151D4DE56133D6837B59766236.taxon	distribution	Distribution. Papua New Guinea: Enga Province. The species is known only from the type locality (Fig. 50).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
CDD3DC151D4DE56133D6837B59766236.taxon	etymology	Etymology. The species is named after Kumul Lodge. The name is an adjective in the nominative singular.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
1159C69C8C01C6557567F405BDAAD44B.taxon	description	Description. Body size and form: Beetle large: TL-H 4.8 - 5.5 mm, TL 5.3 - 5.9 mm, MW 2.6 - 2.75 mm (holotype: TL-H 4.8 mm, TL 5.3 mm, MW 2.6 mm), with broader, oblong-oval habitus. Coloration: Piceous. Head piceous, narrowly brownish anteriorly and sometimes with two brownish spots between eyes. Pronotum piceous, brownish laterally and anteriorly. Elytra piceous, sometime with narrow brownish sutural lines. Head appendages and legs proximally reddish brown, legs distally darker, brownish, especially metathoracic legs (Fig. 24). Surface sculpture: Submatt dorsally. Head with relatively dense punctation (no spaces between punctures or spaces 2 times size of punctures), sparser anteriorly, denser and coarser between eyes; diameter of punctures smaller than or equal to diameter of cells of microreticulation. Pronotum with relatively dense but fine punctation, sparser and finer than on head. Elytra with very fine, sparse punctation. Pronotum and elytra with rather strongly impressed microreticulation. Head with microreticulation stronger. Metaventrite and metacoxae distinctly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles, abdominal ventrites with distinct microreticulation and strioles. Metaventrite medially, metacoxal plates, and abdominal ventrites with very fine, sparse punctation. Structures: Pronotum with distinct lateral bead. Its lateral sides with distinct longitudinal impressions. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively narrow, slightly convex, and smooth, with distinct lateral bead and few lateral setae. Abdominal ventrite 6 slightly truncate. Male: Antennae simple (Fig. 24). Protarsomere 4 with anterior angle slightly expanded, with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 with anterior band of more than 60 and posterior row of 16 relatively long setae (Fig. 48 D). Median lobe in lateral view long, slightly curved, with small, slightly bent downwards, thickened apex; in ventral view, evenly tapering to broadly pointed apex. Paramere slightly concave on dorsal side and with weak dorsal setation, setae on subdistal part stronger and denser than proximal setae (Fig. 48 A-C). Abdominal ventrite 6 with 23 - 27 lateral striae on each side. Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
1159C69C8C01C6557567F405BDAAD44B.taxon	distribution	Distribution. Papua New Guinea: Southern Highlands Province. The species is known only from the type locality (Fig. 50).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
1159C69C8C01C6557567F405BDAAD44B.taxon	etymology	Etymology. The species is named after Mendi Village. The name is an adjective in the nominative singular.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
0FA25962B8B22EAF53ED8A1E28528DA9.taxon	description	Description. Body size and form: Beetle medium-sized: TL-H 4.25 mm, TL 4.8 mm, MW 2.2 mm, with oblong-oval habitus. Coloration: Brown, with reddish pronotum. Head brown, with slightly darker areas posterior to eyes. Pronotum broadly reddish laterally and dark brown medially from anterior to posterior margins. Elytra uniformly brown, in the middle with traces of narrow reddish sutural lines. Head appendages and legs proximally yellowish red, legs distally darker, reddish brown, especially metathoracic legs (Fig. 20). Surface sculpture: Matt dorsally. Head with rather dense, coarse punctation (spaces between punctures 1 - 2 times size of punctures), evidently finer and sparser anteriorly; diameter of punctures smaller than or equal to diameter of cells of microreticulation. Pronotum and elytra with distinct punctation, sparser and finer than on head. Pronotum and elytra with strongly impressed microreticulation. Head with microreticulation stronger. Metaventrite and metacoxae distinctly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal ventrites with distinct microreticulation, strioles, and very fine sparse punctation. Structures: Pronotum with distinct lateral bead. Its lateral sides with longitudinal impressions. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively short, broad, slightly convex and smooth in the middle, with distinct lateral bead and few lateral setae, lateral sides slightly flattened. Abdominal ventrite 6 slightly truncate. Male: Antennae simple (Fig. 20). Protarsomere 4 with large, thick, slightly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior band of more than 50 and posterior row of 13 relatively long, thin setae (Fig. 44 D). Median lobe in lateral view narrowed to apex, with thin, slightly curved upwards apex; in ventral view, narrowed before apex, with apex rounded, of shape of a baby pacifier, with distinct ventral impression. Paramere slightly concave on dorsal side and with long, dense, thin setae situated along dorsal margin, subdistal setae denser, proximal setae much sparser, setae in middle shorter, thinner (Fig. 44 A-C). Abdominal ventrite 6 with 19 - 22 lateral striae on each side. Female: Unknown.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
0FA25962B8B22EAF53ED8A1E28528DA9.taxon	distribution	Distribution. Papua New Guinea: Morobe Province. The species is known only from the type locality (Fig. 50).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
0FA25962B8B22EAF53ED8A1E28528DA9.taxon	etymology	Etymology. The species is named after Menyamya Village. The name is a noun in the nominative singular standing in apposition.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
E66B12B83A0A26924F06AF742F68AA92.taxon	diagnosis	Diagnosis. For complete description, see Balke (1999: 274 - 275). Beetle small: TL-H 3.2 - 3.7 mm; oblong-oval, more strongly attenuated posteriorly; reddish to reddish brown, with head and pronotum slightly paler; shiny, with very fine, sparse punc tation, almost invisible on elytra and weakly impressed microreticulation; pronotum without lateral bead, sometimes with its traces in posterior part; male antennae simple (Fig. 5); male protarsomere 4 with anterolateral seta thin, long, smaller than more laterally situated large setae, slightly curved downwards; male protarsomere 5 long and narrow, with more than 80 relatively long, thin setae, which divided in anterior and posterior ones proximally and mixed up together in distal half of tarsomere (Fig. 29 D); median lobe in lateral view almost straight, its apex rounded and not curved downwards, in ventral view, distally narrowed before apex, apex broad, slightly rounded; paramere slightly concave on dorsal side and with dorsal setae distinctly divided into long, dense subdistal setae and sparser, rather inconspicuous proximal ones, setae in middle part short and fine (Fig. 29 A-C).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
E380B07AE33E715F4245CFAB7BC42A67.taxon	materials_examined	Additional material. 3 females " Papua New Guinea: Eastern Highlands, Hano kotu, Kimiagomo, Okapa, 1661 m, 11. viii. 2006, 06 25.096 S 145 34.556 E, K. Sagata (WB 129) " (ZSM).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
E380B07AE33E715F4245CFAB7BC42A67.taxon	description	Description. Body size and form: Beetle medium-sized: TL-H 3.95 - 4.7 mm, TL 4.3 - 5.05 mm, MW 2.05 - 2.5 mm (holotype: TL-H 4.3 mm, TL 4.7 mm, MW 2.3 mm), with oblong-oval habitus, slightly more attenuated posteriorly. Coloration: Piceous, with reddish brown pronotum. Head reddish brown to piceous, paler anteriorly and darker posterior to eyes. Pronotum dark brown to piceous, with reddish to reddish brown sides narrowly or broadly. Elytra dark brown to piceous, sometime with narrow reddish sutural lines. Head appendages and legs proximally yellowish red, legs distally darker, reddish brown, especially metathoracic legs (Fig. 23). Teneral specimens paler. Surface sculpture: Shiny dorsally. Head mostly with fine, sparse punctation (spaces between punctures 2 - 3 times size of punctures) but punctation denser and coarser between eyes; diameter of punctures smaller than diameter of cells of microreticulation. Pronotum and elytra with very fine, sparse punctation, sometimes inconspicuous on elytra. Pronotum and elytra with weakly impressed microreticulation. Head with microreticulation stronger. Metaventrite and metacoxae distinctly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles, abdominal ventrites with distinct microreticulation and strioles. Metaventrite medially, metacoxal plates, and abdominal ventrites with very fine, sparse, often inconspicuous punctation. Structures: Pronotum with distinct lateral bead. Its lateral sides with distinct longitudinal impressions. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively narrow, slightly convex, and smooth, with distinct lateral bead and few lateral setae. Abdominal ventrite 6 rounded. Male: Antennae simple (Fig. 23). Protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior band of ca 60 and posterior row of 20 relatively long, thin setae (Fig. 47 D). Median lobe in lateral view slightly curved, with apex dully pointed, slightly bent downwards; in ventral view, broadened subdistally, with broad, rounded apex. Paramere slightly concave on dorsal side and with dense setae on subdistal part; proximal setae inconspicuous (Fig. 47 A-C). Abdominal ventrite 6 with 8 - 12 lateral striae on each side. Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
E380B07AE33E715F4245CFAB7BC42A67.taxon	distribution	Distribution. Papua New Guinea: Eastern Highlands Province (Fig. 50).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
E380B07AE33E715F4245CFAB7BC42A67.taxon	etymology	Etymology. The species is named after Okapa Station. The name is a noun in the nominative singular standing in apposition.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
9CBCB5355763015E522E6F3B8AE0D130.taxon	description	Description. Body size and form: Beetle small: TL-H 3.5 - 3.6 mm, TL 3.9 - 4.0 mm, MW 1.95 - 2.0 mm (holotype: TL-H 3.6 mm, TL 4 mm, MW 1.95 mm), with oblong-oval habitus. Coloration: Specimens teneral. Reddish brown, with pronotum slightly paler. Head reddish brown, with small darker areas posterior to eyes. Pronotum yellowish brown, darker on disc. Elytra reddish brown. Head appendages yellowish red, legs reddish, distally darker, especially metathoracic legs (Fig. 9). Surface sculpture: Submatt dorsally. Head with fine, sparse punctation (spaces between punctures 2 - 3 times size of punctures), only with some larger punctures between eyes; diameter of punctures smaller than diameter of cells of microreticulation. Pronotum and elytra with very fine, sparse punctation. Pronotum and elytra with rather strongly impressed microreticulation. Head with microreticulation stronger. Metaventrite and metacoxae distinctly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal ventrites with distinct microreticulation, strioles, and very fine, sparse punctation. Structures: Pronotum with narrow lateral bead. Its lateral sides with inconspicuous, shallow longitudinal impressions. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively broad, slightly convex, with distinct lateral bead and few setae. Abdominal ventrite 6 rounded. Male: Antennae simple (Fig. 9). Protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 long and narrow, with anterior band of ca 40 and posterior row of 6 relatively long, thin setae (Fig. 33 D). Median lobe in lateral view slightly curved, broad, with apex bent downwards, not distinctly thickened, slightly angulate; in ventral view, almost subparallel and distally distinctly narrowed to apex, apex roundly truncate, of shape of a baby pacifier. Paramere slightly concave on dorsal side, with long, dense dorsal setae: subdistal setae strong and dense, setae in middle part shorter and sparser, proximal setae longer and stronger than subdistal ones (Fig. 33 A-C). Abdominal ventrite 6 with 9 - 10 lateral striae on each side. Female: Unknown.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
9CBCB5355763015E522E6F3B8AE0D130.taxon	distribution	Distribution. Papua New Guinea: East Sepik Province (Fig. 50).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
9CBCB5355763015E522E6F3B8AE0D130.taxon	etymology	Etymology. The species is named for Pius, a local collector. The species name is a noun in the genitive case.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
408C1A62210AF103992CD4019BFB43B5.taxon	description	Description. Body size and form: Beetle medium-sized: TL-H 3.3 - 3.6 mm, TL 3.6 - 3.95 mm, MW 1.8 - 2.0 mm (holotype: TL-H 3.3 mm, TL 3.6 mm, MW 1.9 mm), with oblong-oval habitus. Coloration: Reddish to reddish brown, with head and pronotum paler. Head yellowish red to reddish brown, with small darker areas posterior to eyes. Pronotum yellowish red to reddish brown, darker (to brown) on disc. Elytra reddish brown to brown, sometimes with narrow yellowish or reddish sutural lines. Head appendages and legs yellowish red, legs distally darker, especially metathoracic legs (Fig. 7). All specimens are slightly teneral, therefore, the species coloration might be more darker. Surface sculpture: Shiny dorsally. Head with rather dense punctation (spaces between punctures 1 - 2 times size of punctures), evidently finer and sparser anteriorly; diameter of punctures smaller than diameter of cells of microreticulation or equal to it. Pronotum and elytra with very distinct punctation, sparser and slightly finer than on head. Pronotum and elytra with weakly impressed microreticulation. Head with microreticulation stronger. Metaventrite and metacoxae distinctly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal ventrites with distinct microreticulation, strioles, and very fine sparse punctation. Structures: Pronotum without lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively narrow, slightly convex, with distinct lateral bead and few setae. Abdominal ventrite 6 rounded. Male: Antennae simple (Fig. 7). Protarsomere 4 with anterolateral seta thin, long, smaller than more laterally situated large setae, slightly curved downwards. Protarsomere 5 long and narrow, with anterior band of ca 40 and posterior row of 10 relatively long setae (Fig. 32 D). Median lobe in lateral view slightly curved, its apex strongly bent downwards, with visible angle on dorsal side; in ventral view, almost subparallel and distally narrowed to truncate apex. Paramere slightly concave on dorsal side and with long, dense, thin setae, situated along dorsal margin: subdistal setae denser than setae in middle and proximal parts (Fig. 32 A-C). Abdominal ventrite 6 with 5 - 8 lateral striae on each side. Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
408C1A62210AF103992CD4019BFB43B5.taxon	distribution	Distribution. Papua New Guinea: Madang (Fig. 50).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
408C1A62210AF103992CD4019BFB43B5.taxon	etymology	Etymology. The species is named " pseudofume " because shape of its median lobe remains that of E. fume. The name is a noun in the nominative singular standing in apposition.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
3EC65D8496E0B3A1939C22A6FE6ED529.taxon	description	Description. Body size and form: Beetle small: TL-H 3.25 - 3.55 mm, TL 3.65 - 3.95 mm, MW 1.8 - 1.85 mm (holotype: TL-H 3.5 mm, TL 3.85 mm, MW 2.0 mm), with oblong to oblong-oval habitus. Coloration: Reddish brown to dark brown, with head and pronotum paler. Head reddish to reddish brown, with small darker areas posterior to eyes. Pronotum reddish to reddish brown, with dark brown disc. Elytra brown to dark brown, with narrow reddish sutural lines. Head appendages yellowish red, legs reddish, distally darker, especially metathoracic legs (Fig. 18). Teneral specimens paler. Surface sculpture: Submatt dorsally. Head with dense, coarse punctation (no spaces between punctures or spaces of equal size of punctures), evidently finer and sparser anteriorly; diameter of punctures equal to or larger than diameter of cells of microreticulation. Pronotum and elytra with dense and coarse punctation, sparser and finer than on head. Pronotum and elytra with rather strongly impressed microreticulation. Head with microreticulation stronger. Metaventrite and metacoxae distinctly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal ventrites with distinct microreticulation, strioles, and fine sparse punctation. Structures: Pronotum with narrow lateral bead. Its lateral sides with distinct longitudinal impressions. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively broad, slightly convex, with distinct lateral bead and few setae. Abdominal ventrite 6 slightly truncate or very slightly concave. Male: Antennae simple (Fig. 18). Protarsomere 4 with anterior angle slightly expanded, with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 long and narrow, with anterior band of ca 40 and posterior row of 13 relatively long, thin setae (Fig. 42 D). Median lobe in lateral view simple, slightly curved; in ventral view, evenly tapering to broadly pointed apex, side of apex slightly thickened. Paramere slightly concave on dorsal side and with dense setae on subdistal part; proximal setae finer (Fig. 42 A-C). Abdominal ventrite 6 with 9 - 10 lateral striae on each side. Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
3EC65D8496E0B3A1939C22A6FE6ED529.taxon	etymology	Etymology. The species was mistaken for E. pusilla sp. n. due to their similarity. The name is a noun in the nominative singular standing in apposition.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
0E315859AACF9251F923B2F21834DA51.taxon	description	Description. Body size and form: Beetle small: TL-H 2.95 - 3.25 mm, TL 3.2 - 3.6 mm, MW 1.65 - 1.85 mm (holotype: TL-H 2.95 mm, TL 3.25 mm, MW 1.65 mm), with oblong-oval habitus. Coloration: Reddish to reddish brown, with head and pronotum paler. Head yellowish red to reddish, with small darker areas posterior to eyes. Pronotum yellowish red to reddish, with small brownish area on disc. Elytra reddish brown to brown, with narrow reddish sutural lines. Head appendages yellowish red, legs reddish, distally darker, especially metathoracic legs (Fig. 10). Teneral specimens paler. Surface sculpture: Shiny dorsally. Head with dense, coarse punctation (no spaces between punctures or spaces 2 times size of punctures), evidently finer and sparser anteriorly; diameter of punctures equal to or larger than diameter of cells of microreticulation. Pronotum and elytra with dense and coarse punctation, sparser and finer than on head. Pronotum and elytra with weakly impressed microreticulation. Head with microreticulation stronger. Metaventrite and metacoxae distinctly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal ventrites with distinct microreticulation, strioles, and fine sparse punctation. Structures: Pronotum with narrow lateral bead. Its lateral sides with longitudinal impressions. Base of prosternum and neck of prosternal process with distinct ridge, rounded anteriorly. Blade of prosternal process lanceolate, relatively broad, slightly convex, with distinct lateral bead and few setae. Abdominal ventrite 6 slightly truncate or very slightly concave. Male: Antennae simple (Fig. 10). Protarsomere 4 with anterior angle slightly expanded and large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 long and narrow, with anterior band of ca 40 and posterior row of 12 relatively long, thin setae (Fig. 35 D). Median lobe in lateral view simple, slightly curved; in ventral view, evenly tapering to broadly pointed apex, side of apex slightly thickened. Paramere slightly concave on dorsal side and with dense setae on subdistal part; proximal setae finer and much sparser (Fig. 35 A-C). Abdominal ventrite 6 with 5 - 10 lateral striae on each side. Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
0E315859AACF9251F923B2F21834DA51.taxon	distribution	Distribution. Papua New Guinea: Madang and Simbu Provinces (Fig. 50).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
0E315859AACF9251F923B2F21834DA51.taxon	etymology	Etymology. The species name derives from the Latin " pusillus " (small, tiny) to express small size of these beetles. The species name is an adjective in the nominative singular.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
0570DC3AC03825F5BF21C959CD029DCD.taxon	description	Description. Body size and form: Beetle small: TL-H 3.6 mm, TL 4.0 mm, MW 2.0 mm, with broader, oval habitus. Coloration: Reddish brown head and pronotum and piceous elytra. Head reddish in its anterior half and dark brown in posterior one. Pronotum dark brown on disc and gradually paler to yellowish red laterally. Elytra dark brown, paler laterally and almost piceous on disc. Head appendages yellowish red, legs reddish, distally darker, especially metathoracic legs (Fig. 8). Surface sculpture: Shiny dorsally. Head with rather dense punctation (spaces between punctures 1 - 2 times size of punctures) but fine punctation; diameter of punctures smaller than diameter of cells of microreticulation. Pronotum and elytra with distinct punctation, sparser and finer punctation than on head. Pronotum and elytra with weakly impressed microreticulation. Head with microreticulation stronger. Metaven trite and metacoxae distinctly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal ventrites with distinct microreticulation, strioles, and very fine sparse punctation. Structures: Pronotum without lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, rounded anteriorly. Blade of prosternal process lanceolate, elongate, relatively broad, slightly convex, with distinct lateral bead and few. Abdominal ventrite 6 rounded. Male: Antennae simple (Fig. 8). Protarsomere 4 with anterolateral seta long and thin, equal to more laterally situated large setae, slightly curved downwards. Protarsomere 5 long and narrow, with anterior band of more than 40 and posterior row of 12 relatively long, thin setae (Fig. 31 D). Median lobe in lateral view short, slightly curved, with enlarged, rounded, not bent downwards apex; in ventral view, narrow, subparallel, and with truncate apex. Paramere very slightly concave on dorsal side and with long, dense, thin setae, situated along dorsal margin: subdistal setae strong and dense, setae in middle part shorter and sparser, proximal setae long but sparser than subdistal ones (Fig. 31 A-C). Abdominal ventrite 6 without lateral striae on each side, except one with setae. Female: Unknown.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
0570DC3AC03825F5BF21C959CD029DCD.taxon	distribution	Distribution. Papua New Guinea: Simbu and Eastern Highlands Provinces, Crater Mountain. This species is known only from the type locality (Fig. 50).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
0570DC3AC03825F5BF21C959CD029DCD.taxon	etymology	Etymology. The species is named after Sima River. The name is a noun in the nominative singular standing in apposition.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
139897831E9A021F68721075000F3FAC.taxon	description	Description. Body size and form: Beetle small to medium-sized: TL-H 3.3 - 4.15 mm, TL 3.65 - 4.5 mm, MW 1.8 - 2.15 mm (holotype: TL-H 3.9 mm, TL 4.25 mm, MW 2.0 mm), with rather oblong habitus. Coloration: Dark brown to piceous, sometimes with reddish pronotum. Head dark brown to piceous, paler anteriorly. Pronotum dark brown to piceous, with reddish brown lateral sides and sometimes anteriorly and narrowly posteriorly. Elytra dark brown to piceous, sometimes with narrow reddish sutural lines. Head appendages and legs proximally yellowish red, legs distally darker, reddish (Fig. 19). Surface sculpture: Matt dorsally. Head with dense punctation (no spaces between punctures or spaces 2 times size of punctures), finer and sparser anteriorly; diameter of punctures equal to diameter of cells of microreticulation. Pronotum and elytra with slightly sparser and finer punctation than on head. Head, pronotum and elytra with strongly impressed microreticulation. Metaventrite and metacoxae distinctly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal ventrites with distinct microreticulation, strioles, and fine, sparse but distinct punctation. Structures: Pronotum with distinct lateral bead. Its lateral sides with distinct longitudinal impressions. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively narrow, slightly convex, with distinct lateral bead and few setae. Abdominal ventrite 6 broadly rounded. Male: Antennomere simple, slightly stout (Fig. 19). Protarsomere 4 with anterior angle slightly expanded, with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 long and narrow, slightly concave ventrally, with anterior band of ca 40 and irregular posterior row of 17 relatively long setae (Fig. 43 D). Median lobe in lateral view slightly curved, its apex thickened, bent downwards; in ventral view, slightly broadened medially, with broadly pointed apex. Paramere very slightly concave on dorsal side and with dense setae on subdistal part; proximal setae inconspicuous (Fig. 43 A-C). Abdominal ventrite 6 with 7 - 12 lateral striae on each side. Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae. Variability. The specimens from Ineng River are larger: TL-H 3.5 - 4.15 mm; remaining specimens: TL-H 3.3 - 3.7 mm. In addition, the population from Ineng River shows variability in how strongly apex of the median lobe bent downwards in lateral view.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
139897831E9A021F68721075000F3FAC.taxon	distribution	Distribution. Papua New Guinea: Western Highlands Province, near Simbai (Fig. 50).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
139897831E9A021F68721075000F3FAC.taxon	etymology	Etymology. The species is named after Simbai area. The name is an adjective in the nominative singular.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
34DE0731998033B0E14AC5C5B8DF9603.taxon	materials_examined	Additional material. 1 female " Papua New Guinea: Western Highlands, Simbai, 1800 - 2000 m, 25. ii. 2007, 05.16.330 S 144.33.176 E, Kinibel (PNG 133) " (ZSM). 1 female " Papua New Guinea: Western Highlands, Simbai, 1800 - 2000 m, 1. iii. 2007, 05.14.2760 S 144.28.741 E, Kinibel (PNG 138) " (ZSM).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
34DE0731998033B0E14AC5C5B8DF9603.taxon	description	Description. Body size and form: Beetle medium-sized: TL-H 4.05 - 5.0 mm, TL 4.4 - 5.4 mm, MW 2.1 - 2.55 mm (holotype: TL-H 5.0 mm, TL 5.4 mm, MW 2.5 mm), with oblong-oval habitus. Coloration: Piceous, with head and pronotum paler. Head dark brown to piceous, reddish brown to brown anteriorly, with small darker areas posterior to eyes. Pronotum reddish brown to brown laterally and piceous on disc. Elytra brown to piceous, sometimes with narrow reddish sutural lines. Head appendages and legs proximally reddish, legs distally darker, brown, especially metathoracic legs (Fig. 17). Teneral specimens paler. Surface sculpture: Submatt dorsally. Head with rather dense, coarse punctation (spaces between punctures 1 - 2 times size of punctures), evidently finer and sparser anteriorly; diameter of punctures smaller than or equal to diameter of cells of microreticulation. Pronotum with distinct punctation, sparser and finer than on head. Elytra with very fine and sparse punctation. Pronotum and elytra with strongly impressed microreticulation. Head with microreticulation stronger. Metaventrite and metacoxae distinctly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal ventrites with distinct microreticulation, strioles, and fine, sparse punctation. Structures: Pronotum with distinct lateral bead. Its lateral sides with longitudinal impressions. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively narrow, slightly convex, and smooth, with distinct lateral bead and few lateral setae. Abdominal ventrite 6 slightly truncate or broadly rounded. Male: Antennae simple (Fig. 17). Protarsomere 4 with anterior angle slightly expanded, with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 long and narrow, slightly concave ventrally, with anterior band of more than 80 and posterior band of ca 30 relatively long setae (Fig. 41 D). Median lobe in lateral view evenly broad, with rounded, slightly angulated, thickened apex; in ventral view, almost subparallel, with broadly rounded apex, with thickened margins. Paramere slightly concave on dorsal side and with long, dense, thin setae situated along dorsal margin: subdistal setae denser, proximal setae sparser, setae in middle finer (Fig. 41 A-C). Abdominal ventrite 6 with 9 - 14 lateral striae on each side. Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
34DE0731998033B0E14AC5C5B8DF9603.taxon	distribution	Distribution. Papua New Guinea: Western Highlands Province (Fig. 50).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
34DE0731998033B0E14AC5C5B8DF9603.taxon	etymology	Etymology. The species is named after Simbai-Jimi area. The name is a noun in the nominative singular standing in apposition.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
57575C123120C155ADC2ED0D04DD3D95.taxon	description	Description. Body size and form: Beetle small: TL-H 2.7 - 3.2 mm, TL 3.0 - 3.55 mm, MW 1.55 - 1.9 mm (holotype: TL-H 2.95 mm, TL 3.25 mm, MW 1.75 mm), with broader, oval habitus. Coloration: Brownish, with head and pronotum paler. Head yellowish red to reddish brown in anterior half and brown to dark brown in posterior ones. Pronotum yellowish red to reddish brown on sides, brown to dark brown on disc. Elytra brown to dark brown, with narrow reddish sutural lines. Head appendages yellowish red, legs reddish, distally darker, especially metathoracic legs (Fig. 11). Teneral specimens paler. Surface sculpture: Shiny dorsally. As in E. pusilla sp. n. but punctation finer and sparser and microreticulation weakly impressed. Structures: Pronotum with lateral bead. Its lateral sides with shallow longitudinal impressions. Base of prosternum and neck of prosternal process with distinct ridge, rounded anteriorly. Blade of prosternal process lanceolate, relatively broad, slightly convex, with distinct lateral bead and few setae. Abdominal ventrite 6 broadly rounded or slightly truncate. Male: Antennae simple (Fig. 11). Protarsomere 4 with medium-sized, thick, curved anterolateral hook-like seta. Protarsomere 5 long and narrow, with anterior row of 17 and posterior row of 6 relatively short setae (Fig. 34 D). Median lobe in lateral view slightly curved, with thickened, not curved downwards apex; in ventral view, distinctly narrowed subdistally, with roundly truncate apex. Paramere slightly concave on dorsal side and with dense setae on subdistal part; proximal setae finer and sparser (Fig. 34 A-C). Abdominal ventrite 6 with 5 - 8 lateral striae on each side. Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
57575C123120C155ADC2ED0D04DD3D95.taxon	distribution	Distribution. Papua: Puncak Regency. The species is known only from the Iratoi area (Fig. 50).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
57575C123120C155ADC2ED0D04DD3D95.taxon	etymology	Etymology. The species is named for our friend Bob Sumoked (Tomohon, Sulawesi). The species name is a noun in the genitive case.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
0B5149F72140AA3F08E731D92770F49B.taxon	description	Description. Body size and form: Beetle medium-sized: TL-H 3.45 - 4.0 mm, TL 3.85 - 4.45 mm, MW 1.85 - 2.15 mm (holotype: TL-H 3.9 mm, TL 4.3 mm, MW 2.0 mm), with oblong-oval habitus. Coloration: Dark brown to piceous, with paler pronotum. Head reddish to dark brown, paler anteriorly and posteriorly. Pronotum brown to dark brown, with reddish brown lateral sides and sometimes anteriorly and narrowly posteriorly, darker on disc, sometimes to piceous. Elytra brown to piceous, sometimes with narrow reddish sutural lines. Head appendages and legs proximally yellowish red, legs distally darker, reddish brown (Fig. 22). Surface sculpture: Submatt dorsally. Head with rather dense punctation (spaces between punctures 1 - 2 times size of punctures), finer and sparser anteriorly; diameter of punctures smaller than or equal to diameter of cells of microreticulation. Pronotum and elytra with sparser and finer punctation than on head. Head, pronotum and elytra with rather strongly impressed microreticulation. Metaventrite and metacoxae distinctly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal ventrites with distinct microreticulation, strioles, and fine, sparse but distinct punctation. Structures: Pronotum with distinct lateral bead. Its lateral sides with distinct longitudinal impressions. Base of prosternum and neck of prosternal process with distinct ridge, rounded anteriorly. Blade of prosternal process lanceolate, relatively broad, slightly convex, with distinct lateral bead and few setae. Abdominal ventrite 6 rounded. Male: Antennomere simple (Fig. 22). Protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta. Protarsomere 5 long and narrow, with narrow anterior band of ca. 40 and posterior row of 14 relatively long setae (Fig. 46 D). Median lobe in lateral view slightly curved, with apex dully pointed, slightly bent downwards; in ventral view, broadened subdistally, with broad, rounded apex. Paramere very slightly concave on dorsal side and with dense setae on subdistal part; proximal setae inconspicuous (Fig. 46 A-C). Abdominal ventrite 6 with 6 - 8 lateral striae on each side. Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
0B5149F72140AA3F08E731D92770F49B.taxon	distribution	Distribution. Papua New Guinea: Eastern Highlands Province (Fig. 50).	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
0B5149F72140AA3F08E731D92770F49B.taxon	etymology	Etymology. The species is named after Yoginofi Village. The name is a noun in the nominative singular standing in apposition.	en	Shaverdo, Helena, Sagata, Katayo, Balke, Michael (2018): Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae). ZooKeys 803: 7-70, DOI: http://dx.doi.org/10.3897/zookeys.803.28903, URL: http://dx.doi.org/10.3897/zookeys.803.28903
