taxonID	type	description	language	source
AC4D87E08634550F37A46B8E816AF822.taxon	diagnosis	Diagnosis. Small to medium sized Lithobiidae (body length, 13 – 30 mm) with antennae of 17 – 23 articles, generally 20; 4 – 15 ocelli on each side; 2 + 2 moderately small sharp coxosternal teeth; tergites without posterior triangular projections; coxal pores 2 – 7, arranged in one row; females gonopod with 2 + 2 – 5 + 5 spurs, tarsus 1 – tarsus 2 articulation of anterior legs (1 st – 13 th) faint but sometime showing obvious flexion, legs 14 and 15 swollen in males; females gonopod with a more or less massive expansion on the dorsolateral ridge of the second article projecting backwards over the third article; males gonopod short and small, of a single article. According to Zalesskaja (1978), among other genera / subgenera of Lithobiidae Hessebius is characterized chiefly by the massive expansion and projection of the dorsolateral ridge of the females gonopod (see also Eason 1981).	en	Pei, Sujian, Ma, Huiqin, Zapparoli, Marzio, Zhu, Mingsheng (2010): A review of the Chinese species of Hessebius Verhoeff, 1941 (Chilopoda: Lithobiomorpha: Lithobiidae). Zootaxa 2631: 51-61, DOI: 10.5281/zenodo.198338
AC4D87E08634550F37A46B8E816AF822.taxon	materials_examined	Type species. As no type-species was designated, Hessebius Verhoeff, 1941 is actually an unavailable name (Jeekel 2005: 19; Zapparoli 2006). In accordance with the diagnosis of the genus given by Verhoeff (1941), we herewith select Hessebius kosswigi Verhoeff, 1941, one of the two species originally included therein by the author, as type species of Hessebius by subsequent designation, validating here the name. Assigned species. H. armatus Verhoeff, 1943; H. barbipes (Porat, 1893) (= H. hatayensis Ve r h o e ff, 1941?; = H. hatayensis membranaceus Verhoeff, 1941?; = H. kosswigii [sic] Verhoeff, 1941? [non Lithobius kosswigi Chamberlin, 1952]; = H. matsakisi Matic & Stauropulos, 1990; = Lithobius (Archilithobius) pachypus Verhoeff, 1925; = Archeobius phanus Chamberlin, 1952 [non Nadabius phanus Chamberlin, 1941]; = L. platypus Newport, 1845? [non Arebius platypus Chamberlin, 1941]; = H. procurvus Zalesskaja, 1978; = L. (Oligobothrus) vosseleri Verhoeff, 1941?; = L. (A.) vossleri var. propitia Silvestri, 1929); H. jangtseanus (Verhoeff, 1942); H. longispinipes Ma, Pei & Zhu, 2009; H. (?) megapus (Muralevitch, 1907) (= L. megapus bidens Folkmanová & Dobroruka, 1960); H. multicalcaratus Folkmanová, 1958; H. major Folkmanová & Dobroruka, 1960 [non Labrobius major Chamberlin, 1941; non Nampabius major Chamberlin, 1925; non Lithobius porathi major Folkmanov & Dobroruka, 1960]; H. multiforaminis sp. nov. (described below); H. perelae Zalesskaja, 1978; H. pervagatus Zalesskaja, 1978; H. plumatus Zalesskaja, 1978; H. styliferus (Loksa, 1978) n. comb. [synonyms according to Zalesskaja 1978; Eason 1981, 1983, 1992; Zapparoli 1999, 2006].	en	Pei, Sujian, Ma, Huiqin, Zapparoli, Marzio, Zhu, Mingsheng (2010): A review of the Chinese species of Hessebius Verhoeff, 1941 (Chilopoda: Lithobiomorpha: Lithobiidae). Zootaxa 2631: 51-61, DOI: 10.5281/zenodo.198338
AC4D87E08636550A37A4690C8630FE4F.taxon	description	Ma, Pei & Zhu 2009: 195, figs 1 – 7.	en	Pei, Sujian, Ma, Huiqin, Zapparoli, Marzio, Zhu, Mingsheng (2010): A review of the Chinese species of Hessebius Verhoeff, 1941 (Chilopoda: Lithobiomorpha: Lithobiidae). Zootaxa 2631: 51-61, DOI: 10.5281/zenodo.198338
AC4D87E08636550A37A4690C8630FE4F.taxon	diagnosis	Diagnosis. Body length 10.7 – 12.6 mm, antennae composed of 17 – 19 articles, commonly 18 + 18; 6 ocelli on each side arranged in 2 horizontal rows; Tömösváry’s organ moderately large, rounded, almost equal to the adjoining ocelli; 2 + 2 moderately small sharp coxosternal teeth; porodont feebly slender, posterolateral to lateral tooth; all tergites without sharp posterior triangular projections; legs 14 and 15 thicker than anterior ones in female, still more in male; coxal pores 2 – 5, ovate to round, arranged in one row; female gonopods with 2 + 2 moderately thick, bullet – shaped spurs, second article with a long terminal thorn pointing backwards; apical claw simple, with two unconspicuous basal lateral denticles; male gonopods short and small, only a small hemispherical bulge, with 2 long setae on the surface, tip slightly sclerotised. Spinulation. Owing to a misprint, legs spinulation of H. longispinipes was not been included in the original description of this species (Ma et al. 2009). This data is reported here as in Tab. 2. C t P F T C t P F T 1 – 6 – – p amp am – – (a) p ap ap 7 – 8 – – p amp am – – ap ap ap 9 – 10 – – p amp am a – ap ap ap 11 – – p amp am a – amp ap ap 12 – 13 – – p amp am a – amp p p 14 – m amp am a a – amp p p 15 – m amp am – a – amp p – Distribution. Only known from the type locality: China, Barkor County, Hami City, Xinjiang Uygur Autonomous Region, 43 ° 06 ' N 93 ° 00 ' E (Ma et al. 2009). Ecological notes. The species was collected in taiga; it inhabits moderately moist and abundant in organic matter soil; under stones, at 3770 m above sea level.	en	Pei, Sujian, Ma, Huiqin, Zapparoli, Marzio, Zhu, Mingsheng (2010): A review of the Chinese species of Hessebius Verhoeff, 1941 (Chilopoda: Lithobiomorpha: Lithobiidae). Zootaxa 2631: 51-61, DOI: 10.5281/zenodo.198338
AC4D87E08636550A37A4690C8630FE4F.taxon	discussion	Remarks. In a paper on Mongolian centipedes, Loksa (1978) described the new species Schizotergitius styliferus, based on material from “ Bajanchongor aimak [= province]: Cagen Bodg ul Gebirge, Quelle Toorin bulag, 13 km O vom Grenzposten Caganbulag im Cagan Bogd ul Gebirge, 1500 m ” (1 Ƥ, Holotypus) and “ Gobi Altaj aimak [= province]: zwischen Schargyn Gobi und Beger nuur, cca 20 km O von Somon Chalium, Chuural cacran, 1700 m ” (1 3, 1 Ƥ, Paratypes), in south-west Mongolia. This species is known only from the type specimens. Although styliferus has been originally assigned to the Central Asiatic genus Schizotergitius Verhoeff, 1930, it clearly belongs to Hessebius Verhoeff, 1941 (new combination) by the shape of the female gonopods (see Loksa 1978: figs 27, 29). The other characters reported in the original description of this species also agree well with the diagnosis of Hessebius here reported and those of Verhoeff (1941) and Zalesskaja (1978). H. styliferus seems also strictly related to H. longispinipes (Tab. 4) and probably they belong to a same group of species. Both share morphological characters such as the general size, the number and arrangement of the ocelli, the shape of the coxosternum and porodont, the number and shape of the spurs of the female gonopods, the characteristic shape of the dorsal terminal part of the second article of the female gonopods, strongly extending backwards. They differ in characters such as the number of antennal articles, 17 + 17 – 19 + 19 in H. longispinipes, 20 + 20 in H. stilyferus, and in some detail of the spinulation of legs, chiefly the presence of DaC on legs 9 – 15 in H. longispinipes, on legs 11 – 15 in H. styliferus, and the VmF in all the legs in H. longispinipes, only in legs 11 – 14 in H. styliferus. Both characters may show however some variation in Lithobiidae; in Hessebius variation in spinulation has been documented in H. barbipes by Eason (1981) and Negrea & Matic (1995). Worth notice is that the collecting sites of H. styliferus are ca 380 km far one from another and 600 km southwestward from the only locality known of H. longispinipes. Therefore, the relationships between the two taxa deserve further investigation.	en	Pei, Sujian, Ma, Huiqin, Zapparoli, Marzio, Zhu, Mingsheng (2010): A review of the Chinese species of Hessebius Verhoeff, 1941 (Chilopoda: Lithobiomorpha: Lithobiidae). Zootaxa 2631: 51-61, DOI: 10.5281/zenodo.198338
AC4D87E08631550737A46F548679FE94.taxon	materials_examined	Material examined. Holotype: 3 (Fig. 1), adult, 22.9 mm long, Pulan Town, Pulan County, Ali City, Tibet Autonomous Region, 30 ° 18 ' N 80 ° 48 ' E, altitude 4806 m, 17 August 2006, leg. Mingsheng Zhu, Long Liu, Xiaofeng Yang. Paratypes: 2 3, 2 ƤƤ, same date and locality as holotype.	en	Pei, Sujian, Ma, Huiqin, Zapparoli, Marzio, Zhu, Mingsheng (2010): A review of the Chinese species of Hessebius Verhoeff, 1941 (Chilopoda: Lithobiomorpha: Lithobiidae). Zootaxa 2631: 51-61, DOI: 10.5281/zenodo.198338
AC4D87E08631550737A46F548679FE94.taxon	diagnosis	Diagnosis. A Hessebius species with body length 18.9 – 22.9 mm, antennae composed of 20 + 20 articles; 13 – 15 ocelli on each side; Tömösváry’s organ small, rounded, smaller than the adjoining ocelli, posterosuperior ocellus and posterior ocellus comparatively larger than the others; 2 + 2 coxosternal teeth; porodont thickset, posterolateral to lateral tooth; all tergites without sharp posterior triangular projections; a row of comparatively thick setae on the ventral side of tibia of legs 1 – 11, glandular pores on the legs 9 – 15, S 14 and S 15 and genital section; coxal pores 4 – 7, ovate to round, arranged in one row; the first article of female gonopods with 2 + 2 moderately thick, bullet – shaped spurs; apical claw of the third article simple; male gonopods short and small, only a small hemispherical bulge, with 6 – 8 long setae on surface, tip slightly sclerotised. No accessory spurs on leg 15.	en	Pei, Sujian, Ma, Huiqin, Zapparoli, Marzio, Zhu, Mingsheng (2010): A review of the Chinese species of Hessebius Verhoeff, 1941 (Chilopoda: Lithobiomorpha: Lithobiidae). Zootaxa 2631: 51-61, DOI: 10.5281/zenodo.198338
AC4D87E08631550737A46F548679FE94.taxon	description	Description. Body 18.9 – 22.9 mm long, cephalic plate 1.93 – 2.69 mm long, 2.62 – 3.24 mm wide. Antennal articles pale yellow to brown, with gray hue; tergites pale yellow to brown; cephalic plate and T 15 slightly darker; pleural region pale yellow to pale gray; sternites pale yellow – brown to yellow – brown; distal part of forcipules black red, forcipular coxosternite and S 15 yellow – brown, with slightly darker hue; legs pale yellow to pale yellow – brown, tarsi yellow – brown. Cephalic plate longer than wide, rough, convex, anterior part of the cephalic capsule with shallow median sulcus; a moderately large depression in front of the transverse suture, pigment concentrated as close netlike veins; tiny setae inserted in pores scattered very sparsely over the surface; moderately long setae scattered along the marginal ridge; lateral marginal ridge discontinuous, posterior margin straight (Fig. 1). Antennae of 20 + 20 articles (Fig. 1); basal article moderately longer than wide, the second markedly longer than wide; succeeding articles gradually shorten, terminal article typically longer than wide, up to 2.3 – 2.8 times as long as wide. Abundant setae on antennal surface, but fewer setae on outer side and ventral and dorsal side in basal articles, gradual increase in density of setae to about fourth article, then more or less constant. Ocelli 13 – 15 on each side (Fig. 2), oval to rounded, commonly rounded, gently bulging, moderately domed, translucent, usually darkly pigmented in eyeground, arranged in 3 horizontal irregular rows; posterosuperior ocellus and posterior ocellus comparatively bigger, the other ocelli moderately small, approximately equal in size, overhanging the lateral margin of the cephalic plate. Tömösváry’s organ very small (Fig. 2), rounded; close to the adjoining ocelli, placed ventrad of anterolateral margin of cephalic pleurite, moderately larger than the adjoining ocelli. Forcipular coxosternite (Fig. 3) approximately trapezoidal, anterior margin moderately broad with 2 + 2 comparatively sharp teeth, terminal part of each tooth approximately blunt, median diastema relatively deep, V – shaped (Fig. 4); porodont moderately stout, just posterolateral and moderately far from the lateral tooth, without bulge near their base (Fig. 4). Forcipular coxosternite also with moderately long setae sparsely scattered over the surface, comparatively long setae near the anterior margin; surface of forcipular coxosternite and forcipules of male rough as the cephalic plate, but quite smooth in female. All tergites moderately rough, having some wrinkles, backside slightly hunched, tiny setae scattered very sparsely over the surface; T 1 posterolaterally narrower than anterolaterally, generally trapeziform, distinctly broader than T 3 and the cephalic plate, the latter slightly wider than T 3; lateral marginations of all tergites continuous, posterior border of T 1 and T 3 slightly concave, margination continuous, posterior border of TT 5, 8, 10, 12 and 14 concave, margination discontinuous except for T 5, posterior border of TT 2, 4, 6, 7, 9, 11 and 13 straight; all tergites without posterior triangular projections (Fig. 1); setae scattered sparsely along the lateral borders, more numerous near the anterior angles of each tergite. All sternites posterolaterally narrower than anterolaterally, generally trapeziform, moderately smooth, moderately setose on anterior part of each sternite, a pair of longer setae approximately symmetrical on the surface of the anterior and posterior part of each sternite; 2 – 4 longer setae on both anterior lateral borders, 1 – 2 comparatively long setae scattered sparsely on posterior margin of sternites. Male with sparse glandular pores on the surface of S 13, dense glandular pores and wrinkles on the surface of S 14 and S 15 and on genital section. Female S 13 – S 15 moderately smooth than in male; S 15 with a small rounded depression in male, the depressed area moderately larger in female. Tarsus 1 – tarsus 2 articulation not well – defined on legs 1 – 13, defined on legs 14 and 15; claw moderately long and curved ventrad in all legs; accessory spur on both anterior and posterior side of claw of legs 1 – 14, anterior accessory spur moderately long and slender, forming a moderate small angle with the claw; posterior accessory spur slightly strong, forming a comparatively large angle with the claw; no accessory spurs on leg 15; one row of thicker setae regularly arranged on the medial ventral side of tibiae of legs 1 – 10, up to 50 % – 75 % the length of the tibia; only 2 – 3 moderately thicker setae on the medial ventral side of the tibia of leg 11, short to moderately long setae scattered very sparsely over the surface of all articles of legs 1 – 13, thicker setae scattered on the surface of tarsi, almost evenly scattered on anterior, posterior and back sides, two rows of moderately thicker setae on medial ventral side of tibiae; glandular pores on the surface of prefemur, femur, tibia and tarsus of the leg 9, more numerous on the same articles of the leg 10, where glandular pores are also present on coxa and trochanter; highest number of glandular pores on legs 14 and 15; among them, more glandular pores on the anterior and posterior side than ventrad in male; glandular pores only on the surface of prefemur, femur, tibia and tarsus of legs 14 and 15 in the female, the legs more smooth than in the male; legs 14 and 15 slightly thicker and stronger than other legs in female; two comparatively obvious shallow dorsal furrow on the tibia of legs 14 and 15, the male more obvious than female; tarsus 1 5.1 – 5.5 times longer than wide, tarsus 2 60 % – 70 % length of tarsus on legs 15 in female; legs 14 and 15 markedly thicker and stronger than the other legs in male, tarsus 1 4.2 – 5.4 times longer than wide, tarsus 2 50 % – 69 % length of tarsus of male legs 15. Leg spinulation as in Tab. 3. legs ventral dorsal pairs C t P F T C t P F T 1 – 2 – – mp amp am – – ap ap ap Coxal pores arranged in a row, round or slightly ovate, small to moderately large: 5676, 5564, 4554, 66 (7) 6 (5) in the female; usually 6766, 7777, 5666, 5666 in the male; pore-field set in a moderately shallow groove, the fringe of coxal pore-field with eminence, short to moderately long setae scattered sparsely over the surface of the eminence. Female S 15 posterolaterally narrower than anterolaterally, generally trapeziform, straight posteromedially; short to long setae scattered very sparsely over its surface and lateral margins. The sternite of genital segment usually well sclerotised, longer than wide, posterior border moderately deeply concave between condyles of gonopods, except for a small, median approximately rhombic bulge, distally lightly sclerotised, moderately longer setae scattered over the surface, few setae near S 15, regularly fringed with longer setae along the posterior margin. First article of the female gonopods moderately broad, bearing many moderately long setae, with 2 + 2 moderately long, bullet – shaped spurs, inner spur slightly smaller and more anterior than the outer (Fig. 5); second article of the female gonopods with 13 – 15 moderately long setae, arranged in 3 irregular rows; 9 short to long setae along the dorsolateral ridge, the dorsal terminal part strongly extending backwards and forming a thick protuberance (Fig. 6); third article of the female gonopods usually with 4 moderately long setae on the surface, three comparatively long setae along the dorsolateral ridge, apical claw simple and broad (Figs 5, 7). Male S 15 posterolaterally narrower than anterolaterally, generally trapeziform, posteromedially straight, posterior angles rounded, moderately long setae sparsely scattered over its surface; tergite of genital segment wider than long, usually well sclerotised; comparatively long setae about evenly scattered on the ventral surface, slightly fewer near S 15. Posterior margin of the sternite of the genital segment quite deeply concave between gonopods, no bulge medially; gonopods short and small, only a small hemispherical bulge, with 6 – 8 long setae on surface, terminal slightly sclerotised (Fig. 8).	en	Pei, Sujian, Ma, Huiqin, Zapparoli, Marzio, Zhu, Mingsheng (2010): A review of the Chinese species of Hessebius Verhoeff, 1941 (Chilopoda: Lithobiomorpha: Lithobiidae). Zootaxa 2631: 51-61, DOI: 10.5281/zenodo.198338
AC4D87E08631550737A46F548679FE94.taxon	etymology	Etymology. The specific name refers to the large numbers of glandular pores on the legs 9 – 15, ventral of S 14 and S 15.	en	Pei, Sujian, Ma, Huiqin, Zapparoli, Marzio, Zhu, Mingsheng (2010): A review of the Chinese species of Hessebius Verhoeff, 1941 (Chilopoda: Lithobiomorpha: Lithobiidae). Zootaxa 2631: 51-61, DOI: 10.5281/zenodo.198338
AC4D87E08631550737A46F548679FE94.taxon	distribution	Distribution. The new species is to date known only from the type locality.	en	Pei, Sujian, Ma, Huiqin, Zapparoli, Marzio, Zhu, Mingsheng (2010): A review of the Chinese species of Hessebius Verhoeff, 1941 (Chilopoda: Lithobiomorpha: Lithobiidae). Zootaxa 2631: 51-61, DOI: 10.5281/zenodo.198338
AC4D87E08631550737A46F548679FE94.taxon	materials_examined	Ecological notes. The material examined has been collected from Subfrigid Zone and Arid Regions of Plateau; in moderately moist soil, under stones.	en	Pei, Sujian, Ma, Huiqin, Zapparoli, Marzio, Zhu, Mingsheng (2010): A review of the Chinese species of Hessebius Verhoeff, 1941 (Chilopoda: Lithobiomorpha: Lithobiidae). Zootaxa 2631: 51-61, DOI: 10.5281/zenodo.198338
