taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
AC4D691FFFB65E13FF685B9F70A2FDCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268219/files/figure.png	https://doi.org/10.5281/zenodo.3268219	FIGURES 140‒143. Morphology of females (queens). Left lateral aspect of the oculo-malar area of: 140, B. polaris; 141, B. kirbiellus (white lines indicate length and breadth measurements). Left lateral aspect of the outer surface of the left hind tibia (corbicular area): 142, B. polaris; 143, B. neoboreus (white lines indicate highlights in the central area).	FIGURES 140‒143. Morphology of females (queens). Left lateral aspect of the oculo-malar area of: 140, B. polaris; 141, B. kirbiellus (white lines indicate length and breadth measurements). Left lateral aspect of the outer surface of the left hind tibia (corbicular area): 142, B. polaris; 143, B. neoboreus (white lines indicate highlights in the central area).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB65E13FF685B9F70A2FDCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268177/files/figure.png	https://doi.org/10.5281/zenodo.3268177	FIGURES 1‒6. Lateral views of foraging females (queens) of species of the subgenus Alpinobombus (with photo credits): 1, B. alpinus, Norway (A. Staverløkk); 2, B. pyrrhopygus, Sweden (G. Holmström); 3, B. polaris, Canada (B. Heinrich); 4, B. balteatus, Sweden (G. Holmström); 5, B. kirbiellus, USA (D. Wilson); 6, B. hyperboreus, Sweden (L.-I. Larsson). Some images reversed.	FIGURES 1‒6. Lateral views of foraging females (queens) of species of the subgenus Alpinobombus (with photo credits): 1, B. alpinus, Norway (A. Staverløkk); 2, B. pyrrhopygus, Sweden (G. Holmström); 3, B. polaris, Canada (B. Heinrich); 4, B. balteatus, Sweden (G. Holmström); 5, B. kirbiellus, USA (D. Wilson); 6, B. hyperboreus, Sweden (L.-I. Larsson). Some images reversed.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB65E13FF685B9F70A2FDCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268197/files/figure.png	https://doi.org/10.5281/zenodo.3268197	FIGURES 22‒38. 22: Bombus pyrrhopygus global distribution (as for Fig. 15). 23‒38 colour patterns (as for Figs. 16‒21): 23 (#4707) Russia-Arkhangelsk; 24 (#337) Russia-Arkhangelsk; 25 (#273) Sweden; 26 (#272) Sweden; 27 (#250) Sweden; 28 (#249) Sweden; 29 (#258) Sweden; 30 (#4709) Russia-Murmansk; 31 (#257) Sweden; 32 (#4711) Russia-Arkhangelsk; 33 (#4712) Russia-Arkhangelsk; 34 (#4713) Russia-Arkhangelsk; 35 (#242) Norway; 36 (#139) Sweden; 37 (#4710) Norway; 38 (#1375) Russia-Murmansk.	FIGURES 22‒38. 22: Bombus pyrrhopygus global distribution (as for Fig. 15). 23‒38 colour patterns (as for Figs. 16‒21): 23 (#4707) Russia-Arkhangelsk; 24 (#337) Russia-Arkhangelsk; 25 (#273) Sweden; 26 (#272) Sweden; 27 (#250) Sweden; 28 (#249) Sweden; 29 (#258) Sweden; 30 (#4709) Russia-Murmansk; 31 (#257) Sweden; 32 (#4711) Russia-Arkhangelsk; 33 (#4712) Russia-Arkhangelsk; 34 (#4713) Russia-Arkhangelsk; 35 (#242) Norway; 36 (#139) Sweden; 37 (#4710) Norway; 38 (#1375) Russia-Murmansk.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB65E13FF685B9F70A2FDCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268195/files/figure.png	https://doi.org/10.5281/zenodo.3268195	FIGURES 15‒21. 15: Bombus alpinus global distribution shown with: grey spots for specimens examined; black spots for samples with COI barcodes; question marks for uncertain records at low elevation and low latitude; and grey crosses for records of all species of Alpinobombus combined (Alpinobombus species are unknown from the southern hemisphere). Relief map with hill shading, Polar projection, the international boundaries and the Arctic Circle are shown as narrow grey lines, and the northern tree line shown as a broad grey line. Image created in ArcGIS using World_Shaded_Relief basemap which is Copyright: © 2014 Esri. 16‒21: simplified colour-pattern diagrams for the dorsal hair, with olive indicating a mixture of black and yellow. Figure (AL#n) country: 16 (#14) Sweden; 17 (#113) Sweden; 18 (#3785) Austria; 19 (#3787) Austria; 20 (#3788) Austria; 21 (#141) Norway.	FIGURES 15‒21. 15: Bombus alpinus global distribution shown with: grey spots for specimens examined; black spots for samples with COI barcodes; question marks for uncertain records at low elevation and low latitude; and grey crosses for records of all species of Alpinobombus combined (Alpinobombus species are unknown from the southern hemisphere). Relief map with hill shading, Polar projection, the international boundaries and the Arctic Circle are shown as narrow grey lines, and the northern tree line shown as a broad grey line. Image created in ArcGIS using World_Shaded_Relief basemap which is Copyright: © 2014 Esri. 16‒21: simplified colour-pattern diagrams for the dorsal hair, with olive indicating a mixture of black and yellow. Figure (AL#n) country: 16 (#14) Sweden; 17 (#113) Sweden; 18 (#3785) Austria; 19 (#3787) Austria; 20 (#3788) Austria; 21 (#141) Norway.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB65E13FF685B9F70A2FDCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268199/files/figure.png	https://doi.org/10.5281/zenodo.3268199	FIGURES 39‒58. 39: Bombus polaris global distribution (as for Fig. 15). 40‒58 colour patterns (as for Figs. 16‒21): 40 (#4380) Canada-Labrador; 41 (#4396) USA-Alaska; 42 (#3856) Canada-Nunavut; 43 (#59) USA-Alaska; 44 (#43) Denmark- Greenland; 45 (#4740) Denmark-Greenland; 46 (#4407) USA-Alaska; 47 (#334) USA-Alaska; 48 (#285) Canada-Nunavut; 49 (#4383) USA-Alaska; 50 (#44) Denmark-Greenland; 51 (#19) Canada-Northwest Territories; 52 (#4401) USA-Alaska; 53 (#4403) USA-Alaska; 54 (#4884) Canada-Yukon; 55 (#4405) USA-Alaska; 56 (#4384) USA-Alaska; 57 (#4951) USA-Alaska; 58 (#29) Canada-Manitoba.	FIGURES 39‒58. 39: Bombus polaris global distribution (as for Fig. 15). 40‒58 colour patterns (as for Figs. 16‒21): 40 (#4380) Canada-Labrador; 41 (#4396) USA-Alaska; 42 (#3856) Canada-Nunavut; 43 (#59) USA-Alaska; 44 (#43) Denmark- Greenland; 45 (#4740) Denmark-Greenland; 46 (#4407) USA-Alaska; 47 (#334) USA-Alaska; 48 (#285) Canada-Nunavut; 49 (#4383) USA-Alaska; 50 (#44) Denmark-Greenland; 51 (#19) Canada-Northwest Territories; 52 (#4401) USA-Alaska; 53 (#4403) USA-Alaska; 54 (#4884) Canada-Yukon; 55 (#4405) USA-Alaska; 56 (#4384) USA-Alaska; 57 (#4951) USA-Alaska; 58 (#29) Canada-Manitoba.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB65E13FF685B9F70A2FDCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268205/files/figure.png	https://doi.org/10.5281/zenodo.3268205	FIGURES 89‒103. 89: Bombus neoboreus global distribution (as for Fig. 15). 90‒103 colour patterns (as for Figs. 16‒21): 90 (#68) USA-Alaska; 91 (#4706) Canada-Nunavut; 92 (#3816) Canada-Yukon; 93 (#3819) USA-Alaska; 94 (#2055) Canada-Nunavut; 95 (#2058) Canada-Nunavut; 96 (#2071) Canada-Nunavut; 97 (#2051) Canada-Nunavut; 98 (#4406) Canada-Nunavut; 99 (#90) Canada-Nunavut; 100 (#92) Canada-Nunavut; 101 (#93) Canada-Yukon; 102 (#3762) Canada-Nunavut; 103 (#91) Canada-Nunavut.	FIGURES 89‒103. 89: Bombus neoboreus global distribution (as for Fig. 15). 90‒103 colour patterns (as for Figs. 16‒21): 90 (#68) USA-Alaska; 91 (#4706) Canada-Nunavut; 92 (#3816) Canada-Yukon; 93 (#3819) USA-Alaska; 94 (#2055) Canada-Nunavut; 95 (#2058) Canada-Nunavut; 96 (#2071) Canada-Nunavut; 97 (#2051) Canada-Nunavut; 98 (#4406) Canada-Nunavut; 99 (#90) Canada-Nunavut; 100 (#92) Canada-Nunavut; 101 (#93) Canada-Yukon; 102 (#3762) Canada-Nunavut; 103 (#91) Canada-Nunavut.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB65E13FF685B9F70A2FDCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268207/files/figure.png	https://doi.org/10.5281/zenodo.3268207	FIGURES 104‒115. 104: Bombus kluanensis global distribution (as for Fig. 15). 105‒115 colour patterns (as for Figs. 16‒21): 105 (#4473) Canada-Yukon; 106 (#4689) Canada-Yukon; 107 (#16) Canada-Yukon; 108 (#4393) USA-Alaska; 109 (#4466) Canada-Yukon; 110 (#4700) Canada-Yukon; 111 (#3130) Canada-Yukon; 112 (#4876) Canada-Yukon; 113 (#4882) Canada- Yukon; 114 (#4852) Canada-Yukon; 115 (#4881) Canada-Yukon.	FIGURES 104‒115. 104: Bombus kluanensis global distribution (as for Fig. 15). 105‒115 colour patterns (as for Figs. 16‒21): 105 (#4473) Canada-Yukon; 106 (#4689) Canada-Yukon; 107 (#16) Canada-Yukon; 108 (#4393) USA-Alaska; 109 (#4466) Canada-Yukon; 110 (#4700) Canada-Yukon; 111 (#3130) Canada-Yukon; 112 (#4876) Canada-Yukon; 113 (#4882) Canada- Yukon; 114 (#4852) Canada-Yukon; 115 (#4881) Canada-Yukon.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB65E13FF685B9F70A2FDCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268201/files/figure.png	https://doi.org/10.5281/zenodo.3268201	FIGURES 59‒76. 59: Bombus balteatus global distribution (as for Fig. 15). 60‒76 colour patterns (as for Figs.16‒21): 60 (#540) Russia-Krasnoyask; 61 (#119) Sweden; 62 (#541) Russia-Krasnoyarsk; 63 (#542) Russia-Krasnoyarsk; 64 (#117) Mongolia; 65 (#534) Russia-Kamchatka; 66 (#3841) Norway; 67 (#738) Russia-Kamchatka; 68 (#971) Russia- Kamchatka; 69 (#535) Russia-Kamchatka; 70 (#973) Russia-Kamchatka; 71 (#880) Russia-Kamchatka; 72 (#536) Russia-Kamchatka; 73 (#228) Norway; 74 (#392) Sweden; 75 (#3843) Norway; 76 (#538) Russia-Kamchatka.	FIGURES 59‒76. 59: Bombus balteatus global distribution (as for Fig. 15). 60‒76 colour patterns (as for Figs.16‒21): 60 (#540) Russia-Krasnoyask; 61 (#119) Sweden; 62 (#541) Russia-Krasnoyarsk; 63 (#542) Russia-Krasnoyarsk; 64 (#117) Mongolia; 65 (#534) Russia-Kamchatka; 66 (#3841) Norway; 67 (#738) Russia-Kamchatka; 68 (#971) Russia- Kamchatka; 69 (#535) Russia-Kamchatka; 70 (#973) Russia-Kamchatka; 71 (#880) Russia-Kamchatka; 72 (#536) Russia-Kamchatka; 73 (#228) Norway; 74 (#392) Sweden; 75 (#3843) Norway; 76 (#538) Russia-Kamchatka.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB65E13FF685B9F70A2FDCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268203/files/figure.png	https://doi.org/10.5281/zenodo.3268203	FIGURES 77‒88. 77: Bombus kirbiellus global distribution (as for Fig. 15). 78‒88 colour patterns (as for Figs. 16‒21): 78 (#556) Canada-Nunavut; 79 (#294) Canada-Nunavut; 80 (#100) Canada-Nunavut; 81 (#3791) USA-Colorado; 82 (#3790) USA-Colorado; 83 (#3765) Canada-Nunavut; 84 (#3792) USA-Colorado; 85 (#3797) USA-Colorado; 86 (#4915) USA Alaska; 87 (#3768) Canada-Nunavut; 88 (#3812) USA-Colorado.	FIGURES 77‒88. 77: Bombus kirbiellus global distribution (as for Fig. 15). 78‒88 colour patterns (as for Figs. 16‒21): 78 (#556) Canada-Nunavut; 79 (#294) Canada-Nunavut; 80 (#100) Canada-Nunavut; 81 (#3791) USA-Colorado; 82 (#3790) USA-Colorado; 83 (#3765) Canada-Nunavut; 84 (#3792) USA-Colorado; 85 (#3797) USA-Colorado; 86 (#4915) USA Alaska; 87 (#3768) Canada-Nunavut; 88 (#3812) USA-Colorado.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB65E13FF685B9F70A2FDCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268209/files/figure.png	https://doi.org/10.5281/zenodo.3268209	FIGURES 116‒123. 116: Bombus natvigi global distribution (as for Fig. 15). 117‒123 colour patterns (as for Figs. 16‒21): 117 (#3519) Canada-Nunavut; 118 (#184) Canada-Nunavut; 119 (#3551) Canada-Nunavut; 120 (#3546) Canada-Nunavut; 121 (#79) USA-Alaska; 122 (#3557) Canada-Northwest Territories; 123 (#82) Canada-Northwest Territories.	FIGURES 116‒123. 116: Bombus natvigi global distribution (as for Fig. 15). 117‒123 colour patterns (as for Figs. 16‒21): 117 (#3519) Canada-Nunavut; 118 (#184) Canada-Nunavut; 119 (#3551) Canada-Nunavut; 120 (#3546) Canada-Nunavut; 121 (#79) USA-Alaska; 122 (#3557) Canada-Northwest Territories; 123 (#82) Canada-Northwest Territories.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB65E13FF685B9F70A2FDCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268211/files/figure.png	https://doi.org/10.5281/zenodo.3268211	FIGURES 124‒128. 124: Bombus hyperboreus global distribution (as for Fig. 15). 125‒128 colour patterns (as for Figs. 16‒ 21): 125 (#39) Russia-Chukotka (Wrangel Island); 126 (#33) Russia-Magadan; 127 (#1258) Russia-Murmansk; 128 (#533) Russia-Sakha.	FIGURES 124‒128. 124: Bombus hyperboreus global distribution (as for Fig. 15). 125‒128 colour patterns (as for Figs. 16‒ 21): 125 (#39) Russia-Chukotka (Wrangel Island); 126 (#33) Russia-Magadan; 127 (#1258) Russia-Murmansk; 128 (#533) Russia-Sakha.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB65E13FF685B9F70A2FDCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268181/files/figure.png	https://doi.org/10.5281/zenodo.3268181	FIGURE 7. Sample sites and climatic suitability for all Alpinobombus species combined estimated using Maxent from climate variables for the year 2000. Bright yellow and brown areas show where the logistic prediction of occurrence is p> 0.5 (the outlier records with exceptionally low probabilities of suitability were excluded in step-wise iterations); brown spots show all consistent Alpinobombus site records; ‘?’ shows the five records that are in climatically outlying sites. Polar projection, North Pole (starred) at the centre of the map, international boundaries and the Arctic Circle shown as narrow grey lines.	FIGURE 7. Sample sites and climatic suitability for all Alpinobombus species combined estimated using Maxent from climate variables for the year 2000. Bright yellow and brown areas show where the logistic prediction of occurrence is p> 0.5 (the outlier records with exceptionally low probabilities of suitability were excluded in step-wise iterations); brown spots show all consistent Alpinobombus site records; ‘?’ shows the five records that are in climatically outlying sites. Polar projection, North Pole (starred) at the centre of the map, international boundaries and the Arctic Circle shown as narrow grey lines.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB65E13FF685B9F70A2FDCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268185/files/figure.png	https://doi.org/10.5281/zenodo.3268185	FIGURE 8. Diagram representing a corridor-dispersal model, encompassing a set of the short-distance dispersal events permitted (in either direction) between the areas defined in Table 5 (including the outgroup B. ignitus), based on their geographical proximity and the likely disposition of suitable habitat and favourable climates in the past (see the text).	FIGURE 8. Diagram representing a corridor-dispersal model, encompassing a set of the short-distance dispersal events permitted (in either direction) between the areas defined in Table 5 (including the outgroup B. ignitus), based on their geographical proximity and the likely disposition of suitable habitat and favourable climates in the past (see the text).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB65E13FF685B9F70A2FDCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268187/files/figure.png	https://doi.org/10.5281/zenodo.3268187	FIGURE 9. Estimate of the metric gene tree for the 54 longest unique COI-barcode alleles of the subgenus Alpinobombus from MrBayes combined with the Bayesian Poisson-tree-process (PTP) solution with the highest support with spots showing the nodes with the strongest support as coalescents for candidate species (outgroup B. ignitus not shown). Values above the nodes are Bayesian posterior probabilities showing branch support for groups; values below the nodes are PTP Bayesian support values that all daughter alleles are parts of a single species (probabilities within candidate species are all ≤ 0.44). The scale bar is calibrated in substitutions per nucleotide site. Each unique allele is represented by one of the longest available sample sequences, labelled with: the sequence length in number of nucleotides, a taxon name or name describing an unpublished colour pattern, and then a code that consists of a specimen identifier (AL#n) from the project database and (after the hyphen) a sample identifier from BOLD or GenBank, followed with its geographic origin (for simplicity Greenland is included without Denmark).	FIGURE 9. Estimate of the metric gene tree for the 54 longest unique COI-barcode alleles of the subgenus Alpinobombus from MrBayes combined with the Bayesian Poisson-tree-process (PTP) solution with the highest support with spots showing the nodes with the strongest support as coalescents for candidate species (outgroup B. ignitus not shown). Values above the nodes are Bayesian posterior probabilities showing branch support for groups; values below the nodes are PTP Bayesian support values that all daughter alleles are parts of a single species (probabilities within candidate species are all ≤ 0.44). The scale bar is calibrated in substitutions per nucleotide site. Each unique allele is represented by one of the longest available sample sequences, labelled with: the sequence length in number of nucleotides, a taxon name or name describing an unpublished colour pattern, and then a code that consists of a specimen identifier (AL#n) from the project database and (after the hyphen) a sample identifier from BOLD or GenBank, followed with its geographic origin (for simplicity Greenland is included without Denmark).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB65E13FF685B9F70A2FDCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268189/files/figure.png	https://doi.org/10.5281/zenodo.3268189	FIGURE 10. Map of the entire global distribution of records of B. kluanensis s. l.. Grey crosses are records for any Alpinobombus species; black crosses are records of B. neoboreus; overlayed with small white spots for specimens of B. kluanensis s. l.; large green spots for sequenced samples with the kluanensis s. str. alleles (collected 2010‒2016); small red spots for sequenced samples with the ‘unnamed2’ allele (collected 2015‒2017). Relief map with hill shading, with a Polar projection, and with the Arctic Circle and the USA (Alaska) / Canada (Yukon) international boundary shown as grey lines. North pointer and scale bar below, with an inset that shows the location of the detailed area of the global map. Image created in ArcGIS using World_Shad- ed_Relief basemap which is Copyright: © 2014 Esri.	FIGURE 10. Map of the entire global distribution of records of B. kluanensis s. l.. Grey crosses are records for any Alpinobombus species; black crosses are records of B. neoboreus; overlayed with small white spots for specimens of B. kluanensis s. l.; large green spots for sequenced samples with the kluanensis s. str. alleles (collected 2010‒2016); small red spots for sequenced samples with the ‘unnamed2’ allele (collected 2015‒2017). Relief map with hill shading, with a Polar projection, and with the Arctic Circle and the USA (Alaska) / Canada (Yukon) international boundary shown as grey lines. North pointer and scale bar below, with an inset that shows the location of the detailed area of the global map. Image created in ArcGIS using World_Shad- ed_Relief basemap which is Copyright: © 2014 Esri.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB65E13FF685B9F70A2FDCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268191/files/figure.png	https://doi.org/10.5281/zenodo.3268191	FIGURES 11‒13. 11: gene trees for COI; 12: gene trees for 16S; and 13: species trees from *BEAST analysis of data from Thanoosing (2017) displayed using Densitree. Each plot shows the background density of trees from samples of 10,000 Bayesian trees, with the summary ‘root canal’ tree superimposed as a thick line, and the branches exclusively linking samples of B. kluanensis s. l. (#4850, 4869, 4870, 4873) traced in red.	FIGURES 11‒13. 11: gene trees for COI; 12: gene trees for 16S; and 13: species trees from *BEAST analysis of data from Thanoosing (2017) displayed using Densitree. Each plot shows the background density of trees from samples of 10,000 Bayesian trees, with the summary ‘root canal’ tree superimposed as a thick line, and the branches exclusively linking samples of B. kluanensis s. l. (#4850, 4869, 4870, 4873) traced in red.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB65E13FF685B9F70A2FDCA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268193/files/figure.png	https://doi.org/10.5281/zenodo.3268193	FIGURE 14. Assigning names to candidate species in a COI gene tree for the 235 specimens of the subgenus Alpinobombus with sequences. Based on the MrBayes analysis of all of the 54 longest unique alleles in Fig. 9, with the remaining sequences interpolated back into the tree with 0 branch lengths from their matching samples (where a sequence matches more than one longer sequence within a species in the Collapse results, it is arbitrarily added after the first matching sequence). The sequence labels follow the format used in Fig. 9. Lineages with high probabilities of representing candidate species in the Poisson-tree-process (PTP) results (Fig. 9) are shown as thick branches with the most recent common ancestors (coalescents) of each candidate species shown with a black spot, the branches within candidate species shown as thin branches. Asterisks mark samples used as informal proxies for the type specimens of each of the taxon names in Table 2. The proxy sample for the type specimen for the oldest available name (the valid name) for each candidate species (from Fig. 9) is marked in bold.	FIGURE 14. Assigning names to candidate species in a COI gene tree for the 235 specimens of the subgenus Alpinobombus with sequences. Based on the MrBayes analysis of all of the 54 longest unique alleles in Fig. 9, with the remaining sequences interpolated back into the tree with 0 branch lengths from their matching samples (where a sequence matches more than one longer sequence within a species in the Collapse results, it is arbitrarily added after the first matching sequence). The sequence labels follow the format used in Fig. 9. Lineages with high probabilities of representing candidate species in the Poisson-tree-process (PTP) results (Fig. 9) are shown as thick branches with the most recent common ancestors (coalescents) of each candidate species shown with a black spot, the branches within candidate species shown as thin branches. Asterisks mark samples used as informal proxies for the type specimens of each of the taxon names in Table 2. The proxy sample for the type specimen for the oldest available name (the valid name) for each candidate species (from Fig. 9) is marked in bold.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB75E13FF685CF070A2F829.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268221/files/figure.png	https://doi.org/10.5281/zenodo.3268221	FIGURES 144‒147. Morphology of males. Dorsal aspect of the penis valves of: 144, B. natvigi; 145, B. neoboreus (white lines indicate location of the subapical outer tooth of the left penis valve). Dorsal aspect of left gonostylus: 146 B. kirbiellus; 147, B. polaris (white lines indicate anterior, midpoint, and posterior median projections of left gonostylus).	FIGURES 144‒147. Morphology of males. Dorsal aspect of the penis valves of: 144, B. natvigi; 145, B. neoboreus (white lines indicate location of the subapical outer tooth of the left penis valve). Dorsal aspect of left gonostylus: 146 B. kirbiellus; 147, B. polaris (white lines indicate anterior, midpoint, and posterior median projections of left gonostylus).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB75E13FF685CF070A2F829.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268207/files/figure.png	https://doi.org/10.5281/zenodo.3268207	FIGURES 104‒115. 104: Bombus kluanensis global distribution (as for Fig. 15). 105‒115 colour patterns (as for Figs. 16‒21): 105 (#4473) Canada-Yukon; 106 (#4689) Canada-Yukon; 107 (#16) Canada-Yukon; 108 (#4393) USA-Alaska; 109 (#4466) Canada-Yukon; 110 (#4700) Canada-Yukon; 111 (#3130) Canada-Yukon; 112 (#4876) Canada-Yukon; 113 (#4882) Canada- Yukon; 114 (#4852) Canada-Yukon; 115 (#4881) Canada-Yukon.	FIGURES 104‒115. 104: Bombus kluanensis global distribution (as for Fig. 15). 105‒115 colour patterns (as for Figs. 16‒21): 105 (#4473) Canada-Yukon; 106 (#4689) Canada-Yukon; 107 (#16) Canada-Yukon; 108 (#4393) USA-Alaska; 109 (#4466) Canada-Yukon; 110 (#4700) Canada-Yukon; 111 (#3130) Canada-Yukon; 112 (#4876) Canada-Yukon; 113 (#4882) Canada- Yukon; 114 (#4852) Canada-Yukon; 115 (#4881) Canada-Yukon.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB75E13FF685CF070A2F829.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268223/files/figure.png	https://doi.org/10.5281/zenodo.3268223	FIGURES 148‒149. Anterior / left lateral aspect of the lower part of the clypeus, the labrum, and the mandibles of males of: (148) B. neoboreus (#4867), showing the small orange hair patches at the base of the mandibles and the long dense posterior fringing black beard; (149) B. kluanensis (#4876), showing the large dense orange hair pads on the mandibles and the sparse orange hairs in the posterior fringing beard.	FIGURES 148‒149. Anterior / left lateral aspect of the lower part of the clypeus, the labrum, and the mandibles of males of: (148) B. neoboreus (#4867), showing the small orange hair patches at the base of the mandibles and the long dense posterior fringing black beard; (149) B. kluanensis (#4876), showing the large dense orange hair pads on the mandibles and the sparse orange hairs in the posterior fringing beard.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB75E13FF685CF070A2F829.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268209/files/figure.png	https://doi.org/10.5281/zenodo.3268209	FIGURES 116‒123. 116: Bombus natvigi global distribution (as for Fig. 15). 117‒123 colour patterns (as for Figs. 16‒21): 117 (#3519) Canada-Nunavut; 118 (#184) Canada-Nunavut; 119 (#3551) Canada-Nunavut; 120 (#3546) Canada-Nunavut; 121 (#79) USA-Alaska; 122 (#3557) Canada-Northwest Territories; 123 (#82) Canada-Northwest Territories.	FIGURES 116‒123. 116: Bombus natvigi global distribution (as for Fig. 15). 117‒123 colour patterns (as for Figs. 16‒21): 117 (#3519) Canada-Nunavut; 118 (#184) Canada-Nunavut; 119 (#3551) Canada-Nunavut; 120 (#3546) Canada-Nunavut; 121 (#79) USA-Alaska; 122 (#3557) Canada-Northwest Territories; 123 (#82) Canada-Northwest Territories.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB75E13FF685CF070A2F829.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268211/files/figure.png	https://doi.org/10.5281/zenodo.3268211	FIGURES 124‒128. 124: Bombus hyperboreus global distribution (as for Fig. 15). 125‒128 colour patterns (as for Figs. 16‒ 21): 125 (#39) Russia-Chukotka (Wrangel Island); 126 (#33) Russia-Magadan; 127 (#1258) Russia-Murmansk; 128 (#533) Russia-Sakha.	FIGURES 124‒128. 124: Bombus hyperboreus global distribution (as for Fig. 15). 125‒128 colour patterns (as for Figs. 16‒ 21): 125 (#39) Russia-Chukotka (Wrangel Island); 126 (#33) Russia-Magadan; 127 (#1258) Russia-Murmansk; 128 (#533) Russia-Sakha.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB75E13FF685CF070A2F829.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268213/files/figure.png	https://doi.org/10.5281/zenodo.3268213	FIGURES 129‒137. Morphology of the male genitalia for species of the subgenus Alpinobombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 129 B. alpinus (#3824); 130 B pyrrhopygus (#3833); 131 B. polaris (#73); 132 B. balteatus (#3843); 133 B. kirbiellus (#3768); 134 B. neoboreus (#4387); 135 B. kluanensis (#4876); 136 B. natvigi (#81); 137 B. hyperboreus (#3821).	FIGURES 129‒137. Morphology of the male genitalia for species of the subgenus Alpinobombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 129 B. alpinus (#3824); 130 B pyrrhopygus (#3833); 131 B. polaris (#73); 132 B. balteatus (#3843); 133 B. kirbiellus (#3768); 134 B. neoboreus (#4387); 135 B. kluanensis (#4876); 136 B. natvigi (#81); 137 B. hyperboreus (#3821).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB75E13FF685CF070A2F829.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268177/files/figure.png	https://doi.org/10.5281/zenodo.3268177	FIGURES 1‒6. Lateral views of foraging females (queens) of species of the subgenus Alpinobombus (with photo credits): 1, B. alpinus, Norway (A. Staverløkk); 2, B. pyrrhopygus, Sweden (G. Holmström); 3, B. polaris, Canada (B. Heinrich); 4, B. balteatus, Sweden (G. Holmström); 5, B. kirbiellus, USA (D. Wilson); 6, B. hyperboreus, Sweden (L.-I. Larsson). Some images reversed.	FIGURES 1‒6. Lateral views of foraging females (queens) of species of the subgenus Alpinobombus (with photo credits): 1, B. alpinus, Norway (A. Staverløkk); 2, B. pyrrhopygus, Sweden (G. Holmström); 3, B. polaris, Canada (B. Heinrich); 4, B. balteatus, Sweden (G. Holmström); 5, B. kirbiellus, USA (D. Wilson); 6, B. hyperboreus, Sweden (L.-I. Larsson). Some images reversed.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB75E13FF685CF070A2F829.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268181/files/figure.png	https://doi.org/10.5281/zenodo.3268181	FIGURE 7. Sample sites and climatic suitability for all Alpinobombus species combined estimated using Maxent from climate variables for the year 2000. Bright yellow and brown areas show where the logistic prediction of occurrence is p> 0.5 (the outlier records with exceptionally low probabilities of suitability were excluded in step-wise iterations); brown spots show all consistent Alpinobombus site records; ‘?’ shows the five records that are in climatically outlying sites. Polar projection, North Pole (starred) at the centre of the map, international boundaries and the Arctic Circle shown as narrow grey lines.	FIGURE 7. Sample sites and climatic suitability for all Alpinobombus species combined estimated using Maxent from climate variables for the year 2000. Bright yellow and brown areas show where the logistic prediction of occurrence is p> 0.5 (the outlier records with exceptionally low probabilities of suitability were excluded in step-wise iterations); brown spots show all consistent Alpinobombus site records; ‘?’ shows the five records that are in climatically outlying sites. Polar projection, North Pole (starred) at the centre of the map, international boundaries and the Arctic Circle shown as narrow grey lines.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB75E13FF685CF070A2F829.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268185/files/figure.png	https://doi.org/10.5281/zenodo.3268185	FIGURE 8. Diagram representing a corridor-dispersal model, encompassing a set of the short-distance dispersal events permitted (in either direction) between the areas defined in Table 5 (including the outgroup B. ignitus), based on their geographical proximity and the likely disposition of suitable habitat and favourable climates in the past (see the text).	FIGURE 8. Diagram representing a corridor-dispersal model, encompassing a set of the short-distance dispersal events permitted (in either direction) between the areas defined in Table 5 (including the outgroup B. ignitus), based on their geographical proximity and the likely disposition of suitable habitat and favourable climates in the past (see the text).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB75E13FF685CF070A2F829.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268187/files/figure.png	https://doi.org/10.5281/zenodo.3268187	FIGURE 9. Estimate of the metric gene tree for the 54 longest unique COI-barcode alleles of the subgenus Alpinobombus from MrBayes combined with the Bayesian Poisson-tree-process (PTP) solution with the highest support with spots showing the nodes with the strongest support as coalescents for candidate species (outgroup B. ignitus not shown). Values above the nodes are Bayesian posterior probabilities showing branch support for groups; values below the nodes are PTP Bayesian support values that all daughter alleles are parts of a single species (probabilities within candidate species are all ≤ 0.44). The scale bar is calibrated in substitutions per nucleotide site. Each unique allele is represented by one of the longest available sample sequences, labelled with: the sequence length in number of nucleotides, a taxon name or name describing an unpublished colour pattern, and then a code that consists of a specimen identifier (AL#n) from the project database and (after the hyphen) a sample identifier from BOLD or GenBank, followed with its geographic origin (for simplicity Greenland is included without Denmark).	FIGURE 9. Estimate of the metric gene tree for the 54 longest unique COI-barcode alleles of the subgenus Alpinobombus from MrBayes combined with the Bayesian Poisson-tree-process (PTP) solution with the highest support with spots showing the nodes with the strongest support as coalescents for candidate species (outgroup B. ignitus not shown). Values above the nodes are Bayesian posterior probabilities showing branch support for groups; values below the nodes are PTP Bayesian support values that all daughter alleles are parts of a single species (probabilities within candidate species are all ≤ 0.44). The scale bar is calibrated in substitutions per nucleotide site. Each unique allele is represented by one of the longest available sample sequences, labelled with: the sequence length in number of nucleotides, a taxon name or name describing an unpublished colour pattern, and then a code that consists of a specimen identifier (AL#n) from the project database and (after the hyphen) a sample identifier from BOLD or GenBank, followed with its geographic origin (for simplicity Greenland is included without Denmark).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB75E13FF685CF070A2F829.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268189/files/figure.png	https://doi.org/10.5281/zenodo.3268189	FIGURE 10. Map of the entire global distribution of records of B. kluanensis s. l.. Grey crosses are records for any Alpinobombus species; black crosses are records of B. neoboreus; overlayed with small white spots for specimens of B. kluanensis s. l.; large green spots for sequenced samples with the kluanensis s. str. alleles (collected 2010‒2016); small red spots for sequenced samples with the ‘unnamed2’ allele (collected 2015‒2017). Relief map with hill shading, with a Polar projection, and with the Arctic Circle and the USA (Alaska) / Canada (Yukon) international boundary shown as grey lines. North pointer and scale bar below, with an inset that shows the location of the detailed area of the global map. Image created in ArcGIS using World_Shad- ed_Relief basemap which is Copyright: © 2014 Esri.	FIGURE 10. Map of the entire global distribution of records of B. kluanensis s. l.. Grey crosses are records for any Alpinobombus species; black crosses are records of B. neoboreus; overlayed with small white spots for specimens of B. kluanensis s. l.; large green spots for sequenced samples with the kluanensis s. str. alleles (collected 2010‒2016); small red spots for sequenced samples with the ‘unnamed2’ allele (collected 2015‒2017). Relief map with hill shading, with a Polar projection, and with the Arctic Circle and the USA (Alaska) / Canada (Yukon) international boundary shown as grey lines. North pointer and scale bar below, with an inset that shows the location of the detailed area of the global map. Image created in ArcGIS using World_Shad- ed_Relief basemap which is Copyright: © 2014 Esri.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB75E13FF685CF070A2F829.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268191/files/figure.png	https://doi.org/10.5281/zenodo.3268191	FIGURES 11‒13. 11: gene trees for COI; 12: gene trees for 16S; and 13: species trees from *BEAST analysis of data from Thanoosing (2017) displayed using Densitree. Each plot shows the background density of trees from samples of 10,000 Bayesian trees, with the summary ‘root canal’ tree superimposed as a thick line, and the branches exclusively linking samples of B. kluanensis s. l. (#4850, 4869, 4870, 4873) traced in red.	FIGURES 11‒13. 11: gene trees for COI; 12: gene trees for 16S; and 13: species trees from *BEAST analysis of data from Thanoosing (2017) displayed using Densitree. Each plot shows the background density of trees from samples of 10,000 Bayesian trees, with the summary ‘root canal’ tree superimposed as a thick line, and the branches exclusively linking samples of B. kluanensis s. l. (#4850, 4869, 4870, 4873) traced in red.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB75E13FF685CF070A2F829.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268193/files/figure.png	https://doi.org/10.5281/zenodo.3268193	FIGURE 14. Assigning names to candidate species in a COI gene tree for the 235 specimens of the subgenus Alpinobombus with sequences. Based on the MrBayes analysis of all of the 54 longest unique alleles in Fig. 9, with the remaining sequences interpolated back into the tree with 0 branch lengths from their matching samples (where a sequence matches more than one longer sequence within a species in the Collapse results, it is arbitrarily added after the first matching sequence). The sequence labels follow the format used in Fig. 9. Lineages with high probabilities of representing candidate species in the Poisson-tree-process (PTP) results (Fig. 9) are shown as thick branches with the most recent common ancestors (coalescents) of each candidate species shown with a black spot, the branches within candidate species shown as thin branches. Asterisks mark samples used as informal proxies for the type specimens of each of the taxon names in Table 2. The proxy sample for the type specimen for the oldest available name (the valid name) for each candidate species (from Fig. 9) is marked in bold.	FIGURE 14. Assigning names to candidate species in a COI gene tree for the 235 specimens of the subgenus Alpinobombus with sequences. Based on the MrBayes analysis of all of the 54 longest unique alleles in Fig. 9, with the remaining sequences interpolated back into the tree with 0 branch lengths from their matching samples (where a sequence matches more than one longer sequence within a species in the Collapse results, it is arbitrarily added after the first matching sequence). The sequence labels follow the format used in Fig. 9. Lineages with high probabilities of representing candidate species in the Poisson-tree-process (PTP) results (Fig. 9) are shown as thick branches with the most recent common ancestors (coalescents) of each candidate species shown with a black spot, the branches within candidate species shown as thin branches. Asterisks mark samples used as informal proxies for the type specimens of each of the taxon names in Table 2. The proxy sample for the type specimen for the oldest available name (the valid name) for each candidate species (from Fig. 9) is marked in bold.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB75E13FF685CF070A2F829.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268195/files/figure.png	https://doi.org/10.5281/zenodo.3268195	FIGURES 15‒21. 15: Bombus alpinus global distribution shown with: grey spots for specimens examined; black spots for samples with COI barcodes; question marks for uncertain records at low elevation and low latitude; and grey crosses for records of all species of Alpinobombus combined (Alpinobombus species are unknown from the southern hemisphere). Relief map with hill shading, Polar projection, the international boundaries and the Arctic Circle are shown as narrow grey lines, and the northern tree line shown as a broad grey line. Image created in ArcGIS using World_Shaded_Relief basemap which is Copyright: © 2014 Esri. 16‒21: simplified colour-pattern diagrams for the dorsal hair, with olive indicating a mixture of black and yellow. Figure (AL#n) country: 16 (#14) Sweden; 17 (#113) Sweden; 18 (#3785) Austria; 19 (#3787) Austria; 20 (#3788) Austria; 21 (#141) Norway.	FIGURES 15‒21. 15: Bombus alpinus global distribution shown with: grey spots for specimens examined; black spots for samples with COI barcodes; question marks for uncertain records at low elevation and low latitude; and grey crosses for records of all species of Alpinobombus combined (Alpinobombus species are unknown from the southern hemisphere). Relief map with hill shading, Polar projection, the international boundaries and the Arctic Circle are shown as narrow grey lines, and the northern tree line shown as a broad grey line. Image created in ArcGIS using World_Shaded_Relief basemap which is Copyright: © 2014 Esri. 16‒21: simplified colour-pattern diagrams for the dorsal hair, with olive indicating a mixture of black and yellow. Figure (AL#n) country: 16 (#14) Sweden; 17 (#113) Sweden; 18 (#3785) Austria; 19 (#3787) Austria; 20 (#3788) Austria; 21 (#141) Norway.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB75E13FF685CF070A2F829.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268197/files/figure.png	https://doi.org/10.5281/zenodo.3268197	FIGURES 22‒38. 22: Bombus pyrrhopygus global distribution (as for Fig. 15). 23‒38 colour patterns (as for Figs. 16‒21): 23 (#4707) Russia-Arkhangelsk; 24 (#337) Russia-Arkhangelsk; 25 (#273) Sweden; 26 (#272) Sweden; 27 (#250) Sweden; 28 (#249) Sweden; 29 (#258) Sweden; 30 (#4709) Russia-Murmansk; 31 (#257) Sweden; 32 (#4711) Russia-Arkhangelsk; 33 (#4712) Russia-Arkhangelsk; 34 (#4713) Russia-Arkhangelsk; 35 (#242) Norway; 36 (#139) Sweden; 37 (#4710) Norway; 38 (#1375) Russia-Murmansk.	FIGURES 22‒38. 22: Bombus pyrrhopygus global distribution (as for Fig. 15). 23‒38 colour patterns (as for Figs. 16‒21): 23 (#4707) Russia-Arkhangelsk; 24 (#337) Russia-Arkhangelsk; 25 (#273) Sweden; 26 (#272) Sweden; 27 (#250) Sweden; 28 (#249) Sweden; 29 (#258) Sweden; 30 (#4709) Russia-Murmansk; 31 (#257) Sweden; 32 (#4711) Russia-Arkhangelsk; 33 (#4712) Russia-Arkhangelsk; 34 (#4713) Russia-Arkhangelsk; 35 (#242) Norway; 36 (#139) Sweden; 37 (#4710) Norway; 38 (#1375) Russia-Murmansk.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB75E13FF685CF070A2F829.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268199/files/figure.png	https://doi.org/10.5281/zenodo.3268199	FIGURES 39‒58. 39: Bombus polaris global distribution (as for Fig. 15). 40‒58 colour patterns (as for Figs. 16‒21): 40 (#4380) Canada-Labrador; 41 (#4396) USA-Alaska; 42 (#3856) Canada-Nunavut; 43 (#59) USA-Alaska; 44 (#43) Denmark- Greenland; 45 (#4740) Denmark-Greenland; 46 (#4407) USA-Alaska; 47 (#334) USA-Alaska; 48 (#285) Canada-Nunavut; 49 (#4383) USA-Alaska; 50 (#44) Denmark-Greenland; 51 (#19) Canada-Northwest Territories; 52 (#4401) USA-Alaska; 53 (#4403) USA-Alaska; 54 (#4884) Canada-Yukon; 55 (#4405) USA-Alaska; 56 (#4384) USA-Alaska; 57 (#4951) USA-Alaska; 58 (#29) Canada-Manitoba.	FIGURES 39‒58. 39: Bombus polaris global distribution (as for Fig. 15). 40‒58 colour patterns (as for Figs. 16‒21): 40 (#4380) Canada-Labrador; 41 (#4396) USA-Alaska; 42 (#3856) Canada-Nunavut; 43 (#59) USA-Alaska; 44 (#43) Denmark- Greenland; 45 (#4740) Denmark-Greenland; 46 (#4407) USA-Alaska; 47 (#334) USA-Alaska; 48 (#285) Canada-Nunavut; 49 (#4383) USA-Alaska; 50 (#44) Denmark-Greenland; 51 (#19) Canada-Northwest Territories; 52 (#4401) USA-Alaska; 53 (#4403) USA-Alaska; 54 (#4884) Canada-Yukon; 55 (#4405) USA-Alaska; 56 (#4384) USA-Alaska; 57 (#4951) USA-Alaska; 58 (#29) Canada-Manitoba.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB75E13FF685CF070A2F829.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268201/files/figure.png	https://doi.org/10.5281/zenodo.3268201	FIGURES 59‒76. 59: Bombus balteatus global distribution (as for Fig. 15). 60‒76 colour patterns (as for Figs.16‒21): 60 (#540) Russia-Krasnoyask; 61 (#119) Sweden; 62 (#541) Russia-Krasnoyarsk; 63 (#542) Russia-Krasnoyarsk; 64 (#117) Mongolia; 65 (#534) Russia-Kamchatka; 66 (#3841) Norway; 67 (#738) Russia-Kamchatka; 68 (#971) Russia- Kamchatka; 69 (#535) Russia-Kamchatka; 70 (#973) Russia-Kamchatka; 71 (#880) Russia-Kamchatka; 72 (#536) Russia-Kamchatka; 73 (#228) Norway; 74 (#392) Sweden; 75 (#3843) Norway; 76 (#538) Russia-Kamchatka.	FIGURES 59‒76. 59: Bombus balteatus global distribution (as for Fig. 15). 60‒76 colour patterns (as for Figs.16‒21): 60 (#540) Russia-Krasnoyask; 61 (#119) Sweden; 62 (#541) Russia-Krasnoyarsk; 63 (#542) Russia-Krasnoyarsk; 64 (#117) Mongolia; 65 (#534) Russia-Kamchatka; 66 (#3841) Norway; 67 (#738) Russia-Kamchatka; 68 (#971) Russia- Kamchatka; 69 (#535) Russia-Kamchatka; 70 (#973) Russia-Kamchatka; 71 (#880) Russia-Kamchatka; 72 (#536) Russia-Kamchatka; 73 (#228) Norway; 74 (#392) Sweden; 75 (#3843) Norway; 76 (#538) Russia-Kamchatka.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB75E13FF685CF070A2F829.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268203/files/figure.png	https://doi.org/10.5281/zenodo.3268203	FIGURES 77‒88. 77: Bombus kirbiellus global distribution (as for Fig. 15). 78‒88 colour patterns (as for Figs. 16‒21): 78 (#556) Canada-Nunavut; 79 (#294) Canada-Nunavut; 80 (#100) Canada-Nunavut; 81 (#3791) USA-Colorado; 82 (#3790) USA-Colorado; 83 (#3765) Canada-Nunavut; 84 (#3792) USA-Colorado; 85 (#3797) USA-Colorado; 86 (#4915) USA Alaska; 87 (#3768) Canada-Nunavut; 88 (#3812) USA-Colorado.	FIGURES 77‒88. 77: Bombus kirbiellus global distribution (as for Fig. 15). 78‒88 colour patterns (as for Figs. 16‒21): 78 (#556) Canada-Nunavut; 79 (#294) Canada-Nunavut; 80 (#100) Canada-Nunavut; 81 (#3791) USA-Colorado; 82 (#3790) USA-Colorado; 83 (#3765) Canada-Nunavut; 84 (#3792) USA-Colorado; 85 (#3797) USA-Colorado; 86 (#4915) USA Alaska; 87 (#3768) Canada-Nunavut; 88 (#3812) USA-Colorado.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB75E13FF685CF070A2F829.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268205/files/figure.png	https://doi.org/10.5281/zenodo.3268205	FIGURES 89‒103. 89: Bombus neoboreus global distribution (as for Fig. 15). 90‒103 colour patterns (as for Figs. 16‒21): 90 (#68) USA-Alaska; 91 (#4706) Canada-Nunavut; 92 (#3816) Canada-Yukon; 93 (#3819) USA-Alaska; 94 (#2055) Canada-Nunavut; 95 (#2058) Canada-Nunavut; 96 (#2071) Canada-Nunavut; 97 (#2051) Canada-Nunavut; 98 (#4406) Canada-Nunavut; 99 (#90) Canada-Nunavut; 100 (#92) Canada-Nunavut; 101 (#93) Canada-Yukon; 102 (#3762) Canada-Nunavut; 103 (#91) Canada-Nunavut.	FIGURES 89‒103. 89: Bombus neoboreus global distribution (as for Fig. 15). 90‒103 colour patterns (as for Figs. 16‒21): 90 (#68) USA-Alaska; 91 (#4706) Canada-Nunavut; 92 (#3816) Canada-Yukon; 93 (#3819) USA-Alaska; 94 (#2055) Canada-Nunavut; 95 (#2058) Canada-Nunavut; 96 (#2071) Canada-Nunavut; 97 (#2051) Canada-Nunavut; 98 (#4406) Canada-Nunavut; 99 (#90) Canada-Nunavut; 100 (#92) Canada-Nunavut; 101 (#93) Canada-Yukon; 102 (#3762) Canada-Nunavut; 103 (#91) Canada-Nunavut.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB25E16FF685FF47141F976.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268177/files/figure.png	https://doi.org/10.5281/zenodo.3268177	FIGURES 1‒6. Lateral views of foraging females (queens) of species of the subgenus Alpinobombus (with photo credits): 1, B. alpinus, Norway (A. Staverløkk); 2, B. pyrrhopygus, Sweden (G. Holmström); 3, B. polaris, Canada (B. Heinrich); 4, B. balteatus, Sweden (G. Holmström); 5, B. kirbiellus, USA (D. Wilson); 6, B. hyperboreus, Sweden (L.-I. Larsson). Some images reversed.	FIGURES 1‒6. Lateral views of foraging females (queens) of species of the subgenus Alpinobombus (with photo credits): 1, B. alpinus, Norway (A. Staverløkk); 2, B. pyrrhopygus, Sweden (G. Holmström); 3, B. polaris, Canada (B. Heinrich); 4, B. balteatus, Sweden (G. Holmström); 5, B. kirbiellus, USA (D. Wilson); 6, B. hyperboreus, Sweden (L.-I. Larsson). Some images reversed.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB25E16FF685FF47141F976.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268195/files/figure.png	https://doi.org/10.5281/zenodo.3268195	FIGURES 15‒21. 15: Bombus alpinus global distribution shown with: grey spots for specimens examined; black spots for samples with COI barcodes; question marks for uncertain records at low elevation and low latitude; and grey crosses for records of all species of Alpinobombus combined (Alpinobombus species are unknown from the southern hemisphere). Relief map with hill shading, Polar projection, the international boundaries and the Arctic Circle are shown as narrow grey lines, and the northern tree line shown as a broad grey line. Image created in ArcGIS using World_Shaded_Relief basemap which is Copyright: © 2014 Esri. 16‒21: simplified colour-pattern diagrams for the dorsal hair, with olive indicating a mixture of black and yellow. Figure (AL#n) country: 16 (#14) Sweden; 17 (#113) Sweden; 18 (#3785) Austria; 19 (#3787) Austria; 20 (#3788) Austria; 21 (#141) Norway.	FIGURES 15‒21. 15: Bombus alpinus global distribution shown with: grey spots for specimens examined; black spots for samples with COI barcodes; question marks for uncertain records at low elevation and low latitude; and grey crosses for records of all species of Alpinobombus combined (Alpinobombus species are unknown from the southern hemisphere). Relief map with hill shading, Polar projection, the international boundaries and the Arctic Circle are shown as narrow grey lines, and the northern tree line shown as a broad grey line. Image created in ArcGIS using World_Shaded_Relief basemap which is Copyright: © 2014 Esri. 16‒21: simplified colour-pattern diagrams for the dorsal hair, with olive indicating a mixture of black and yellow. Figure (AL#n) country: 16 (#14) Sweden; 17 (#113) Sweden; 18 (#3785) Austria; 19 (#3787) Austria; 20 (#3788) Austria; 21 (#141) Norway.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB25E16FF685FF47141F976.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268213/files/figure.png	https://doi.org/10.5281/zenodo.3268213	FIGURES 129‒137. Morphology of the male genitalia for species of the subgenus Alpinobombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 129 B. alpinus (#3824); 130 B pyrrhopygus (#3833); 131 B. polaris (#73); 132 B. balteatus (#3843); 133 B. kirbiellus (#3768); 134 B. neoboreus (#4387); 135 B. kluanensis (#4876); 136 B. natvigi (#81); 137 B. hyperboreus (#3821).	FIGURES 129‒137. Morphology of the male genitalia for species of the subgenus Alpinobombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 129 B. alpinus (#3824); 130 B pyrrhopygus (#3833); 131 B. polaris (#73); 132 B. balteatus (#3843); 133 B. kirbiellus (#3768); 134 B. neoboreus (#4387); 135 B. kluanensis (#4876); 136 B. natvigi (#81); 137 B. hyperboreus (#3821).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB25E16FF685FF47141F976.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268187/files/figure.png	https://doi.org/10.5281/zenodo.3268187	FIGURE 9. Estimate of the metric gene tree for the 54 longest unique COI-barcode alleles of the subgenus Alpinobombus from MrBayes combined with the Bayesian Poisson-tree-process (PTP) solution with the highest support with spots showing the nodes with the strongest support as coalescents for candidate species (outgroup B. ignitus not shown). Values above the nodes are Bayesian posterior probabilities showing branch support for groups; values below the nodes are PTP Bayesian support values that all daughter alleles are parts of a single species (probabilities within candidate species are all ≤ 0.44). The scale bar is calibrated in substitutions per nucleotide site. Each unique allele is represented by one of the longest available sample sequences, labelled with: the sequence length in number of nucleotides, a taxon name or name describing an unpublished colour pattern, and then a code that consists of a specimen identifier (AL#n) from the project database and (after the hyphen) a sample identifier from BOLD or GenBank, followed with its geographic origin (for simplicity Greenland is included without Denmark).	FIGURE 9. Estimate of the metric gene tree for the 54 longest unique COI-barcode alleles of the subgenus Alpinobombus from MrBayes combined with the Bayesian Poisson-tree-process (PTP) solution with the highest support with spots showing the nodes with the strongest support as coalescents for candidate species (outgroup B. ignitus not shown). Values above the nodes are Bayesian posterior probabilities showing branch support for groups; values below the nodes are PTP Bayesian support values that all daughter alleles are parts of a single species (probabilities within candidate species are all ≤ 0.44). The scale bar is calibrated in substitutions per nucleotide site. Each unique allele is represented by one of the longest available sample sequences, labelled with: the sequence length in number of nucleotides, a taxon name or name describing an unpublished colour pattern, and then a code that consists of a specimen identifier (AL#n) from the project database and (after the hyphen) a sample identifier from BOLD or GenBank, followed with its geographic origin (for simplicity Greenland is included without Denmark).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB25E16FF685FF47141F976.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268193/files/figure.png	https://doi.org/10.5281/zenodo.3268193	FIGURE 14. Assigning names to candidate species in a COI gene tree for the 235 specimens of the subgenus Alpinobombus with sequences. Based on the MrBayes analysis of all of the 54 longest unique alleles in Fig. 9, with the remaining sequences interpolated back into the tree with 0 branch lengths from their matching samples (where a sequence matches more than one longer sequence within a species in the Collapse results, it is arbitrarily added after the first matching sequence). The sequence labels follow the format used in Fig. 9. Lineages with high probabilities of representing candidate species in the Poisson-tree-process (PTP) results (Fig. 9) are shown as thick branches with the most recent common ancestors (coalescents) of each candidate species shown with a black spot, the branches within candidate species shown as thin branches. Asterisks mark samples used as informal proxies for the type specimens of each of the taxon names in Table 2. The proxy sample for the type specimen for the oldest available name (the valid name) for each candidate species (from Fig. 9) is marked in bold.	FIGURE 14. Assigning names to candidate species in a COI gene tree for the 235 specimens of the subgenus Alpinobombus with sequences. Based on the MrBayes analysis of all of the 54 longest unique alleles in Fig. 9, with the remaining sequences interpolated back into the tree with 0 branch lengths from their matching samples (where a sequence matches more than one longer sequence within a species in the Collapse results, it is arbitrarily added after the first matching sequence). The sequence labels follow the format used in Fig. 9. Lineages with high probabilities of representing candidate species in the Poisson-tree-process (PTP) results (Fig. 9) are shown as thick branches with the most recent common ancestors (coalescents) of each candidate species shown with a black spot, the branches within candidate species shown as thin branches. Asterisks mark samples used as informal proxies for the type specimens of each of the taxon names in Table 2. The proxy sample for the type specimen for the oldest available name (the valid name) for each candidate species (from Fig. 9) is marked in bold.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB25E1BFF685951704FFC48.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268177/files/figure.png	https://doi.org/10.5281/zenodo.3268177	FIGURES 1‒6. Lateral views of foraging females (queens) of species of the subgenus Alpinobombus (with photo credits): 1, B. alpinus, Norway (A. Staverløkk); 2, B. pyrrhopygus, Sweden (G. Holmström); 3, B. polaris, Canada (B. Heinrich); 4, B. balteatus, Sweden (G. Holmström); 5, B. kirbiellus, USA (D. Wilson); 6, B. hyperboreus, Sweden (L.-I. Larsson). Some images reversed.	FIGURES 1‒6. Lateral views of foraging females (queens) of species of the subgenus Alpinobombus (with photo credits): 1, B. alpinus, Norway (A. Staverløkk); 2, B. pyrrhopygus, Sweden (G. Holmström); 3, B. polaris, Canada (B. Heinrich); 4, B. balteatus, Sweden (G. Holmström); 5, B. kirbiellus, USA (D. Wilson); 6, B. hyperboreus, Sweden (L.-I. Larsson). Some images reversed.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB25E1BFF685951704FFC48.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268197/files/figure.png	https://doi.org/10.5281/zenodo.3268197	FIGURES 22‒38. 22: Bombus pyrrhopygus global distribution (as for Fig. 15). 23‒38 colour patterns (as for Figs. 16‒21): 23 (#4707) Russia-Arkhangelsk; 24 (#337) Russia-Arkhangelsk; 25 (#273) Sweden; 26 (#272) Sweden; 27 (#250) Sweden; 28 (#249) Sweden; 29 (#258) Sweden; 30 (#4709) Russia-Murmansk; 31 (#257) Sweden; 32 (#4711) Russia-Arkhangelsk; 33 (#4712) Russia-Arkhangelsk; 34 (#4713) Russia-Arkhangelsk; 35 (#242) Norway; 36 (#139) Sweden; 37 (#4710) Norway; 38 (#1375) Russia-Murmansk.	FIGURES 22‒38. 22: Bombus pyrrhopygus global distribution (as for Fig. 15). 23‒38 colour patterns (as for Figs. 16‒21): 23 (#4707) Russia-Arkhangelsk; 24 (#337) Russia-Arkhangelsk; 25 (#273) Sweden; 26 (#272) Sweden; 27 (#250) Sweden; 28 (#249) Sweden; 29 (#258) Sweden; 30 (#4709) Russia-Murmansk; 31 (#257) Sweden; 32 (#4711) Russia-Arkhangelsk; 33 (#4712) Russia-Arkhangelsk; 34 (#4713) Russia-Arkhangelsk; 35 (#242) Norway; 36 (#139) Sweden; 37 (#4710) Norway; 38 (#1375) Russia-Murmansk.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB25E1BFF685951704FFC48.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268213/files/figure.png	https://doi.org/10.5281/zenodo.3268213	FIGURES 129‒137. Morphology of the male genitalia for species of the subgenus Alpinobombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 129 B. alpinus (#3824); 130 B pyrrhopygus (#3833); 131 B. polaris (#73); 132 B. balteatus (#3843); 133 B. kirbiellus (#3768); 134 B. neoboreus (#4387); 135 B. kluanensis (#4876); 136 B. natvigi (#81); 137 B. hyperboreus (#3821).	FIGURES 129‒137. Morphology of the male genitalia for species of the subgenus Alpinobombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 129 B. alpinus (#3824); 130 B pyrrhopygus (#3833); 131 B. polaris (#73); 132 B. balteatus (#3843); 133 B. kirbiellus (#3768); 134 B. neoboreus (#4387); 135 B. kluanensis (#4876); 136 B. natvigi (#81); 137 B. hyperboreus (#3821).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB25E1BFF685951704FFC48.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268187/files/figure.png	https://doi.org/10.5281/zenodo.3268187	FIGURE 9. Estimate of the metric gene tree for the 54 longest unique COI-barcode alleles of the subgenus Alpinobombus from MrBayes combined with the Bayesian Poisson-tree-process (PTP) solution with the highest support with spots showing the nodes with the strongest support as coalescents for candidate species (outgroup B. ignitus not shown). Values above the nodes are Bayesian posterior probabilities showing branch support for groups; values below the nodes are PTP Bayesian support values that all daughter alleles are parts of a single species (probabilities within candidate species are all ≤ 0.44). The scale bar is calibrated in substitutions per nucleotide site. Each unique allele is represented by one of the longest available sample sequences, labelled with: the sequence length in number of nucleotides, a taxon name or name describing an unpublished colour pattern, and then a code that consists of a specimen identifier (AL#n) from the project database and (after the hyphen) a sample identifier from BOLD or GenBank, followed with its geographic origin (for simplicity Greenland is included without Denmark).	FIGURE 9. Estimate of the metric gene tree for the 54 longest unique COI-barcode alleles of the subgenus Alpinobombus from MrBayes combined with the Bayesian Poisson-tree-process (PTP) solution with the highest support with spots showing the nodes with the strongest support as coalescents for candidate species (outgroup B. ignitus not shown). Values above the nodes are Bayesian posterior probabilities showing branch support for groups; values below the nodes are PTP Bayesian support values that all daughter alleles are parts of a single species (probabilities within candidate species are all ≤ 0.44). The scale bar is calibrated in substitutions per nucleotide site. Each unique allele is represented by one of the longest available sample sequences, labelled with: the sequence length in number of nucleotides, a taxon name or name describing an unpublished colour pattern, and then a code that consists of a specimen identifier (AL#n) from the project database and (after the hyphen) a sample identifier from BOLD or GenBank, followed with its geographic origin (for simplicity Greenland is included without Denmark).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB25E1BFF685951704FFC48.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268199/files/figure.png	https://doi.org/10.5281/zenodo.3268199	FIGURES 39‒58. 39: Bombus polaris global distribution (as for Fig. 15). 40‒58 colour patterns (as for Figs. 16‒21): 40 (#4380) Canada-Labrador; 41 (#4396) USA-Alaska; 42 (#3856) Canada-Nunavut; 43 (#59) USA-Alaska; 44 (#43) Denmark- Greenland; 45 (#4740) Denmark-Greenland; 46 (#4407) USA-Alaska; 47 (#334) USA-Alaska; 48 (#285) Canada-Nunavut; 49 (#4383) USA-Alaska; 50 (#44) Denmark-Greenland; 51 (#19) Canada-Northwest Territories; 52 (#4401) USA-Alaska; 53 (#4403) USA-Alaska; 54 (#4884) Canada-Yukon; 55 (#4405) USA-Alaska; 56 (#4384) USA-Alaska; 57 (#4951) USA-Alaska; 58 (#29) Canada-Manitoba.	FIGURES 39‒58. 39: Bombus polaris global distribution (as for Fig. 15). 40‒58 colour patterns (as for Figs. 16‒21): 40 (#4380) Canada-Labrador; 41 (#4396) USA-Alaska; 42 (#3856) Canada-Nunavut; 43 (#59) USA-Alaska; 44 (#43) Denmark- Greenland; 45 (#4740) Denmark-Greenland; 46 (#4407) USA-Alaska; 47 (#334) USA-Alaska; 48 (#285) Canada-Nunavut; 49 (#4383) USA-Alaska; 50 (#44) Denmark-Greenland; 51 (#19) Canada-Northwest Territories; 52 (#4401) USA-Alaska; 53 (#4403) USA-Alaska; 54 (#4884) Canada-Yukon; 55 (#4405) USA-Alaska; 56 (#4384) USA-Alaska; 57 (#4951) USA-Alaska; 58 (#29) Canada-Manitoba.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB25E1BFF685951704FFC48.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268193/files/figure.png	https://doi.org/10.5281/zenodo.3268193	FIGURE 14. Assigning names to candidate species in a COI gene tree for the 235 specimens of the subgenus Alpinobombus with sequences. Based on the MrBayes analysis of all of the 54 longest unique alleles in Fig. 9, with the remaining sequences interpolated back into the tree with 0 branch lengths from their matching samples (where a sequence matches more than one longer sequence within a species in the Collapse results, it is arbitrarily added after the first matching sequence). The sequence labels follow the format used in Fig. 9. Lineages with high probabilities of representing candidate species in the Poisson-tree-process (PTP) results (Fig. 9) are shown as thick branches with the most recent common ancestors (coalescents) of each candidate species shown with a black spot, the branches within candidate species shown as thin branches. Asterisks mark samples used as informal proxies for the type specimens of each of the taxon names in Table 2. The proxy sample for the type specimen for the oldest available name (the valid name) for each candidate species (from Fig. 9) is marked in bold.	FIGURE 14. Assigning names to candidate species in a COI gene tree for the 235 specimens of the subgenus Alpinobombus with sequences. Based on the MrBayes analysis of all of the 54 longest unique alleles in Fig. 9, with the remaining sequences interpolated back into the tree with 0 branch lengths from their matching samples (where a sequence matches more than one longer sequence within a species in the Collapse results, it is arbitrarily added after the first matching sequence). The sequence labels follow the format used in Fig. 9. Lineages with high probabilities of representing candidate species in the Poisson-tree-process (PTP) results (Fig. 9) are shown as thick branches with the most recent common ancestors (coalescents) of each candidate species shown with a black spot, the branches within candidate species shown as thin branches. Asterisks mark samples used as informal proxies for the type specimens of each of the taxon names in Table 2. The proxy sample for the type specimen for the oldest available name (the valid name) for each candidate species (from Fig. 9) is marked in bold.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFB25E1BFF685951704FFC48.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268181/files/figure.png	https://doi.org/10.5281/zenodo.3268181	FIGURE 7. Sample sites and climatic suitability for all Alpinobombus species combined estimated using Maxent from climate variables for the year 2000. Bright yellow and brown areas show where the logistic prediction of occurrence is p> 0.5 (the outlier records with exceptionally low probabilities of suitability were excluded in step-wise iterations); brown spots show all consistent Alpinobombus site records; ‘?’ shows the five records that are in climatically outlying sites. Polar projection, North Pole (starred) at the centre of the map, international boundaries and the Arctic Circle shown as narrow grey lines.	FIGURE 7. Sample sites and climatic suitability for all Alpinobombus species combined estimated using Maxent from climate variables for the year 2000. Bright yellow and brown areas show where the logistic prediction of occurrence is p> 0.5 (the outlier records with exceptionally low probabilities of suitability were excluded in step-wise iterations); brown spots show all consistent Alpinobombus site records; ‘?’ shows the five records that are in climatically outlying sites. Polar projection, North Pole (starred) at the centre of the map, international boundaries and the Arctic Circle shown as narrow grey lines.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFBF5E1EFF685A707683FAD4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268177/files/figure.png	https://doi.org/10.5281/zenodo.3268177	FIGURES 1‒6. Lateral views of foraging females (queens) of species of the subgenus Alpinobombus (with photo credits): 1, B. alpinus, Norway (A. Staverløkk); 2, B. pyrrhopygus, Sweden (G. Holmström); 3, B. polaris, Canada (B. Heinrich); 4, B. balteatus, Sweden (G. Holmström); 5, B. kirbiellus, USA (D. Wilson); 6, B. hyperboreus, Sweden (L.-I. Larsson). Some images reversed.	FIGURES 1‒6. Lateral views of foraging females (queens) of species of the subgenus Alpinobombus (with photo credits): 1, B. alpinus, Norway (A. Staverløkk); 2, B. pyrrhopygus, Sweden (G. Holmström); 3, B. polaris, Canada (B. Heinrich); 4, B. balteatus, Sweden (G. Holmström); 5, B. kirbiellus, USA (D. Wilson); 6, B. hyperboreus, Sweden (L.-I. Larsson). Some images reversed.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFBF5E1EFF685A707683FAD4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268199/files/figure.png	https://doi.org/10.5281/zenodo.3268199	FIGURES 39‒58. 39: Bombus polaris global distribution (as for Fig. 15). 40‒58 colour patterns (as for Figs. 16‒21): 40 (#4380) Canada-Labrador; 41 (#4396) USA-Alaska; 42 (#3856) Canada-Nunavut; 43 (#59) USA-Alaska; 44 (#43) Denmark- Greenland; 45 (#4740) Denmark-Greenland; 46 (#4407) USA-Alaska; 47 (#334) USA-Alaska; 48 (#285) Canada-Nunavut; 49 (#4383) USA-Alaska; 50 (#44) Denmark-Greenland; 51 (#19) Canada-Northwest Territories; 52 (#4401) USA-Alaska; 53 (#4403) USA-Alaska; 54 (#4884) Canada-Yukon; 55 (#4405) USA-Alaska; 56 (#4384) USA-Alaska; 57 (#4951) USA-Alaska; 58 (#29) Canada-Manitoba.	FIGURES 39‒58. 39: Bombus polaris global distribution (as for Fig. 15). 40‒58 colour patterns (as for Figs. 16‒21): 40 (#4380) Canada-Labrador; 41 (#4396) USA-Alaska; 42 (#3856) Canada-Nunavut; 43 (#59) USA-Alaska; 44 (#43) Denmark- Greenland; 45 (#4740) Denmark-Greenland; 46 (#4407) USA-Alaska; 47 (#334) USA-Alaska; 48 (#285) Canada-Nunavut; 49 (#4383) USA-Alaska; 50 (#44) Denmark-Greenland; 51 (#19) Canada-Northwest Territories; 52 (#4401) USA-Alaska; 53 (#4403) USA-Alaska; 54 (#4884) Canada-Yukon; 55 (#4405) USA-Alaska; 56 (#4384) USA-Alaska; 57 (#4951) USA-Alaska; 58 (#29) Canada-Manitoba.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFBF5E1EFF685A707683FAD4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268213/files/figure.png	https://doi.org/10.5281/zenodo.3268213	FIGURES 129‒137. Morphology of the male genitalia for species of the subgenus Alpinobombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 129 B. alpinus (#3824); 130 B pyrrhopygus (#3833); 131 B. polaris (#73); 132 B. balteatus (#3843); 133 B. kirbiellus (#3768); 134 B. neoboreus (#4387); 135 B. kluanensis (#4876); 136 B. natvigi (#81); 137 B. hyperboreus (#3821).	FIGURES 129‒137. Morphology of the male genitalia for species of the subgenus Alpinobombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 129 B. alpinus (#3824); 130 B pyrrhopygus (#3833); 131 B. polaris (#73); 132 B. balteatus (#3843); 133 B. kirbiellus (#3768); 134 B. neoboreus (#4387); 135 B. kluanensis (#4876); 136 B. natvigi (#81); 137 B. hyperboreus (#3821).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFBF5E1EFF685A707683FAD4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268219/files/figure.png	https://doi.org/10.5281/zenodo.3268219	FIGURES 140‒143. Morphology of females (queens). Left lateral aspect of the oculo-malar area of: 140, B. polaris; 141, B. kirbiellus (white lines indicate length and breadth measurements). Left lateral aspect of the outer surface of the left hind tibia (corbicular area): 142, B. polaris; 143, B. neoboreus (white lines indicate highlights in the central area).	FIGURES 140‒143. Morphology of females (queens). Left lateral aspect of the oculo-malar area of: 140, B. polaris; 141, B. kirbiellus (white lines indicate length and breadth measurements). Left lateral aspect of the outer surface of the left hind tibia (corbicular area): 142, B. polaris; 143, B. neoboreus (white lines indicate highlights in the central area).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFBF5E1EFF685A707683FAD4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268221/files/figure.png	https://doi.org/10.5281/zenodo.3268221	FIGURES 144‒147. Morphology of males. Dorsal aspect of the penis valves of: 144, B. natvigi; 145, B. neoboreus (white lines indicate location of the subapical outer tooth of the left penis valve). Dorsal aspect of left gonostylus: 146 B. kirbiellus; 147, B. polaris (white lines indicate anterior, midpoint, and posterior median projections of left gonostylus).	FIGURES 144‒147. Morphology of males. Dorsal aspect of the penis valves of: 144, B. natvigi; 145, B. neoboreus (white lines indicate location of the subapical outer tooth of the left penis valve). Dorsal aspect of left gonostylus: 146 B. kirbiellus; 147, B. polaris (white lines indicate anterior, midpoint, and posterior median projections of left gonostylus).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFBF5E1EFF685A707683FAD4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268187/files/figure.png	https://doi.org/10.5281/zenodo.3268187	FIGURE 9. Estimate of the metric gene tree for the 54 longest unique COI-barcode alleles of the subgenus Alpinobombus from MrBayes combined with the Bayesian Poisson-tree-process (PTP) solution with the highest support with spots showing the nodes with the strongest support as coalescents for candidate species (outgroup B. ignitus not shown). Values above the nodes are Bayesian posterior probabilities showing branch support for groups; values below the nodes are PTP Bayesian support values that all daughter alleles are parts of a single species (probabilities within candidate species are all ≤ 0.44). The scale bar is calibrated in substitutions per nucleotide site. Each unique allele is represented by one of the longest available sample sequences, labelled with: the sequence length in number of nucleotides, a taxon name or name describing an unpublished colour pattern, and then a code that consists of a specimen identifier (AL#n) from the project database and (after the hyphen) a sample identifier from BOLD or GenBank, followed with its geographic origin (for simplicity Greenland is included without Denmark).	FIGURE 9. Estimate of the metric gene tree for the 54 longest unique COI-barcode alleles of the subgenus Alpinobombus from MrBayes combined with the Bayesian Poisson-tree-process (PTP) solution with the highest support with spots showing the nodes with the strongest support as coalescents for candidate species (outgroup B. ignitus not shown). Values above the nodes are Bayesian posterior probabilities showing branch support for groups; values below the nodes are PTP Bayesian support values that all daughter alleles are parts of a single species (probabilities within candidate species are all ≤ 0.44). The scale bar is calibrated in substitutions per nucleotide site. Each unique allele is represented by one of the longest available sample sequences, labelled with: the sequence length in number of nucleotides, a taxon name or name describing an unpublished colour pattern, and then a code that consists of a specimen identifier (AL#n) from the project database and (after the hyphen) a sample identifier from BOLD or GenBank, followed with its geographic origin (for simplicity Greenland is included without Denmark).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFBF5E1EFF685A707683FAD4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268197/files/figure.png	https://doi.org/10.5281/zenodo.3268197	FIGURES 22‒38. 22: Bombus pyrrhopygus global distribution (as for Fig. 15). 23‒38 colour patterns (as for Figs. 16‒21): 23 (#4707) Russia-Arkhangelsk; 24 (#337) Russia-Arkhangelsk; 25 (#273) Sweden; 26 (#272) Sweden; 27 (#250) Sweden; 28 (#249) Sweden; 29 (#258) Sweden; 30 (#4709) Russia-Murmansk; 31 (#257) Sweden; 32 (#4711) Russia-Arkhangelsk; 33 (#4712) Russia-Arkhangelsk; 34 (#4713) Russia-Arkhangelsk; 35 (#242) Norway; 36 (#139) Sweden; 37 (#4710) Norway; 38 (#1375) Russia-Murmansk.	FIGURES 22‒38. 22: Bombus pyrrhopygus global distribution (as for Fig. 15). 23‒38 colour patterns (as for Figs. 16‒21): 23 (#4707) Russia-Arkhangelsk; 24 (#337) Russia-Arkhangelsk; 25 (#273) Sweden; 26 (#272) Sweden; 27 (#250) Sweden; 28 (#249) Sweden; 29 (#258) Sweden; 30 (#4709) Russia-Murmansk; 31 (#257) Sweden; 32 (#4711) Russia-Arkhangelsk; 33 (#4712) Russia-Arkhangelsk; 34 (#4713) Russia-Arkhangelsk; 35 (#242) Norway; 36 (#139) Sweden; 37 (#4710) Norway; 38 (#1375) Russia-Murmansk.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFBA5E21FF685BC47176FB23.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268177/files/figure.png	https://doi.org/10.5281/zenodo.3268177	FIGURES 1‒6. Lateral views of foraging females (queens) of species of the subgenus Alpinobombus (with photo credits): 1, B. alpinus, Norway (A. Staverløkk); 2, B. pyrrhopygus, Sweden (G. Holmström); 3, B. polaris, Canada (B. Heinrich); 4, B. balteatus, Sweden (G. Holmström); 5, B. kirbiellus, USA (D. Wilson); 6, B. hyperboreus, Sweden (L.-I. Larsson). Some images reversed.	FIGURES 1‒6. Lateral views of foraging females (queens) of species of the subgenus Alpinobombus (with photo credits): 1, B. alpinus, Norway (A. Staverløkk); 2, B. pyrrhopygus, Sweden (G. Holmström); 3, B. polaris, Canada (B. Heinrich); 4, B. balteatus, Sweden (G. Holmström); 5, B. kirbiellus, USA (D. Wilson); 6, B. hyperboreus, Sweden (L.-I. Larsson). Some images reversed.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFBA5E21FF685BC47176FB23.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268201/files/figure.png	https://doi.org/10.5281/zenodo.3268201	FIGURES 59‒76. 59: Bombus balteatus global distribution (as for Fig. 15). 60‒76 colour patterns (as for Figs.16‒21): 60 (#540) Russia-Krasnoyask; 61 (#119) Sweden; 62 (#541) Russia-Krasnoyarsk; 63 (#542) Russia-Krasnoyarsk; 64 (#117) Mongolia; 65 (#534) Russia-Kamchatka; 66 (#3841) Norway; 67 (#738) Russia-Kamchatka; 68 (#971) Russia- Kamchatka; 69 (#535) Russia-Kamchatka; 70 (#973) Russia-Kamchatka; 71 (#880) Russia-Kamchatka; 72 (#536) Russia-Kamchatka; 73 (#228) Norway; 74 (#392) Sweden; 75 (#3843) Norway; 76 (#538) Russia-Kamchatka.	FIGURES 59‒76. 59: Bombus balteatus global distribution (as for Fig. 15). 60‒76 colour patterns (as for Figs.16‒21): 60 (#540) Russia-Krasnoyask; 61 (#119) Sweden; 62 (#541) Russia-Krasnoyarsk; 63 (#542) Russia-Krasnoyarsk; 64 (#117) Mongolia; 65 (#534) Russia-Kamchatka; 66 (#3841) Norway; 67 (#738) Russia-Kamchatka; 68 (#971) Russia- Kamchatka; 69 (#535) Russia-Kamchatka; 70 (#973) Russia-Kamchatka; 71 (#880) Russia-Kamchatka; 72 (#536) Russia-Kamchatka; 73 (#228) Norway; 74 (#392) Sweden; 75 (#3843) Norway; 76 (#538) Russia-Kamchatka.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFBA5E21FF685BC47176FB23.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268213/files/figure.png	https://doi.org/10.5281/zenodo.3268213	FIGURES 129‒137. Morphology of the male genitalia for species of the subgenus Alpinobombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 129 B. alpinus (#3824); 130 B pyrrhopygus (#3833); 131 B. polaris (#73); 132 B. balteatus (#3843); 133 B. kirbiellus (#3768); 134 B. neoboreus (#4387); 135 B. kluanensis (#4876); 136 B. natvigi (#81); 137 B. hyperboreus (#3821).	FIGURES 129‒137. Morphology of the male genitalia for species of the subgenus Alpinobombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 129 B. alpinus (#3824); 130 B pyrrhopygus (#3833); 131 B. polaris (#73); 132 B. balteatus (#3843); 133 B. kirbiellus (#3768); 134 B. neoboreus (#4387); 135 B. kluanensis (#4876); 136 B. natvigi (#81); 137 B. hyperboreus (#3821).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFBA5E21FF685BC47176FB23.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268187/files/figure.png	https://doi.org/10.5281/zenodo.3268187	FIGURE 9. Estimate of the metric gene tree for the 54 longest unique COI-barcode alleles of the subgenus Alpinobombus from MrBayes combined with the Bayesian Poisson-tree-process (PTP) solution with the highest support with spots showing the nodes with the strongest support as coalescents for candidate species (outgroup B. ignitus not shown). Values above the nodes are Bayesian posterior probabilities showing branch support for groups; values below the nodes are PTP Bayesian support values that all daughter alleles are parts of a single species (probabilities within candidate species are all ≤ 0.44). The scale bar is calibrated in substitutions per nucleotide site. Each unique allele is represented by one of the longest available sample sequences, labelled with: the sequence length in number of nucleotides, a taxon name or name describing an unpublished colour pattern, and then a code that consists of a specimen identifier (AL#n) from the project database and (after the hyphen) a sample identifier from BOLD or GenBank, followed with its geographic origin (for simplicity Greenland is included without Denmark).	FIGURE 9. Estimate of the metric gene tree for the 54 longest unique COI-barcode alleles of the subgenus Alpinobombus from MrBayes combined with the Bayesian Poisson-tree-process (PTP) solution with the highest support with spots showing the nodes with the strongest support as coalescents for candidate species (outgroup B. ignitus not shown). Values above the nodes are Bayesian posterior probabilities showing branch support for groups; values below the nodes are PTP Bayesian support values that all daughter alleles are parts of a single species (probabilities within candidate species are all ≤ 0.44). The scale bar is calibrated in substitutions per nucleotide site. Each unique allele is represented by one of the longest available sample sequences, labelled with: the sequence length in number of nucleotides, a taxon name or name describing an unpublished colour pattern, and then a code that consists of a specimen identifier (AL#n) from the project database and (after the hyphen) a sample identifier from BOLD or GenBank, followed with its geographic origin (for simplicity Greenland is included without Denmark).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFBA5E21FF685BC47176FB23.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268203/files/figure.png	https://doi.org/10.5281/zenodo.3268203	FIGURES 77‒88. 77: Bombus kirbiellus global distribution (as for Fig. 15). 78‒88 colour patterns (as for Figs. 16‒21): 78 (#556) Canada-Nunavut; 79 (#294) Canada-Nunavut; 80 (#100) Canada-Nunavut; 81 (#3791) USA-Colorado; 82 (#3790) USA-Colorado; 83 (#3765) Canada-Nunavut; 84 (#3792) USA-Colorado; 85 (#3797) USA-Colorado; 86 (#4915) USA Alaska; 87 (#3768) Canada-Nunavut; 88 (#3812) USA-Colorado.	FIGURES 77‒88. 77: Bombus kirbiellus global distribution (as for Fig. 15). 78‒88 colour patterns (as for Figs. 16‒21): 78 (#556) Canada-Nunavut; 79 (#294) Canada-Nunavut; 80 (#100) Canada-Nunavut; 81 (#3791) USA-Colorado; 82 (#3790) USA-Colorado; 83 (#3765) Canada-Nunavut; 84 (#3792) USA-Colorado; 85 (#3797) USA-Colorado; 86 (#4915) USA Alaska; 87 (#3768) Canada-Nunavut; 88 (#3812) USA-Colorado.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFFBA5E21FF685BC47176FB23.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268181/files/figure.png	https://doi.org/10.5281/zenodo.3268181	FIGURE 7. Sample sites and climatic suitability for all Alpinobombus species combined estimated using Maxent from climate variables for the year 2000. Bright yellow and brown areas show where the logistic prediction of occurrence is p> 0.5 (the outlier records with exceptionally low probabilities of suitability were excluded in step-wise iterations); brown spots show all consistent Alpinobombus site records; ‘?’ shows the five records that are in climatically outlying sites. Polar projection, North Pole (starred) at the centre of the map, international boundaries and the Arctic Circle shown as narrow grey lines.	FIGURE 7. Sample sites and climatic suitability for all Alpinobombus species combined estimated using Maxent from climate variables for the year 2000. Bright yellow and brown areas show where the logistic prediction of occurrence is p> 0.5 (the outlier records with exceptionally low probabilities of suitability were excluded in step-wise iterations); brown spots show all consistent Alpinobombus site records; ‘?’ shows the five records that are in climatically outlying sites. Polar projection, North Pole (starred) at the centre of the map, international boundaries and the Arctic Circle shown as narrow grey lines.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF855E24FF685A8876EBFA8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268177/files/figure.png	https://doi.org/10.5281/zenodo.3268177	FIGURES 1‒6. Lateral views of foraging females (queens) of species of the subgenus Alpinobombus (with photo credits): 1, B. alpinus, Norway (A. Staverløkk); 2, B. pyrrhopygus, Sweden (G. Holmström); 3, B. polaris, Canada (B. Heinrich); 4, B. balteatus, Sweden (G. Holmström); 5, B. kirbiellus, USA (D. Wilson); 6, B. hyperboreus, Sweden (L.-I. Larsson). Some images reversed.	FIGURES 1‒6. Lateral views of foraging females (queens) of species of the subgenus Alpinobombus (with photo credits): 1, B. alpinus, Norway (A. Staverløkk); 2, B. pyrrhopygus, Sweden (G. Holmström); 3, B. polaris, Canada (B. Heinrich); 4, B. balteatus, Sweden (G. Holmström); 5, B. kirbiellus, USA (D. Wilson); 6, B. hyperboreus, Sweden (L.-I. Larsson). Some images reversed.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF855E24FF685A8876EBFA8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268203/files/figure.png	https://doi.org/10.5281/zenodo.3268203	FIGURES 77‒88. 77: Bombus kirbiellus global distribution (as for Fig. 15). 78‒88 colour patterns (as for Figs. 16‒21): 78 (#556) Canada-Nunavut; 79 (#294) Canada-Nunavut; 80 (#100) Canada-Nunavut; 81 (#3791) USA-Colorado; 82 (#3790) USA-Colorado; 83 (#3765) Canada-Nunavut; 84 (#3792) USA-Colorado; 85 (#3797) USA-Colorado; 86 (#4915) USA Alaska; 87 (#3768) Canada-Nunavut; 88 (#3812) USA-Colorado.	FIGURES 77‒88. 77: Bombus kirbiellus global distribution (as for Fig. 15). 78‒88 colour patterns (as for Figs. 16‒21): 78 (#556) Canada-Nunavut; 79 (#294) Canada-Nunavut; 80 (#100) Canada-Nunavut; 81 (#3791) USA-Colorado; 82 (#3790) USA-Colorado; 83 (#3765) Canada-Nunavut; 84 (#3792) USA-Colorado; 85 (#3797) USA-Colorado; 86 (#4915) USA Alaska; 87 (#3768) Canada-Nunavut; 88 (#3812) USA-Colorado.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF855E24FF685A8876EBFA8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268213/files/figure.png	https://doi.org/10.5281/zenodo.3268213	FIGURES 129‒137. Morphology of the male genitalia for species of the subgenus Alpinobombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 129 B. alpinus (#3824); 130 B pyrrhopygus (#3833); 131 B. polaris (#73); 132 B. balteatus (#3843); 133 B. kirbiellus (#3768); 134 B. neoboreus (#4387); 135 B. kluanensis (#4876); 136 B. natvigi (#81); 137 B. hyperboreus (#3821).	FIGURES 129‒137. Morphology of the male genitalia for species of the subgenus Alpinobombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 129 B. alpinus (#3824); 130 B pyrrhopygus (#3833); 131 B. polaris (#73); 132 B. balteatus (#3843); 133 B. kirbiellus (#3768); 134 B. neoboreus (#4387); 135 B. kluanensis (#4876); 136 B. natvigi (#81); 137 B. hyperboreus (#3821).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF855E24FF685A8876EBFA8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268219/files/figure.png	https://doi.org/10.5281/zenodo.3268219	FIGURES 140‒143. Morphology of females (queens). Left lateral aspect of the oculo-malar area of: 140, B. polaris; 141, B. kirbiellus (white lines indicate length and breadth measurements). Left lateral aspect of the outer surface of the left hind tibia (corbicular area): 142, B. polaris; 143, B. neoboreus (white lines indicate highlights in the central area).	FIGURES 140‒143. Morphology of females (queens). Left lateral aspect of the oculo-malar area of: 140, B. polaris; 141, B. kirbiellus (white lines indicate length and breadth measurements). Left lateral aspect of the outer surface of the left hind tibia (corbicular area): 142, B. polaris; 143, B. neoboreus (white lines indicate highlights in the central area).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF855E24FF685A8876EBFA8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268221/files/figure.png	https://doi.org/10.5281/zenodo.3268221	FIGURES 144‒147. Morphology of males. Dorsal aspect of the penis valves of: 144, B. natvigi; 145, B. neoboreus (white lines indicate location of the subapical outer tooth of the left penis valve). Dorsal aspect of left gonostylus: 146 B. kirbiellus; 147, B. polaris (white lines indicate anterior, midpoint, and posterior median projections of left gonostylus).	FIGURES 144‒147. Morphology of males. Dorsal aspect of the penis valves of: 144, B. natvigi; 145, B. neoboreus (white lines indicate location of the subapical outer tooth of the left penis valve). Dorsal aspect of left gonostylus: 146 B. kirbiellus; 147, B. polaris (white lines indicate anterior, midpoint, and posterior median projections of left gonostylus).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF855E24FF685A8876EBFA8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268187/files/figure.png	https://doi.org/10.5281/zenodo.3268187	FIGURE 9. Estimate of the metric gene tree for the 54 longest unique COI-barcode alleles of the subgenus Alpinobombus from MrBayes combined with the Bayesian Poisson-tree-process (PTP) solution with the highest support with spots showing the nodes with the strongest support as coalescents for candidate species (outgroup B. ignitus not shown). Values above the nodes are Bayesian posterior probabilities showing branch support for groups; values below the nodes are PTP Bayesian support values that all daughter alleles are parts of a single species (probabilities within candidate species are all ≤ 0.44). The scale bar is calibrated in substitutions per nucleotide site. Each unique allele is represented by one of the longest available sample sequences, labelled with: the sequence length in number of nucleotides, a taxon name or name describing an unpublished colour pattern, and then a code that consists of a specimen identifier (AL#n) from the project database and (after the hyphen) a sample identifier from BOLD or GenBank, followed with its geographic origin (for simplicity Greenland is included without Denmark).	FIGURE 9. Estimate of the metric gene tree for the 54 longest unique COI-barcode alleles of the subgenus Alpinobombus from MrBayes combined with the Bayesian Poisson-tree-process (PTP) solution with the highest support with spots showing the nodes with the strongest support as coalescents for candidate species (outgroup B. ignitus not shown). Values above the nodes are Bayesian posterior probabilities showing branch support for groups; values below the nodes are PTP Bayesian support values that all daughter alleles are parts of a single species (probabilities within candidate species are all ≤ 0.44). The scale bar is calibrated in substitutions per nucleotide site. Each unique allele is represented by one of the longest available sample sequences, labelled with: the sequence length in number of nucleotides, a taxon name or name describing an unpublished colour pattern, and then a code that consists of a specimen identifier (AL#n) from the project database and (after the hyphen) a sample identifier from BOLD or GenBank, followed with its geographic origin (for simplicity Greenland is included without Denmark).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF855E24FF685A8876EBFA8C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268201/files/figure.png	https://doi.org/10.5281/zenodo.3268201	FIGURES 59‒76. 59: Bombus balteatus global distribution (as for Fig. 15). 60‒76 colour patterns (as for Figs.16‒21): 60 (#540) Russia-Krasnoyask; 61 (#119) Sweden; 62 (#541) Russia-Krasnoyarsk; 63 (#542) Russia-Krasnoyarsk; 64 (#117) Mongolia; 65 (#534) Russia-Kamchatka; 66 (#3841) Norway; 67 (#738) Russia-Kamchatka; 68 (#971) Russia- Kamchatka; 69 (#535) Russia-Kamchatka; 70 (#973) Russia-Kamchatka; 71 (#880) Russia-Kamchatka; 72 (#536) Russia-Kamchatka; 73 (#228) Norway; 74 (#392) Sweden; 75 (#3843) Norway; 76 (#538) Russia-Kamchatka.	FIGURES 59‒76. 59: Bombus balteatus global distribution (as for Fig. 15). 60‒76 colour patterns (as for Figs.16‒21): 60 (#540) Russia-Krasnoyask; 61 (#119) Sweden; 62 (#541) Russia-Krasnoyarsk; 63 (#542) Russia-Krasnoyarsk; 64 (#117) Mongolia; 65 (#534) Russia-Kamchatka; 66 (#3841) Norway; 67 (#738) Russia-Kamchatka; 68 (#971) Russia- Kamchatka; 69 (#535) Russia-Kamchatka; 70 (#973) Russia-Kamchatka; 71 (#880) Russia-Kamchatka; 72 (#536) Russia-Kamchatka; 73 (#228) Norway; 74 (#392) Sweden; 75 (#3843) Norway; 76 (#538) Russia-Kamchatka.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF805E26FF685B3B742AFC24.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268205/files/figure.png	https://doi.org/10.5281/zenodo.3268205	FIGURES 89‒103. 89: Bombus neoboreus global distribution (as for Fig. 15). 90‒103 colour patterns (as for Figs. 16‒21): 90 (#68) USA-Alaska; 91 (#4706) Canada-Nunavut; 92 (#3816) Canada-Yukon; 93 (#3819) USA-Alaska; 94 (#2055) Canada-Nunavut; 95 (#2058) Canada-Nunavut; 96 (#2071) Canada-Nunavut; 97 (#2051) Canada-Nunavut; 98 (#4406) Canada-Nunavut; 99 (#90) Canada-Nunavut; 100 (#92) Canada-Nunavut; 101 (#93) Canada-Yukon; 102 (#3762) Canada-Nunavut; 103 (#91) Canada-Nunavut.	FIGURES 89‒103. 89: Bombus neoboreus global distribution (as for Fig. 15). 90‒103 colour patterns (as for Figs. 16‒21): 90 (#68) USA-Alaska; 91 (#4706) Canada-Nunavut; 92 (#3816) Canada-Yukon; 93 (#3819) USA-Alaska; 94 (#2055) Canada-Nunavut; 95 (#2058) Canada-Nunavut; 96 (#2071) Canada-Nunavut; 97 (#2051) Canada-Nunavut; 98 (#4406) Canada-Nunavut; 99 (#90) Canada-Nunavut; 100 (#92) Canada-Nunavut; 101 (#93) Canada-Yukon; 102 (#3762) Canada-Nunavut; 103 (#91) Canada-Nunavut.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF805E26FF685B3B742AFC24.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268213/files/figure.png	https://doi.org/10.5281/zenodo.3268213	FIGURES 129‒137. Morphology of the male genitalia for species of the subgenus Alpinobombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 129 B. alpinus (#3824); 130 B pyrrhopygus (#3833); 131 B. polaris (#73); 132 B. balteatus (#3843); 133 B. kirbiellus (#3768); 134 B. neoboreus (#4387); 135 B. kluanensis (#4876); 136 B. natvigi (#81); 137 B. hyperboreus (#3821).	FIGURES 129‒137. Morphology of the male genitalia for species of the subgenus Alpinobombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 129 B. alpinus (#3824); 130 B pyrrhopygus (#3833); 131 B. polaris (#73); 132 B. balteatus (#3843); 133 B. kirbiellus (#3768); 134 B. neoboreus (#4387); 135 B. kluanensis (#4876); 136 B. natvigi (#81); 137 B. hyperboreus (#3821).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF805E26FF685B3B742AFC24.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268219/files/figure.png	https://doi.org/10.5281/zenodo.3268219	FIGURES 140‒143. Morphology of females (queens). Left lateral aspect of the oculo-malar area of: 140, B. polaris; 141, B. kirbiellus (white lines indicate length and breadth measurements). Left lateral aspect of the outer surface of the left hind tibia (corbicular area): 142, B. polaris; 143, B. neoboreus (white lines indicate highlights in the central area).	FIGURES 140‒143. Morphology of females (queens). Left lateral aspect of the oculo-malar area of: 140, B. polaris; 141, B. kirbiellus (white lines indicate length and breadth measurements). Left lateral aspect of the outer surface of the left hind tibia (corbicular area): 142, B. polaris; 143, B. neoboreus (white lines indicate highlights in the central area).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF805E26FF685B3B742AFC24.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268221/files/figure.png	https://doi.org/10.5281/zenodo.3268221	FIGURES 144‒147. Morphology of males. Dorsal aspect of the penis valves of: 144, B. natvigi; 145, B. neoboreus (white lines indicate location of the subapical outer tooth of the left penis valve). Dorsal aspect of left gonostylus: 146 B. kirbiellus; 147, B. polaris (white lines indicate anterior, midpoint, and posterior median projections of left gonostylus).	FIGURES 144‒147. Morphology of males. Dorsal aspect of the penis valves of: 144, B. natvigi; 145, B. neoboreus (white lines indicate location of the subapical outer tooth of the left penis valve). Dorsal aspect of left gonostylus: 146 B. kirbiellus; 147, B. polaris (white lines indicate anterior, midpoint, and posterior median projections of left gonostylus).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF805E26FF685B3B742AFC24.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268223/files/figure.png	https://doi.org/10.5281/zenodo.3268223	FIGURES 148‒149. Anterior / left lateral aspect of the lower part of the clypeus, the labrum, and the mandibles of males of: (148) B. neoboreus (#4867), showing the small orange hair patches at the base of the mandibles and the long dense posterior fringing black beard; (149) B. kluanensis (#4876), showing the large dense orange hair pads on the mandibles and the sparse orange hairs in the posterior fringing beard.	FIGURES 148‒149. Anterior / left lateral aspect of the lower part of the clypeus, the labrum, and the mandibles of males of: (148) B. neoboreus (#4867), showing the small orange hair patches at the base of the mandibles and the long dense posterior fringing black beard; (149) B. kluanensis (#4876), showing the large dense orange hair pads on the mandibles and the sparse orange hairs in the posterior fringing beard.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF805E26FF685B3B742AFC24.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268187/files/figure.png	https://doi.org/10.5281/zenodo.3268187	FIGURE 9. Estimate of the metric gene tree for the 54 longest unique COI-barcode alleles of the subgenus Alpinobombus from MrBayes combined with the Bayesian Poisson-tree-process (PTP) solution with the highest support with spots showing the nodes with the strongest support as coalescents for candidate species (outgroup B. ignitus not shown). Values above the nodes are Bayesian posterior probabilities showing branch support for groups; values below the nodes are PTP Bayesian support values that all daughter alleles are parts of a single species (probabilities within candidate species are all ≤ 0.44). The scale bar is calibrated in substitutions per nucleotide site. Each unique allele is represented by one of the longest available sample sequences, labelled with: the sequence length in number of nucleotides, a taxon name or name describing an unpublished colour pattern, and then a code that consists of a specimen identifier (AL#n) from the project database and (after the hyphen) a sample identifier from BOLD or GenBank, followed with its geographic origin (for simplicity Greenland is included without Denmark).	FIGURE 9. Estimate of the metric gene tree for the 54 longest unique COI-barcode alleles of the subgenus Alpinobombus from MrBayes combined with the Bayesian Poisson-tree-process (PTP) solution with the highest support with spots showing the nodes with the strongest support as coalescents for candidate species (outgroup B. ignitus not shown). Values above the nodes are Bayesian posterior probabilities showing branch support for groups; values below the nodes are PTP Bayesian support values that all daughter alleles are parts of a single species (probabilities within candidate species are all ≤ 0.44). The scale bar is calibrated in substitutions per nucleotide site. Each unique allele is represented by one of the longest available sample sequences, labelled with: the sequence length in number of nucleotides, a taxon name or name describing an unpublished colour pattern, and then a code that consists of a specimen identifier (AL#n) from the project database and (after the hyphen) a sample identifier from BOLD or GenBank, followed with its geographic origin (for simplicity Greenland is included without Denmark).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF805E26FF685B3B742AFC24.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268181/files/figure.png	https://doi.org/10.5281/zenodo.3268181	FIGURE 7. Sample sites and climatic suitability for all Alpinobombus species combined estimated using Maxent from climate variables for the year 2000. Bright yellow and brown areas show where the logistic prediction of occurrence is p> 0.5 (the outlier records with exceptionally low probabilities of suitability were excluded in step-wise iterations); brown spots show all consistent Alpinobombus site records; ‘?’ shows the five records that are in climatically outlying sites. Polar projection, North Pole (starred) at the centre of the map, international boundaries and the Arctic Circle shown as narrow grey lines.	FIGURE 7. Sample sites and climatic suitability for all Alpinobombus species combined estimated using Maxent from climate variables for the year 2000. Bright yellow and brown areas show where the logistic prediction of occurrence is p> 0.5 (the outlier records with exceptionally low probabilities of suitability were excluded in step-wise iterations); brown spots show all consistent Alpinobombus site records; ‘?’ shows the five records that are in climatically outlying sites. Polar projection, North Pole (starred) at the centre of the map, international boundaries and the Arctic Circle shown as narrow grey lines.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF825E29FF685D947775FEC1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268189/files/figure.png	https://doi.org/10.5281/zenodo.3268189	FIGURE 10. Map of the entire global distribution of records of B. kluanensis s. l.. Grey crosses are records for any Alpinobombus species; black crosses are records of B. neoboreus; overlayed with small white spots for specimens of B. kluanensis s. l.; large green spots for sequenced samples with the kluanensis s. str. alleles (collected 2010‒2016); small red spots for sequenced samples with the ‘unnamed2’ allele (collected 2015‒2017). Relief map with hill shading, with a Polar projection, and with the Arctic Circle and the USA (Alaska) / Canada (Yukon) international boundary shown as grey lines. North pointer and scale bar below, with an inset that shows the location of the detailed area of the global map. Image created in ArcGIS using World_Shad- ed_Relief basemap which is Copyright: © 2014 Esri.	FIGURE 10. Map of the entire global distribution of records of B. kluanensis s. l.. Grey crosses are records for any Alpinobombus species; black crosses are records of B. neoboreus; overlayed with small white spots for specimens of B. kluanensis s. l.; large green spots for sequenced samples with the kluanensis s. str. alleles (collected 2010‒2016); small red spots for sequenced samples with the ‘unnamed2’ allele (collected 2015‒2017). Relief map with hill shading, with a Polar projection, and with the Arctic Circle and the USA (Alaska) / Canada (Yukon) international boundary shown as grey lines. North pointer and scale bar below, with an inset that shows the location of the detailed area of the global map. Image created in ArcGIS using World_Shad- ed_Relief basemap which is Copyright: © 2014 Esri.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF825E29FF685D947775FEC1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268207/files/figure.png	https://doi.org/10.5281/zenodo.3268207	FIGURES 104‒115. 104: Bombus kluanensis global distribution (as for Fig. 15). 105‒115 colour patterns (as for Figs. 16‒21): 105 (#4473) Canada-Yukon; 106 (#4689) Canada-Yukon; 107 (#16) Canada-Yukon; 108 (#4393) USA-Alaska; 109 (#4466) Canada-Yukon; 110 (#4700) Canada-Yukon; 111 (#3130) Canada-Yukon; 112 (#4876) Canada-Yukon; 113 (#4882) Canada- Yukon; 114 (#4852) Canada-Yukon; 115 (#4881) Canada-Yukon.	FIGURES 104‒115. 104: Bombus kluanensis global distribution (as for Fig. 15). 105‒115 colour patterns (as for Figs. 16‒21): 105 (#4473) Canada-Yukon; 106 (#4689) Canada-Yukon; 107 (#16) Canada-Yukon; 108 (#4393) USA-Alaska; 109 (#4466) Canada-Yukon; 110 (#4700) Canada-Yukon; 111 (#3130) Canada-Yukon; 112 (#4876) Canada-Yukon; 113 (#4882) Canada- Yukon; 114 (#4852) Canada-Yukon; 115 (#4881) Canada-Yukon.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF825E29FF685D947775FEC1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268213/files/figure.png	https://doi.org/10.5281/zenodo.3268213	FIGURES 129‒137. Morphology of the male genitalia for species of the subgenus Alpinobombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 129 B. alpinus (#3824); 130 B pyrrhopygus (#3833); 131 B. polaris (#73); 132 B. balteatus (#3843); 133 B. kirbiellus (#3768); 134 B. neoboreus (#4387); 135 B. kluanensis (#4876); 136 B. natvigi (#81); 137 B. hyperboreus (#3821).	FIGURES 129‒137. Morphology of the male genitalia for species of the subgenus Alpinobombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 129 B. alpinus (#3824); 130 B pyrrhopygus (#3833); 131 B. polaris (#73); 132 B. balteatus (#3843); 133 B. kirbiellus (#3768); 134 B. neoboreus (#4387); 135 B. kluanensis (#4876); 136 B. natvigi (#81); 137 B. hyperboreus (#3821).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF825E29FF685D947775FEC1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268223/files/figure.png	https://doi.org/10.5281/zenodo.3268223	FIGURES 148‒149. Anterior / left lateral aspect of the lower part of the clypeus, the labrum, and the mandibles of males of: (148) B. neoboreus (#4867), showing the small orange hair patches at the base of the mandibles and the long dense posterior fringing black beard; (149) B. kluanensis (#4876), showing the large dense orange hair pads on the mandibles and the sparse orange hairs in the posterior fringing beard.	FIGURES 148‒149. Anterior / left lateral aspect of the lower part of the clypeus, the labrum, and the mandibles of males of: (148) B. neoboreus (#4867), showing the small orange hair patches at the base of the mandibles and the long dense posterior fringing black beard; (149) B. kluanensis (#4876), showing the large dense orange hair pads on the mandibles and the sparse orange hairs in the posterior fringing beard.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF825E29FF685D947775FEC1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268205/files/figure.png	https://doi.org/10.5281/zenodo.3268205	FIGURES 89‒103. 89: Bombus neoboreus global distribution (as for Fig. 15). 90‒103 colour patterns (as for Figs. 16‒21): 90 (#68) USA-Alaska; 91 (#4706) Canada-Nunavut; 92 (#3816) Canada-Yukon; 93 (#3819) USA-Alaska; 94 (#2055) Canada-Nunavut; 95 (#2058) Canada-Nunavut; 96 (#2071) Canada-Nunavut; 97 (#2051) Canada-Nunavut; 98 (#4406) Canada-Nunavut; 99 (#90) Canada-Nunavut; 100 (#92) Canada-Nunavut; 101 (#93) Canada-Yukon; 102 (#3762) Canada-Nunavut; 103 (#91) Canada-Nunavut.	FIGURES 89‒103. 89: Bombus neoboreus global distribution (as for Fig. 15). 90‒103 colour patterns (as for Figs. 16‒21): 90 (#68) USA-Alaska; 91 (#4706) Canada-Nunavut; 92 (#3816) Canada-Yukon; 93 (#3819) USA-Alaska; 94 (#2055) Canada-Nunavut; 95 (#2058) Canada-Nunavut; 96 (#2071) Canada-Nunavut; 97 (#2051) Canada-Nunavut; 98 (#4406) Canada-Nunavut; 99 (#90) Canada-Nunavut; 100 (#92) Canada-Nunavut; 101 (#93) Canada-Yukon; 102 (#3762) Canada-Nunavut; 103 (#91) Canada-Nunavut.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF825E29FF685D947775FEC1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268187/files/figure.png	https://doi.org/10.5281/zenodo.3268187	FIGURE 9. Estimate of the metric gene tree for the 54 longest unique COI-barcode alleles of the subgenus Alpinobombus from MrBayes combined with the Bayesian Poisson-tree-process (PTP) solution with the highest support with spots showing the nodes with the strongest support as coalescents for candidate species (outgroup B. ignitus not shown). Values above the nodes are Bayesian posterior probabilities showing branch support for groups; values below the nodes are PTP Bayesian support values that all daughter alleles are parts of a single species (probabilities within candidate species are all ≤ 0.44). The scale bar is calibrated in substitutions per nucleotide site. Each unique allele is represented by one of the longest available sample sequences, labelled with: the sequence length in number of nucleotides, a taxon name or name describing an unpublished colour pattern, and then a code that consists of a specimen identifier (AL#n) from the project database and (after the hyphen) a sample identifier from BOLD or GenBank, followed with its geographic origin (for simplicity Greenland is included without Denmark).	FIGURE 9. Estimate of the metric gene tree for the 54 longest unique COI-barcode alleles of the subgenus Alpinobombus from MrBayes combined with the Bayesian Poisson-tree-process (PTP) solution with the highest support with spots showing the nodes with the strongest support as coalescents for candidate species (outgroup B. ignitus not shown). Values above the nodes are Bayesian posterior probabilities showing branch support for groups; values below the nodes are PTP Bayesian support values that all daughter alleles are parts of a single species (probabilities within candidate species are all ≤ 0.44). The scale bar is calibrated in substitutions per nucleotide site. Each unique allele is represented by one of the longest available sample sequences, labelled with: the sequence length in number of nucleotides, a taxon name or name describing an unpublished colour pattern, and then a code that consists of a specimen identifier (AL#n) from the project database and (after the hyphen) a sample identifier from BOLD or GenBank, followed with its geographic origin (for simplicity Greenland is included without Denmark).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF825E29FF685D947775FEC1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268191/files/figure.png	https://doi.org/10.5281/zenodo.3268191	FIGURES 11‒13. 11: gene trees for COI; 12: gene trees for 16S; and 13: species trees from *BEAST analysis of data from Thanoosing (2017) displayed using Densitree. Each plot shows the background density of trees from samples of 10,000 Bayesian trees, with the summary ‘root canal’ tree superimposed as a thick line, and the branches exclusively linking samples of B. kluanensis s. l. (#4850, 4869, 4870, 4873) traced in red.	FIGURES 11‒13. 11: gene trees for COI; 12: gene trees for 16S; and 13: species trees from *BEAST analysis of data from Thanoosing (2017) displayed using Densitree. Each plot shows the background density of trees from samples of 10,000 Bayesian trees, with the summary ‘root canal’ tree superimposed as a thick line, and the branches exclusively linking samples of B. kluanensis s. l. (#4850, 4869, 4870, 4873) traced in red.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF8D5E29FF685FE77685F879.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268209/files/figure.png	https://doi.org/10.5281/zenodo.3268209	FIGURES 116‒123. 116: Bombus natvigi global distribution (as for Fig. 15). 117‒123 colour patterns (as for Figs. 16‒21): 117 (#3519) Canada-Nunavut; 118 (#184) Canada-Nunavut; 119 (#3551) Canada-Nunavut; 120 (#3546) Canada-Nunavut; 121 (#79) USA-Alaska; 122 (#3557) Canada-Northwest Territories; 123 (#82) Canada-Northwest Territories.	FIGURES 116‒123. 116: Bombus natvigi global distribution (as for Fig. 15). 117‒123 colour patterns (as for Figs. 16‒21): 117 (#3519) Canada-Nunavut; 118 (#184) Canada-Nunavut; 119 (#3551) Canada-Nunavut; 120 (#3546) Canada-Nunavut; 121 (#79) USA-Alaska; 122 (#3557) Canada-Northwest Territories; 123 (#82) Canada-Northwest Territories.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF8D5E29FF685FE77685F879.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268213/files/figure.png	https://doi.org/10.5281/zenodo.3268213	FIGURES 129‒137. Morphology of the male genitalia for species of the subgenus Alpinobombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 129 B. alpinus (#3824); 130 B pyrrhopygus (#3833); 131 B. polaris (#73); 132 B. balteatus (#3843); 133 B. kirbiellus (#3768); 134 B. neoboreus (#4387); 135 B. kluanensis (#4876); 136 B. natvigi (#81); 137 B. hyperboreus (#3821).	FIGURES 129‒137. Morphology of the male genitalia for species of the subgenus Alpinobombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 129 B. alpinus (#3824); 130 B pyrrhopygus (#3833); 131 B. polaris (#73); 132 B. balteatus (#3843); 133 B. kirbiellus (#3768); 134 B. neoboreus (#4387); 135 B. kluanensis (#4876); 136 B. natvigi (#81); 137 B. hyperboreus (#3821).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF8D5E29FF685FE77685F879.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268221/files/figure.png	https://doi.org/10.5281/zenodo.3268221	FIGURES 144‒147. Morphology of males. Dorsal aspect of the penis valves of: 144, B. natvigi; 145, B. neoboreus (white lines indicate location of the subapical outer tooth of the left penis valve). Dorsal aspect of left gonostylus: 146 B. kirbiellus; 147, B. polaris (white lines indicate anterior, midpoint, and posterior median projections of left gonostylus).	FIGURES 144‒147. Morphology of males. Dorsal aspect of the penis valves of: 144, B. natvigi; 145, B. neoboreus (white lines indicate location of the subapical outer tooth of the left penis valve). Dorsal aspect of left gonostylus: 146 B. kirbiellus; 147, B. polaris (white lines indicate anterior, midpoint, and posterior median projections of left gonostylus).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF8D5E29FF685FE77685F879.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268187/files/figure.png	https://doi.org/10.5281/zenodo.3268187	FIGURE 9. Estimate of the metric gene tree for the 54 longest unique COI-barcode alleles of the subgenus Alpinobombus from MrBayes combined with the Bayesian Poisson-tree-process (PTP) solution with the highest support with spots showing the nodes with the strongest support as coalescents for candidate species (outgroup B. ignitus not shown). Values above the nodes are Bayesian posterior probabilities showing branch support for groups; values below the nodes are PTP Bayesian support values that all daughter alleles are parts of a single species (probabilities within candidate species are all ≤ 0.44). The scale bar is calibrated in substitutions per nucleotide site. Each unique allele is represented by one of the longest available sample sequences, labelled with: the sequence length in number of nucleotides, a taxon name or name describing an unpublished colour pattern, and then a code that consists of a specimen identifier (AL#n) from the project database and (after the hyphen) a sample identifier from BOLD or GenBank, followed with its geographic origin (for simplicity Greenland is included without Denmark).	FIGURE 9. Estimate of the metric gene tree for the 54 longest unique COI-barcode alleles of the subgenus Alpinobombus from MrBayes combined with the Bayesian Poisson-tree-process (PTP) solution with the highest support with spots showing the nodes with the strongest support as coalescents for candidate species (outgroup B. ignitus not shown). Values above the nodes are Bayesian posterior probabilities showing branch support for groups; values below the nodes are PTP Bayesian support values that all daughter alleles are parts of a single species (probabilities within candidate species are all ≤ 0.44). The scale bar is calibrated in substitutions per nucleotide site. Each unique allele is represented by one of the longest available sample sequences, labelled with: the sequence length in number of nucleotides, a taxon name or name describing an unpublished colour pattern, and then a code that consists of a specimen identifier (AL#n) from the project database and (after the hyphen) a sample identifier from BOLD or GenBank, followed with its geographic origin (for simplicity Greenland is included without Denmark).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF8D5E29FF685FE77685F879.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268211/files/figure.png	https://doi.org/10.5281/zenodo.3268211	FIGURES 124‒128. 124: Bombus hyperboreus global distribution (as for Fig. 15). 125‒128 colour patterns (as for Figs. 16‒ 21): 125 (#39) Russia-Chukotka (Wrangel Island); 126 (#33) Russia-Magadan; 127 (#1258) Russia-Murmansk; 128 (#533) Russia-Sakha.	FIGURES 124‒128. 124: Bombus hyperboreus global distribution (as for Fig. 15). 125‒128 colour patterns (as for Figs. 16‒ 21): 125 (#39) Russia-Chukotka (Wrangel Island); 126 (#33) Russia-Magadan; 127 (#1258) Russia-Murmansk; 128 (#533) Russia-Sakha.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF8D5E29FF685FE77685F879.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268207/files/figure.png	https://doi.org/10.5281/zenodo.3268207	FIGURES 104‒115. 104: Bombus kluanensis global distribution (as for Fig. 15). 105‒115 colour patterns (as for Figs. 16‒21): 105 (#4473) Canada-Yukon; 106 (#4689) Canada-Yukon; 107 (#16) Canada-Yukon; 108 (#4393) USA-Alaska; 109 (#4466) Canada-Yukon; 110 (#4700) Canada-Yukon; 111 (#3130) Canada-Yukon; 112 (#4876) Canada-Yukon; 113 (#4882) Canada- Yukon; 114 (#4852) Canada-Yukon; 115 (#4881) Canada-Yukon.	FIGURES 104‒115. 104: Bombus kluanensis global distribution (as for Fig. 15). 105‒115 colour patterns (as for Figs. 16‒21): 105 (#4473) Canada-Yukon; 106 (#4689) Canada-Yukon; 107 (#16) Canada-Yukon; 108 (#4393) USA-Alaska; 109 (#4466) Canada-Yukon; 110 (#4700) Canada-Yukon; 111 (#3130) Canada-Yukon; 112 (#4876) Canada-Yukon; 113 (#4882) Canada- Yukon; 114 (#4852) Canada-Yukon; 115 (#4881) Canada-Yukon.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF8F5E2EFF685EF471A3FDE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268177/files/figure.png	https://doi.org/10.5281/zenodo.3268177	FIGURES 1‒6. Lateral views of foraging females (queens) of species of the subgenus Alpinobombus (with photo credits): 1, B. alpinus, Norway (A. Staverløkk); 2, B. pyrrhopygus, Sweden (G. Holmström); 3, B. polaris, Canada (B. Heinrich); 4, B. balteatus, Sweden (G. Holmström); 5, B. kirbiellus, USA (D. Wilson); 6, B. hyperboreus, Sweden (L.-I. Larsson). Some images reversed.	FIGURES 1‒6. Lateral views of foraging females (queens) of species of the subgenus Alpinobombus (with photo credits): 1, B. alpinus, Norway (A. Staverløkk); 2, B. pyrrhopygus, Sweden (G. Holmström); 3, B. polaris, Canada (B. Heinrich); 4, B. balteatus, Sweden (G. Holmström); 5, B. kirbiellus, USA (D. Wilson); 6, B. hyperboreus, Sweden (L.-I. Larsson). Some images reversed.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF8F5E2EFF685EF471A3FDE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268211/files/figure.png	https://doi.org/10.5281/zenodo.3268211	FIGURES 124‒128. 124: Bombus hyperboreus global distribution (as for Fig. 15). 125‒128 colour patterns (as for Figs. 16‒ 21): 125 (#39) Russia-Chukotka (Wrangel Island); 126 (#33) Russia-Magadan; 127 (#1258) Russia-Murmansk; 128 (#533) Russia-Sakha.	FIGURES 124‒128. 124: Bombus hyperboreus global distribution (as for Fig. 15). 125‒128 colour patterns (as for Figs. 16‒ 21): 125 (#39) Russia-Chukotka (Wrangel Island); 126 (#33) Russia-Magadan; 127 (#1258) Russia-Murmansk; 128 (#533) Russia-Sakha.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF8F5E2EFF685EF471A3FDE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268213/files/figure.png	https://doi.org/10.5281/zenodo.3268213	FIGURES 129‒137. Morphology of the male genitalia for species of the subgenus Alpinobombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 129 B. alpinus (#3824); 130 B pyrrhopygus (#3833); 131 B. polaris (#73); 132 B. balteatus (#3843); 133 B. kirbiellus (#3768); 134 B. neoboreus (#4387); 135 B. kluanensis (#4876); 136 B. natvigi (#81); 137 B. hyperboreus (#3821).	FIGURES 129‒137. Morphology of the male genitalia for species of the subgenus Alpinobombus from the dorsal aspect, anterior to the left of the image, posterior to the right: 129 B. alpinus (#3824); 130 B pyrrhopygus (#3833); 131 B. polaris (#73); 132 B. balteatus (#3843); 133 B. kirbiellus (#3768); 134 B. neoboreus (#4387); 135 B. kluanensis (#4876); 136 B. natvigi (#81); 137 B. hyperboreus (#3821).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF8F5E2EFF685EF471A3FDE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268187/files/figure.png	https://doi.org/10.5281/zenodo.3268187	FIGURE 9. Estimate of the metric gene tree for the 54 longest unique COI-barcode alleles of the subgenus Alpinobombus from MrBayes combined with the Bayesian Poisson-tree-process (PTP) solution with the highest support with spots showing the nodes with the strongest support as coalescents for candidate species (outgroup B. ignitus not shown). Values above the nodes are Bayesian posterior probabilities showing branch support for groups; values below the nodes are PTP Bayesian support values that all daughter alleles are parts of a single species (probabilities within candidate species are all ≤ 0.44). The scale bar is calibrated in substitutions per nucleotide site. Each unique allele is represented by one of the longest available sample sequences, labelled with: the sequence length in number of nucleotides, a taxon name or name describing an unpublished colour pattern, and then a code that consists of a specimen identifier (AL#n) from the project database and (after the hyphen) a sample identifier from BOLD or GenBank, followed with its geographic origin (for simplicity Greenland is included without Denmark).	FIGURE 9. Estimate of the metric gene tree for the 54 longest unique COI-barcode alleles of the subgenus Alpinobombus from MrBayes combined with the Bayesian Poisson-tree-process (PTP) solution with the highest support with spots showing the nodes with the strongest support as coalescents for candidate species (outgroup B. ignitus not shown). Values above the nodes are Bayesian posterior probabilities showing branch support for groups; values below the nodes are PTP Bayesian support values that all daughter alleles are parts of a single species (probabilities within candidate species are all ≤ 0.44). The scale bar is calibrated in substitutions per nucleotide site. Each unique allele is represented by one of the longest available sample sequences, labelled with: the sequence length in number of nucleotides, a taxon name or name describing an unpublished colour pattern, and then a code that consists of a specimen identifier (AL#n) from the project database and (after the hyphen) a sample identifier from BOLD or GenBank, followed with its geographic origin (for simplicity Greenland is included without Denmark).	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
AC4D691FFF8F5E2EFF685EF471A3FDE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/3268209/files/figure.png	https://doi.org/10.5281/zenodo.3268209	FIGURES 116‒123. 116: Bombus natvigi global distribution (as for Fig. 15). 117‒123 colour patterns (as for Figs. 16‒21): 117 (#3519) Canada-Nunavut; 118 (#184) Canada-Nunavut; 119 (#3551) Canada-Nunavut; 120 (#3546) Canada-Nunavut; 121 (#79) USA-Alaska; 122 (#3557) Canada-Northwest Territories; 123 (#82) Canada-Northwest Territories.	FIGURES 116‒123. 116: Bombus natvigi global distribution (as for Fig. 15). 117‒123 colour patterns (as for Figs. 16‒21): 117 (#3519) Canada-Nunavut; 118 (#184) Canada-Nunavut; 119 (#3551) Canada-Nunavut; 120 (#3546) Canada-Nunavut; 121 (#79) USA-Alaska; 122 (#3557) Canada-Northwest Territories; 123 (#82) Canada-Northwest Territories.	2019-07-03	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.		Zenodo	biologists	Williams, Paul H.;Berezin, Mikhail V.;Cannings, Sydney G.;Cederberg, Björn;Ødegaard, Frode;Rasmussen, Claus;Richardson, Leif L.;Rykken, Jessica;Sheffield, Cory S.;Thanoosing, Chawatat;Byvaltsev, Alexandr M.			
