taxonID	type	description	language	source
AC58878FFFF7FFDAFF452892DA98FF7C.taxon	description	Oedaspis is characterized by a usually very short proboscis, posterior orbital setae usually stubby white and 2 – 5 frontal setae, frequently variable even within the same species and often differing either side. Variation in the number of setae also occurs on the thorax and presutural dorsocentral setae are occasionally present. The scutellum is shiny or subshiny and often swollen, wings with 2 distinct setae on Sc break, cell c broadly hyaline medially without a dark spot or band, and a distinct band or band like pattern resembling λ or H pattern in middle at least in females. Abdomen subshining brownish or shiny black, with microtrichia. Aculeus often with serrations and glans of phallus with elongated flap-like vesica, rounded at apex.	en	Singh, Maneesh Pal, Sneha, Sharma, Sharma, Isha, Gupta, Divender, Hancock, David Lawrence, Prabhakar, Chandra Shekhar (2024): A new species of genus Oedaspis Loew and new distribution records of Acidiostigma Hendel & Goniurellia Hendel from India (Diptera: Tephritidae). Zootaxa 5514 (5): 469-480, DOI: 10.11646/zootaxa.5514.5.4, URL: http://dx.doi.org/10.11646/zootaxa.5514.5.4
AC58878FFFF4FFD0FF452B6ADB76FED2.taxon	description	(Figure 1 – 5)	en	Singh, Maneesh Pal, Sneha, Sharma, Sharma, Isha, Gupta, Divender, Hancock, David Lawrence, Prabhakar, Chandra Shekhar (2024): A new species of genus Oedaspis Loew and new distribution records of Acidiostigma Hendel & Goniurellia Hendel from India (Diptera: Tephritidae). Zootaxa 5514 (5): 469-480, DOI: 10.11646/zootaxa.5514.5.4, URL: http://dx.doi.org/10.11646/zootaxa.5514.5.4
AC58878FFFF4FFD0FF452B6ADB76FED2.taxon	diagnosis	Diagnosis: A medium-sized (5.1 mm) species with 3 – 5 frontal setae and brownish head, scutum black distributed with pale microtrichia, pale yellowish orange at lateral margins and above scuto-scutellar suture, scutellum convex, light fulvous, shiny with 4 distinct black spots around bases of each scutellar seta, pleura, notopleuron and postpronotal lobes yellowish orange. Presutural dorsocentral setae absent. Wing with 5 pale brownish, darker edged bands. Cubital and discal bands joined in cell r 2 + 3 leaving hyaline indentation in cell r 1 above them and forming a λ-like pattern. Pterostigma with 2 – 3 irregular-shaped black spots. Abdomen brownish fulvous in males with variable black patches on tergites I, III, IV & V, tergite II with median, small, roundish black spot. Female abdomen shiny black, oviscape black with reddish fulvous band in middle without any variations except oviscape entirely black in some specimen.	en	Singh, Maneesh Pal, Sneha, Sharma, Sharma, Isha, Gupta, Divender, Hancock, David Lawrence, Prabhakar, Chandra Shekhar (2024): A new species of genus Oedaspis Loew and new distribution records of Acidiostigma Hendel & Goniurellia Hendel from India (Diptera: Tephritidae). Zootaxa 5514 (5): 469-480, DOI: 10.11646/zootaxa.5514.5.4, URL: http://dx.doi.org/10.11646/zootaxa.5514.5.4
AC58878FFFF4FFD0FF452B6ADB76FED2.taxon	description	Description Head: Fulvous to brownish and even darker over face and parafacial area, head broader than high, postgena slightly bulged, antennae with scape brown with 4 – 5 dark black setulae, pedicel pale yellowish with numerous dark setulae on dorsal side and few setulae slightly longer than others, flagellomere pale yellowish, slightly curved dorsally and bean-shaped but not forming sharp point, arista lightly plumose (Figure 1 A & B). Compound eyes 1.45 times as high as long. Gena, postgena and orbital region on frons with numerous pale yellowish setulae. Lunule horseshoe-shaped and parafacial lightly bulgy. Ocellar triangle black and area around it light coloured as compared to rest of head. Five frontal setae (variable in numbers), 2 orbital setae (anterior black and posterior pale but this character variable, some specimens with both setae blackish), 1 ocellar setae (some specimen with 2 pair), median vertical setae pale and lateral vertical setae stubby white, 6 postocellar setae stubby white (median setae longer), postocular setulae stubby white with black in alternate fashion and 1 genal setae pale; palpi with 6 – 8 black setulae and labella with slightly longer whitish setulae. Proboscis reduced, slightly shorter than flagellomere. Thorax: Setose with white setulae, mostly dull grayish black except area just below prescutellar acrosichals slightly reddish brown in some males. Postpronotal lobe, notopleuron, anepisternal stripe, katepisternum, meron dark brownish or greyish. Anepimeron, katatergite with light dark patch and anatergite darker. Subcutellum and mediotergite black, postalar calli slightly bulged and reddish brown. Scutellum lightly convex, yellowish with 4 distinct black spots around scutellar setae, almost equal in size. Full complement of thoracal setae, all almost equal in size except posterior notopleural setae smaller and lighter in colour than the others. Postsutural dorscocentral setae variable in this species, some specimens having multiple setae. Anepisternal setae 2 (anterior black and posterior whitish) with 2 – 3 smaller pale setae. Halter creamy white. All legs yellowish except femora slightly darker than the rest of leg, setose with numerous black setulae. Fore femora with 3 rows of setae, two on dorsolateral side and one on ventrolateral side. Setae on ventrolateral side are slightly longer and black while the rest are stubby. Mid tibia with 1 apical spine but two slightly smaller spines also present on either side of longer spine. Hind femora with row of minute black setulae over entire length. Wing 5 mm long with light brownish distinct bands, cell bc fulvous except hyaline spot in middle. Subbasal, discal, cubital, preapical and apical bands present. Entire costal cell with brownish infuscation except broadly hyaline indentation in middle, cell sc brownish except thin hyaline line on Sc break and with 3 distinct black spots, the latter variable in most specimens, connected in some and distinct in others (Figure 5 A – F). Discal and cubital bands connected over r-m crossvein, forming a λ-shaped joint, covering area of cells br and r 4 + 5 either side of r-m crossvein. Subbasal band connected to discal band apically in cells r 1 and r 2 + 3. Cubital band connected to apical band in cell r 1 and with 2 hyaline spots just below vein R 1 and above vein R 2 + 3, these hyaline spots variable in shape. Bands are light brownish with orange hue but margins and bases of all bands black. Hyaline indentation in between preapical and apical bands is variable (Figure 4 G – I); in some specimens preapical and apical bands connected forming a hyaline spot in cell r 2 + 3, but separate in other specimens. Hyaline indentations at apex veins R 2 + 3 and R 4 + 5 also variable; these are absent, narrow and broader in some of paratypes. Hyaline indentations in front of vein R 1 also variable, triangular in some specimens or trapezoidal in others. Abdomen: Shiny black, shiny and setose, black setae on lateral margins of tergite II, III, IV, V and VI, 5 – 6 setae on posterior rim of tergite VI. Abdomen orange yellow in males with narrow transverse black bands in middle of tergite I, III, IV and V often interrupted in middle and a median black spot often present below transverse black band on tergite I. Tergite II yellowish orange in most specimens examined. Black bands on abdominal tergites of males are variable in shape & colour (Figure 4 D – F). Sternite V of male with V-shaped deep posterior concavity (Figure 2 F). Genitalia: Oviscape black with reddish fulvous band in middle, 1.2 mm long, aculeus 1.45 mm long, its lateral flanges with 9 – 12 serrations on margins (Figure 3 A & B), tip of aculeus chisel-shaped, sharply pointed at apex and with 3 preapical setulae. Eversible membrane 1.54 mm long with finger-like roundish blunt spicules on distal end, sharp triangular spicules in middle and similar spicules like middle eversible membrane present proximally with irregular round gritty spicules on lateral margins (Figure 3 D & E). Two spermathecae, circular and striated (Figure 3 F). Epandrium oval in posterior view (Figure 4 A), rectangular in lateral view (Figure 4 B), both lobes of lateral surstylus almost equal in length and blunt. Median surstylus shorter than lateral surstylus, each end with two black sharp-ended spiny prensisetae. Proctiger and posterior end of epandrium setose. Phallus 1.87 mm long excluding glans (0.24 mm), latter with narrow preputium, elongate saddle-like acrophallus and broad, apical membranous lobe roundish (Figure 4 C).	en	Singh, Maneesh Pal, Sneha, Sharma, Sharma, Isha, Gupta, Divender, Hancock, David Lawrence, Prabhakar, Chandra Shekhar (2024): A new species of genus Oedaspis Loew and new distribution records of Acidiostigma Hendel & Goniurellia Hendel from India (Diptera: Tephritidae). Zootaxa 5514 (5): 469-480, DOI: 10.11646/zootaxa.5514.5.4, URL: http://dx.doi.org/10.11646/zootaxa.5514.5.4
AC58878FFFF4FFD0FF452B6ADB76FED2.taxon	materials_examined	Type material: Holotype ♀, ex Artemisia sp., INDIA, Himachal Pradesh, Solan, Nauni, 30 ° 51 ’ 47 ” N 77 ° 10 ’ 7 ” E. 15. iv. 2024, Maneesh Leg. at Zoological Survey of India (High Altitude Regional Centre (HARC), Solan, Himachal Pradesh). Paratypes 2 ♂ & 1 ♀, on Prunus sp., INDIA, Himachal Pradesh, Solan, Nauni, 30 ° 51 ’ 47 ” N 77 ° 10 ’ 7 ” E. 21. iii. 2023, Sneha & Maneesh Leg. with first author. Paratypes 3 ♂ & 4 ♀, same data as holotype.	en	Singh, Maneesh Pal, Sneha, Sharma, Sharma, Isha, Gupta, Divender, Hancock, David Lawrence, Prabhakar, Chandra Shekhar (2024): A new species of genus Oedaspis Loew and new distribution records of Acidiostigma Hendel & Goniurellia Hendel from India (Diptera: Tephritidae). Zootaxa 5514 (5): 469-480, DOI: 10.11646/zootaxa.5514.5.4, URL: http://dx.doi.org/10.11646/zootaxa.5514.5.4
AC58878FFFF4FFD0FF452B6ADB76FED2.taxon	etymology	Etymology: The species is named after its type locality “ Nauni ” (Himachal Pradesh, India).	en	Singh, Maneesh Pal, Sneha, Sharma, Sharma, Isha, Gupta, Divender, Hancock, David Lawrence, Prabhakar, Chandra Shekhar (2024): A new species of genus Oedaspis Loew and new distribution records of Acidiostigma Hendel & Goniurellia Hendel from India (Diptera: Tephritidae). Zootaxa 5514 (5): 469-480, DOI: 10.11646/zootaxa.5514.5.4, URL: http://dx.doi.org/10.11646/zootaxa.5514.5.4
AC58878FFFF4FFD0FF452B6ADB76FED2.taxon	biology_ecology	Host plant: Artemisia sp. (Asteraceae). O. nauniensis sp. nov. was reared from subterranean stem galls on Artemisia sp. (Figs 5 J – L). It is impossible to detect the galls without uprooting the plant and we did not observe any kind of symptoms on infested plants except they showed more branching from the crown region compared with healthy plants. Artemisia sp. is well distributed in the Himalayas and its density indicates abundance of O. nauniensis sp. nov. in Himachal Pradesh. Artemisia species are, so far, the only recorded host plants for Eurasian species of Oedaspis (Korneyev, 2002) except O. quinquefasciata Becker (from Canary Islands) on Schizogyne sp.	en	Singh, Maneesh Pal, Sneha, Sharma, Sharma, Isha, Gupta, Divender, Hancock, David Lawrence, Prabhakar, Chandra Shekhar (2024): A new species of genus Oedaspis Loew and new distribution records of Acidiostigma Hendel & Goniurellia Hendel from India (Diptera: Tephritidae). Zootaxa 5514 (5): 469-480, DOI: 10.11646/zootaxa.5514.5.4, URL: http://dx.doi.org/10.11646/zootaxa.5514.5.4
AC58878FFFF4FFD0FF452B6ADB76FED2.taxon	discussion	Remarks: Oedaspis nauniensis sp. nov. possesses high variability in wing pattern and frontal as well as thoracic dorsocentral setal distribution. In general appearance it is similar to O. dichotoma but can be differentiated by darker scutum, black abdomen and aculeus having serrations only on lateral flanges. It is also similar to O. japonica but can be differentiated by lack of presutural dorsocentral setae, pterostigma with 3 black spots and shiny black abdomen of female. Males of O. japonica have orange abdomens without any transverse black bands while O. nauniensis males have orange-yellow abdomens with transverse light black bands on tergites III – V.	en	Singh, Maneesh Pal, Sneha, Sharma, Sharma, Isha, Gupta, Divender, Hancock, David Lawrence, Prabhakar, Chandra Shekhar (2024): A new species of genus Oedaspis Loew and new distribution records of Acidiostigma Hendel & Goniurellia Hendel from India (Diptera: Tephritidae). Zootaxa 5514 (5): 469-480, DOI: 10.11646/zootaxa.5514.5.4, URL: http://dx.doi.org/10.11646/zootaxa.5514.5.4
AC58878FFFFEFFD0FF452F2EDA71F960.taxon	discussion	[Note: A third species, G. persignata Freidberg (= ceylonensis Freidberg), is known from North Africa and Ethiopia to Turkmenistan and from Sri Lanka and possibly occurs in India. It is similar to G. tridens but the hyaline indentation in cell r 1 beyond apex of pterostigma does not cross vein R 2 + 3 or enter cell r 4 + 5; both species were illustrated by Freidberg (1980)]	en	Singh, Maneesh Pal, Sneha, Sharma, Sharma, Isha, Gupta, Divender, Hancock, David Lawrence, Prabhakar, Chandra Shekhar (2024): A new species of genus Oedaspis Loew and new distribution records of Acidiostigma Hendel & Goniurellia Hendel from India (Diptera: Tephritidae). Zootaxa 5514 (5): 469-480, DOI: 10.11646/zootaxa.5514.5.4, URL: http://dx.doi.org/10.11646/zootaxa.5514.5.4
AC58878FFFFEFFD0FF452B89DD08FAB0.taxon	description	(Figure 6 A, B, E & F)	en	Singh, Maneesh Pal, Sneha, Sharma, Sharma, Isha, Gupta, Divender, Hancock, David Lawrence, Prabhakar, Chandra Shekhar (2024): A new species of genus Oedaspis Loew and new distribution records of Acidiostigma Hendel & Goniurellia Hendel from India (Diptera: Tephritidae). Zootaxa 5514 (5): 469-480, DOI: 10.11646/zootaxa.5514.5.4, URL: http://dx.doi.org/10.11646/zootaxa.5514.5.4
AC58878FFFFEFFD0FF452B89DD08FAB0.taxon	diagnosis	Diagnosis: A small sized (2.97 mm) dark greyish species with well developed chaetotaxy on head and thorax. Three frontal setae and 2 orbital setae, posterior orbital stubby white and shorter than anterior, ocellar setae crossed forming X-pattern, outer vertical stubby white and postocular thick stubby white. Thorax dark grayish, setose, with 3 darker lyrate vittae, full complement of thoracic setae except lacking apical scutellar setae. Entire scutum and abdomen covered in whitish setulae. All legs yellowish fulvous. Wings (2.75 mm) hyaline except reticulate pattern on wing apex. Dark black spot present in cell r 2 + 3 confluent with vein R 4 + 5, circular oval spot present in cell r 4 + 5 beside 6 irregular shaped hyaline spots (Figure 6 B). Apical Y-shaped fork present in cell r 4 + 5 and outer arms confluent with vein R 4 + 5 and M. Abdomen black except lateral margins of tergites I and II brownish. Narrow brownish band present on posterior end of tergites II – V, 5 – 6 long setae present on posterior margin of tergite V. Genitalia: Epandrium elongated oval in posterior view, epandrium higher than proctiger, lateral surstylus longer than median surstylus, latter with dark spiny prensisetae, aedeagus 0.29 mm long excluding glans (0.12 mm) and apical lobe (1.28 mm), the latter unusually elongate (Figure 6 E & F).	en	Singh, Maneesh Pal, Sneha, Sharma, Sharma, Isha, Gupta, Divender, Hancock, David Lawrence, Prabhakar, Chandra Shekhar (2024): A new species of genus Oedaspis Loew and new distribution records of Acidiostigma Hendel & Goniurellia Hendel from India (Diptera: Tephritidae). Zootaxa 5514 (5): 469-480, DOI: 10.11646/zootaxa.5514.5.4, URL: http://dx.doi.org/10.11646/zootaxa.5514.5.4
AC58878FFFFEFFD0FF452B89DD08FAB0.taxon	materials_examined	Material examined: 1 ♂, on Bidens pilosa L., INDIA, Himachal Pradesh, Solan, Nauni, 30 ° 51 ’ 47 ” N 77 ° 10 ’ 7 ”. 5. vi. 2024, Maneesh Leg. at Zoological Survey of India (HARC, Solan).	en	Singh, Maneesh Pal, Sneha, Sharma, Sharma, Isha, Gupta, Divender, Hancock, David Lawrence, Prabhakar, Chandra Shekhar (2024): A new species of genus Oedaspis Loew and new distribution records of Acidiostigma Hendel & Goniurellia Hendel from India (Diptera: Tephritidae). Zootaxa 5514 (5): 469-480, DOI: 10.11646/zootaxa.5514.5.4, URL: http://dx.doi.org/10.11646/zootaxa.5514.5.4
AC58878FFFFEFFD0FF452B89DD08FAB0.taxon	biology_ecology	Host plant: Not recorded in the region (possibly Pulicaria sp.).	en	Singh, Maneesh Pal, Sneha, Sharma, Sharma, Isha, Gupta, Divender, Hancock, David Lawrence, Prabhakar, Chandra Shekhar (2024): A new species of genus Oedaspis Loew and new distribution records of Acidiostigma Hendel & Goniurellia Hendel from India (Diptera: Tephritidae). Zootaxa 5514 (5): 469-480, DOI: 10.11646/zootaxa.5514.5.4, URL: http://dx.doi.org/10.11646/zootaxa.5514.5.4
AC58878FFFFEFFD0FF452B89DD08FAB0.taxon	distribution	Distribution: Egypt: Sinai (Freidberg & Kugler, 1989), Iran (Mohamadzade Namin et al., 2018), and India: Himachal Pradesh (first record).	en	Singh, Maneesh Pal, Sneha, Sharma, Sharma, Isha, Gupta, Divender, Hancock, David Lawrence, Prabhakar, Chandra Shekhar (2024): A new species of genus Oedaspis Loew and new distribution records of Acidiostigma Hendel & Goniurellia Hendel from India (Diptera: Tephritidae). Zootaxa 5514 (5): 469-480, DOI: 10.11646/zootaxa.5514.5.4, URL: http://dx.doi.org/10.11646/zootaxa.5514.5.4
AC58878FFFFEFFD2FF452D5EDECFFF4E.taxon	description	(Figure 6 C & D)	en	Singh, Maneesh Pal, Sneha, Sharma, Sharma, Isha, Gupta, Divender, Hancock, David Lawrence, Prabhakar, Chandra Shekhar (2024): A new species of genus Oedaspis Loew and new distribution records of Acidiostigma Hendel & Goniurellia Hendel from India (Diptera: Tephritidae). Zootaxa 5514 (5): 469-480, DOI: 10.11646/zootaxa.5514.5.4, URL: http://dx.doi.org/10.11646/zootaxa.5514.5.4
AC58878FFFFEFFD2FF452D5EDECFFF4E.taxon	diagnosis	Diagnosis: Species well described by Wang (1998) and our specimen fits the description. It is medium sized (6.1 mm) brown-red species having well developed chaetotaxy on head and thorax. Three frontal setae and 2 orbital setae. Occiput pale brown with row of thin black setae. Gena pale brown with two strong setae and row of setulae. Row of setulae present on either side of face. Scutum red-brown, covered with minute black dots, dorsocentral setae situated behind the line of postsutural supra-alar setae. Two anepisternal setae, no yellow marking on thorax. All leg segments pale brownish and fore femora with 7 – 8 posteroventral setae. Wings largely dimidiate (6.3 mm), cell c slightly longer than cell sc, broad dark band present, covering anterior half of wing above vein M except cells bc and c hyaline and a hyaline spot in cell r 1 at apex of pterostigma. Abdomen oval, brownish, setose, tergites IV – VI blackish. Terminal area of female abdomen black with dark and flattened oviscape. Genitalia not dissected.	en	Singh, Maneesh Pal, Sneha, Sharma, Sharma, Isha, Gupta, Divender, Hancock, David Lawrence, Prabhakar, Chandra Shekhar (2024): A new species of genus Oedaspis Loew and new distribution records of Acidiostigma Hendel & Goniurellia Hendel from India (Diptera: Tephritidae). Zootaxa 5514 (5): 469-480, DOI: 10.11646/zootaxa.5514.5.4, URL: http://dx.doi.org/10.11646/zootaxa.5514.5.4
AC58878FFFFEFFD2FF452D5EDECFFF4E.taxon	materials_examined	Material examined: 1 ♀, Shimla, Himachal Pradesh, India, 29. VIII. 2020, Isha Sharma Leg. at Zoological Survey of India (HARC, Solan).	en	Singh, Maneesh Pal, Sneha, Sharma, Sharma, Isha, Gupta, Divender, Hancock, David Lawrence, Prabhakar, Chandra Shekhar (2024): A new species of genus Oedaspis Loew and new distribution records of Acidiostigma Hendel & Goniurellia Hendel from India (Diptera: Tephritidae). Zootaxa 5514 (5): 469-480, DOI: 10.11646/zootaxa.5514.5.4, URL: http://dx.doi.org/10.11646/zootaxa.5514.5.4
AC58878FFFFEFFD2FF452D5EDECFFF4E.taxon	biology_ecology	Host plant: Unknown.	en	Singh, Maneesh Pal, Sneha, Sharma, Sharma, Isha, Gupta, Divender, Hancock, David Lawrence, Prabhakar, Chandra Shekhar (2024): A new species of genus Oedaspis Loew and new distribution records of Acidiostigma Hendel & Goniurellia Hendel from India (Diptera: Tephritidae). Zootaxa 5514 (5): 469-480, DOI: 10.11646/zootaxa.5514.5.4, URL: http://dx.doi.org/10.11646/zootaxa.5514.5.4
AC58878FFFFCFFD2FF452B3CDE24FBC6.taxon	discussion	[Note: A. harmandi, described from a male, and A. lucens, described from a female, both from Darjeeling, are possibly synonymous]	en	Singh, Maneesh Pal, Sneha, Sharma, Sharma, Isha, Gupta, Divender, Hancock, David Lawrence, Prabhakar, Chandra Shekhar (2024): A new species of genus Oedaspis Loew and new distribution records of Acidiostigma Hendel & Goniurellia Hendel from India (Diptera: Tephritidae). Zootaxa 5514 (5): 469-480, DOI: 10.11646/zootaxa.5514.5.4, URL: http://dx.doi.org/10.11646/zootaxa.5514.5.4
